Acarorum Catalogus: Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae) (Faunistica)

The first volume of the series “Acarorum Catalogus” is focused to the subfamilies Calyptostomatoidea and Erythraeoidea. ...

Author: P. Beron

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The first volume of the series “Acarorum Catalogus” is focused to the subfamilies Calyptostomatoidea and Erythraeoidea. More than 750 species are listed with their synonyms and the citations in various papers. The book contains also the list by country and authors having published data on these families for all species known. There are other lists including hosts of the larvae of these mites, species inquirendae, list of the many new combinations and bibliography of more than 1000 titles. This first volume of the series represents the format of the next volumes of other groups of mites.

ACARORUM CATALOGUS

ACARORUM CATALOGUS I

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ACARORUM CATALOGUS I ACARIFORMES:

Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae)

Petar Beron

Acarorum Catalogus I

ACARORUM CATALOGUS Editor Petar Beron

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ACARORUM CATALOGUS I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae)

Petar Beron

Edition of Pensoft Publishers and the National Museum of Natural History, Sofia Bulgarian Academy of Sciences Sofia–Moscow 2008

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ACARORUM CATALOGUS I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae) Petar Beron

First published 2008 ISBN 978-954-642-308-5

Beron P. 2008. Acarorum Catalogus 1. Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae). Pensoft & Natn. Mus. Nat. Hist., Sofia, 272 pp.

© PENSOFT Publishers © National Museum of Natural History, Sofia All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted in any form by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior written permission of the copyright owner.

Pensoft Publishers Geo Milev Str. 13a, Sofia 1111, Bulgaria Fax: +359-2-870-42-82 [email protected] www.pensoft.net

Printed in Bulgaria, March 2008


Contents Acarorum Catalogus – the Idea and the Construction The Acari – what are they? Systems of Acari

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The System of the Acari Accepted by the Present Author Selected References on Acari

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The Mite Subcohort Erythraeina

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The Composition of Erythraeina

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Catalogue of the Mite Superfamily Calyptostomatoidea Superfamily Calyptostomatoidea Southcott Family Calyptostomatidae Oudemans Distribution of Calyptostomatoidea Index of Calyptostomatoidea

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Bibliography of Calyptostomatoidea

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Catalogue of the Mite Superfamily Erythraeoidea Superfamily Erythraeoidea Grandjean Family Smarididae Vitzthum

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Subfamily Smaridinae Southcott

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Subfamily Hirstiosomatinae Southcott Family Erythraeidae Robineau-Desvoidy Subfamily Erythraeinae Southcott

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Subfamily Leptinae Billberg, sensu Southcott Subfamily Callidosomatinae Southcott Tribe Callidosomatini Southcott Tribe Charletoniini Southcott

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Subfamily Phanolophinae Southcott 174 Subfamily Balaustiinae Southcott

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Subfamily Myrmicotrombiinae Southcott

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Species Incertae Sedes, Species Inquirendae Doubtful or excluded names

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Other names, published in Erythraeoidea Fossil Erythraeoidea

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Taxonomical Changes in this Catalogue Distribution of Erythraeoidea Hosts of Erythraeoidea

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Index of Erythraeoidea

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Bibliography of Erythraeoidea

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Acarorum Catalogus – the Idea and the Construction The Acari is a huge, heterogeneous (=polyphyletic) group of arachnid orders containing more than 50 000 species globally. The many scattered papers on the subject published in specialized and other editions are difficult to find and encompass, most groups have not been revised and the need for a general catalogue for all Acari is evident. Such catalogues have been made for other Arachnida (Pseudoscorpions, etc.), and they have proved extremely useful. The idea of the editor is to publish a long series of catalogues of Acari made by specialists in different superfamilies, subcohorts, cohorts, or other higher taxa. All qualified colleagues are invited to contact P. Beron (NMNH, 1000 Sofia, Bld Tsar Osvoboditel 1).

The Acari – what are they? Systems of Acari The position of Acari in the system of Chelicerata has been viewed differently by the authors and it is difficult to say which system is better and reflects the reality – these contradicting systems have resulted from the work of the most experienced specialists of the XXth Century. If we check the classifications proposed in the last 50 years, we shall observe an unbelievable lack of congruence concerning the higher categories of mites. We further enumerate some of the views on this group of Arthropods, which is nonetheless accepted unanimously as one entity called Acari, or mites. The prevailing opinion is that the Arachnids form a class and the mites are part of it. An exception is the system of Dubinin (1956, 1959), followed by Feider (1965). Dubinin understands the Chelicerata as a superclass and the mites as a class, formed by three orders. Van der Hammen (1961) and Evans, Sheals and Macfarlane (1961) introduced the idea that the mites form a subclass within the Arachnida. The system of van der Hammen (several papers) is rather radical. He elevated many groups usually considered as suborders to full ordinal rank (v.d. Hammen, 1968 – seven orders, 1972 – eight orders). In the famous Traité de Zoologie of Grassé, M. André (1949) considers six of them as orders, however in some pages before that (in the same paper!) he considers them as suborders of a single order Acari. The fast evolution of thinking, even of a single author, is evident. Zachvatkin (1952) proposed a revolutionary system, in which the three orders of Acari (Acariformes, Parasitiformes and Opilioacarina) are ascribed to different super orders of Arachnida. These three orders are now accepted by most Acarologists. An exception is the system of Krantz (1978), in which Opilioacarida are included in the Parasitiformes. The three orders are often united (according to some authors artificially), in the subclass Acari, Acarida or Acaromorpha. Many authors accept the biphyletic origin of Acari, based mostly on the views of F. Grandjean (1935). He discovered the fact that some mites (Acariformes) contain an optically active substance called actinochitin, or actinopylin,

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whereas some others (Opilioacarida and Parasitiformes) do not contain this substance. Accordingly, he divided the mites into Actinochitinosi and Anactinochitinosi. These names have been changed into Anactinotrichida and Actinotrichida by v.d. Hammen (1968), as super-ordinal names. In the recent decades, however, most authors consider the system of v.d. Hammen (1968) as too clumsy and complicated, and use simply the three generally accepted ordinal names with different endings (-ina, -ida, etc.). The general definition of mites as a group, or subclass is: Acari – Arachnids possessing a gnathosoma and (with very few exceptions) a hexapod larva. Somatic segmentation is inconspicuous or absent. The podosoma is broadly joined to the opisthosoma to form an idiosoma. The definitions of the three orders (extracted mostly from v.d. Hammen, 1968) are: Acariformes - the ordinary setae have an axis of actinopyline - a small numbers of “lyrifissures” (maximum seven on the idiosoma and one on the appendages) - generally neither free coxae nor sternum - presence of a sejugal groove - no dorsal lyrifissure on the chelicerae - no claws on the palp - trichobothria are often present on the legs and on the body - often with regressive phases (calyptostases or elattostases) Parasitiformes - Stigmata belonging to opisthosomal homologous series (two pairs in Holothyrida, one in Gamasida and Ixodida) - considerable number of “lyrifissures” on the idiosoma - often with claws (or their homologues) on the palp - often with sternum and free coxae - no sejugal groove between legs 2 and 3 - no trichobothrias - femur divided into basi – and telofemur by secondary articulation - often with many secondary articulations on the leg tarsi Opilioacarida – 4 pairs of stigmata The views of different authors (mainly after 1950) concerning the system of Acari The Acari as a class Canestrini (1891) – Acaroidea – class, divided into 6 orders: Astigmata, Hydracarina, Prostigmata, Cryptostigmata, Metastigmata, Mesostigmata Dubinin (1956b, 1957) – Acaromorpha (class in Chelicerophora)


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Dubinin (1959) – Acaromorpha (class in subtype Chelicerophora, superclass Chelicerata). Orders: Acaroformes, Parasitiformes and Opilioacarina Feider (1965) – Acaromorpha (class) Wharton (1976) – Acari (class of the subphylum Chelicerata, containing 3 orders: Parasitiformes, Opilioacariformes and Acariformes) Hoffmann (1979) – Acarida (class) The Acari as a subclass of Arachnida Hammen van der (1961) – Acarida (subclass of Arachnida, containing 3 orders: Opilioacarida, Anactinotrichida and Actinotrichida) Evans, Sheals & Macfarlane (1961) – Acari – subclass of two superorders (Anactinochaeta and Actinochaeta) and 7 orders (Notostigmata, Tetrastigmata, Prostigmata, Metastigmata, Mesostigmata, Astigmata and Cryptostigmata) Hammen van der (1968) – Acarida – subclass of Arachnida with two superorders (Anactinotrichida and Actinotrichida) and 7 orders: Opilioacarida, Holothyrida, Gamasida, Ixodida (Anactinotrichida); Actinedida (= Trombidiformes), Oribatida, Acaridida (Actinotrichida). Hammen van der (1972) – Acarida – subclass of Arachnida with two superorders (Anactinotrichida and Actinotrichida) and 8 orders: Opilioacarida, Holothyrida, Gamasida, Ixodida (Anactinotrichida); Actinedida, Oribatida, Acaridida, Tarsonemida (Actinotrichida). Wilson (1974) – Acarina – subclass of Arachnida with three orders: Opilioacariformes, Parasitiformes, and Acariformes, seven suborders and 333 families. Lindquist (1984) – Acarida – subclass of Arachnida Evans (1992) – Acari – subclass of Arachnida with two superorders Anactinotrichida and Actinotrichida and seven orders: Notostigmata, Holothyrida, Ixodida, Mesostigmata, Prostigmata, Astigmata, Oribatida. The Acari as one order Trägårdh (1932) – introduced a new suborder Palaeacariformes Wharton (1938) – Acarina (order with suborders Trombidina, Oribatina, Parasitina, Ixodina, etc.) André (1949) – Acari (order of class Arachnida with 6 suborders); further in the same paper as orders (of the heterogenous group of Acari) Baker & Wharton (1952) – Acarina (order of class Arachnoidea) Turk (1953) – Acari (order) Dusbabek (1998) – Acari (order with suborders Metastigmata, Mesostigmata, Prostigmata and Astigmata)

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The Acari as two (?) biphyletic orders within different classes Zoological Record (2006) – Acari as two different (biphyletic?) orders within Arachnida Krantz (1978) – Parasitiformes (includes Opilioacarida, Holothyrida, Gamasida and Ixodida as suborders) and Acariformes (includes Actinedida, Acaridida and Oribatida as suborders) The Acari as two biphyletic orders within different classes Van der Hammen (1977) – Chelicerata are subdivided into 7 classes. The Actinotrichida are included in class Epimerata, the Anactinotrichida – in class Cryptognomae The Acari as three different (polyphyletic?) orders within Arachnida Zachvatkin (1952) – orders Acariformes, Parasitiformes and Opilioacarina within class Chelicerata Dubinin (1956) – orders Acariformes, Parasitiformes and Opilioacarina within class Acaromorpha Gilyarov & Krivolutzkyi (1975) – orders Opilioacarina, or Notostigmata, Parasitiformes (includes suborders Ixodides and Mesostigmata) and Acariformes (includes suborders Sarcoptiformes and Trombidiformes). Sarcoptiformes includes the groups Acaridiae and Oribatei. The position and volume of Opilioacarida With (1904) – Notostigmata – suborder of Acari André (1949) – Notostigmata (= Opilioacaroidea) – suborder of Acari; further in the same paper as order Baker & Wharton (1952) – Notostigmata (group within suborder Onychopalpida of order Acarina). One family Opilioacaridae Zachvatkin (1952) – Opilioacarina (order of superorder Actinoderma, class Chelicerata) Dubinin (1956, 1959) – Opilioacarina (order of class Acaromorpha) Evans, Sheals & Macfarlane (1961) – Notostigmata (order of superorder Anactinochaeta, subclass Acari) Hammen v.d. (1961) – Opilioacarida (order of subclass Acarida), with one suborder Notostigmatina Hammen v.d. (1966) – Opilioacarida Johnston (1968) – Opilioacariformes (order of Arachnida) Hammen v.d. (1968) – Opilioacarida (order of subclass Acarida) Hammen v.d. (1972) – Opilioacarida (order of subclass Acarida) Krivoluckyi (1975) – Opilioacarina (order) Kranz (1978) – Opilioacarida (suborder of Parasitiformes) Evans (1992) – Notostigmata (order of subclass Acari)


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The position and volume of Acariformes André (1949) – 3 suborders of Acari: Thrombidiformes, Sarcoptiformes, Tetrapodili; further in the same paper as orders Baker & Wharton (1952) – Sarcoptiformes (suborder of order Acarina), divided into Acaridiae and Oribatei (groups) Zachvatkin (1952) – Acariformes (order of superorder Actinochaeta, class Chelicerata) Dubinin (1956, 1959) – Acariformes (order of class Acaromorpha) Knülle (1959) – “monophyletic group”, containing Endeostigmata, Oribatei, Prostigmata, Tarsonemini, Tetrapodili, and Acaridiae. Hammen van der (1961) – Actinotrichida (order of subclass Acarida) Hammen van der (1968) – Actinotrichida (superorder of subclass Acarida) Hammen van der (1972) – Actinotrichida (superorder of subclass Acarida) Hammen van der (1977) – Actinotrichida (subclass of class Epimerata of phylum Chelicerata) Flechtmann (1977) – Trombidiformes (order of subclass Acari) Krantz (1978) – Acariformes (order). Includes three suborders: Actinedida, Acaridida and Oribatida) Kethley (1982) – Acariformes (= Actinotrichida)(order of subclass Acari). Subdivided into 3 suborders : Astigmata, Oribatei and Prostigmata. Evans (1992) – Actinotrichida (superorder of subclass Acari). Halliday (1998) – order Acariformes Within Order Acariformes: The position and volume of Acaridida André (1949) – Acaridiae (suborder of order Sarcoptiformes with 20 families) Baker & Wharton (1952) – Sarcoptiformes (suborder of order Acarina). Divided into Acaridiae and Oribatei (as groups) Dubinin (1959) – Sarcoptiformes (suborder of order Acariformes) Hammen van der (1961) – Acarina (suborder of order Actinotrichida) Evans et al. (1961) – Astigmata (order) Hammen van der (1968) – Acaridida (order of subclass Acarida) Hammen van der (1972) – Acaridida (order of subclass Acarida) Flechtmann (1977) – Acaridiae (suborder of Sarcoptiformes) Krantz (1978) – Acaridida (suborder of Acariformes) O’Connor (1982) – Astigmata (suborder of order Acariformes) Evans (1992) – Astigmata (order of superorder Actinotrichida) Halliday (1998) – Astigmata (suborder of order Acariformes)

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The position and volume of Prostigmata André (1949) – Trombidiformes – suborder of Acari, divided into Tarsonemini and Prostigmata; further in the same paper as order with three superfamilies: Tarsonemini, Endeostigmata and Prostigmata. Tetrapodili (Eriophyidae and Phyllocoptidae) are treated first as suborder (p.794), than as order (p.884) Baker & Wharton (1952) – Trombidiformes (suborder of order Acarina). Divided into Tetrapodili, Tarsonemini and Prostigmata. Turk (1953) – Trombidiformes (suborder of order Acari). Includes supercohorts Tarsonemini, Endeostigmata, Prostigmata. Cunliffe (1955) – Trombidiformes (suborder of order Acarina). Includes two supercohorts (Heterostigmata, Prostigmata and Parasitenini), 5 cohorts (Tarsonemina, Endeostigmatina, Promatina, Parasitengina and Hydrachnellae) and 45 families. Dubinin (1959) – Trombidiformes (suborder of order Acariformes) Hammen van der (1961) – Prostigmatina (suborder of order Actinitrichida) Evans, Sheals & Macfarlane (1961) – Prostigmata (order of superorder Actinochaeta, subclass Acari) Wainstein (1965) – Trombidiformes (suborder) Hammen van der (1968) – Actinedida (= Trombidiformes) – order of subclass Acarida Gilyarov, Wainstein & Krivolutzkyi (1978) – Trombidiformes (suborder). Contains 6 cohorts: Tetrapodili, Tarsonemina, Endeostigmata, Parasitengona, Pterygosomata and Prostigmata). Krantz (1978) – Actinedida (suborder of Acariformes). Includes 5 cohorts (Pachygnathina, Adamystina, Labidostommatina, Eupodina and Eleutherogonina), 28 superfamilies and 111 families. Kethley (1982) – Prostigmata (= Trombidiformes = Actinedida). Subdivision: 8 cohorts, 31 superfamilies, 133 families, more than 1100 genera and 14000 nominate species. Kethley (1990) – Prostigmata (Actinedida). Subdivision: 6 cohorts, 31 superfamilies, 136 families, 1100 genera and 14000 or more species. Evans (1992) – Prostigmata (order of superorder Actinotrichida). Follows Kethley (1982) Halliday (1998) – Prostigmata (suborder of order Acariformes) The position and volume of Oribatida André (1949) – Oribatei – suborder of order Sarcoptiformes Baker & Wharton (1952) – Oribatei (group of suborder Sarcoptiformes) Grandjean (1953) – divided into 11 groups (10 groups of inferior Oribatids and 1 group of superior Oribatids, called “Circumdehiscentiae”. Dubinin (1959) – Oribatei (suborder of order Acariformes) Hammen van der (1961) – Oribatina (suborder of order Actinotrichida) Evans, Sheals & Macfarlane (1961) – Cryptostigmata (order of superorder Actinochaeta, class Acari)


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Hammen van der (1968) – Oribatida (order of subclass Acarida) Balogh (1972) – Oribatei Gilyarov & Krivolutzkyi (1975) – Oribatei (group within suborder Sarcoptiformes) Flechtmann (1977) – Oribatei, or Cryptostigmata (suborder of Sarcoptiformes) Krantz (1978) – Oribatida (suborder of Acariformes). Includes 2 supercohorts (Macropylides and Brachypylides), 7 cohorts (Bifemoratina, Ptyctimina, Arthronotina, Holonotina, Polytrichae, Oligotrichae, Pterogasterina), 44 superfamilies and families. Johnston (1982) – Oribatida (= Oribatei or Cryptostigmata) – suborder of order Acariformes, 145 fam. Schatz (1983) – Oribatei (Oribatida; Cryptostigmata) – suborder of order Acariformes, subclass Acari Tarman (1983) – Oribatei (suborder of order Acariformes) Norton (1990) – Oribatida Evans (1992) – Oribatida (order of superorder Actinotrichida) Halliday (1998) – Oribatida (suborder of order Acariformes) The position and volume of Parasitiformes André (1949) – Parasitiformes (= Gamasiformes) – suborder of Acari; further in the same paper as order with Mesostigmata and Ixodoidea as suborders Baker & Wharton (1952) – Mesostigmata and Ixodides (suborders of order Acarina) Zachvatkin (1952) – Parasitiformes (order of superorder Actinoderma, class Chelicerata) Turk (1953) – Parasitiformes (suborder of order Acari). Contains supercohorts Mesostigmata and Ixodides Dubinin (1956b) – Parasitiformes (order of class Acaromorpha). Dubinin (1959) – Parasitiformes (order of class Acaromorpha). Contains suborders Holothyroidea, Monogynaspida, Trigynaspida and Ixodides Hammen van der (1961) – Anactinotrichida (order of subclass Acarida) Hammen van der (1968) – Anactinotrichida (superorder of subclass Acarida) Karg (1971) – Anactinochaeta (Parasitiformes) – suborder of order Acarina Hammen van der (1972) – Anactinotrichida (superorder of subclass Acari) Hammen van der (1977) – Anactinotrichida (subclass of class Cryptognomae of phylum Chelicerata) Flechtmann (1977) – Parasitiformes (order of subclass Acari) Krantz (1978) – Parasitiformes (order) Lindquist (1984) – Anactinotrichida Karg (1989) – Parasitiformes (= Anactinotrichida)(suborder of order Acari) Evans (1992) – Anactinotrichida (superorder of subclass Acari). Subdivided into four orders: Notostigmata, Holothyrida, Ixodida, Mesostigmata Schmölzer (1995) – Anactinochaeta (Parasitiformes) – suborder of order Acarina Halliday (1998) – order Parasitiformes

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Within Order Parasitiformes : The position of Holothyrida André (1949) – Holothyroidea (suborder of Acari; further in the same paper as order) Baker & Wharton (1952) – Holothyroidea (group within the suborder Onychopalpida, together with Notostigmata = Opilioacarida) Dubinin (1959) – Holothyroidea (suborder of order Parasitiformes) Hammen van der (1961) – Holothyrina (suborder of order Anactinotrichida) Evans, Sheals & Macfarlane (1961) – Tetrastigmata (order of superorder Anactinochaeta, subclass Acari) Johnston (1968) – Holothyrina (suborder) Hammen van der (1968) – Holothyrida (order of subclass Acarida) Hammen van der (1972) – Holothyrida (order of subclass Acarida) Krantz (1978) – Holothyrida (suborder of Parasitiformes) Johnston (1982) – Holothyrina (suborder of Parasitiformes) Evans (1992) – Holothyrida (order of superorder Anactinotrichida) Lehtinen (1995) – Holothyrina (within Anactinotrichida) Halliday (1998) – Holothyrida (suborder of order Parasitiformes) The position of Gamasida André (1949) – Mesostigmata – suborder of order Parasitiformes Baker & Wharton (1952) – Mesostigmata (suborder of order Acarina) Turk (1953) – Mesostigmata (supercohort of suborder Parasitiformes) Camin & Gorirossi (1955) – Mesostigmata (suborder of order Acarina). Subdivided into two new supercohorts (Monogynaspida: cohorts Liroaspina, Gamasina and Uropodina – and Trigynaspida: cohorts Antennophorina and Cercomegistina) and 12 superfamilies. Dubinin (1959) – two suborders of order Parasitiformes (Monogynaspida and Trigynaspida) Evans, Sheal & Macfarlane (1961) – Mesostigmata (order of superorder Anactinochaeta, class Acari) Hammen van der (1961) – Mesostigmatina (suborder of order Anacinotrichida) Hammen van der (1968) – Gamasida (order of subclass Acarida) Krantz (1970) – Mesostigmata (suborder) Hammen van der (1972) – Gamasida (order of subclass Acarida) Gilyarov & Bregetova (1977) – Mesostigmata (supercohort of Parasitiformes) Flechtmann (1977) – Mesostigmata (suborder of Parasitiformes) Krantz (1978) – Gamasida (suborder of Parasitiformes) Johnston (1982) – Mesostigmata (= Gamasida)(suborder of Parasitiformes) Domrow (1987) – Mesostigmata (order) with suborders Monogynaspida and Trigynaspida Krantz & Ainscough (1990) – Mesostigmata (Gamasida)


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Evans (1992) – Mesostigmata (order of superorder Anactinotrichida) Halliday (1998) – Mesostigmata (suborder of order Parasitiformes) The position of Ixodida André (1949) – Ixodoidea – suborder of order Parasitiformes Baker & Wharton (1952) – Ixodides (suborder of order Acarina) Turk (1953) – Ixodides (supercohort of suborder Parasitiformes) Dubinin (1959) – Ixodides (suborder of order Parasitiformes) Hammen van der (1961) – Ixodina (suborder of order Anactinotrichida) Evans et al. (1961) – Metastigmata Feider (1965) – Ixodides (suborder of Parasitiformes) Hammen van der (1968) – Ixodida (order of subclass Acarida) Krantz (1970) – Metastigmata (suborder of Parasitiformes) Hammen van der (1972) – Ixodida (order of subclass Acarida) Gilyarov & Bregetova (1977) – Ixodides, or Metastigmata (supercohort of Parasitiformes) Flechtmann (1977) – Metastigmata, or Ixodides (suborder of Parasitiformes) Krantz (1978) – Ixodida (suborder of Parasitiformes) Johnston (1982) – Ixodida (= Ixodides, Metastigmata)( suborder of Parasitiformes) Halliday (1998) – Ixodida (suborder of Parasitiformes) Camicas et al. (1998) – Ixodida (within Acarida)

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The System of the Acari Accepted by the Present Author The system used at the start of this Catalogue is a modern compilation of the existing, often contradictory, views. The choice is a subjective one of the Editor. It could be changed during the preparation of the next volumes, with the development of the system of Acari. The main sources (after André, 1949) for compiling this list were the papers of Zachvatkin (1952), Baker & Wharton (1952), Cunliffe (1955), Camin & Gorirossi. (1955), Yunker (1955), Dubinin (1956b), Hughes (1959), Bekker (1960), Southcott (1961a), Wooley (1961), Grandjean (1969), Hammen L. van der (1968, 1971, 1972), Balogh (1972), Vercammen-Grandjean (1973), Krantz (1978), Gilyarov, Wainstein & Krivolutzkyi (1978), O’Connor (1982), Johnston (1982), Krantz & Ainscough (1990), Kethley (1982, 1990), Bernini (1986), Norton (1990), Welbourne (1991), Evans (1992), Camicas et al. (1998). Class Arachnida Subclass Acari (= Acarina; Acarida; Acaromorpha) Order Opilioacarida (= Opilioacarina) Order Acariformes (= Actinotrichida) Suborder Prostigmata (= Trombidiformes; Actinedida) Cohort Endeostigmata Superfamily Pachygnathoidea Superfamily Adamystoidea Cohort Eupodina Superfamily Bdelloidea Superfamily Eupodoidea Superfamily Halacaroidea Superfamily Nematalycoidea Superfamily Tydeoidea Cohort Labidostommatina Superfamily Labidostommatoidea Cohort Anystina Superfamily Anystoidea Superfamily Caeculoidea Superfamily Paratydeoidea Superfamily Pomerantzioidea Superfamily Pterygosomatoidea Cohort Parasitengona Subcohort Erythraeina Superfamily Calyptostomatoidea Superfamily Erythraeoidea Subcohort Trombidiina

The System of the Acari Accepted by the Present Author

Superfamily Tanaupodoidea Superfamily Chyzerioidea Superfamily Trombiculoidea Superfamily Trombidioidea Superfamily Stygothrombioidea Superfamily Hydryphantoidea Superfamily Eylaioidea Superfamily Hydrovolzioidea Superfamily Lebertioidea Superfamily Hygrobatoidea Superfamily Arrenuroidea Cohort Eleutherengona Superfamily Raphignathoidea Superfamily Iolinoidea Superfamily Cheyletoidea Superfamily Tetranychoidea Superfamily Tarsocheyloidea Superfamily Heterocheyloidea Superfamily Pyemotoidea Superfamily Pygmephoroidea Superfamily Tarsonemoidea Suborder Acaridida (= Astigmata; Acaridei) Superfamily Schizoglyphoidea Superfamily Histiostomatoidea Superfamily Canestrinioidea Superfamily Hemisarcoptoidea Superfamily Glycyphagoidea Superfamily Acaroidea Superfamily Hypoderoidea Superfamily Pterolichoidea Superfamily Pyroglyphoidea Superfamily Analgoidea Superfamily Psoroptoidea Suborder Oribatida (= Oribatei, Oribatina, Cryptostigmata) Cohort Macropylina Superfamily Archeonothroidea Superfamily Palaeacaroidea Superfamily Ctenacaroidea Superfamily Hypochthonioidea Superfamily Protoplophoroidea Superfamily Brachychthonioidea

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Superfamily Atopochthonioidea Superfamily Phthiracaroidea Superfamily Euphthiracaroidea Superfamily Lohmannioidea Superfamily Collohmannioidea Superfamily Crotonoidea Superfamily Nanhermannioidea Superfamily Hermannioidea Cohort Brachypylina Superfamily Hermannielloidea Superfamily Liodoidea Superfamily Gymnodamaeoidea Superfamily Damaeoidea Superfamily Cepheoidea Superfamily Microzetoidea Superfamily Amerobelboidea Superfamily Eremaeoidea Superfamily Zetorchestoidea Superfamily Gustavioidea Superfamily Plateremaeoidea Superfamily Carabodoidea Superfamily Oppioidea Superfamily Hydrozetoidea Superfamily Cymbaeremaeoidea Superfamily Licneremaeoidea Superfamily Oripodoidea Superfamily Ceratozetoidea Superfamily Phenopelopoidea Superfamily Oribatelloidea Superfamily Achipterioidea Superfamily Galumnoidea Order Parasitiformes (= Anactinotrichida) Suborder Gamasida (= Mesostigmata) Cohort Sejina Superfamily Sejoidea Cohort Microgyniina Superfamily Microgynioidea Cohort Arctacarina Fam. Arctacaridae Cohort Uropodina Superfamily Thinozerconoidea

The System of the Acari Accepted by the Present Author

Superfamily Polyaspidoidea Superfamily Uropodoidea Cohort Diarthrophallina Superfamily Diarthrophaloidea Cohort Cercomegistina Superfamily Cercomegistoidea Cohort Antennophorina Superfamily Aenictequoidea Superfamily Antennophoroidea Superfamily Celaenopsoidea Superfamily Fedrizzioidea Superfamily Megisthanoidea Superfamily Parantennuloidea Cohort Parasitina Superfamily Parasitoidea Cohort Dermanyssina Superfamily Rhodacaroidea Superfamily Veigaioidea Superfamily Eviphidoidea Superfamily Ascoidea Superfamily Dermanyssoidea Suborder Ixodida Superfamily Argasoidea Superfamily Nuttallielloidea Superfamily Ixodoidea

19

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Selected References on Acari Amrine J. W., Jr., T. B. Stasny. 1994. Catalog of the Eriophyoidea (Acarina: Prostigmata) of the World. – Indira Publ. House, I-IX, 1-798. Amrine J. W., Jr. 1996. Keys to the world genera of the Eriophyoidea (Acari: Prostigmata). – Indira, Michigan, 1-186 André M. 1949. Ordre des Acariens. – Traité de Zoologie (Ed. P.P. Grassé). – 6: 794-892. Arthur D. R. 1960. A Monograph of the Ixodoidea. Part 5. On the genera Dermacentor, Anocentor, Cosminomma, Boophilus and Margaropus. – Cambridge Univ. Press: 251 p. Atyeo W. T. 1960. A Revision of the Mite Family Bdellidae in North and Central America (Acarina, Prostigmata). – Univ. Kansas Science Bull., 40(8): 345-499. Baker E.W. 1965. A Review of the Genera of the Family Tydeidae (Acarina). – Advances in Acarology, Cornell Univ. Press, N.Y., 2: 95-133. Baker E.W., T.M. Evans, D.J.Gould, W.B.Hull, H.L. Keegan. 1956. A Manual of Parasitic Mites of Medical or Economic Importance. – Natl. Pest Control Assoc. Tech. Publ.: 170 p. Baker E.W., A.E. Pritchard. 1960. The tetranychoid mites of Africa. – Hilgardia, 29(11): 455-574. Baker E.W., G.W. Wharton. 1952. An Introduction to Acarology. – Macmillan Co., New York, 465 p.+XIII. Balogh J. 1972. The Oribatid genera of the World. – Akadémiai Kiadó, Budapest. Balogh J., P. Balogh. 1988. Oribatid mites of the Neotropical Region I. – J. Balogh (Ed.) The Soil Mites of the World, 2, Akadémiai Kiadó, Budapest, 335 p. Balogh J., P. Balogh. 1990. Oribatid mites of the Neotropical Region II. – J. Balogh (Ed.) The Soil Mites of the World, 3, Akadémiai Kiadó, Budapest, 333 p. Berlese A. 1904. Acari mirmecofili. – Redia, 1: 299-474. Bekker E.G. 1959. Concerning the Acarina as a natural grouping. – Trud. Inst. Morf. Zhiv., 27: 151-162. Bekker E.G. 1960. Systematics and comparative anatomy in solving the problem of the phylogeny of ticks and mites (Acarina). – Rept. No 1. Critical discussion of the views of acarologists – taxonomists on the polyphylogeny of the order Acarina. – Vest. Mosk. Univ. Ser. 6 – Biol., 15(4): 13-20. Berlese A. 1882-1903. Acari, Myriapoda et Pseudoscorpiones [or Scorpiones] (“A.M.S.”) hucusque in Italia reperta. Fasciculi 1-101. Bernini F. 1986. Current ideas on the phylogeny and the adaptive radiations of Acarida. – Boll. Zool., 53: 279-313. Bertrand M. 1990. La famille des Labidostomidae Oudemans, 1904 (Acari: Actinedida). Révision des genres et sous-genres et catalogue des espèces décrites. – Acarologia, 31(1): 31-38. Bolland H.R., J. Gutierrez, C. Flechtmann. 1998. World catalogue of the spider mite family (Acari: Tetranychidae). – Brill, Leiden, Boston & Köln, 1-392. Camicas J.-L., J.-P. Hervy, F. Adam, P.-C. Morel. 1998. The ticks of the world (Acarida, Ixodida). Nomenclature, described stages, hosts, distribution (including new species described before 1/01/96. – Orstom Editions, Paris, 1-233.

Selected References on Acari

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Camin J.H., F.E. Gorirossi. 1955. A revision of the suborder Mesostigmata (Acarina), based on new interpretations of comparative morphological data. – Chicago Acad. Sci., Spec. Publ., 11: 1-70. Camin J.H., P.C. Morel. 1977. Position systématique et classification des tiques (Acarida: Ixodida). – Acarologia, 18(3): 410-420. Canestrini G. 1891. Abbozzo del sistema Acarologico. – Atti Istit. Ven., 38: 699-725. Canestrini G., P. Kramer. 1899. Demodicidae und Sarcoptidae. – Das Tierreich, 7: 193 pp. Coineau Y. 1974. Elements pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). – Mém. Mus. Nat. d’Hist. nat., (N.S.)A, 81: 299 p. Cook D.R. 1974. Water mite genera and subgenera. – Mem. Amer. Ent. Inst., 21: 860 p. Cross E.A. 1965. The generic relationships of the family Pyemotidae (Acarina:Trombidiformes). – Univ. Kansas Sci. Bull., 45(2): 29-275. Cunliffe F. 1955. A proposed classification of the Trombidiforme mites (Acarina). – Ent. Soc. Washington, 57(5): 209-218. Domrow R. 1987. Acari Mesostigmata Parasitic on Australian Vertebrates: an Annotated Checklist, Keys and Bibliography. – Invertebr. Taxon., 1: 817-948. Domrow R., L.N. Lester. 1985. Chiggers of Australia (Acari: Trombiculidae): an Annotated Checklist, Keys and Bibliography. – Australian J. Zool., Suppl. Ser. No 114: 1-111. Dubinin V.B. 1951. Feather mites (Analgesoidea). Part I. Introduction to their study. – Fauna U.S.S.R., 6(5): 1-363. Dubinin V.B. 1953. Feather mites (Analgesoidea). Part II. Epidermoptidae and Freyanidae. – Fauna U.S.S.R., 6(6): 1-411. Dubinin V.B. 1956a. Feather mites (Analgesoidea). Part III. Pterolichidae. – Fauna U.S.S.R., 6(7): 1-814. [Dubinin V.B.] ÄóáèíèíÂ.Á. 1956b. [The orders of mites and their position in the system of Chelicerata] ÎòðÿäûêëåùåéèïîëîæåíèåèõâñèñòåìåChelicerata. –Òð.IIíàó÷í. êîíô.ïàðàçèòîë.ÓÑÑÐ,Èçä.Èíñò.çîîë.ÀÍÓÑÑÐ,Êèåâ. [Papers of the IIth Sci. Conf. of Parasitology, Ukraine, Kiev]. Dubinin V.B. 1957. [On a new classification of mites Class Acaromorpha W.Dubinin and its position in the system of chelicerate animals, Chelicerophora W. Dubinin]. – Ninth Conf. parasit. Probl.: 82-85 (In Russian). Dubinin V.B. 1959. Subtype Chelicerophora W. Dubinin nom. nov. and its position in the system. 1. Structural peculiarities of Chelicerophora and their classification. – Zool. Zhurnal, 38(8): 1163-1189. Dusbabek F. 1998. Acari Parasiti. – In: Juberthie Ch. and V. Decu (eds), Encyclopaedia biospeologica, 2: 920-928. Estrada-Pena A. 1989. Index-catalog of the ticks (Acarina: Ixodoidea) in the world. Volume 1: genus Haemaphysalis. – Univ. de Zaragoza, Zaragoza: i -IV: 1-932. Evans G.O. 1992. Principles of Acarology. – C.A.B. International, Vol. 1 - 262 p., Vol. 2 - 301 p. Evans G.O., J. G. Sheals & D. Macfarlane.1961. The terrestrial Acari of the British Isles. An introduction to their morphology, biology and classification, I. Introduction and biology. – London, 219 p.

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Evans G.O., J. G. Sheals & D. Macfarlane.1968. The terrestrial Acari of the British Isles. An introduction to their morphology, biology and classification, Vol. I. Alden & Moubray Ltd., 219 p. Fain A. 1957. Les Acariens des familles Epidermoptidae et Rhinonyssidae parasites des fosses nasales d’Oiseaux au Ruanda-Urundi et au Congo belge. – Ann. Mus. r. Congo belge, Tervuren, Série in-8o, Sci. Zool., 60: 176 p. Fain A. 1961. Les acariens parasites endopulmonaires des serpentes. – Bull. Inst. Roy. Sci. natur. Belg., 37(6): 1-135. Fain A. 1962. Les acariens mesostigmatiques ectoparasites des serpentes. – Bull. Inst. Roy. Sci. natur. Belg., 38(18): 1-149. Fain A. 1963a. Chaetotaxie et classification des Speleognathinae. – Bull. Inst. Roy. Sci. nat. Belg., 39: 1-80. Fain A. 1963b. Les Acariens producteurs de gale chez les Lemuriens et les singes, avec une étude des Psoroptidae (Sarcoptiformes). – Bull. Inst. Roy. Sci. Nat, de Belgique, 39(32): 1-125. Fain A. 1964. Les Ophioptidae acariens parasites des écailles des serpents (Trombidiformes). – Bull. Inst. Roy. Sci. nat. Belg., 40(15): 1-57. Fain A. 1965. A review of the family Epidermoptidae Trouessart parasitic on the skin of birds. Parts I – II. – Kon. Acad. Wettensch., Lett. Schone Kunsten Belg. (Wetensch.), 27(84): 1-176; 5-144. Fain A. 1967a. Les acariens de la famille Knemidokoptidae producteurs de gale chez les oiseaux (Sarcoptiformes). – Acta Zool. Path. Antverp., 45: 3-145. Fain A. 1967b. Les hypopes parasites des tissus cellulaires des oiseaux (Hypodectidae: Sarcoptiformes). – Bull. Inst. R. Sci. Nat. Belg., 43 (4): 1-139. Fain A. 1967c. Les hypopes des Glycyphagidae nidicoles en Afrique au sud du Sahara (Acarina: Sarcoptiformes). – Ann. Mus. r. de l’Afrique Centrale – Tervuren, Sér. in-8o, Sci. Zoologiques, 156: 89 p. Fain A. 1968. Etude de la variabilité de Sarcoptes scabiei avec une révision des Sarcoptidae. – Acta Zool. et Pathol. Antverpiensia, 47: 1-196. Fain A. 1969. Les deutonymphs hypopiales vivant en association phorétique sur les Mammifères (Acarina: Sarcoptiformes). – Bull. Inst. r. Sci. nat. Belg., 45(33): 1-262. Fain A. 1971. Les Listrophorides en Afrique au sud du Sahara (Acarina: Sarcoptiformes) II. Familles Listrophoridae et Chirodiscidae. – Acta Zool. et Pathol. Antverpiensia, 54: 231 p. Fain A. 1972a. Les Listrophorides en Afrique au sud du Sahara (Acarina: Sarcoptiformes) III. Famille Atopomelidae. – Ann. Mus. r. de l’Afrique Centrale – Tervuren, Sér. in-8o, Sci. Zoologiques, 197: 200 p. Fain A. 1972b. Les Listrophorides d’Australie et de Nouvelle-Guinée (Acarina: Sarcoptiformes). – Bull. Inst. r. Sci. nat. Belg., 48(5): 149 p. Fain A. 1973. Les Listrophorides d’Amérique néotropicale (Acarina: Sarcoptiformes) I. Familles Listrophoridae et Chirodiscidae. – Bull. Inst. r. Sci. nat. Belg., 49(6): 149 p. Fain A. 1976. Arachnides Acariens Astigmata Listrophoroidea. – Faune de Madagascar, Orstom, Paris, 42: 131 p.


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Fain A. 1978. Les Myobiidae d’Afrique au sud du Sahara et de Madagascar (Acarina – Prostigmata). – Mus. R. Afr. Centrale, Tervuren, Annales, Serie in-8o, Sci. Zool., 224: 186 p. Fain A. 1979. Les Listrophorides d’Amérique néotropicale (Acarina: Astigmates) II. Famille Atopomelidae. – Bull. Inst. r. Sci. nat. Belg., 51(7): 158 p. Fain A., P. Elsen. 1967. Les Acariens de la famille Knemidokoptidae producteurs de gale chez les oiseaux (Sarcoptiformes). – Acta Zool. et Pathol. Antverpiensia, 45: 142 p. Feider Z. 1955. Fauna Republicii Populare Romine. Arachnida, Acarina, Trombidoidea. – Acad. Rep. Pop. Rom., 5(1): 1-186. Feider Z. 1965. Fauna Republicii Populare Romine. Arachnida, Acaromorpha, Suprafamilia Ixodoidea (Capuşe). – Acad. Rep. Pop. Rom., 5(2): 1-404. Garrett L.E., F.H. Haramoto. 1967. A Catalog of Hawaiian Acarina. – 19(3): 381-414. Gabryś G. 1999. The world genera of Microtrombidiidae (Acari, Actinedida, Trombidioidea). – Monographs of the Upper Silesian Museum, 2, pp. 361. Gaud J., W.T. Atyeo. 1996. Feather mites of the world (Acarina: Astigmata): the supraspecific taxa. – Kon. Mus. V. Midden – Afrika Tervuren, Belgie, Annalen Zool. Wetenschappen, 277(1): 1-193. Idem, Part 2. Illustrations. – Kon. Mus. V. Midden – Afrika Tervuren, Belgie, 277(2): 1-436. Gilyarov M.S., D.A. Krivolutzkyi.1975. Sarcoptiformes. – In: Key for identification of soil – inhabiting mites. – Ed. Nauka, Moskow: 491 pp. Gilyarov M.S., N.G. Bregetova. 1977. Mesostigmata. – In: Key for identification of soil – inhabiting mites. – Ed. Nauka, Moskow: 718 pp. Gilyarov M.S., B. A. Wainstein & D.A. Krivolutzkyi.1978. Trombidiformes. – In: Key for identification of soil – inhabiting mites. – Ed. Nauka, Moskow: 271 pp. Grandjean F. 1935. Observations sur les Acariens. – Bull. Mus. Nat. Hist. Natur., Paris, ser. 2, 7: 201-208. Grandjean F. 1953. Essai de classification des Oribates (Acariens). – Bull. Soc. Zool. de France, 78(5-6): 421-446. Grandjean F. 1969. Considérations sur le classement des Oribates. Leur division en 6 groupes majeurs. – Acarologia, 11: 127-153. Gregson J.D. 1956. The Ixodoidea of Canada. – Canada Dept. of Agriculture, Ottawa, Ont., Publ. 930: 1-92. Hahn C.W. 1835. Arachniden, v. 2, fa. 5. Hallas Th. E. 1978. Fortegnelse over danske mider (Acari)[Check list of Danish mites]. – Ent. Meddr., 46: 27-45. Halliday B. 1998. Mites of Australia. A Checklist and Bibliography. – Monographs on Invertebrate Taxonomy, 5: 317 pp. Hammen L. van der. 1961. Description of Holothyrus grandjeani nov. sp., and notes on the classification of the mites. – Nova Guinea, Zool., 9: 173-194. Hammen L. van der. 1966. Studies on Opilioacarida (Arachnids) I. Description of Opilioacarus texanus (Chamberlin et Mulaik) and revised classification of the genera. – Zool. Verh., 86: 1-80. Hammen L. van der. 1968. Introduction générale, la classification, la terminologie morphologique, l’ontogenèse et l’évolution des Acariens. – Acarologia, 10: 401-412.

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Hammen L. van der. 1971. Classification and phylogeny of mites. – Proc. 3rd Int. Congr. Acarology, Prague: 275-282. Hammen L. van der. 1972. A revised classification of the mites (Arachnidea, Acarida) with diagnoses, a key, and notes on phylogeny. – Zool. Meded., 47(22): 273-292. Hammen L. van der. 1977. A new classification of Chelicerata. – Zool. Meded. Leiden, 51: 307-319. Hammen L. van der. 1989 . An introduction to comparative arachnology. – SPB Academie Publishing b.v., The Hague, I-X, 1-576. Hoffmann A. 1979. Razones por las cuales deben elevarse los ácaros a la categoría de Clase Acarida. – Folia Ent. Mexicana, 42: 49. Hoogstraal H. 1953. Ticks (Ixodoidea) of the Malagasy Faunal Region (excepting the Seychelles). – Bull. Mus. Comp. Zoology at Harvard College, 111(2): 37-113. Hoogstraal H. 1956. African Ixodoidea. Ticks of the Sudan. – Bur. Med. Surg., U.S. Navy 1: 1101 p. Hoogstraal H. (Ed). 1989. Bibliography of ticks and tickborne diseases from Homer (about 800 BC to December 1984). – Special publications, NAMRU – 3, Cairo, 8 vols. Hoogstraal H., M.N. Kaiser. 1958. The ticks (Ixodoidea) of Egypt. A brief review and keys. – The J. of the Egyptian Public Health Association, 33(3): 51-85. Horak I.G., J.-L. Camicas, J. E. Keirans. 2002. The Argasidae, Ixodidae and Nuttalliellidae (Acari: Ixodida): a Worls List of valid tick names. – Experim. and Applied Acarology, 28(1-4): 27-54. Hughes T.E. 1959. Mites, or the Acari. – Univ. of London, Athlone Press, 225 pp. Hughes R.D., C.G. Jackson. 1958. A review of the Anoetidae (Acari). – Virginia J. Sci., 9, N.S.(1): 5-198 p. Jeppson L.R., H.H. Keifer, E.W. Baker. 1975. Mites Injurious to Economic Plants. – Univ. Calif. Press, Berkeley: 614 pp. Jin Dao-chao. 2000. Cladistics and phylogeny of superfamily relationships within Hydrachnellae (Acari, Acariformes). – Acta Entomologica Sinica, 43(3): 309-317. Johnston D.E. 1968. An Atlas of Acari I. The Families of Parasitiformes and Opilioacariformes. – Acarology Lab. Publ. 172, Acarology Publ., Columbus: 110 pp. Johnston D.E. 1982a. Oribatida. – In: S.P. Parker (Ed.) Synopsis and Classification of Living Organisms, McGraw – Hill, Inc., 2: 145-146. Johnston D.E. 1982b. Parasitiformes. – In: S.P. Parker (Ed.) Synopsis and Classification of Living Organisms, McGraw – Hill, Inc., 2: 111-117. Karafiat H. 1959. Systematik und Ökologie der Scutacariden. – Beiträge zur Systematik und Ökologie mitteleuropäischer Acarina,1 (4): 627-712. Karg W. 1971. Acari (Acarina), Milben. Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. – In: Die Tierwelt Deutschlands, 59. Teil, 475 pp. Karg W. 1989. Acari (Acari), Milben – Unterordnung Parasitiformes (Anactinochaeta). Uropodina Kramer, Schildkrotenmilben. – Tierwelt Dtschl., 67: 1-203.


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Kethley J.B. 1970. A revision of the family Syringophilidae (Prostigmata: Acarina). – Contrib. Amer. Ent. Inst., 5(6): 1-76. Kethley J.B. 1979. An Index to the Acarina Collections of the World (to the family level). – Special Publ., NAMRU 3, Cairo: 87 pp. Kethley J.B. 1982. Acarifomes. Prostigmata. – In: S.P. Parker (Ed.) Synopsis and Classification of Living Organisms, McGraw – Hill, Inc., 2: 117-145. Kethley J. 1990. Acarina: Prostigmata (Actinedida). – In: D.L. Dindal (Ed.): Soil Biology Guide, John Wiley & Sons, New York etc., Prostigmata: 667-756. Knülle W. 1959. Morphologische und entwicklungsgeschichtliche Untersuchungen zum phylogenetischen System der Acari: Acariformes Zachv. II. Acaridiae: Acaridae. Grundlagen der Getreidemilben – Systematik. – Mitt. Zool. Mus. Berlin, 35(2): 347-416. Koch C.L. 1836-1841. Deutschlands Crustaceen, Myriapoden und Arachniden. Koch C.L. 1842. Übersicht des Arachnidensystems. – Nürnberg. Kolebinova M. 1992. Fauna bulgarica. 21. Acariformes, Trombidioidea, Trombiculidae, Leeuwenhoekiidae. – Ed. Ac. Sci. bulg., Sofia, 172 pp. (in Bulgarian). Kramer P. 1877. Grundzüge zur Systematik der Milben. – Arch. F. Naturg., 2: 215-247. Krantz G.W. 1970. A Manual of Acarology, First Ed. – Oregon State Univ. Bookstores, Corvallis: 335 p. Krantz G.W. 1978. A Manual of Acarology. Second Edition. – Oregon State Univ. Book Stores, Inc., Corvallis, 509 p. Krantz G.W., B.D. Ainscough. 1990. Acarina: Mesostigmata (Gamasida). – In: D.L. Dindal (Ed.): Soil Biology Guide, John Wiley & Sons, New York etc., Gamasida: p. 583-665. Krczal H. 1959. Systematik und Ökologie der Pyemotiden. –Beitr. Syst. Ökol. Mitteleurop. Acarina, 1(3): 385-839. Latreille P.A. 1802. Histoire naturelle, générale et particulière, des Crustacés et des Insectes. – Paris, 3: 63-69. Latreille P.A. 1806. Genera crustaceorum et insectorum secundum ordinem naturalem in Familias disposita, Iconibus Exemplisque plurimus explicata. Parisiis et Argentorati – 1: 146. Lee D.C. 1970. The Rhodacaridae (Acari: Mesostigmata): classification, external morphology and distribution of genera. – Rec. S. Austral. Mus., 16(3): 1-219. Lehtinen P. T. 1995. Revision of the old world Holothyridae (Arachnida: Anactinotrichida: Holothyrina). – Invertebrate Taxonomy, 9(4): 767-826. Lindquist E.E. 1976. Transfer of the Tarsocheylidae to the Heterostigmata, and reassignement of the Tarsonemina and Heterostigmata to lower hierarchic status in the Prostigmata (Acari). – Can. Ent., 108: 23-48. Lindquist E.E. 1984. Current theories on the evolution of major groups of Acari and on their relationships with other groups of Arachnida, with consequent implications for their classification. – In: D.A. Griffiths and C.E. Brown (Eds), Acarology VI, 1. John Wiley, N.Y.: 28-62. Lindquist E.E. 1986. The world genera of Tarsonemidae (Acari. Heterostigmata): a morphological, phylogenetic, and systematic revision, with a reclassification of family-group taxa in the Heterostigmata. – Mem. Entomol. Soc. Canada, 136: 1-517.

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Lombardini G. 1938. Chiave analitica ai sottordini alle superfamiglie ed alle famiglie degli Acari. – Redia, Firenze, 24: 199-217. Lucas H. 1840. Histoire naturelle des Crustacés, des Arachnides et des Myriapodes. – Paris, 600 pp., P. Dumeril Ed. Mąkol J. 2000. Catalogue of the World Trombidiidae (Acari: Actinotrichida: Trombidioidea). – Annales Zoologici (Warszawa), 50(4): 599-625. Mąkol J. 2005. Trombidiidae (Acari: Actinotrichida: Trombidioidea) of Poland. – Fauna Poloniae Vol 1 New Series. Warszawa, 259 pp. Manilla G. 1998. Ixodida. – Fauna d’Italia, 36, Calderini, Bologna, 280 pp. Marshall V.G., R.M. Reeves, R.A. Norton. 1987. Catalogue of the Oribatida of continental United States and Canada. – Mem. Ent. Soc. Can., 1-418. Meyer M.K.P. et al. 1973. Acari of the Ethiopian Region. – Dept. of Agricult. Techn. Services, Entomology Memoir No 29: 1-45. Michael A.D. 1884. British Oribatidae, Vol.1. – Ray Society, London. Micherdzinski W. 1969. Die Familie Parasitidae Oudemans 1901 (Acarina, Mesostigmata). – Zak. Zool. Syst. Pol. Acad. Nauk, Krakow: 690 pp. Moraes G.J.de, J.A.McMurthy, H.A.Denmark & C.B.Campos. 2004. A revised catalog of the mite family Phytoseiidae. – Zootaxa, 434: 494 pp. Nitzsch C.L. 1818. Acarina, Milben, Mieten. – In: J.S. Ersch & J.G. Gruber, Allgemeine Encyclopädie der Wissenschaften, etc. I: 249-253. Norton R.A. 1990. Acarina: Oribatida – In: D.L. Dindal (Ed.): Soil Biology Guide, John Wiley & Sons, New York etc., Oribatida: p. 779-803. Norton R.A., J.B.Kethley, D.E.Johnston & B.M.O’Connor. 1993. Phylogenetic perspectives on genetic systems and reproductive modes of mites. p.8-99 In: D.L. Wrensch and M.A.Ebbert (Eds). Evolution and diversity of sex ratio in insects and mites. Chapman & Hall, New York. O’Connor B.M. 1979. A review of the family Heterocoptidae. – In: J.G. Rodriguez (ed.) Recent Advnces in Acarology, 2: 429-433. O’Connor B.M. 1982. Astigmata. – In: S.P. Parker (Ed.) Synopsis and Classification of Living Organisms, McGraw – Hill, Inc., 2: 146-169. O’Connor B.M. 1984. Phylogenetic relationships among higher taxa in the Acariformes, with particular reference to the Astigmata. pp.19-27 in D.A.Griffiths and C.E.Bowman.1984. Acarology VI, Vol.1. Ellis-Horwood Ltd., Chichester. Oudemans A.C. 1906. Nieuwe classificatie der Acari. – Ent. Ber., 2: 43-46. Oudemans A.C. 1923. Studie over de sedert 1877 ontworpen systemen der Acari; nieuwe classificatie; phylogenerische beschouwingen. – Tijdschr. Ent., 66: 49-85. Oudemans A.C. 1926. Kritisch Historisch Oversicht der Acarologie. 1 (850 V.C. – 1758), E. J. Brill: 500 pp. Oudemans A.C. 1929. Kritisch Historisch Oversicht der Acarologie. 2 (1759-1804), E. J. Brill: 1097 pp. Oudemans A.C. 1936-37. Kritisch Historisch Oversicht der Acarologie. 3 (1805-1850), E. J. Brill: 3379 pp.


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Yamaguti N., V.J. Tipton, H.L. Keegan, S. Toshioka. 1971. Ticks of Japan, Korea, and the Ryukyu Islands. – Brigham Young Univ. Science Bull., Biol. Series, 15: 1-226. Yunker C.E. 1955. A proposed classification of the Acaridiae (Acarina: Sarcoptiformes). – Proc. Helm. Soc. Wash., 22: 98-105. Yunker C.E. 1973. Mites. – In: Parasites of Laboratory Animals. R.J. Flynn, ed. Iowa State Univ. Press, Ames: 425-492. Zacharda M. 1980. Soil mites of the family Rhagidiidae (Actinedida: Eupodoidea), morphology, systematics, ecology. – Acta Univ. Carol. Biol. 1978: 489-785. Zachvatkin A.A. 1941. Fauna of U.S.S.R. Arachnoidea 6(1), Tyroglyphoidea (Acari). – Zool. Inst. Acad. Sci. U.S.S.R., N.S. 28 [translated by Amer. Inst. Biol. Sci.: 573 p.]. Zachvatkin A.A. 1952. [Subdivision of the mites (Acarina) into orders and the position of these in the system of the Chelicerata]. – Parazit. Sbornik, 14: 5-46.


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The Mite Subcohort Erythraeina Concepts concerning the place and composition of Erythraeina The comprehensive revision of Southcott (1961a) is the base of the modern concept of Erythraeina. With the first papers of the distinguished Australian acarologists R.V. Southcott and H. Womersley (Womersley & Southcott, 1941, Southcott, 1946), the relationships among the many confused generic and even family names were made clear and are widely accepted since. Welbourn (1991) is the author of a modern system of phylogenetic relationships of the terrestrial Parasitengona. Based on cladistic analysis, “three subcohorts are recognized in the Parasitengona: Erythraeina, Hydracarina and Trombidiina”. The system of Erythraeina follows Southcott (1961a), except of the subfamily Phanolophinae, which was moved to Erythraeidae, following Welbourn (1991).

The studies on Erythraeina Calyptostomatoidea. This superfamily consists of a single family and genus with very few species. The first (and, apparently the only one in West Europe) is Calyptostoma velutinus (Müller, 1776). Longtime the names Smaris and Smaridia were applied by authors, such as C.L. Koch (1837, 1842), L. Koch (1878), Trägårdh (1904a) and Schweizer (1922), for Calyptostomatid mites. Cambridge (1875) described the genus Calyptostoma and the authors after André (1933), with rare exceptions (Starkoff & Starkoff, 1950), use this name for the mites in the present family Calyptostomatidae Oudemans, 1923. The superfamily status of the Calyptostomatid mites was recognized as late as 1955 by Feider (under “Calyptostomidoidea”), and was confirmed and amended by Southcott (1961a). The detailed studies of Vistorin -Theis (1974-1977), completed by Wainstein (1976, 1980), presently provide a good idea about these strangely conservative mites in Europe. With regard to the representatives outside Europe, more research is to be done, as very few papers have been published about these mites in Africa (Lawrence, 1944), Asia (Berlese, 1905; Vitzthum, 1926; Shiba, 1969; Beron, in prep.) and Australia (Womersley, 1943; Southcott, 1961a). Erythraeoidea. In the XVIIIth century only several (now hardly usable) descriptions appear of mites belonging to Erythraeoidea by J.A.Scopoli, O.F.Müller, C. De Geer, F. de P. Schrank, P. Rossi. The first fundamental contributions to the systematic of Erythraeoidea are credited to P. Latreille in his publications of 1796-1806. There are described the genera Leptus, Erythraeus and Smaris. The name of the family Erythraeidae was given by Robineau-Desvoidy (1828). This name is antedated by Leptidae Billberg, 1820, but for

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various reasons Erythraeidae is preferred and widely used (see Southcott, 1961a). Von Heyden (1826) described the other large genus Balaustium. In the XIXth century, several authors described mites belonging to Erythraeoidea (C.L. Koch, L. Koch, Karsch, Pavesi, Lucas, Cambridge, Birula, Kramer, Stoll). Dugès, and especially Canestrini, tried to find a place for the Erythroid mites in the system of Acari. The important leap forward in the research on Erythraeoidea came with the papers of the Italian A. Berlese and the Dutch A.C. Oudemans, who not only described many new taxa, but created the modern system of Acari. Until 1900, only nine valid genera of Erythraeoidea were described (Smaris Latreille, Fessonia von Heyden, Eatoniana Cambridge, Erythraeus Latreille, Leptus Latreille, Caeculisoma Berlese, Abrolophus Berlese, Cecidopus Karsch, Balaustium von Heyden), and the system of the group remained very confused. From the beginning of the XXth century, up to W.W.II the number of described taxa increased rapidly. Prominent Acarologists like Banks, Thor, Trägårdh, Ewing, Willmann, Halbert, Hirst, André and others contributed to better understanding of these Acari. In Australia, H. Womersley and R.V. Southcott carried out important studies and started the work on creating the modern system of Erythraeoidea. In the same time, in Europe other Acarologists (Oudemans, Vitzthum, Grandjean) also worked on this system, something clearly needed. Partial new revision of Smarididae was published in the last paper of Oudemans (1941), however, according to Southcott (1961a), the quality of this paper is so much inferior to the papers of the famous Dutch Acarologist, that it would have been better to have remained unwritten. The many descriptions of new taxa are primitive and unusable. However, in 43 years (1900-1943) only 12 valid genera were added to the system of Erythraeoidea by Oudemans, Trägårdh, Hirst, Womersley and André). In 1941, in Australia, Womersley and Southcott published their serious and detailed revision of Smarididae, the actual basis of our knowledge on this family. The difficult communications during the war were the cause of a parallel revision of the same family by Grandjean. His very detailed revision was published only in 1947 and the names given by Grandjean became junior synonyms of the names, given by the Australian authors. The champion of the modern research on Erythraeoidea, however, was R. V. Southcott (1919-1998). Since his first paper in 1941, this most prominent specialist on Erythraeoidea published several important revisions and laid the foundations of the modern system of this superfamily. The most used is the revision of the genera of Erythraeoidea (1961), forming a base for all subsequent workers. In the last 40 years, besides Southcott, important data have published Schweizer (Switzerland), Shiba (Japan), Meyer (South Africa), Smiley (North America), Hoffmann (Mexico), Khot (India), Haitlinger and Gabryś (Poland), etc. Large areas remain unexplored. In 1972, Vercammen-Grandjean described the fossil species Proterythraeus southcotti, representing new genus and new (fossil) family Proterythraeidae. The genus was accepted by the further researchers, although the family was not. Now Proterythraeus is a member of Erythraeidae.


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Since 1980, a number of authors (Ahsan & Anwarullah, Beron, Dhiman & Sharma, Fain, Gabryś, Goldarazena, Gupta, Haitlinger, Haliday, Iravanlou, Mąkol, Rack, Ramaraju & Mohanasundaram, Rawat, Ripka, Saboori, Southcott until his death in April 1998, Treat, Tsai & Chow, Ueckermann, Vishnupuriya, Welbourn, Zhang, Zheng and others) have published papers on Erythraeoidea. There are almost no contributions on their biology, zoogeography, economic importance and role in the ecosystems. One of the most important collections of these mites (still under study) is housed in the Natural Museum of Natural History in Sofia (Collection Beron). Presently, in the family Erythraeidae are accepted by the present author 55 recent genera, 39 of which described after 1940. The number of species exceeds 750, although many of the older descriptions are to be redone.

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The composition of Erythraeina Order Acariformes Suborder Prostigmata Cohort Parasitengona Subcohort Erythraeina Superfamily Calyptostomatoidea Family Calyptostomatidae – 1 genus, 6 (?) species Superfamily Erythraeoidea Family Erythraeidae – 55 genera, 750 more or less recognizable species Family Smarididae – 12 genera, 52 species Genera within Erythraeoidea Smarididae Calorema Southcott, 1963-1 sp. Clavismaris Southcott, 1963-2 Clipeosoma Southcott, 1948-3 Fessonia von Heyden, 1826-10 Hirstiosoma Womersley, 1934-7 Kraussiana Southcott, 1961-2

Neophanolophus Shiba, 1976-4 Pilosoma Southcott, 1961-1 Smaris Latreille, 1796-12 Sphaerotarsus Womersley, 1936-6 Surasmaris Southcott, 1995-1 Trichosmaris Southcott, 1963-3

Erythraeidae Abalakeus Southcott, 1994-3 sp. Abrolophus Berlese, 1891-89 Andrevella Southcott, 1961-2 Augustsonella Southcott, 1961-2 Balaustium von Heyden, 1826-39 Bursaustium Haitlinger, 2000-2 Caeculisoma Berlese, 1888-21 Callidosoma Womersley, 1936-24 Carastrum Southcott, 1988-2 Cecidopus Karsch, 1879-7 Charletonia Oudemans, 1910-98 Claverythraeus Trägårdh, 1937-1 Curteria Southcott, 1961-5 (= Pedroerythraeus Haitlinger, 2004) Dambullaeus Haitlinger, 2001-1 Eatoniana Cambridge, 1898-3 Erythraeus Latreille, 1806-98 Erythraxus Southcott, 1961-1 Erythrellus Southcott, 1946-1

Erythrites Southcott, 1961-5 Erythroides Southcott, 1946-5 Forania Southcott, 1961-1 Fozustium Haitlinger, 2005-1 Grandjeanella Southcott, 1961-9 Guatustium Haitlinger, 2000-1 Harpagella Southcott, 1996-1 Hauptmannia Oudemans, 1910-19 Helladerythraeus Beron, 1988-1 Italustium Haitlinger, 2000-1 Kakamasia Lawrence, 1944-2 Kamertonia Gabryś, 1999-1 Lasioerythraeus Welbourn et Young, 1987-4 Leptus Latreille, 1796-269 Lomeustium Haitlinger, 2007-1 Microsmarialla Khot, 1963-1 Microsmaris Hirst, 1926-6 Moldoustium Haitlinger, 2007-1

The composition of Erythraeina

Momorangia Southcott, 1972-4 Mypongia Southcott, 1961-1 Myrmycotrombium Womersley, 1934-2 Neoabrolophus Khot, 1965-4 Neobalaustium Willmann, 1951-2 Neosmaris Hirst, 1926-1 Opserythraeus Fain, 1996-1 Palenqustium Haitlinger, 2000-1 Paraphanolophus Smiley, 1968-2 Phanolophus André, 1927-1 Podosmaridia Trägårdh, 1937-1

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Pollux Southcott, 1961-3 [Proterythraeus Vercammen-Grandjean, 1972-1 – fossil] Pussardia Southcott, 1961-1 Rainbowia Southcott, 1961-2 Ramsayella Zhang, 2000-1 Rudaemannia Haitlinger, 2000-1 Taranakia Southcott, 1988-1 Tepotztlana Hoffmann et Mendez, 1973-1 Wartookia Southcott, 1961-1

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Catalogue of the Mite Superfamily Calyptostomatoidea Order Acariformes Suborder Prostigmata (Actinedida) Cohort Parasitengona Subcohort Erythraeina Superfamily Calyptostomatoidea Southcott Calyptostomidoidea Feider, 1955: 17 Calyptostomatoidea Southcott, 1961a: 404 Calyptostomata Wainstein, 1965: 77 (Cohort) Calyptostomoidea Wainstein, 1978: 242 Calyptostomaoidea Robaux, 1973: 121

Family Calyptostomatidae Oudemans Calyptostomidae Oudemans, 1923: 77 Calyptostomatidae: Southcott, 1961a: 402

Type genus: Calyptostoma Cambridge, 1875

Genus Calyptostoma Cambridge (ad., larv.) Calyptostoma Cambridge, 1875: 384

Type species: Calyptostoma hardii Cambridge, 1875

Calyptostoma brevirostris Wainstein (ad) Calyptostoma brevirostris Wainstein, 1976: 158

Type locality: Borok, Yaroslavskaja oblast Distribution: Russia

Calyptostoma caelatum (Berlese)(ad.) Smaris caelata Berlese, 1905: 156 Calyptostoma caelatum Vitzthum, 1926: 177; 1931b: 84

Type locality: Bogor, Tijbodas, Java Distribution: Java, Sumatra (Indonesia)

Calyptostoma extranea (L. Koch)(ad.) Smaridia extranea: L. Koch, 1867: 70; Rainbow, 1960; Oudemans, 1941: 180 Calyptostoma extranea (L. Koch): Halliday, 1998: 44

Type locality: Brisbane Distribution: Australia

Superfamily Calyptostomatoidea

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Calyptostoma longirostris Wainstein (ad.) Calyptostoma longirostris Wainstein, 1976: 161

Type locality: Borok, Yaroslavskaja oblast Distribution: Russia

Calyptostoma velutinum (Müller) (ad., larval) Acarus velutinus Müller, 1776: 187 Trombidium expalpe Hermann, 1804: 40 Acarus expalpis Latreille, 1817: 120; 1829: 286; Dugès, 1836: 303 Smaridia expalpis: Dugès, 1834: 16; Lamarck, 1838: 287 Smaris impressa C.L.Koch, 1837: Fasc. XV, fig.1; 1842:38; Haller, 1882: 314; Canestrini, 1885: 142 Smaris expalpis (Hermann): C.L. Koch, 1842: 37; Thor, 1900: 3; Berlese, 1887: Fasc. XXXIX, 2; Trägårdh, 1904: 54; Hull, 1918: 23; Jablonowski, 1918: 3; Schweizer, 1922: 83 Smaris plana L. Koch, 1878: 127 Calyptostoma expalpe (Hermann): Sellnick, 1931: 696; André, 1933: 486; Irk, 1939: 165; Dudich et al., 1940: 55; Willmann, 1935: 29, 1950: 191, 1951b: 151; 1951c: 154; 1954: 245; Schweizer, 1951:166; Mihelčič, 1958a:108; 1971:144; Schweizer & Bader, 1963: 315; Schmölzer, 1993: 61 Fessonia prominens Banks, 1916b: 225 Calyptostoma prominens (Banks): Womersley, 1936: 118 Calyptostoma velutinum (Müller): Willmann, 1956: 248; Gabryś & Mąkol, 1996: 487 Calyptostoma velutinus (Müller): Oudemans, 1929: 596; Rack, 1976: 75; Shiba, 1969: 106; VistorinTheis, 1977a: 55; Gabryś & Haitlinger, 1986: 473; Gabryś, 1988: 22; Gabryś & Mąkol, 1991: 708; 1994: 126; Beron, 1995: 6; Mąkol & Gabryś, 1999: 29; Haitlinger, 2003e: 9 Smaris lyncaea Berlese, 1887: f. 39,6; Starkoff & Starkoff, 1950: 96 Calyptostoma lyncaeum: Willmann, 1939a: 17; Haarlov, 1957: 26; 1960: 162; Mihelčič, 1958a: 108; 1958d: 108; 1971: 144; Schweizer & Bader, 1963: 315; Hallas, 1978: 30 Calyptostoma latiseta Shiba, 1976: 195 Calyptostoma simplexa Shiba, 1976: 198

Type locality of Calyptostoma latiseta Shiba, 1976: Malaya Type locality of Calyptostoma simplexa Shiba, 1976: Malaya Distribution: Albania, Austria, Belgium, Bulgaria, Czech Rep., Denmark, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Japan, Malaya, Norway, Poland, Portugal (Madeira), Russia (Siberia), Sweden, Switzerland Calyptostoma sp. – Georgia (Rekk, 1976), Java (Vitzthum, 1931b) Species inquirendae: Calyptostoma hardyi Cambridge, 1875 (described as “hardii”) – England, syn. of Calyptostoma velutinus?

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Smaris lyncaea Berlese, 1887 (Calyptostoma lyncaeum, C. lincaeum in Wainstein, 1976) – Austria, Denmark, Germany, Hungary, Italy, Switzerland – syn. of Calyptostoma velutinus? Smaris neoguineana G. Canestrini, 1897 (= Calyptostoma neoguineanum, fide Vitzthum, 1931b) – New Guinea Calyptostoma leegei Oudemans, 1903 – Holland Smaris exculpta Berlese, 1917 – East Africa Calyptostoma stellatum Willmann, 1953 – Austria Calyptostoma tracheatum Lawrence, 1944 – South Africa


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Distribution of Calyptostomatoidea EUROPE Albania – Calyptostoma velutinum Austria (Franz, 1943, 1954; Irk, 1939; Mihelčič, 1958d, 1971; Vistorin – Theis, 1976, 1977a; Willmann, 1951b, 1951c, 1953) – Calyptostoma velutinum Belgium (Cooreman, 1943, 1945; Fain & Baugnée, 1996; Fain, Baugnée & Hidvegi, 1992; Willmann, 1935) – Calyptostoma velutinum Bulgaria (Beron, 1995) – Calyptostoma velutinum Croatia (Vistorin – Theis, 1976) – Calyptostoma velutinum Czech Rep. (Haitlinger, 2003f; Willmann, 1954) – Calyptostoma velutinum Denmark (Haarløv, 1957, 1960; Hallas, 1978; Müller, 1776) – Calyptostoma velutinum (= lyncaeum) France (contin.)(André, 1929, 1933, 1953; Cooreman, 1955b; Fain & D’Amico, 1997; Hermann, 1804; Latreille, 1796; Schmölzer, 1956; Southcott, 1961a; Theodoridès, 1955) – Calyptostoma velutinum Germany (Franke, 1940; Haitlinger, 1995, 1996, 1997; Haller, 1882; C.L. Koch, 1842; Münchberg, 1935, 1936, 1956; Oudemans, 1903; Rack, 1976; Southcott, 1961a; Vistorin – Theis, 1975, 1976; Weidner, 1954; Willmann, 1955) – Calyptostoma velutinum Great Britain (Cambridge, 1875; Evans, 1952; Evans & Browning, 1953; Hull, 1918; Killington & Bathe, 1946; Rack, 1974; Smith, 1951; Turk, 1945a, 1945b, 1953) – Calyptostoma velutinum (= C. hardyi) Greece (Berlese, 1916; Cooreman, 1955a; Haitlinger, 1984, 1993; Sellnick, 1931) – Calyptostoma velutinum Holland (Oudemans, 1903) – Calyptostoma velutinum (= C. leegei) Hungary (Dudich, Kolosvary & Szalay, 1940; Gabryś & Mąkol, 1991, 1996; Jablonowski, 1918) – Calyptostoma velutinum Italy (contin.)(Berlese, 1885, 1887, 1905,1910, 1918, ; Canestrini G., 1884, 1885; Starkoff O. & I. Starkoff, 1950; Cooreman, 1955c; Haitlinger, 1996, 1998; Lombardini, 1958; Oudemans, 1901) – Calyptostoma velutinum Faroe Is. (Trägårdh, 1931) – Calyptostoma velutinum Norway (Thor, 1897, 1900) – Calyptostoma velutinum (= plana = expalpis) Poland (Gabryś, 1988; Gabryś & Haitlinger, 1986; Gabryś & Mąkol, 1995; Haitlinger (1991), Mąkol & Gabryś, 1999; Oudemans, 1912; Southcott, 1992; Willmann, 1933, 1939, 1949, 1950, 1956) – Calyptostoma velutinum Portugal (cont.) (Vistorin – Theis, 1976) – Calyptostoma velutinum Madeira (Southcott, 1992; Vistorin -Theis, 1976; Willmann, 1939a) – Calyptostoma velutinum (=lyncaeum) Romania (Cooreman, 1951; Feider, 1948, 1954, 1955, 1956, 1967; Szalay, 1931; Vistorin -Theis, 1976) – Calyptostoma velutinum Russia (all, continent.)(Koch L., 1879; Wainstein, 1976) – Calyptostoma brevirostris, C.

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longirostris; C. velutinum Slovenia (Vistorin – Theis, 1976; Haitlinger, 2003d) – Calyptostoma velutinum Sweden (Danell, 1987; Rack, 1977; Southcott, 1992; Willmann, 1943) – Calyptostoma velutinum Switzerland (Haller, 1885; Schweizer, 1922, 1951, 195 ; Schweizer & Bader, 1963; Vistorin – Theis, 1976) – Calyptostoma velutinum AFRICA East Africa (Berlese, 1916) – Inc. sedis: “Smaris exculptum” South Africa (Lawrence, 1944) – Calyptostoma tracheatum ASIA Indonesia (Berlese, 1905, 1910; Haitlinger, 1990, 1998; Vitzthum, 1926) – Calyptostoma caelatum Japan (Shiba, 1969) – Calyptostoma velutinum Nepal (Beron, in prep.) – Calyptostoma sp. New Guinea (G. Canestrini, 1897) – Inc. sedis: “Smaris neoguineana” OCEANIA Australia (L. Koch, 1867; Southcott, 1961a; Womersley, 1943) – Calyptostoma extranea


Index of Calyptostomatoidea The names, accepted in the present Catalogue, are in bold brevirostris, Calyptostoma 36 caelata, Smaris 36 caelatum, Calyptostoma 36 Calyptostoma 36 Calyptostomaoidea 36 Calyptostomatidae 36 Calyptostomatoidea 36 Calyptostomidae 36 Calyptostomidoidea 36 Calyptostomoidea 36 exculpta, Smaris 38 expalpe, Trombidium 37 expalpis, Smaridia 37 expalpis, Smaris 37 expalpe, Calyptostoma 37 expalpe, Trombidium 37 extranea, Calyptostoma 36 extranea, Smaridia 36 hardii, Calyptostoma 37

hardyi, Calyptostoma 37 impressa, Smaris 37 latiseta, Calyptostoma 37 leegei, Calyptostoma 38 longirostris, Calyptostoma 37 lincaeum, Calyptostoma 38 lyncaea, Smaris 38 lyncaeum, Calyptostoma 38 neoguineana, Smaris 38 neoguineanum, Calyptostoma 38 plana, Smaris 37 prominens, Fessonia 37 simplexa, Calyptostoma 37 stellatum, Calyptostoma 38 tracheatum, Calyptostoma 38 velutinus, Acarus 37 velutinum, Calyptostoma 37 velutinus, Calyptostoma 37

41

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Shiba M. 1969. Taxonomic investigations on free living mites in the subalpine forest on Shiga Heights, IBP area. II. Prostigmata. – Bull. Nat. Sci. Mus. Tokyo, 12(1): 65-115. Shiba M. 1976. Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. – Nature and Life in South-east Asia, 7: 83-229. Smith K.G.V. 1951. Acarine larvae on Limonia tripunctata F. (Dipt., Tipulidae) and on a species of Chironomidae (Dipt.). – Ent. mon. mag., 87: 63. Southcott R.V. 1961a. Studies on the systematics and biology of the Erythraeoidea (Acarina), with a critical revision of the genera and subfamilies. – Aust. J. Zool., 9(3): 367-610. Starkoff O., I. Starkoff. 1950. Contributo alla conoscenza dell’acarofauna di Roma e dintorni IIa nota: sottordini Sarcoptiformes e Prostigmata. – Rivista di Parassitologia, 11(2): 85-101. Szalay L. 1931. Beiträge zur Kenntnis der Afterskorpion und Milbenfauna des Retyezat – Gebirges. – Ann. Mus. Nat. Hung., 17: 24 Theis G., R. Schuster. 1974. Gestielte Tropfchenspermatophoren bei Calyptostomiden (Acari, Trombidiformes) – Mitt. naturwiss. Ver. Steiermark, 104: 183-185. Theodoridès J. 1955. Contribution à l’étude des parasites et phorétiques de Coléoptères terrestres. – Actualités scient. et industrielles, Paris (suppl. 4 “Vie et Milieu”): 155-257. Thor S. 1897. Andet bidrag til kundeskaben om Norges hydrachnider. – Archiv f. M.&N., 20: 3. Thor S. 1900. Første undersøgelse af Norges Rhyncholophidae. – Forh. Vidensk Selsk. Krist, 1900(2): 1-18. Trägårdh I. 1904. Monographie der arktischen Acariden. – Fauna Arctica, 4(1): 1-77. Trägårdh I. 1931a. Acarina from Juan Fernandez Islands. – Nat. Hist. Juan Fernandez and Easter Islands, 3, 4, Uppsala: 553-628. Trägårdh I. 1931b. Terrestrial Acarina. – Zoology of the Faroes, Kopenhagen, 49: 23-69. Turk F. A. 1945a. Studies of Acari. Second series: Descriptions of new species and notes on established forms of parasitic mites. – Parasitology, 36(3-4): Turk F.A. 1945b. Studies of Acari, V. – Notes on and descriptions of new and little-known British Acari. – Ann. Mag. Nat. Hist. (11)12: 795-820. Turk F.A. 1953. A Synonymic Catalogue of British Acari: Part I. – Ann. Mag. Nat. Hist., 12th Ser., 61: 1-26, 81-99. Vistorin-Theis G. 1975. Entwicklungszyklus der Calyptostomiden (Acari, Trombidiformes). – Acarologia, 17(4): 683-692. Vistorin-Theis G. 1976. Morphologische Analyse der Calyptostomiden-Larve (Acari, Trombidiformes). – Sitzungsber. Österr. Akad. Wiss., Mathem.-naturwiss. Kl, Abt. I, 185 (8-10): 283-294. Vistorin-Theis G. 1977a. Morphologisch-taxonomische Studien an der Milbenfamilie Calyptostomidae (Acari, Trombidiformes). – Sb. Österr. Akad. Wiss., Math.-Nat. Kl, Abt. I, 185: 55-89. Vistorin-Theis G. 1977b. Ernährungsbiologische Beobachtungen an Calyptostomiden (Acari, Trombidiformes). – Zool. Anz., Jena, 199(5-6): 381-385. Vitzthum H.G. 1926. Malayische Acari. – Treubia, 8: 1-198. Vitzthum H. 1929. Ordnung Milben, Acari. – In: Die Tierwelt Mitteleuropas, 3(7): 1-112. Vitzthum H.G. 1931a. Ordnung der Arachnida: Acari – Milben. – In: W. Kükenthal, Handbuch der Zoologie, 3(2), 1: 1-160.


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Vitzthum H.G. 1931b. Terrestrische Acarinen (unter Ausschluss der Oribatiden und Ixodiden) der Deutschen Limnologischen Sunda-Expedition. – Arch. Hydrobiol. (Plankt.)(Suppl.), 9(1): 59-134. Wainstein B. A. 1965. [About the system of water mites and their place in the suborder Trombidiformes]. – Trudy Inst. Biol. vnutrennih vod, 8(11): 66-83 (in Russian). Wainstein B. A. 1976. The genus Calyptostoma (Acariformes, Calyptostomatidae) – Zool. J., 56: 158-159. Wainstein B. A. 1978. [System, Evolution and Phylogeny of Trombidiiform Mites], p. 228-245. – In: M.S. Gilyarov (Ed.) Key for identification of Soil-Inhabiting Mites. Trombidiformes. – Ed. “Nauka”, Moskow, 271 p. (in Russian). Wainstein B. A. 1979. Ontogenesis and the systematic position of mites of the family Calyptostomidae (Acariformes). – Revue d’Entomol. de I’URSS, 58(1): 211-216. Weidner H. 1954. Die Pseudoskorpione, Weberknechte und Milben der Umgebung von Hamburg mit besonderer Berücksichtigung der für den Menschen wichtigen Arten. – Ent. Mitt. Zool. St. Zool. Mus. Hamburg, 4: 54 p. Willmann C. 1933. Acari aus dem Moosebruch. – Zeitschr. Morph. Ökol. Tiere, 27: 373-383. Willmann C. 1935. Exploration biologique des cavernes de la Belgique...XXVe contribution, Acari. – Bull. Mus. Roy. Hist. Nat. Belgique, 11(29): 1-41. Willmann C. 1939a. Die Arthropodenfauna von Madeira nach den Ergebnissen der Reise von Prof. Dr. O. Lundblad Juli-August 1935. XIV. Terrestrische Acari (exkl. Ixodidae). – Ark. Zool., 31A(10): 1-42. Willmann C. 1943. Terrestrische Milben aus Swedisch Lappland. – Arch. Hydrobiol., 40(1): 208-239. Willmann C. 1950. Milben aus Mineralquellen (2 Mitteilung). – Zool. Anz., 145(7-8): 186-195. Willmann C. 1951a. Die hochalpine Milbenfauna der mittleren Hohen Tauern insbesondere des Grossglockner-Gebietes (Acari). – Bonner Zool. Beitr, 2(1-2): 141-176. Willmann C. 1951b. Untersuchungen über die terrestrische Milbenfauna im pannonischen Klimagebiet Österreichs. – Sitzungsber. Österr. Akad. Wiss., Math. – naturwiss. Kl, Abt.l, 160 (1-2): 91-176. Willmann C. 1953. Neue Milben aus den ostlichen Alpen. – Sizber. Österr. Akad. Wiss. Wien, math.-naturwiss. Kl., Abt.I, 162: 449-519. Willmann C. 1954. Mährische Acari hauptsachlich aus dem Gebiete des Mährischen Karstes. – Česk. paras., 1: 213-272. Willmann C. 1955. Milben aus dem Sudwestlichen Sachsen. – Abh. Mus. Tierk. Dresden, 22(2): 207-225. Willmann C. 1956. Milben aus dem Naturschtzgebiet auf dem Spieglitzer (Glatzer) Schneeberg. – Českoslov. Parasitol., 3: 211-273. Womersley H. 1936. Additions to the Trombidiid and Erythraeid Acarine Fauna of Australia and New Zealand. – Linn. Soc. J. – Zoology, 40 (269): 107-121. Womersley H. 1942. Additions to the Acarina of Australia (Trombidiidae and Calyptostomidae). – Rec. South Australian Mus., 7: 169-181.

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Catalogue of the Mite Superfamily Erythraeoidea Superfamily Erythraeoidea Grandjean Apobolostigmata Oudemans, 1909: 59 (in part); 1923 (in part); Vitzthum, 1929: 60; 1931: 148 Erythraeoidea Grandjean, 1947a: 2; Southcott, 1961a: 402

Family Smarididae Vitzthum Smarididae Vitzthum, 1929: 69; Southcott, 1961a: 413 Smarisidae Grandjean, 1943: 242; 1947a: 1 Smaridiidae Baker & Wharton, 1952: 241; Mayer & Ryke, 1959: 305

Type genus: Smaris Latreille, 1796

Subfamily Smaridinae Southcott Smaridinae Southcott, 1946: 143 Fessoniinae Southcott, 1946: 143

Type genus: Smaris Latreille, 1796

Genus Calorema Southcott (ad.) Calorema Southcott, 1962: 185

Type species: Calorema azteka Southcott, 1962

Calorema azteka Southcott (ad.) Calorema azteka Southcott, 1962: 185

Type locality: Panama, Mexico Distribution: Panama, Mexico

Genus Fessonia von Heyden (ad.) Fessonia von Heyden, 1826: 609 Veithia Oudemans, 1941: 182; revision: Zhang, 2003: 1 Oecosmaris Grandjean, 1943d: 242 (nomen nudum), 1947a: 16

Type species of Fessonia von Heyden: Trombidium papillosum Hermann, 1804 sensu Berlese (1884) Type species of Veithia Oudemans: Veithia schneideri Oudemans, 1941 Type species of Oecosmaris Grandjean: Oecosmaris callitricha Grandjean, 1947

Fessonia assmuthi (Oudemans)(ad.) Veithia assmuthi Oudemans, 1941: 182; Buitendijk, 1945: 347 Fessonia assmuthi (Oudemans): Zhang, 2003: 1

Type locality: Bombay (Chandala), on a bat (chance association) Distribution: India

Superfamily Erythraeoidea

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Fessonia australiensis Southcott (ad.) Fessonia australiensis Southcott, 1946: 176

Type locality: Mataranka, Northern Territory, Australia Distribution: Australia, Burma, China, India, Mexico

Fessonia lacrimosa Southcott (ad.) Fessonia lacrimosa Southcott, 1963: 208

Type locality: Tamazunchale, San Luis Potosi, Mexico Distribution: Mexico

Fessonia lappacea Southcott (ad.) Fessonia lappacea Southcott, 1963: 201

Type locality: California, South Carolina, Mexico Distribution: U.S.A., Mexico

Fessonia nigeriae Beron (ad.) Fessonia nigeriae Beron, in print

Type locality: Jos, Plateau State Distribution: Nigeria

Fessonia papillosa (Hermann) Berlese (ad.) Trombidium papillosum Hermann, 1804: 29; von Heyden, 1826: 609; Gervais, 1844: 174 Smaris papillosus: Latreille, 1817: 121; de Lamarck, 1818: 54 Smaridia papillosa (Hermann): Latreille, 1819: 351; 1829: 287; Dugès, 1834:12; Lucas, 1840: 491; C.L. Koch: 1842: 52 Erythraeus papilosus: Latreille, 1823: 168 Rhyncholophus papillosus (Herm.): C.L.Koch, 1837: f.16, f.17; 1841: 430; Haller, 1882: 314; Canestrini & Fanzago, 1877: 72; Canestrini, 1885: 152; Jablonowski, 1918: 3 Smaris papillosum (Herm., 1804): Oudemans, 1937: 954 Fessonia papillosa: von Heyden, 1826: 609; Southcott, 1946: 177

Type locality: France Distribution: Croatia, France, Germany, Greece (incl. Chios), Hungary, Italy (incl. Sicily and Sardinia)

Fessonia schneideri (Oudemans)(ad.) Veithia schneideri Oudemans, 1941: 182; Buitendijk, 1945: 347 Fessonia schneideri (Oudemans): Zhang, 2003: 1

Type locality: San Remo Distribution: Italy

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Fessonia scobina Southcott (ad.) Fessonia scobina Southcott, 1963: 206

Type locality: Guerrero, Mexico Distribution: Mexico

Fessonia serrata Southcott (ad.) Fessonia serrata Southcott, 1963: 196

Type locality: Guatemala Distribution: Guatemala

Fessonia taylori Southcott (ad.) Fessonia taylori Southcott, 1961c: 146

Type locality: New South Wales Distribution: Australia (New South Wales)

Genus Kraussiana Southcott (ad.) Fessonia Meyer & Ryke, 1959: 322 (in part) Kraussiana Southcott, 1961a: 442

Type species: Fessonia brevicristata Meyer et Ryke, 1959

Kraussiana brevicristata (Meyer et Ryke) (ad.) Fessonia brevicristata Meyer & Ryke, 1959: 322 Kraussiana brevicristata (Meyer et Ryke): Southcott, 1961a: 442

Type locality: Bathurst Distribution: South Africa

Kraussiana mitsukoae Shiba (ad.) Kraussiana mitsukoae Shiba, 1976: 189

Type locality: Pasoh Forest, Malaya Distribution: Malaysia

Kraussiana sp. Distribution: Aurapushekaru I., Palau group, Caroline Islands (Southcott,1961a: 442)

Genus Neophanolophus Shiba (larval) Neophanolophus Shiba, 1976: 191

Type species: Neophanolophus morii Shiba, 1976

Neophanolophus chinensis Zheng (larval) Neophanolophus chinensis Zheng, 2001: 491

Type locality: Hunan Distribution: China


51

Neophanolophus liui Zheng (larval) Neophanolophus liui Zheng, 2001: 493


Neophanolophus morii Shiba (larval) Neophanolophus morii Shiba, 1976: 193

Type locality: Pasoh Forest, Malaya Distribution: Malaysia Host: unknown (in litter)

Neophanolophus shibai Zhang (larval) Neophanolophus shibai Zhang, 1995: 373

Type locality: Simla, Himachal Pradesh Distribution: India (Himachal Pradesh) Host: Erythroneura sp. (Cicadellidae, Homoptera)

Genus Pilosoma Southcott (larval, may be larva and junior synonym of Fessonia, see Zhang, 1995: 373) Oecosmaris Grandjean, 1947a: 39; 1947b (larva only) Pilosoma Southcott, 1961a: 454; Zhang, 1995: 373

Type species: Pilosoma pluto Southcott, 1961 (pro Oecosmaris callitricha (larva only) Grandjean, 1947

Pilosoma pluto Southcott (larva, adult) Oecosmaris callitricha (larva only) Grandjean, 1947 Pilosoma pluto Southcott, 1961a: 454

Type locality: France Distribution: France

Genus Smaris Latreille (larval and ad.) Smaris Latreille, 1796: 180 Smaridia Latreille, 1817: 121 Sclerosmaris Grandjean, 1947a: 16 ?Samouellea Oudemans, 1941: 181 ?Leuchsia Oudemans, 1941: 182 ?Dugesia Berlese, 1883: A.M.S., 5,4 (full synonymy in Southcott, 1961a: 439)

Type species of Smaris Latreille: Trombidium squamatum Hermann, 1804 (Smaris squamata sensu Berlese, 1883) Type species of Sclerosmaris Grandjean: Sclerosmaris philopempta Grandjean, 1947

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Type species of Samouella Oudemans: Smaridia depilata Berlese, 1888 Type species of Leuchsia Oudemans: Leuchsia grandjeani Oudemans, 1941

Smaris arenicola Southcott (larval) Smaris arenicola Southcott, 1997: 2

Type locality: Loveday, South Australia Distribution: Australia (S. Australia)

Smaris bipartita Mihelčič (ad.) Smaris bipartita Mihelčič, 1958a: 284

Type locality: Casa de Campo, Spain Distribution: Spain

Smaris boneti Southcott (ad.) Smaris boneti Southcott, 1963: 182

Type locality: Nevado de Colima, Jalisco, Mexico Distribution: Mexico

Smaris cooperi Southcott (ad.) Smaris cooperi Southcott, 1961c

Type locality: Australia Distribution: Australia

Smaris grandjeani grandjeani (Oudemans)(ad.) Leuchsia grandjeani Oudemans, 1941: 182 Smaris grandjeani Southcott, 1961a: 434 Smaris grandjeani grandjeani (Oudemans): Southcott, 1963: 176; Zhang, 2001: 17

Type locality: Colon Distribution: Panama

Smaris grandjeani christensoni Southcott (ad.) Smaris grandjeani christensoni Southcott, 1963: 178

Type locality: California, Texas, New Mexico, Guatemala, Mexico, Distribution: Guatemala, Mexico, U.S.A.(California, Texas, New Mexico)

Smaris lanceolata Southcott (ad.) Smaris lanceolata Southcott, 1963: 170

Type locality: Panama, Mexico Distribution: Panama, Mexico


53

Smaris nicoleti (Gervais) (ad) Smaridia nicoletii Gervais, 1849: 36 Smaris nicoletii (Gervais): Oudemans, 1937: 953; 1941: 180

Type locality: Chile (San Cárlos de Chiloe) Distribution: Chile

Smaris philopempta (Grandjean) (ad.) Sclerosmaris philopempta Grandjean, 1947a: 53; Radford, 1950: 148; Baker & Wharton, 1952: 243, 460 Sclerosmaris philopempta: Southcott, 1961a: 440; Beron (in prep.)

Type locality: Gruissan (Aude, France) Distribution: Bulgaria, France

Smaris pinus Zhang (ad.) Smaris pinus Zhang, 1995a: 70

Type locality: Cyprus Distribution: Cyprus Host: in association with Matsucoccus josephi (Homoptera: Margarodidae)

Smaris prominens (Banks)(ad. and larv.) Fessonia prominens Banks, 1916: 225 Calyptostoma prominens (Banks): Womersley, 1934: 235 Smaris prominens (Banks): Womersley & Southcott, 1941: 63; Southcott, 1946: 177; 1997: 2

Type locality: Victoria Distribution: Australia (Victoria, New S. Wales, S. Australia) Host: ant’s nest; larvae on Psocoptera (Liposcelis = Troctes divinatorius, Liposcelididae)

Smaris squamata (Hermann)(ad.) Trombidium squamatum Hermann, 1804: 29 Smaridia squamata (Hermann): Latreille, 1823: 173; Dugès, 1834: 12; C.L.Koch, 1842: 52 Dugesia squamata (Herm.) Smaris squamatus: Berlese, 1883: A.M.S. 5,4 Rhyncholophus squamatus Herm.: Canestrini, 1885: 153 Smaris squamatum (Hermann): Southcott, 1946: 177 Smaris squamata (Hermann): Sellnick, 1931: 696; Feider, 1948: 209; Bottazzi, 1956: 44

Type locality: Strassburg, France Distribution: Bulgaria, France, Germany, Greece (incl. Creta and Rhodos), Italy (incl. Sardinia and Sicily), Romania, Spain

54


Smaris zeteki Southcott (ad.) Smaris zeteki Southcott, 1963: 165

Type species: Baro Colorado Is., Panama Distribution: Panama

(?) Smaris heteropoda (Berlese) (ad.) Smaridia heteropoda Berlese, 1910: 348

Type locality: “Capetown” Distribution: South Africa

Genus Surasmaris Southcott (ad.) Surasmaris Southcott, 1995: 57

Type species: Surasmaris longirostris Southcott, 1995

Surasmaris longirostris Southcott Surasmaris longirostris Southcott, 1995: 57

Type locality: Costa Rica Distribution: Costa Rica

Subfamily Hirstiosomatinae Southcott Hirstiosominae Southcott, 1946a: 173 Hirstiosomatinae Southcott, 1961a: 442 (nom. emend.)

Type genus: Hirstiosoma Womersley, 1934

Genus Clavismaris Southcott (ad.) Clavismaris Southcott, 1963: 232

Type species: Clavismaris conifera Southcott, 1963

Clavismaris conifera Southcott (ad.) Calvismaris conifera Southcott, 1963: 233

Type locality: Gurrero, Mexico Distribution: Mexico

Clavismaris cybaea Southcott (ad.) Clavismaris cybaea Southcott, 1963: 235

Type locality: Costa Rica, Texas, ? France Distribution: Costa Rica, Texas (U.S.A.), ? France


55

Genus Clipeosoma Southcott (larval) Clipeosoma Southcott, 1948: 254

Type species: Clipeosoma copiolarum Southcott, 1948

Clipeosoma copiolarum Southcott (larval) Clipeosoma copiolarum Southcott, 1948: 254

Type locality: Babiang in the Aitape region Distribution: New Guinea Host: “on the floor of the rain-forest”

Clipeosoma jupiter Southcott (larval) Smaris latreillei Grandjean, 1947a: 20 – larva Clipeosoma jupiter Southcott, 1961a: 445; Gabryś, 1989: 165; Gabrys & Mąkol, 1991: 709

Type locality: Perigueux (SW France) Distribution: France, Hungary, Poland

Clipeosoma furtadoi Shiba (larval) Clipeosoma furtadoi Shiba, 1976: 191

Type locality: Kuala Pilah, Negeri Sembilan Distribution: Malaysia Host: found free living

Genus Hirstiosoma Womersley (ad.) Hirstiosoma Womersley, 1934: 242 Niemannia Oudemans, 1941: 181 (full synonymy in Southcott, 1961a: 443)

Type species of Hirstiosoma Womersley: Hirstiosoma scalaris Womersley, 1934 (emend. to scalare by Southcott, 1961a: 443) Type species of Niemannia Oudemans: Smaridia ampulligera Berlese, 1887

Hirstiosoma ampulligera (Berlese) (ad.) Smaridia ampulligera Berlese, 1887: Fasc. XXXIX, 10; 1888a: 185; Thor, 1900: 5; Leonardi in Berlese & Leonardi, 1901: 16; Hull, 1918: 23 Niemannia ampulligera (Berlese): Oudemans, 1941: 181 Smaris ampulligera (Berlese): Starkoff & Starkoff, 1950: 96; Schweizer, 1951: 165; Schweizer & Bader, 1963: 314; Willmann, 1951c: 154; Turk, 1953: 26; Schmölzer, 1956: 561; 1993: 61; Bottazzi, 1956: 44 Hirstiosoma ampulligera (Berlese): Southcott, 1961a: 421

Type locality: Italy, Florence (“in agri Florentinii muscis”) Distribution: Austria, Bulgaria, France, Great Britain, Italy, Norway, Spain, Switzerland; ? Argentina, Brasil, Paraguay, Chile

56


Hirstiosoma bolivari Southcott (ad.) Hirstiosoma bolivari Southcott, 1963: 211

Type locality: Palmira, Morelos, Mexico Distribution: Mexico

Hirstiosoma glacialis (Schmölzer)(comb. nov.)(ad.) Smaris glacialisSchmölzer, 1956: 562

Type locality: Refuge Temple-Ecrins, 2740 m, Dauphiné Distribution: France

Hirstiosoma latreillei (Grandjean)(ad.) Smaris latreillei Grandjean, 1947a: 19 Hirstiosoma latreillei (Grandjean): Gabrys & Mąkol, 1991: 709; 1994: 131

Type locality: France Distribution: France, Hungary, Poland

Hirstiosoma novaehollandiae Womersley (ad.) Hirstiosoma novae – hollandiae Womersley, 1936: 118; Womersley & Southcott, 1941: 71

Type locality: Manurewa, Auckland Distribution: New Zealand

Hirstiosoma scalare Womersley (ad.) Hirstiosoma scalare Womersley, 1934: 242; Southcott, 1961a: 443 Hirstiosoma scalaris (Womersley): Womersley & Southcott, 1941: 69

Type locality: Victor Harbour (S. Australia) Distribution: Australia

Hirstiosoma tasmaniensis Womersley et Southcott (ad.) Hirstiosoma tasmaniensis Womersley & Southcott, 1941: 69

Type locality: Mount Welington, Hobart (Tasmania) Distribution: Tasmania

Genus Sphaerotarsus Womersley (larval and ad.) Sphaerotarsus Womersley, 1936: 119

Type species: Sphaerotarsus allmani Womersley, 1936

Sphaerotarsus allmani Womersley (ad.) Sphaerotarsus allmani Womersley, 1936: 119; Womersley & Southcott, 1941: 76

Type locality: Bathurst, New South Wales Distribution: Australia (New South Wales)


Sphaerotarsus claviger Womersley et Southcott (ad.) Sphaerotarsus claviger Womersley & Southcott, 1941: 76

Type locality: Bathurst (New South Wales) Distribution: Australia

Sphaerotarsus leptopilus Womersley et Southcott (ad.) Sphaerotarsus leptopilus Womersley & Southcott, 1941: 75

Type locality: Victor Harbour (S. Australia) Distribution: Australia (S. Australia)

Sphaerotarsus monticolus Southcott (larval) Sphaerotarsus monticolus Southcott, 1997: 6

Type locality: Mt. Lofty, S. Australia Distribution: Australia (South Australia)

Sphaerotarsus ripicolus (Womersley) (Ny.) Caeculisoma ripicola: Womersley, 1934: 239 (Ny.) Sphaerotarsus allmani: Womersley, 1936: 119 (part) Sphaerotarsus ripicolus Womersley & Southcott, 1941: 73

Type locality: Victor Harbour, S. Australia Distribution: Australia (S. Australia, Victoria)

Sphaerotarsus womersleyi Southcott (ad.) Caeculisoma ripicola: Womersley, 1934: 238 (ad.) Sphaerotarsus allmani: Womersley, 1936: 120 (part) Sphaerotarsus ripicolus Womersley & Southcott, 1941: 73 (nom. invalid.) Sphaerotarsus womersleyi Southcott, 1946: 177

Type locality: Victor Harbour (S.Australia) Distribution: Australia (S. Australia, Victoria)

Genus Trichosmaris Southcott (ad.) Trichosmaris Southcott, 1963: 215

Type species: Trichosmaris dispar Southcott, 1963

Trichosmaris dispar dispar Southcott (ad.) Trichosmaris dispar Southcott, 1963: 215

Type locality: Guatemala, Mexico, U.S.A. (many places) Distribution: Guatemala, Mexico, U.S.A.

57

58


Trichosmaris dispar dentella Southcott (ad.) Trichosmaris dispar dentella Southcott, 1963: 223

Type locality: San Jose de Guatemala, Guatemala Distribution: Guatemala

Trichosmaris jacoti (Southcott) (ad.) Smaris sericea Jacot, 1938: 123 Hirstiosoma sericea (Jacot): Womersley & Southcott, 1941: 63, 78 Hirstiosoma jacoti Southcott, 1946a: 177 Trichosmaris jacoti (Southcott): Southcott, 1963: 226

Type locality: Darien, Georgia, U.S.A. Distribution: U.S.A. (Georgia)

Trichosmaris papuana Beron (ad.) Trichosmaris papuana Beron, 2002: 73

Type locality: Mt. Fugilil, 3100 m, Western Province, Papua New Guinea Distribution: Papua New Guinea

Family Erythraeidae Robineau-Desvoidy Leptides: Billberg, 1820: 126 Erythreidae [sic] Robineau-Desvoidy, 1828: 144 (in part) Rhyncholophides: C.L. Koch, 1842: 49 (in part) Leptoidae: Agassiz, 1846: 206 (in part); 1848: 595 Rhyncholophoidae: Agassiz, 1846: 206 (in part) Rhyncholophidae: Kramer, 1877: 227 Rhyncholophini: Canestrini, 1885 (in part) Erythraeinae: Oudemans, 1902 (subfam. of Thrombidiidae [sic]): 60 Erythreidae Oudemans, 1909: 60 Rhyncholophinae (subfam. of Thrombidiidae [sic]) Hull, 1918: 22 Leptidae: Oudemans, 1941: 178; Feider, 1954: 1017; Feider & Suciu, 1959: 439 Balaustiidae: Grandjean, 1947a: 12

Type genus of Leptidae: Leptus Latreille, 1796 Type genus of Erythraeidae: Erythraeus Latreille, 1806 Type genus of Balaustiidae: Balaustium v. Heyden, 1826

Subfamily Erythraeinae Southcott Erythraeinae Southcott, 1957: 97

Type genus: Erythraeus Latreille, 1806


59

Genus Abalakeus Southcott (larval, Ny) Abalakeus Southcott, 1994b: 81

Type species: Abalakeus chekei Southcott, 1994

Abalakeus bambusae Zhang Zhi-Qiang, Zhang Yanxuan et Lin Jianzhen (larval) Abalakeus bambusae Zhang Zhi-Qiang, Zhang Yanxuan & Lin Jianzhen, 2000: 69

Type locality: Nanping County, Fujian, China Distribution: China Host: on Phyllostachys pubescens (bamboo, Poaceae)

Abalakeus chekei Southcott (larval) Abalakeus chekei Southcott, 1994b: 81

Type locality: Tagalak, SW of Abalak Distribution: Niger Host: locust (Hieroglyphus daganensis, Acrididae, Orthoptera)

Abalakeus lorestanicus Saboori et Lachinani (larval, Ny) Abalakeus lorestanicus Saboori & Lachinani, 2003: 155; Mayoral & Barranco, 2004: 217; 2006: 125

Type locality: Khorram Abad (Iran) Distribution: Iran, Spain Host: Steropleurus andalusius (Acrididae, Orthoptera)

Genus Augustsonella Southcott (ad.) Augustsonia Southcott, 1961a: 464 Augustsonella Southcott, 1961a: 505

Type species: Erythraeus tuberculatus Augustson, 1940

Augustsonella southcotti Hoffmann et Mendez (ad.) Augustsonella southcotti Hoffmann & Mendez, 1973: 125

Type locality: Tepoztlan, Morelos, Mexico Distribution: Mexico

Augustsonella tuberculata (Augustson) (ad.) Erythraeus tuberculatus Augustson, 1940: 192 Augustsonella tuberculata (Augustson): Southcott, 1961a: 507; Hoffmann & Mendez, 1973: 125

Type locality: El Segundo, California Distribution: U.S.A. (California)

Genus Claverythraeus Trägårdh (ad.) Claverythraeus Trägårdh, 1937: 10

Type species: Claverythraeus mongolicus Trägårdh, 1937

60


Claverythraeus mongolicus Trägårdh (ad.) Claverythraeus mongolicus Trägårdh, 1937: 10

Type locality: Urumchi (China) Distribution: China (Sinziang Uygur Ziziqi)

Genus Curteria Southcott (ad.) Morieria sensu Willmann, 1951b: 150 Curteria Southcott, 1961a: 488 Pedroerythraeus Haitlinger, 2004g: 220 (synonymized by Wohltmann et al., 2007: 573)

Type species: Morieria (?) curticristata Willmann, 1951 Type species of Pedroerythraeus Haitlinger, 2004: P. ernesti Haitlinger, 2004

Curteria curticristata (Willmann) (ad.) Morieria (?) curticristata Willmann, 1951b: 165 Morieria curticristata (Willmann): Schweizer, 1951: 133; Cooreman, 1955a: 126; Schweizer & Bader, 1963: 311 Curteria curticristata: Southcott, 1961a: 489; Gabryś, 1992: 257

Type locality: “Glockner-Gruppe”, 2000-2300 m, Austria Distribution: Austria, Switzerland, ? Greece

Curteria episcopalis (C.L. Koch) (ad.) Rhyncholophus episcopalis C.L.Koch, 1837: fa.16, t.9; 1841: 414; 1842: 50 Erythraeus flavus Oudemans, 1903c: 92 Balaustium episcopalis (Koch): Oudemans, 1937: 1939 Erythraeus episcopale (Koch): Buitendijk, 1945: 349 Curteria episcopalis (C.L.Koch): Gabryś, 1992: 256; Gabryś & Mąkol, 1994: 127; Wohltmann, Mąkol & Gabryś, 2007: 574

Type locality of Rhyncholophus episcopalis: “Neumarkt in der Oberpfalz” Type locality of Erythraeus flavus: “Eiland Borkum”, Holland Distribution: Germany, Holland, Poland

Curteria ernesti (Haitlinger) (larval) Pedroerythraeus ernesti Haitlinger, 2004g: 220 Curteria ernesti (Haitlinger): Wohltmann, Mąkol & Gabryś, 2007: 577

Type locality: Don Pedro, La Palma Distribution: Canary Is. (La Palma) Host: unknown


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Curteria fageli (Cooreman) (ad.) Morieria fageli Cooreman, 1956: 4 Curteria fageli (Cooreman): Gabryś, 1992: 257

Type locality: El Kantara and Biskra, Algeria Distribution: Algeria

Curteria graeca Beron (ad.) Curteria graeca Beron, 1988: 10; Gabryś, 1992: 257

Type locality: Lindos (Rhodes) Distribution: Greece (Creta, Rhodes, Kythnos)

Curteria southcotti Gabryś (ad.) Curteria southcotti Gabryś, 1992: 247; Gabryś & Mąkol, 1994: 127

Type locality: Biala Gora near Tomaszow Lubelski, Zamosc District, Poland Distribution: Poland, ? Hungary

Curteria sp. Distribution: Northern Germany (Wendt, 1996)

Genus Eatoniana Cambridge (ad.) Rhyncholophus L. Koch, 1856: 412 (in part) Macropus (as subgenus of Rhyncholophus) Birula, 1893: 388 (praeocc.) Eatonia Cambridge, 1897: 939 (praeocc.) Eatoniana Cambridge, 1898: 348 Lucasiella Banks, 1900: 32 ? Erythraeus (Ptilolophus) Berlese, 1916 : 292 NB. For detailed synonymy see Southcott, 1961a: 486

Type species (original designation): Eatonia scopulifera Cambridge, 1897

Eatoniana plumipes (L. Koch) (ad.) Rhyncholophus plumipes L. Koch, 1856: 412; George, 1896: 153 Rhyncholophus (Macropus) plumifer Birula, 1893: 388 Eatonia scopulifera Cambridge, 1897: 939 Lucasiella plumipes: Banks, 1900: 32; Trägårdh, 1904a: 68 Erythraeus (Lucasiella) plumipes: André, 1931: 354 Eatoniana plumipes (L. Koch): Southcott, 1961a: 487 Erythraeus (Erythraeus) plumipes: André, 1964: 1 Erythraeus plumipes: Long, 1980: 377

Type locality: Malaga, Spain Distribution: Algeria, Egypt, France, Greece (Corfu), ? Jersey, Spain, Switzerland, Transcaucasia (Nahitchevan), Transcaspia, Tunisia (may be different forms)

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? Eatoniana claviger (Berlese)(ad.) Erythraeus (Ptilolophus) claviger Berlese, 1916: 292 (not Ptilophus, as in Southcott, 1961a: 486) ? Eatoniana claviger (Berlese): André, 1964: 2

Type locality: La Plata Distribution: Argentina

Eatoniana namaquensis (Lawrence) (ad.) Erythraeus namaquensis Lawrence, 1937: 273 Eatoniana namaquamensis [sic] (Lawrence): Southcott, 1961a: 486

Type locality: Kamieskroon, Namaqualand Distribution: South Africa Names used for plume – footed taxa, conspecific or not with Eatoniana plumipes (L. Koch): Rhyncholophus plumipes L. Koch, 1856 – Malaga, Spain Rhyncholophus plumipes L. Koch – Corfu (Frauenfeld, 1868) Rhyncholophus ? plumipes Lucas, 1864 – Algeria, Tunisia Lucasiella plumipes: Trägårdh, 1904a – Egypt, Cairo Rhyncholophus plumipes Lucas – Switzerland (Haller, 1885) Rhyncholophus (Macropus) plumifer Birula, 1893 – Nachitchevan, Aschabad Rhyncholophus plumipes George, 1896 – Jersey (Channel Islands) Eatonia scopulifera Cambridge, 1897 – Algeria Rhyncholophus Halleri Banks, 1900 – Switzerland (for the species of Haller) Eatoniana plumifer: George, 1907 – England Eatoniana georgei Oudemans, 1941 (for Rhyncholophus plumipes George, 1896 non Rhyncholophus plumipes C.L. Koch, 1856) – Jersey Eatoniana plumifer forma transcaspica: Oudemans, 1941 – Ashkhabad

Genus Erythraeus Latreille (ad. and larv.) Erythraeus Latreille, 1806: 146 (adult) Rhyncholophus Dugès, 1834: 31 Rhyncholophus (Apectolophus) Berlese, 1891, A.M.S. 59 (1)2 Bochartia Oudemans, 1910b: 49 (larval); Southcott, 1961a: 491 (as genus) Bockartia [sic]: Womersley, 1934: 251 Parerythraeus Southcott, 1946: 11 (adult) Paraerythraeus Southcott: Khot, 1963: 233 (misspelling)

Type species of Rhyncholophus Dugès, 1834, Apectolophus Berlese, 1891 and Erythraeus Latreille, 1806: Acarus phalangoides de Geer, 1778 Type species of Bochartia: Bochartia kuyperi Oudemans, 1910 Type species of Parerythraeus: Parerythraeus gregoryi Southcott, 1946


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Subgenus Erythraeus s.str. Erythraeus: Latreille, 1806: 146 (adult)

Type species: Acarus phalangoides de Geer, 1778

Erythraeus (E.) acis (Berlese) (ad.) Rhyncholophus phalangioides var. Acis Berlese, 1882 Erythraeus acis (Berlese): Berlese, 1882; Mihelčič, 1958b: 44; Schweizer, 1951: 131; Schweizer & Bader, 1963: 274; Schmölzer, 1993: 61; Gabryś & Mąkol, 1996: 488; Bernini et al., 1994-96: 40

Type locality: Sicily (Italy) Distribution: Austria, Hungary, Italy (incl. Sicily), Poland, Spain, Switzerland

Erythraeus (E.) acutipilis Mihelčič (ad.) Erythraeus acutipilis Mihelčič, 1958a: 279

Type locality: Madrid Distribution: Spain

Erythraeus (E.) adpendiculatus (Schrank) (ad.) Acarus adpendiculatus Schrank, 1781: 518 Rhyncholophus rubescens Dugès: 1834: 15; 1836: 492 Rhyncolophus rubescens: Lucas, 1840: 437; 1850: 473 Rhyncholophus adpendiculatus: Oudemans, 1937: 1962 Erythraeus appendiculatus (Schrank): Buitendijk, 1945: 349 Erythraeus adpendiculatus (Schrank): Karppinen, 1958: 45; Gabryś & Mąkol, 1991: 709; 1996: 488; Gabryś, 1997: 225

Type locality: Austria Distribution: Austria, Finland, France, Germany, Holland, Hungary, Italy, Poland

Erythraeus (E.) adrastus (Southcott) (larval) Bochartia adrastus Southcott, 1961a: 492; Fain & Baugnée, 1996: 22 Erythraeus adrastus (Southcott): Haitlinger, 1987: 725

Type locality: Eremitagegesletten, Denmark Distribution: Belgium, Denmark Hosts: Lasius flavus (Hym., Formicidae), Cicadellidae (Homoptera)

Erythraeus (E.) akbariani Haitlinger et Saboori (larval) Erythraeus akbariani Haitlinger & Saboori, 1996: 125

Type locality: Ardebil Distribution: Iran Host: Aphididae (Homoptera)

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Erythraeus (E.) ankaraicus Saboori, Cobanoglu et Bayram (larval) Erythraeus ankaraicus: Saboori, Cobanoglu & Bayram, 2004: 137; Bayram & Cobanoglu, 2005: 163

Type locality: Ankara Distribution: Turkey Host: Hyalopterus pruni (Aphididae, Homoptera)

Erythraeus (E.) aokii Shiba (ad.) Erythraeus aokii Shiba, 1972: 52

Type locality: summit of Mt. Poroshiri, Hidaka, Hokkaido Distribution: Japan (Hokkaido, Honshu)

Erythraeus (E.) berninensis Haitlinger (larval) Erythraeus (E.) berninensis Haitlinger, 2007b: 62

Type locality: Col Bernina, ca. 2300 m Distribution: Switzerland Host: from plants and from undetermined Cicadellidae (Homoptera, Auchenorhyncha)

Erythraeus (E.) bolivianus Beron (ad.) Erythraeus (E.) bolivianus Beron, in prep.

Type locality: above La Paz, 4480 m Distribution: Bolivia

Erythraeus (E.) bulgaromontanus Beron (larval) Erythraeus bulgaromontanus Beron, 1982: 50

Type locality: Rila (2250-2925 m) Distribution: Bulgaria Host: under stones

Erythraeus (E.) campestri Mihelčič (ad) Erythraeus campestri Mihelčič, 1958a: 279

Type locality: Madrid Distribution: Spain

Erythraeus (E.) capillatus Lawrence (ad) Erythraeus capillatus Lawrence, 1944: 452

Type locality: World’s View, Pietermaritzburg Distribution: South Africa

Erythraeus (E.) chinensis (Zheng) (comb. nov.) (larval) Bochartia chinensis Zheng, 2002: 91



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Erythraeus (E.) cinereus (Dugès) (ad.) Rhyncholophus cinereus Dugès, 1834: 15; 1836: 495; Canestrini, 1885: 143; Jablonowski, 1918: 3 Rhyncolophus cinereus: Lucas, 1840: 473; 1850: 473 Trombidium cinereum: Gervais, 1844: 183 Erythraeus cinereus (Dugès): Buitendijk, 1945: 349; Gabryś & Mąkol, 1991: 709; 1994: 127; Bernini et al., 1994-96: 40; Mąkol & Gabryś, 1999: 30; Haitlinger, 2002e: 148

Type locality: Montpellier (France) Distribution: France, Holland, Hungary, Italy (incl. Sicily and Sardinia), Poland

Erythraeus (E.) crocatus (C.L. Koch)(ad.) Rhyncholophus crocatus C.L.Koch, 1837: fa.16, t.15; 1841: 430; 1842: 50; Canestrini, 1885: 151; Jablonowski, 1918: 3 Balaustium crocatus: Oudemans, 1937: 1938 Erythraeus crocarus: Bernini et al., 1994-96: 40

Type locality: Germany Distribution: Croatia, Germany, Hungary, Italy

Erythraeus (E.) decalvatus Berlese (ad.) Erythraeus decalvatus Berlese, 1920: 210

Type locality: Germany Distribution: Germany

Erythraeus (E.) delhiensis (Khot) (ad.) Paraerythraeus delhiensis Khot, 1963: 234 Erythraeus delhiensis (Khot): Gabryś, 1989: 60

Type locality: Delhi, India Distribution: India

Erythraeus (E.) electoralis (C.L. Koch)(ad.) Rhyncholophus electoralis C.L.Koch, 1842: 50; Canestrini & Fanzago, 1877: 70; Canestrini, 1885: 147; Jablonowski, 1918: 3 Erythraeus electoralis (C.L. Koch): Bernini et al., 1994-96: 40

Type locality: Germany Distribution: Germany, Hungary, Italy (incl. Sardinia), Romania

Erythraeus (E.) elegans (Khot)(ad.) Paraerythraeus elegans Khot, 1963: 238 Erythraeus elegans (Khot): Gabryś, 1989: 60

Type locality: Delhi Distribution: India

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Erythraeus (E.) elwirae Haitlinger (larval) Erythraeus elwirae Haitlinger, 1987a: 727; 2002e: 148; Gabryś, 1988: 23 Type locality: Jelenia Gora, Poland Distribution: Poland Host: “from plants”

Erythraeus (E.) exilipes Banks (ad.) Erythraeus exilipes Banks, 1947: 110

Type locality: Duke Forest, Durham Distribution: USA (North Carolina)

Erythraeus (E.) garmsaricus Saboori Goldarazena et Khajeali (larval) Erythraeus garmsaricus Saboori Goldarazena & Khajeali, 2004: 168

Type locality: Garmsar, Semnan Prov. (Iran) Distribution: Iran Host: “undetermined aphids”

Erythraeus (E.) germanicus Oudemans (ad.) Erythraeus germanicus Oudemans, 1903b: 88; Buitendijk, 1945: 350

Type locality: Germany Distribution: Germany

Erythraeus (E.) gertrudae Haitlinger (larval) Erythraeus gertrudae Haitlinger, 1987a: 726; 2000b: 191; 2002e: 148; Gabryś, 1988: 23

Type locality: Wielecin n. Wyszkow (Voiv. Ostroleka) Distribution: Finland, Norway, Poland, Slovakia, Slovenia Host: “from plants”; undet. larva of Homoptera (Slovenia)

Erythraeus (E.) ghilarovi Schuster et Turk (ad.) Erythraeus ghilarovi Schuster & Turk, 1987: 15

Type locality: “Andes of southern Peru, puna region along the main road between Arequipa and Juliaca; pass NE Pati, altitude about 4700 m” Distribution: Peru

Erythraeus (E.) glabrisetosus Willmann (ad.) Erythraeus glabrisetosus Willmann, 1953: 493

Type locality: Pfalten Kogel, Graz Distribution: Austria

Erythraeus (E.) gregoryi (Southcott) (ad.) Parerythraeus gregoryi Southcott, 1946b: 11 Erythraeus gregoryi (Southcott): Gabryś, 1989: 60


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Type locality: Coomalie Creek, Northern Territory, Australia Distribution: Australia (N. Territory)

Erythraeus (E.) hibernans Oudemans (ad.) Erythraeus hibernans Oudemans, 1901: 219; 1902: 36; Buitendijk, 1945: 350; Bernini et al., 1994-96: 40

Type locality: San Remo Distribution: Italy

Erythraeus (E.) hirsti Beron [nom. nov. pro Erythraeus pilosus (Hirst, 1928), nec Erythraeus pilosus Banks, 1894] Leptus pilosus Hirst, 1928: 569 Erythraeus pilosus (Hirst): Womersley, 1934: 220; Southcott, 1946: 34

Type locality: Glen Osmond (larva), Dubbo, N.S.Wales (adult) Distribution: Australia (New South Wales) Host: Psocoptera (Myopsocus sp., Myopsocidae), Heteroptera

Erythraeus (E.) hypertrichotus Saboori, Goldarazena et Khajeali (larval) Erythraeus (E.) hypertrichotus Saboori, Goldarazena & Khajeali, 2004: 163

Type locality: Tabris Distribution: Iran Host: “undetermined aphids”

Erythraeus (E.) imperfectus Mihelčič (ad.) Erythraeus imperfectus Mihelčič, 1958a: 280

Type locality: Caceres, Spain Distribution: Spain

Erythraeus (E.) indicus (Khot) (ad.) Paraerythraeus indicus Khot, 1963: 236 Erythraeus indicus (Khot): Gabryś, 1989: 60

Type locality: Gwalior, Madhya Pradesh Distribution: India (Madhya Pradesh)

Erythraeus (E.) italicus Berlese (ad.) Erythraeus italicus Berlese, 1920: 209; Bernini et al., 1994-96: 40

Type locality: “Italia centrale (Pisano)” Distribution: Italy

Erythraeus (E.) jacoti Goosmann (ad.) Erythraeus Jacoti Goosmann, 1925: 97

Type locality: “near Peking” Distribution: China

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Erythraeus (E.) johnstoni Oudemans (ad.) Erythraeus johnstoni Oudemans, 1937: 1967

Type locality: Berwickshire Distribution: England

Erythraeus (E.) jowitae Haitlinger (larval) Erythraeus jowitae Haitlinger, 1987a: 725; 2007b: 62; Gabryś, 1988: 22; Gabryś & Mąkol, 1991: 709; 1996: 488

Type locality: Voiv. Szczecin, Poland Distribution: France, Hungary, Poland Host: “from plants”

Erythraeus (E.) kacperi Haitlinger (larval) Erythraeus (E.) kacperi Haitlinger, 2004b: 158

Type locality: Phnom Penh Distribution: Cambodia Host: from plants

Erythraeus (E.) kresnensis Beron (larval) Erythraeus kresnensis Beron, 1982: 47; Haitlinger, 2000d: 384

Type locality: Kresna, SW Bulgaria Distribution: Bulgaria, Turkey Host: Machilidae (Thysanura)

Erythraeus (E.) kuyperi (Oudemans)(larval) Bochartia kuyperi Oudemans, 1910b: 49; Vitzthum, 1925; Pussard & André, 1929: 295; André, 1929b: 1; 1951: 253; Irk, 1939b: 168; Willmann, 1951c: 154; Schweizer, 1951: 161; Sellnick, 1958: 43; Southcott, 1961a: 491 Erythraeus kuyperi (Oudemans): Buitendijk, 1945: 350; Haitlinger, 1987a: 731; 2003f: 7; Gabryś, 1988: 23; Gabryś & Mąkol, 1994: 127

Type locality: De Bilt, Holland Distribution: Austria, Czech Rep., France, Germany, Holland, Iceland, Israel, Italy, Poland, Sweden, Switzerland, Ukraine Host: Aphis papaveris, Macrosiphum fragariae, etc. (Homoptera, Aphididae)

Erythraeus (E.) longisetosus Mihelčič (ad) Erythraeus longisetosus Mihelčič, 1958b: 44

Type locality: Sierra Nevada Distribution: Spain


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Erythraeus (E.) malwinae Haitlinger (larval) Erythraeus malwinae Haitlinger, 1995: 262; 2002b: 95; 2002e: 148; 2005a: 74

Type locality: Garmisch-Partenkirchen Distribution: Andorra, Germany, Poland Host: from plants

Erythraeus (E.) mariolae Haitlinger (larval) Erythraeus mariolae Haitlinger, 1994a: 408

Type locality: Bad Ragaz, Switzerland Distribution: Switzerland Host: “from plants”

Erythraeus (E.) mirjavehi Gabryś (ad.) Erythraeus mirjavehi Gabryś, 1989: 60

Type locality: Mirjaveh (near Zahedan, E. Iran) Distribution: Iran

Erythraeus (E.) moeritzensis Haitlinger (larval) Erythraeus moeritzensis Haitlinger, 2007b: 66

Type locality: Le Punt Chamnes Ch. N. Sankt Möritz, ca. 1950 m Distribution: Switzerland Host: “from herbaceous plants”

Erythraeus (E.) monikae Haitlinger (larval) Erythraeus monikae Haitlinger, 1987a: 728; 1995: 263; 2000b: 191; 2002d: 554; 2003d: 3; Gabryś, 1988: 23

Type locality: Glogówko n. Glogów (Voiv. Legnica), Poland Distribution: Finland, Germany, Slovakia, Slovenia, Poland Host: “from plants”

Erythraeus (E.) nasalis Evans (ad.) Erythraeus nasalis Evans, 1953: 279

Type locality: Kilimanjaro, 3810 m Distribution: Tanzania

Erythraeus (E.) nigritiensis André (ad.) Erythraeus nigritiensis André, 1962: 307

Type locality: Djebel Merkhiyat nr. Omdurman Distribution: Sudan

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Erythraeus (E.) nivalis Schweizer (ad.) Erythraeus nivalis Schweizer, 1951: 123; Schmölzer, 1956: 557, 1993: 60; Franz, 1954: 381; Mihelčič, 1958d: 108 (= ? Rhyncholophus nivalis Heer, 1845: 6; Erichson, 1847: 67; Erythraeus nivalis Heer: Oudemans, 1937: 1979)

Type locality of Rhyncholophus nivalis Heer: Switzerland, “Bündnerberg, im Kanton Glarus, auf der obersten Spitze des P.Levarone, im Engadin (9580 F.) und auf der Spitze des Umbrail 9100 F. ü.M.”. Type locality of Erythraeus nivalis Schweizer: Blais and other localities in the Swiss Alps, up to 3207 m; in Dauphiné up to 3770 m (Schmölzer, 1956) Distribution: Austria, France, Switzerland

Erythraeus (E.) opilionoides (C.L. Koch) (ad.) Rhyncholophus opilionoides C.L. Koch, 1837: 16, tab. 3; 1842: 51; Haller, 1882: 314 Erythraeus opilionoides (C.L. Koch): Gabryś & Mąkol, 1991: 709; 1994: 127; 1996: 488; Bernini et al., 1994-96: 40; Haitlinger, 2002e: 148 Erythraeus opilionides: Willmann, 1951c: 148; Franz, 1954: 381

Type locality: Germany Distribution: Austria, Germany, Holland, Hungary, Norway, Poland, ex-Yugoslavia

Erythraeus (E.) orientalis Khot (ad.) Erythraeus orientalis Khot, 1963: 240

Type locality: Poona, Maharashtra Distribution: India (Maharashtra)

Erythraeus (E.) osmondensis Southcott (larval) Erythraeus osmondensis Southcott, 1946: 31

Type locality: Mt. Osmond, S. Australia Distribution: Australia (S. Australia) Host: thrips (Thysanoptera)

Erythraeus (E.) peruvianus Beron (ad.) Erythraeus (E.) peruvianus Beron, in prep.

Type locality: Huaraz Distribution: Peru

Erythraeus (E.) phalangoides (de Geer) (ad.) Acarus phalangoides de Geer, 1778: 134; 1783: 205 Trombidium phalangioides (de Geer): Hermann, 1804: 33 Rhyncholophus phalangioides (De Geer): Dugès, 1834: 30; C.L. Koch, 1837: 16, 4; 1842: fa. 3: 51; Kramer, 1897: 78; Jablonowski, 1918: 3; Hull, 1918: 24-


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Rhyncholophus phalangoides (De Geer): Schweizer, 1922: 83 Rhyncholophus opilionoides: C.L. Koch, 1837: fa.16, t. 3; 1842: fa. 3: 51 Rhyncolophus degeerii Lucas, 1840: 472 Apectolophus phalangioides (de Geer): Berlese, 1891: fasc. LIX, 1 Erythraeus phalangoides (de Geer): Thor & Ushida, 1933: 138; Buitendijk, 1945: 350; Willmann, 1951c: 148; Turk, 1953: 26; Franz, 1954: 381; Shulov, 1957: 236; Schmölzer, 1993: 60; Mihelčič, 1958a: 279, 1958b: 44, 1971: 145; Lindroth et al., 1973: 130; Witalinski, 1985: 43; Gabryś, 1988: 22; Gabryś & Mąkol, 1994: 127; Haitlinger, 2002e: 148 Erythraeus phalangioides (de Geer): Latreille, 1806: 146; 1817: 423; Leach, 1816: 443; Contarini, 1847: 162; Trägårdh, 1904b: 58; André, 1933: 487; Sellnick, 1940: 36; 1949: 129; Willmann, 1949: 128; Starkoff & Starkoff, 1950: 96; Schweizer, 1951: 124; Bottazzi, 1956: 43; Karppinen, 1958: 44; Schweizer & Bader, 1963: 272; Bernini et al., 1994-96: 40 Rhyncholophus gracilipes: Kramer, 1897: 77 Erythraeus phalangiodes var. gracilipes (Kram.): Trägårdh, 1904b: 59 (really Erythraeus!) Erythraeus phalangioides (de Geer) var. gracilipes (Kram.): Trägårdh, 1910: 485. Note: The description and picture are of Leptus sp. larva ! Erythraeus phalangiodes gracilipes: Schmölzer, 1993: 61 Rhyncholophus intermedius Trägårdh, 1902: 4 Erythraeus dubiosus Schweizer, 1951: 133; Franz, 1954: 381; Schweizer & Bader, 1963: 133; Gabryś & Haitlinger, 1986: 472

Type locality of Erythraeus dubiosus Schweizer: Buffalora, La Drosa, Val Muschauns, Switzerland Type locality of Acarus phalangoides de Geer: Sweden Type locality of Rhyncholophus opilionoides C.L. Koch, 1837: Zweibrücken, Germany Type locality of Rhyncholophus intermedius Trägårdh: Lappland Type locality of Erythraeus dubiosus Schweizer: Buffalora, La Drosa, Val Muschauns, Switzerland Distribution: Austria, Denmark, Finland, France, Germany, Great Britain, Greenland, Holland, Hungary, Iceland, Italy, Kuril Is., Norway, Poland, Romania, Spain, Sweden, Switzerland

Erythraeus (E.) phalangoides nipponicus Kawashima (larval and adult) Erythraeus nipponicus Kawashima, 1961b: 23 (larval) Erythraeus flavopictus Hara & Hanada, 1962: 268 (larval); Morikawa, 1963: 104 (564)

Type locality of E. nipponicus Kawashima: Japan Type locality of E. flavopictus Hara et Hanada: Narashino, Chiba Pref. Distribution: Japan

Erythraeus (E.) picaforticus Haitlinger (larval) Erythraeus picaforticus Haitlinger, 2002f: 192

Type locality: Can Picafort, Mallorca Distribution: Mallorca (Balearic Is., Spain)

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Erythraeus (E.) plumatus Beron, nom. nov. (larval) Erythraeus plumosus Tseng, Yang & Pan, 1976: 68 (nomen praeoccupatum from E. plumosus Khot, 1963

Type locality: Taiwan Distribution: Taiwan Host: Hemiptera

Erythraeus (E.) plumosus Khot (ad.) Erythraeus plumosus Khot, 1963: 240


Erythraeus (E.) principalis (C.L. Koch)(ad) Rhyncholophus principalis C.L. Koch, 1837: fa.16, t.8; 1842: 50 Balaustium principalis: Oudemans, 1937: 1947 Erythraeus principalis (C.L. Koch)

Type locality: “Fasanerie bei Zweibrücken”, Germany Distribution: Germany

Erythraeus (E.) pseudonivalis Schmölzer (ad.) Erythraeus pseudonivalis Schmölzer, 1956: 559

Type locality: Temple-Ecrins, 2740 m, Alps Distribution: France

Erythraeus (E.) regalis (C.L. Koch) (ad.) Rhyncholophus regalis C.L. Koch, 1837: fa. 16, tab.5 ; 1841: 430; 1842: 50; Canestrini, 1885: 146; Berlese, 1892: Fasc. LXII, 1900: 164; Halbert, 1915: 114; Hull, 1918: 24; Schweizer, 1922: 82 Belaustium regale (L. Koch): Oudemans, 1897: 120 Rhyncholophus signatus L. Koch, 1879: 124 – Krasnojarsk (Russia) Rhyncolophus regalis: Tafner, 1905 Rhyncholophus communis George, 1910: 427 Rhyncholophus niger George, 1912: 252 Erythraeus regalis: Trägårdh, 1904b: 61; Berlese, 1920: 19; André, 1927: 169; Irk, 1939b: 167; Szalay, 1940: 198; Sellnick, 1940: 36; Buitendijk, 1945: 350; Feider, 1948: 13; Starkoff & Starkoff, 1950: 96; Willmann, 1951c: 148; 1954: 241; 1955: 218; Schweizer, 1951: 121; Franz, 1954: 382; Schmölzer, 1993: 60; Schweizer & Bader, 1963: 269; Bottazzi, 1956: 43; Mihelčič, 1957: 107; 1958a: 279; 1958b: 44; 1971: 145; Karppinen, 1958: 45; Cooreman, 1955c: 14; Lindroth et al., 1973: 130; Gabryś & Haitlinger, 1986: 472; Gabryś, 1988: 22; Gabryś & Mąkol, 1994: 127; 1996: 488; Bernini et al., 1994-96: 40; Haitlinger, 2002e: 148

Type locality of Rhyncholophus regalis C.L. Koch, 1837: Germany Distribution: Albania, Algeria, Austria, Belgium, Bulgaria, Finland, Germany, Great Britain, Holland, Hungary, Iceland, Ireland, Italy (incl. Sardinia), Poland, Romania, Russia, Spain, Switzerland


Erythraeus (E.) remotus Banks (ad.) Erythraeus remotus Banks, 1924: 93

Type locality: Indefatigable Distribution: Galapagos

Erythraeus (E.) rilensis Beron (larval) Erythraeus rilensis Beron, 1982: 55

Type locality: Rila, Grančar (2250 m) Distribution: Bulgaria

Erythraeus (E.) rupestris (Linnaeus) (ad.) Acarus rupestris Linnaeus, 1758: 618; Dwigubski, 1802: 194 Rhyncholophus imperialis C.L.Koch, 1837: fa.16, t.6; 1842: 50; L.Koch, 1879: 125 Erythraeus imperialis (C.L. Koch): Willmann, 1949: 128; Franz, 1954: 381; Gabryś, 1988: 22 Gamasus rupestris: Fabricius, 1822: 364 Erythraeus rupestris: Brebisson, 1827: 265; Karppinen, 1958: 45

Type locality: Europe (? Sweden – listed in Fauna suecica, 1746) Type locality of Rhyncholophus imperialis C.L.Koch: Krasnoyarsk (Russia) Distribution: Austria, Finland, Poland, Russia, ? Sweden

Erythraeus (E.) rutgeri Haitlinger (larval) Erythraeus rutgeri Haitlinger, 2003a: 142

Type locality: Rhodes, Eleousa (28o03’ N, 36o18’ E) Distribution: Greece (Rhodes) Host: from plants

Erythraeus (E.) sabrinae Haitlinger et Saboori (larval) Erythraeus sabrinae Haitlinger & Saboori, 1996: 128

Type locality: Tehran Distribution: Iran Host: Aphididae (Homoptera)

Erythraeus (E.) serratociliatus (Khot)(ad.) Paraerythraeus serratociliatus Khot, 1963: 236 Erythraeus serratociliatus (Khot): Gabryś, 1989: 60


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Erythraeus (E.) shojaii Babolmorad et Saboori (larval) Erythraeus shojaii Babolmorad & Saboori, 2000: 119

Type locality: Karaj, Iran Distribution: Iran Host: Monosteira unicostata (Tingidae, Hemiptera)

Erythraeus (E.) sifi Haitlinger (larval) Erythraeus sifi Haitlinger, 2000c: 381

Type locality: Cavusin Distribution: Turkey Host: unknown

Erythraeus (E.) similis (G. Canestrini) (ad.) Rhyncholophus similis G. Canestrini, 1885: 144 Erythraeus similis: Bernini et al., 1994-96: 40

Type locality: Sardinia Distribution: Italy (Sardinia)

Erythraeus (E.) simplex Banks (ad.) Erythraeus simplex Banks, 1915: 40

Type locality: U.S.A. Distribution: U.S.A.

Erythraeus (E.) southcotti Goldarazena et Zhang (larval) Erythraeus southcotti Goldarazena & Zhang, 1998b: 149; Haitlinger, 2002f: 19; 2006e

Type locality: Tafalla (Navarra, Spain) Distribution: Spain, Mallorca (Balearic Is.) Host: Therioaphis luteola (Aphididae, Homoptera)

Erythraeus (E.) spatulatopilis Mihelčič (ad) Erythraeus spatulatopilis Mihelčič, 1958b: 44


Erythraeus (E.) stuarti Southcott (larval) Erythraeus stuarti Southcott, 1946b: 29

Type locality: Attack Creek, Northern Territory Distribution: Australia (N.T.) Host: “running free over the ground”


Erythraeus (E.) styriacus Turk (ad., larval) Erythraeus styriacus Turk, 1981: 208; Haitlinger, 2007a: 72 Type locality: Weinzodl near Graz (Austria) Distribution: Austria, Bosnia and Herzegovina, Croatia, Macedonia, Slovenia

Erythraeus (E.) tinnae Haitlinger (larval) Erythraeus (E.) tinnae Haitlinger, 1997a: 124

Type locality: Tenerife, Igueste, 1500 m Distribution: Canary Is. Host: unknown

Erythraeus (E.) titikaka Beron (larval) Erythraeus (E.) titikaka Beron, in prep.

Type locality: Puno, 3900 m Distribution: Peru Host: unknown

Erythraeus (E.) tragardhi (Southcott)(ad.) Erythraeus Dugèsi Nicolet: Trägårdh, 1904: 65, non E. Dugesii (Lucas, 1846) Parerythraeus tragardhi Southcott, 1961a: 512 Erythraeus tragardhi (Southcott): Gabryś, 1989: 60

Type locality: Cairo, Egypt Distribution: Algeria, Egypt

Erythraeus (E.) yangshuonicus Haitlinger (larval) Erythraeus (E.) yangshuonicus Haitlinger, 2006a: 86

Type locality: Yangshuo n. Guilin Distribution: China Host: from herbaceous plants

Erythraeus (E.) zhangi Haitlinger (larval) Erythraeus (E.) zhangi Haitlinger, 2006a: 88

Type locality: park in Beijing Distribution: China Host: from herbaceous plants

Subgenus Zaracarus Southcott (larval) Erythraeus (Zaracarus) Southcott, 1995b: 223 Type species: Erythraeus (Zaracarus) lancifer Southcott, 1995

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Erythraeus (Zaracarus) aydinicus Saboori, Cakmak et Nouri-Gonbalani (larval) Erythraeus (Zaracarus) aydinicus Saboori, Cakmak & Nouri-Gonbalani, 2004: 131

Type locality: Aydin (Turkey) Distribution: Turkey Host: Empoasca sp. (Cicadellidae, Homoptera)

Erythraeus (Zaracarus) budapestensis Fain et Ripka (larval) Erythraeus (Zaracarus) budapestensis Fain & Ripka, 1998: 44; Haitlinger, 2004d: 145; 2006f: 37; 2007a: 74

Type locality: park in Budapest Distribution: Croatia, Greece, Hungary, Bosnia and Hercegovina, Montenegro Host: Hop-tree, Ptelea trifoliata (Rutaceae), infested with Aphis fabae, from herbaceous plants.

Erythraeus (Zaracarus) didonae Haitlinger (larval) Erythraeus (Zaracarus) didonae Haitlinger, 2000c: 383

Type locality: Bursa, Turkey Distribution: Turkey Host: unknown

Erythraeus (Zaracarus) eleonorae Haitlinger (larval) Erythraeus eleonorae Haitlinger, 1987a: 729; Gabryś, 1988: 23 Erythraeus (Zaracarus) eleonorae Haitlinger, 1997: 128

Type locality: Voiv. Lublin, Koszalin and Pila, Poland Distribution: Poland

Erythraeus (Zaracarus) fabiolae Haitlinger (larval) Erythraeus (Zaracarus) fabiolae Haitlinger, 1997a: 126

Type locality: Tenerife, Igueste, 1400 m Distribution: Canary Is.

Erythraeus (Zaracarus) iranicus Saboori et Akrami (larval) Erythraeus (Zaracarus) iranicus Saboori & Akrami, 2001: 159

Type locality: Karaj (Iran) Distribution: Iran Host: Homoptera, Aphidodea

Erythraeus (Zaracarus) jinkaensis Haitlinger (larval) Erythraeus (Zaracarus) jinkaensis Haitlinger, 2006h: 80

Type locality: Ethiopia Distribution: Ethiopia


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Erythraeus (Zaracarus) kastaniensis Haitlinger (larval) Erythraeus (Zaracarus) kastaniensis Haitlinger, 2006b: 111

Type locality: Kastania, Pagondas (Samos) Distribution: Greece (Samos) Host: from plants

Erythraeus (Zaracarus) kharrazii Saboori (larval) Erythraeus (Zaracarus) kharrazii Saboori, 2000: 125

Type locality: Karaj Distribution: Iran Host: in pitfall trap

Erythraeus (Zaracarus) kurdistaniensis Khanjani et Ueckermann (larval) Erythraeus (Zaracarus) kurdistaniensis Khanjani & Ueckermann, 2005: 123

Type locality: Kurdistan (Iran) Distribution: Iran

Erythraeus (Zaracarus) lancifer Southcott (larval) Erythraeus (Zaracarus) lancifer Southcott, 1995: 223

Type locality: Puerta de Pina, Pina, Zaragoza Prov. Distribution: Spain Host: Dolichopodidae (Diptera)

Erythraeus (Zaracarus) longipedus Saboori et Nowzari (larval) Erythraeus (Zaracarus) longipedus Saboori & Nowzari, 2001:127

Type locality: Shahryar Distribution: Iran Host: Aphidodea (Homoptera)

Erythraeus (Zaracarus) passidonicus Haitlinger (larval) Erythraeus (Zaracarus) passidonicus Haitlinger, 2006b: 114

Type locality: Passidonia (Samos) Distribution: Greece (Samos) Host: from plants

Erythraeus (Zaracarus) preciosus Goldarazena et Zhang (larval) Erythraeus preciosus Goldarazena & Zhang, 1998b: 153 Erythraeus (Zaracarus) preciosus Goldarazena et Zhang: Haitlinger, 2002f: 194; 2006e: 106; 2007c

Type locality: Navarra, Tafalla (Spain) Distribution: Greece, Spain (Navarra, Aragon), Mallorca (Balearic Is.), Ukraine (Crimea Host: Therioaphis luteola (Homoptera, Aphididae); herbaceous plants

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Erythraeus (Zaracarus) rajabii Saboori (larval) Erythraeus (Zaracarus) rajabii Saboori, 2000:127

Type locality: Karaj Distribution: Iran Host: Cicadellidae (Homoptera)

Erythraeus (Zaracarus) sibuljinicus Haitlinger (larval) Erythraeus (Zaracarus) sibuljinicus Haitlinger, 2004d: 145

Type locality: Šibuljina n. Starigrad Distribution: Croatia Host: from plants

Erythraeus (Zaracarus) tehranicus Haitlinger et Saboori (larval) Erythraeus (Zaracarus) tehranicus Haitlinger & Saboori, 1996: 128

Type locality: Tehran Distribution: Iran Host: from plants

Erythraeus (Zaracarus) ueckermanni Saboori, Nowzari et Bagheri-Zenouz (larval) Erythraeus (Zaracarus) ueckermanni Saboori, Nowzari & Bagheri-Zenouz, 2004: 78

Type locality: Shahryar (Iran) Distribution: Iran Host: unknown

Genus Erythraxus Southcott (ad.) Erythraxus Southcott, 1961a: 509

Type species: Erythraeus bisetosa Wharton, 1938

Erythraxus bisetosus (Wharton) (ad.) Erythraeus bisetosa Wharton, 1938: 143 Erythraxus bisetosus (Wharton): Southcott, 1961a: 509

Type locality: San Isidro Cave, Yucatan, Mexico Distribution: Mexico (Yucatan)

Genus Erythrellus Southcott (ad.) Erythrellus Southcott, 1946b: 10

Type species: Erythrellus imbricatus Southcott, 1946

Erythrellus imbricatus Southcott (ad.) Erythrellus imbricatus Southcott, 1946b: 10

Type locality: Glen Osmond, South Australia Distribution: Australia (South Australia)


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Genus Erythrites Southcott (ad.) Erythrites Southcott, 1961a: 466

Type species: Leptus reginae Hirst, 1928 Bockartia [sic] Womersley, 1934: 251 (in part)

Subgenus Erythrites Southcott (ad.) Erythrites (Erythrites) Southcott, 1961a: 467

Type species: Leptus reginae Hirst, 1928

Erythrites (E.) reginae (Hirst)(ad.) Leptus reginae Hirst, 1928: 569 Erythraeus reginae (Hirst): Womersley, 1934: 219 Erythrites (E.) reginae (Hirst): Southcott, 1961a: 467 Leptus antepodianus Hirst, 1928: 570 (synonymized with E. reginae by Womersley, 1934: 219 sub “antipodianus”)

Type locality: Adelaide, South Australia Type locality of Leptus antepodianus Hirst: Tanunda, S. Australia Distribution: Australia (S.A.)

Subgenus Gerites Southcott (ad.) Erythrites (Gerites) Southcott, 1961a: 468

Type species: Erythraeus guttatus Southcott, 1946

Erythrites (Gerites) guttatus (Southcott) (ad.) Erythraeus guttatus Southcott, 1946b: 31 Erythrites (Gerites) guttatus Southcott, 1961a: 468

Type locality: Glen Osmond, S. Australia Distribution: Australia

Subgenus Urrbrites Southcott (ad. and larv.) Erythrites (Urrbrites) Southcott, 1961a: 468

Type species: Erythraeus urrbrae Womersley, 1934

Erythrites (Urrbrites) urrbrae (Womersley) (ad.) Erythraeus urrbrae Womersley, 1934: 221; Southcott, 1946b: 36 Erythrites (Urrbrites) urrbrae Southcott, 1961a: 468

Type locality: Urrbrae, South Australia Distribution: Australia Hosts: Myopsocus sp. (Psocoptera, Myopsocidae)

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Erythrites (? Urrbrites) jacksoni Southcott (larval) Erythrites jacksoni Southcott, 1988: 227

Type locality: Edwards-Mingha Ridge, Arthur’s Pass, 1830 m, New Zealand Distribution: New Zealand Host: Brachaspis collinus (Acrididae, Orthoptera)

Subgenus Terebrites Southcott (larval) Erythrites (Terebrites) Southcott, 1961a: 469

Type species: Erythraeus womersleyi Southcott, 1946

Erythrites (Terebrites) womersleyi (Southcott)(larval) Erythraeus womersleyi Southcott, 1946b: 40 Erythrites (Terebrites) womersleyi Southcott, 1961a: 469

Type locality: Glen Osmond, S. Australia Distribution: Australia (S. Australia) Host: adult Erythroides serratus (Erythraeidae, Acariformes)

Genus Erythroides Southcott (ad. and larval) Erythroides Southcott, 1946b: 12

Type species: Erythraeus serratus Womersley, 1936

Erythroides clavatus Southcott (larval) Erythroides clavatus Southcott, 1946b: 14


Erythroides darwini Southcott (ad.) Erythroides darwini Southcott, 1946b: 17

Type locality: Adelaide River, Northern Territory, Australia Distribution: Australia (N. Territory)

Erythroides macdonnelli Southcott (ad.) Erythroides macdonnelli Southcott, 1946b: 17

Type locality: Alice Springs, Northern Territory, Australia Distribution: Australia (N. Territory)

Erythroides neoserratus Southcott (ad.) Erythroides neoserratus Southcott, 1946b: 13



Erythroides serratus (Womersley) (ad.) Erythraeus serratus Womersley, 1936: 117 Erythroides serratus (Womersley): Southcott, 1946: 12

Type locality: Bathurst, New South Wales Distribution: Australia (N.S.W., South Australia)

Genus Forania Southcott (larval) Forania Southcott, 1961a: 489

Type species (original designation): Bochartia mentonensis André, 1929

Forania mentonensis (André) (larval) Bochartia mentonensis André, 1929: 255; 1930: 544 Forania mentonensis (André): Southcott, 1961a: 490

Type locality: Ubac Foran between Ubi and Sospel, Alpes-Maritimes, 800 m Distribution: France Hosts: Plagiolepis pygmaea, Pheidole palidula (Formicidae, Hymenoptera)

Genus Helladerythraeus Beron (stat. nov.)(ad.) Erythraeus (Helladerythraeus) Beron, 1988: 3

Type species: Erythraeus (Helladerythraeus) pachytibialis Beron, 1988

Helladerythraeus pachytibialis (Beron)(stat. nov.)(ad.) Erythraeus (Helladerythraeus) pachytibialis Beron, 1988: 3 Type locality: Lindos (Rhodes) Distribution: Greece (Rhodes and Kythnos Is.)

Genus Kakamasia Lawrence (ad.) Kakamasia Lawrence, 1944: 440

Type species: Kakamasia polygona Lawrence, 1944

Kakamasia pilosa Lawrence (ad.) Kakamasia pilosa Lawrence, 1944: 442

Type locality: Montagu, Cape Province, South Africa Distribution: South Africa

Kakamasia polygona Lawrence (ad.) Kakamasia polygona Lawrence, 1944: 440

Type locality: Kakamas, Little Namaqualand, South Africa Distribution: South Africa

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Genus Lasioerythraeus Welbourn et Young (ad. and larv.) Lasioerythraeus Welbourn & Young, 1987: 231

Type species: Lasioerythraeus johnstoni Welbourn et Young, 1987

Lasioerythraeus johnstoni Welbourn et Young (ad. and larval) Lasioerythraeus johnstoni Welbourn & Young, 1987: 232; Haitlinger, 2004g: 130; Young & Welbourn, 1988: 269

Type locality: Washington Co., Mississippi, U.S.A. Distribution: Dominican Republic, U.S.A. (Mississippi) Hosts: Lygus lineolaris and Holcostethus sp. (Heteroptera, Miridae, Pentatomidae) and whitefly (Homoptera, Aleyrodidae)

Lasioerythraeus shirleyanneae (McDaniel et Bolen)(larval) Bochartia shirleyanneae McDaniel & Bolen, 1981: 40 Lasioerythraeus shirleyanneae (McDaniel et Bolen): Welbourn & Young, 1987: 238

Type locality: 10 miles N Sinton, San Patricio Co., U.S.A. Distribution: Texas (U.S.A.)

Lasioerythraeus welbourni Beron (larval) Lasioerythraeus welbourni Beron, in prep.

Type locality: La Paz, 4480 m, Bolivia Distribution: Bolivia

Lasioerythraeus whitcombi (Smiley) (ad.) Erythraeus whitcombi Smiley, 1964: 110 Lasioerythraeus whitcombi (Smiley): Welbourn & Young, 1987: 238

Type locality: Conway Co., Arkansas, U.S.A. Distribution: Arkansas, U.S.A.

Genus Kamertonia Gabryś (ad) Kamertonia Gabryś, 1999: 57

Type species: Kamertonia polonica Gabryś, 1999

Kamertonia polonica Gabryś (ad.) Kamertonia polonica Gabryś, 1999: 57

Type locality: Orzechovo near Ustka (Pomorskie Province, N Poland), sand dune Distribution: Poland

Genus Neosmaris Hirst (ad.) Neosmaris Hirst, 1926: 612

Type species: Neosmaris novaezealandiae Hirst, 1926


Neosmaris novaezealandiae Hirst (ad.) Neosmaris novaezealandiae Hirst, 1926: 612

Type locality: Routeburn, Otago, New Zealand Distribution: New Zealand

Genus Opserythraeus Fain (larval) Opserythraeus Fain, 1996: 252

Type species: Opserythraeus hoffmannae Fain, 1996

Opserythraeus hoffmannae Fain (larval) Opserythraeus hoffmannae Fain, 1996: 253

Type locality: Rugege Forest, 2500 m Distribution: Rwanda Host: “from moss”

Genus Paraphanolophus Smiley (larval) Paraphanolophus Smiley, 1968: 20

Type species: Paraphanolophus metcalfei Smiley, 1968

Paraphanolophus halffteri Beron (larval) Paraphanolophus halffteri Beron, 1996: 25

Type locality: Teapa, Tabasco Distribution: Mexico (Tabasco)

Paraphanolophus metcalfei Smiley (larval) Paraphanolophus metcalfei Smiley, 1968: 20

Type locality: Belize Distribution: Belize Host: Saccharosydne saccharivora (Homoptera, Auchenorhyncha, Delphacidae)

Genus Podosmaridia Trägårdh (ad.) Podosmaridia Trägårdh, 1937: 12

Type species: Podosmaridia longipes Trägårdh, 1937

Podosmaridia longipes Trägårdh (ad.) Podosmaridia longipes Trägårdh, 1937: 13

Type locality: Urumchi, Sinjan – Uigur Autonomous Region of China, not Mongolia! Distribution: China (Sinjan-Uigur Autonomous Region)

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Genus Rainbowia Southcott (adult) Rhyncholophus Rainbow, 1906: 156 (in part) Erythraeus Womersley, 1934: 218 (adult. in part); Southcott, 1946b: 18-22 (ad.), p. 19, 22-25 (larva), in part; Meyer & Ryke, 1959: 306-307 (in part) Bochartia: Womersley, 1936: 120-121 (larva, in part)

Type species: Leptus imperator Hirst, 1928

Rainbowia celeripes (Rainbow)(ad.) Rhyncholophus celeripes Rainbow, 1906: 156 Erythraeus celeripes Womersley, 1934: 218 (in part); Southcott, 1946b: 20 Rainbowia celeripes (Rainbow): Southcott, 1961a: 470

Type locality: Enfield, New South Wales Distribution: Australia (N.S.W.)

Rainbowia imperator (Hirst) (ad. and larva) Leptus imperator Hirst, 1928: 570 Erythraeus celeripes Womersley, 1934: 218 (in part) Bochartia oudemansi: Womersley, 1936: 121 (larva) Erythraeus imperator: Southcott, 1946b: 20 (adult) Erythraeus oudemansi: Southcott, 1946b: 22 (larva) Rainbowia imperator (Hirst): Southcott, 1961a: 470

Type locality: Lucindale, South Australia Type locality of Bochartia oudemansi Womersley (larva): Adelaide, S.A. (under bark) Distribution: Australia (S.A.)

Genus Ramsayella Zhang (larval) Ramsayella Zhang, 2000: 36

Type species: Ramsayella rangitata Zhang, 2000

Ramsayella rangitata Zhang (larval) Ramsayella rangitata Zhang, 2000: 36

Type locality: Lawrence Valley, Hodder Valley (1000-1830 m) Distribution: New Zealand Host: Brachaspis sp. (Acrididae, Orthoptera)

Genus Taranakia Southcott (larval) Taranakia Southcott, 1988: 229

Type species: Taranakia lambi Southcott, 1988


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Taranakia lambi Southcott (larval) Taranakia lambi Southcott, 1988: 230; Clark, 2004: 19

Type locality: Manaia, Taranaki and Mayor Is., Owairaka, New Zealand Distribution: New Zealand Hosts: Melanchra mutans (Noctuidae, Lep.), Epyaxa rosearia (Geometridae, Lep.) and “phasmid”

Genus Tepoztlana Hoffmann et Mendez (ad.) Tepoztlana Hoffmann & Mendez, 1973: 131 Type species: Tepoztlana sandovali Hoffman et Mendez, 1973

Tepoztlana sandovali Hoffmann et Mendez (ad.) Tepoztlana sandovali Hoffmann & Mendez, 1973: 131

Type locality: Tepoztlan, Morelos, Mexico Distribution: Mexico

Subfamily Leptinae Billberg (sensu Southcott, 1961a) Leptides Billberg, 1820: 126 (in part) Leptidae Oudemans, 1941: 178 Leptinae Southcott, 1957: 98

Type genus: Leptus Latreille, 1796

Genus Leptus Latreille (ad. and larval) Leptus: Latreille, 1796: 177 Ritteria: Kramer, 1887: 228 (synonymized by Southcott, 1961a: 573) Achorolophus: Berlese, 1891: 2 (subgenus of Rhyncholophus) Erythraeus: Oudemans, 1903: 92 (part); non Latreille, 1806: 146 Atomus Banks, 1915: 39, nec Atomus Latreille, 1795, nec Atomus Augustson, 1940

Type species of Leptus Latreille (original designation): Acarus phalangii de Geer, 1778 Type species of Rhyncholophus Berlese: Trombidium trimaculatum Hermann, 1804 Type species of Achorolophus Berlese: Rhyncholophus nemorum C.L. Koch, 1836 Type species of Ritteria Kramer: Leptus nemorum (C.L. Koch) Schweizer, 1951

Subgenus Leptus Latreille (ad. and larval) Leptus: Latreille, 1796: 177

Type species (original designation): Acarus phalangii de Geer, 1778

Leptus (L.) abrafoicus Haitlinger (larval) Leptus (Leptus) abrafoicus: Haitlinger, 2006g: 113

Type locality: Abrafo Distribution: Ghana Host: “from herbaceous plants”

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Leptus (L.) absoloni Willmann (ad.) Leptus absoloni: Willmann, 1954: 242

Type locality: Sloupské údolí, Pusti Žleb, Moravia Distribution: Czech Rep.

Leptus (L.) acuminatus Shiba (ad.) Leptus acuminatus: Shiba, 1976: 175


Leptus (L.) adaminae Haitlinger (larval) Leptus adaminae: Haitlinger, 2004a: 150

Type locality: Foz do Iguacu Distribution: Brazil Host: from plants

Leptus (L.) addari Haitlinger (larval) Leptus addari Haitlinger, 1999: 61

Type locality: Bangkok Distribution: Thailand Host: “from plants”

Leptus (L.) admeti Haitlinger (larval) Leptus admeti: Haitlinger, 1998: 105

Type locality: Soekaranda, Sumatra Distribution: Sumatra (Indonesia) Host: Airathus ferialis (Fulgoridae, Homoptera)

Leptus (L.) aeolopae Chinniah et Mohanasundaram (larval) Leptus aeolopae Chinniah & Mohanasundaram, 1998: 1

Type locality: Tamil Nadu, Coimbatore Distribution: India (Tamil Nadu) Host: Aeolopus tamulus (Acrididae, Orthoptera)

Leptus (L.) agenori Haitlinger (larval) Leptus agenori : Haitlinger, 1999: 57

Type locality: Kota Bahru (Malaysia), Noparattara n. Krabi (Thailand) Distribution: Malaysia, Thailand Host: undetermined Orthoptera, on wings


Leptus (L.) aggoratus Haitlinger (larval) Leptus aggoratus: Haitlinger, 1990a: 29

Type locality: Zambia Distribution: Zambia Host: Tenebrionidae indet.

Leptus (L.) agilis (Canestrini)(ad.) Rhyncholophus agilis Canestrini, 1884: 2 Grognieria agilis: Oudemans, 1941: 179 Leptus agilis (Canestrini)

Type locality: Trentino, Porretta (Italy) Distribution: Italy

Leptus (L.) agrotis Southcott (larva, Dny, ad.) Leptus agrotis: Southcott, 1993: 1487

Type locality: Australian Capital Territory Distribution: Australia (ACT) Host: Agrotis infusa (Noctuidae, Lepidoptera)

Leptus (L.) akkus Haitlinger (larval) Leptus akkus: Haitlinger, 1990c: 46

Type locality: Spain Distribution: Spain Host: Akis sunci (Tenebrionidae, Coleoptera)

Leptus (L.) albertensis Southcott (larval) Leptus albertensis: Southcott, 1992a: 50

Type locality: Sunnydale, Lloydminster, Alberta Distribution: Canada (Alberta) Host: Hypocoena rufistrigata (Noctuidae, Lepidoptera)

Leptus (L.) alberti Haitlinger (larval) Leptus alberti: Haitlinger, 1991: 270

Type locality: Brazil Distribution: Brazil Host: Homophoeta personata (Chrysomelidae, Alticinae, Col.)

Leptus (L.) aldonae Haitlinger (larval) Leptus aldonae: Haitlinger, 1987g: 702

Type locality: Antananarivo Distribution: Madagascar Host: “collected from plants”.

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Leptus (L.) alkmenae Haitlinger (larval) Leptus alkmenae: Haitlinger, 1998: 103

Type locality: Uttar Pradesh Distribution: India Host: Cassida syrtica (Col., Chrysomelidae)

Leptus (L.) alopecurus Southcott (larval) Leptus alopecurus: Southcott, 1991: 738

Type locality: Queensland (Innisfail) Distribution: Australia Host: Cydistomyia doddi (Tabanidae, Diptera)

Leptus (L.) alvimordax Southcott (larval) Leptus alvimordax: Southcott, 1991: 734

Type locality: Queensland (Alma Bay, Kuranda) Distribution: Australia Host: Cydistomyia ? avida (Tabanidae, Diptera)

Leptus (L.) andae Haitlinger (larval) Leptus andae Haitlinger, 2003a: 136

Type locality: Rhodes (Gennadi) Distribution: Greece (Rhodes) Host: Orthoptera, Coleoptera (Tenebrionidae indet.)

Leptus (L.) annikae Haitlinger (larval) Leptus annikae Haitlinger, 2000c: 154

Type locality: “30 km east from Puerto Maldonado, at river Madre de Dios” Distribution: Peru Host: Crotonotus sp. (Pseudophyllidae, Orthoptera)

Leptus (L.) anomalus Southcott (larval) Leptus anomalus: Southcott, 1946b: 41; 1991: 748; Haitlinger, 1998: 107

Type locality: Glen Osmond, South Australia Distribution: Australia (S.A.) Host: Colibe bicarinatus (Col., Tenebrionidae)

Leptus (L.) ariel Southcott (larval) Leptus sp. : Wilson et al., 1987: 853 Leptus ariel: Southcott, 1989a: 124; 1992: 29; Haitlinger, 2000c: 156

Type locality: Guatemala Distribution: Guatemala, Peru


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Host: Apis mellifera (Apidae, Hymenoptera); unident. Eumolpinae (Chrysomelidae, Coleoptera)

Leptus (L.) asahinai Kawashima (larval) Leptus asahinai: Kawashima, 1958: 201; Treat, 1975: 218

Type locality: Tanashimachi, Kitatama-gun, Tokyo, Japan Distribution: Japan Host: Lymantria mathura aurora (Lymantriidae, Lepidoptera)

Leptus (L.) assagasicus Haitlinger (larval) Leptus assagasicus Haitlinger, 2001b: 474

Type locality: Assagasi n. Durban (S 29o39’, E 31o05’) Distribution: Republic of South Africa Host: “from plants in meadows”

Leptus (L.) astrubali Haitlinger (larval) Leptus astrubali Haitlinger, 1999a: 61; 2002a: 183; 2004b: 158

Type locality: Ayutthaya Distribution: India, Thailand, Myanmar (Burma) Host: “from plants”

Leptus (L.) atticolus Lawrence (larval) Leptus atticolus: Lawrence, 1940: 405

Type locality: Pietermaritzburg, Natal, South Africa Distribution: South Africa Host: Saitis sp. (Attidae, Araneae)

Leptus (L.) augusti Haitlinger (larval) Leptus augusti Haitlinger, 1999b: 54

Type locality: Vangvieng Distribution: Laos Host: Hypomeces squamosus (Curculionidae, Tanymecinae, Col.)

Leptus (L.) auliacus Haitlinger (larval) Leptus auliacus: Haitlinger, 1998: 98

Type locality: Dzambul (Kazakhstan) Distribution: Kazakhstan Host: Coleoptera, Tenebrionidae undet.

90


Leptus (L.) aureliani Fain et Elsen (larval) Leptus aureliani: Fain & Elsen, 1987: 107

Type locality: Butare, Rwanda Distribution: Rwanda Host: “dans le nid d’un pigeon Streptopelia semitorquata”.

Leptus (L.) bakeri Southcott (larval) Leptus bakeri: Southcott, 1992a: 126

Type locality: North Farm, Oktibbeha Co., Mississippi Distribution: USA (Mississippi) Host: Tettigidea lateralis, Paratettix cucullatus (Tetrigidae, Orthoptera)

Leptus (L.) balicus Haitlinger (larval) Leptus balicus: Haitlinger, 2006d: 227

Type locality: Sanur (Bali) Distribution: Indonesia (Bali) Host: from undetermined Orthoptera and from herbaceous plants

Leptus (L.) bankensis Southcott (larval) Leptus bankensis: Southcott, 1999: 222

Type locality: Banka Banka, Northern Territory Distribution: Australia (N. Territory) Host: “locust”(Orthoptera)

Leptus (L.) barmeedius Southcott (larval) Leptus barmeedius: Southcott, 1999: 226

Type locality: Loveday, South Australia Distribution: Australia (S. Australia) Host: Urodacus cf. yaschenkoi (Scorpionidae, Scorpiones)

Leptus (L.) bathypogonus Womersley (larval) Leptus bathypogonus: Womersley, 1934: 247; Feider, 1956: 150; Southcott, 1991: 727; 1999: 208

Type locality: Perth (West Australia) Distribution: Australia (West Australia, South Australia) Host: Bathypogonus [sic] for Bathypogon sp. (Diptera, Asilidae); grasshopper (Orthoptera)

Leptus (L.) batjallus Southcott (larval) Leptus batjallus: Southcott, 1999: 242

Type locality: Fraser Island, Queensland Distribution: Australia (Queensland) Host: Nethra annulipes (Heteroptera, Gelastocoridae)


91

Leptus (L.) batoricus Haitlinger (larval) Leptus batoricus: Haitlinger, 1998: 99

Type locality: Mongolia, unknown loc. Distribution: Mongolia Hosts: Pimelia cephalotes (Col., Tenebrionidae)

Leptus (L.) baudini Southcott (larval) Leptus baudini: Southcott, 1999: 137

Type locality: Kangaroo Island, S.A. Distribution: Australia (South Australia) Host: Urodacus manicatus (Scorpionidae, Scorpiones)

Leptus (L.) belicolus Southcott (larval) Leptus belicolus: Southcott, 1993: 1517

Type locality: South Australia Distribution: Australia (S. Australia) Host: Belus filiformis (Col., Belidae)

Leptus (L.) benzaliensis Fain et Elsen (larval) Leptus benzaliensis: Fain & Elsen, 1972: 66

Type locality: Congo (Kinshassa) Distribution: Ivory Coast, Kenya, Congo (Kinshassa) Hosts: Glossina spp. (Glossinidae, Diptera), Scarabaeidae (Coleoptera)

Leptus (L.) berlesei Oudemans (ad.) Leptus berlesei: Oudemans, 1913b: 19 (nom. nov. pro Rhyncholophus nemorum var. vertex Berlese, 1887 nec vertex Kramer) Achorolophus vertex Oudemans, 1913a: 16 (nec vertex Kramer)

Type locality: Italy Distribution: Holland, Italy

Leptus (L.) beroni Fain (larval) Achorolophus ignotus Oudemans, 1912: 156 (part); non Erythraeus ignotus Oudemans, 1903: 92; 1904: 94 Leptus ignotus: Beron, 1975: 57 (part, non Oudemans, 1903); Haitlinger, 1987: 340 (part) Leptus beroni Fain, 1992: 108; Wohltmann, 1995: 447; Haitlinger, 2002e: 149; 2003d: 4; 2007b: 61 Leptus holmiae: Southcott, 1992: 60; Stasiov, 2003: 23

Type locality of Leptus beroni Fain: Forêt de Soignes, near Bruxelles, Belgium Type locality of Leptus holmiae Southcott: Stockholm, Sweden Distribution: Belgium, Bulgaria, Denmark, France, Germany, Great Britain, Holland, Hungary, Iceland, Ireland, Norway, Poland, Russia, Slovakia, Slovenia, Sweden, Switzerland

92


Hosts: Mitopus morio, Opilio ruzickai, Megabunus diadema, Oligolophus hanseni (Phalangiidae, Opiliones), Homoptera, Diptera, Lepidoptera

Leptus (L.) bertoldi Haitlinger (larval) Leptus bertoldi: Haitlinger, 1990: 166

Type loc.: North Ghana, Daboya Distribution: Ghana Host: Tenebrionidae undet. (Coleoptera)

Leptus (L.) bicristatus Fain et Elsen (larval) Leptus bicristatus: Fain & Elsen, 1987: 115; Cokendolpher, 1993: 134

Type locality: Chowo Rocks, Nyika Plateau, Malawi Distribution: Malawi Host: Opiliones (Cristina lettowi, Phalangiidae)

Leptus (L.) boggohoranus Haitlinger (larval) Leptus boggohoranus: Haitlinger, 1990b: 50; 1993: 1481

Type locality: New Guinea Distribution: New Guinea Host: Cicindelidae (Coleoptera), ? Curculionidae

Leptus (L.) bogoriacus Haitlinger (larval) Leptus bogoriacus Haitlinger, 2001b: 474

Type locality: Bogoria (N 0o25’, E 36o25’) Distribution: Kenya Host: Tenebrionidae indet.

Leptus (L.) brachypodos Zheng (larval) Leptus brachypodos: Zheng, 1996: 232

Type locality: Mt. Hupingshan, Shimen Co., Hunan Prov. Distribution: China (Hunan Prov.) Host: Insecta indet.

Leptus (L.) cabareticus Haitlinger (larval) Leptus cabareticus Haitlinger, 2004g: 232

Type locality: Cabarete Distribution: Dominican Republic Host: from plants


Leptus (L.) calcar Southcott (larval) Leptus calcar: Southcott, 1999: 196

Type locality: Townswille, Queensland Distribution: Australia (Queensland), Papua New Guinea Hosts: Heteroptermis obscurella, Stenocantops vitripennis, etc. (Acrididae, Orthoptera)

Leptus (L.) calidus Shiba (larval) Leptus calidus: Shiba, 1976: 178

Type locality: Pasoh Forest, Malaya Distribution: Malaysia Host: unknown

Leptus (L.) californicus Southcott (larval) Leptus californicus: Southcott, 1992: 43

Type locality: Smith Creek, Mt. Hamilton, Santa Clara Co., California Distribution: U.S.A. (California) Host: Dissosteira pictipennis (Acrididae, Orthoptera)

Leptus (L.) calix Southcott (larval) Leptus calix: Southcott, 1992a: 91

Type locality: Tyringham, Massachusetts Distribution: USA (Massachusetts) Host: Ochropleura plecta (Noctuidae, Lepidoptera)

Leptus (L.) calvatus Willmann (ad.) Leptus calvatus: Willmann, 1951c: 151; Franz, 1954: 382

Type locality: Ebreichsdorf, Zurndorf, Austria Distribution: Austria

Leptus (L.) calvescens (Berlese)(ad.) Rhyncholophus calvescens: Berlese, 1888: 184; Canestrini, 1898: 193 Achorolophus calvescens: Oudemans, 1906b: 108 Abrolophus calvescens: Berlese, 1913: 131 Leptus calvescens (Berlese)

Type locality: Brasil Distribution: Brasil, Paraguay, Argentina; ? New Guinea

Leptus (L.) cameronensis Shiba (larval) Leptus cameronensis: Shiba, 1976: 180


93

94


Leptus (L.) carduus Southcott (larval) Leptus carduus: Southcott, 1999: 260

Type locality: Glen Osmond, South Australia Distribution: Australia (S. Australia) Host: Urodacus manicatus (Scorpionidae, Scorpiones)

Leptus (L.) carpenteri Fain et Elsen (larval) Leptus carpenteri: Fain & Elsen, 1972: 62

Type locality: Lukunga near Kinshasa, Zaire Distribution: DR Congo (Zaire) Host: Glossina fuscipes quanzensis (Glossinidae, Diptera)

Leptus (L.) cavernicola Fain et Elsen (larval) Leptus cavernicola: Fain & Elsen, 1987: 116

Type locality: lava cave at Ruhengeri, Rwanda Distribution: Rwanda Host: Nematocera (Diptera)

Leptus (L.) cerambycius Southcott (larval) Leptus cerambycius: Southcott, 1993: 1502; 1999: 222

Type locality: Glen Osmond, South Australia Distribution: Australia (S. Australia) Host: ex Cerambycidae (Col.) and caught free-living

Leptus (L.) cercopius Southcott (larval) Leptus cercopius: Southcott, 1992a: 73

Type locality: Daufin Island, Alabama Distribution: U.S.A. (Alabama) Host: Lepyronia quadrangularis (Cercopidae, Homoptera)

Leptus (L.) charanyca Fain (larval) Leptus charanyca: Fain, 1991: 148

Type locality: Linkebeek (Bruxelles) Distribution: Belgium Host: Charanyca trigrammica (Noctuidae, Lepidoptera)

Leptus (L.) charon Southcott (larval) Leptus charon: Southcott, 1991: 752; 1993: 1525; 1999: 238; Seeman & Miller, 2002: 49

Type locality: Australian Capital Territory (Hall) Distribution: Australia (ACT, NSW, SA, Tasmania)


95

Host: Calliphora stygia (Calliphoridae, Diptera); Cercophonius squama (Bothriuridae, Scorpiones); Isopeda sp. (Sparassidae, Araneae), Anthela sp. (Anthelidae, Lepidoptera), Coleoptera

Leptus (L.) cheesmanae Southcott (larval) Leptus cheesmanae: Southcott, 1999:130

Type locality: Torricelli Mts., 200-1000 ft. (PNG), paratype from Wondecla (Queensland) Distribution: Papua New Guinea, Australia (Queensland) Host: cricket, phasmid

Leptus (L.) chelonethus Womersley (larval) Leptus chelonethus: Womersley, 1934: 246; Southcott, 1991: 721 Leptus chelonchus (Wom.): Feider, 1956: 150

Type locality: Rottnest Island, West Australia Distribution: West Australia (Australia) Host: Pseudoscorpion

Leptus (L.) chibai Shiba (ad.) Leptus chibai: Shiba, 1976: 173


Leptus (L.) chrotogonae Chinniah et Mohanasundaram (larval) Leptus chrotogonae Chinniah & Mohanasundaram, 1998: 3

Type locality: Tamil Nadu, Coimbatore Distribution: India (Tamil Nadu) Host: Chrotogonus sp. (Pyrgomorphidae, Orthoptera)

Leptus (L.) clarki Southcott (larval) Leptus clarki: Southcott, 1989b: 243; 1992a: 75

Type locality: Howe, Butt Co., Idaho Distribution: U.S.A. (Idaho) Host: Pogonomyrmex salicinus (Formicidae, Hymenoptera)

Leptus (L.) clavatus Southcott (larval) Leptus clavatus: Southcott, 1999:170

Type locality: Myponga, South Australia Distribution: Australia (S. Australia) Host: under bark of Eucalyptus sp.

96


Leptus (L.) clelandi Southcott (larval) Leptus clelandi: Southcott, 1999:167

Type locality: Encounter Bay, South Australia Distribution: Australia (S. Australia) Host: cicada (Homoptera)

Leptus (L.) clethrionomydis Haitlinger (larval) Leptus clethrionomydis: Haitlinger, 1987d: 345; Southcott, 1992: 504; Gabryś, 1988: 23; Gabryś & Mąkol, 1991: 709

Type locality: Wroclaw-Redzin, Poland Distribution: Hungary, Poland Host: Clethrionomys glareolus (Rodentia, Mammalia, chance host)

Leptus (L.) coccineus (Scopoli) (larval) Pediculus coccineus: Scopoli, 1763: 386

Leptus coccineus (Scopoli) – auct. Type locality: Slovenia (Ljubljana) Distribution: Slovenia

Leptus (L.) coloanensis Haitlinger (larval) Leptus coloanensis Haitlinger, 2006a: 92

Type locality: Coloane Island, Macao Distribution: China Host: from herbaceous plants

Leptus (L.) comosus Southcott (larval) Leptus comosus: Southcott, 1991: 743

Type locality: Adelaide (South Australia) Distribution: Australia (South Australia) Host: Stomoxys calcitrans (Stomoxydae, Diptera)

Leptus (L.) cooremani Fain (larval) Leptus cooremani: Fain, 1992: 107

Type locality: Marche-en-Famenne, Belgium Distribution: Belgium Host: Lepidoptera indet.

Leptus (L.) coreophilus Southcott (larval) Leptus coreophilus: Southcott, 1992a: 98

Type locality: Fontainebleau St., Pk. St. Tammany Parish, Louisiana Distribution: U.S.A. (Louisiana) Host: Acanthocephala terminalis (Coreidae, Heteroptera)


97

Leptus (L.) cultellus Southcott (larval) Leptus cultellus: Southcott, 1999: 216

Type locality: Blinman, South Australia Distribution: Australia (S. Australia) Host: Chortoicetes terminifera (Acrididae, Orthoptera)

Leptus (L.) curtipes Schweizer (ad.) Leptus curtipes Schweizer, 1951: 145; Franz, 1954: 382; Cooreman, 1955c: 15; Mihelčič, 1958a: 280; 1958b: 45; Schweizer & Bader, 1963: 308

Type locality: Blais, 2410 m; Murtaröl, 2587 m, Swiss Alps Distribution: Austria, Belgium, Italy, Spain, Switzerland

Leptus (L.) cyrili Haitlinger (larval) Leptus cyrili Haitlinger, 1991: 270

Type locality: Sao Paulo Distribution: Brazil Host: Lycidae (Col.)

Leptus (L.) dalicus Haitlinger (larval) Leptus dalicus Haitlinger, 2006h: 85

Type locality: Dali Distribution: Ethiopia Host: undetermined small Carabidae (Col.)

Leptus (L.) debeauforti (Oudemans)(larval) Erythraeus debeauforti: Oudemans, 1905a: 217 Erythraeus de Beauforti: Oudemans, 1906b: 108 Achorolophus debeauforti (Oudemans): Oudemans, 1905c: 9; 1912: 181 Leptus debeauforti (Oudemans): Buitendijk, 1945: 347

Type locality: Humboldt Bay, West New Guinea Distribution: Irian Jaja, or West New Guinea (Indonesia) Host: Formicidae (Hymenoptera)

Leptus (L.) dinekaicus Haitlinger (larval) Leptus dinekaicus Haitlinger, 2006h: 83

Type locality: Dineka n. Turmi, South Ethiopia Distribution: Ethiopia Host: from plants

98


Leptus (L.) diversus Mihelčič (ad.) Leptus diversus Mihelčič, 1958a: 281

Type locality: El Escorial, 50 km N Madrid, Spain Distribution: Spain

Leptus (L.) dolichopodos Zheng (larval) Leptus dolichopodos: Zheng, 1996: 237

Type locality: Hunan Distribution: China (Hunan) Host: unknown

Leptus (L.) draco Southcott (larval) Leptus draco: Southcott, 1984: 351; 1991: 719; 1993: 1475; 1999: 120

Type locality: Buso (New Guinea) Distribution: Papua New Guinea Hosts: Neopromachus sp. (Phasmatidae, Phasmida), Odonata, Orthoptera, Homoptera (Cicadidae)

Leptus (L.) droozi Southcott (larval) Leptus droozi: Southcott, 1992a: 46

Type locality: Olustree, Florida Distribution: U.S.A. (Florida) Host: Epiblema strenuana (Tortricidae, Lepidoptera)

Leptus (L.) dubius (Paoli) (larval) Achorolophus dubius: Paoli, 1937: 185 Leptus dubius: Southcott; 1961a: 520, 1992a: 17; Beron, 1975: 45

Type locality: Foggia Distribution: Italy Host: Prionotropis hystrix appula (Pamphagidae), Euchorthippus declivus, Stauroderus sp., Dociostaurus maroccanus, Oedipoda miniata, Calliptamus barbarus (Acrididae, Orthoptera), Cytherea obscura (Bombyliidae, Diptera)

Leptus (L.) elderi Southcott (larval) Leptus elderi: Southcott, 1999: 175

Type locality: Rockhampton, Queensland Distribution: Australia (Queensland) Host: Heteropsylla cubana (Psyllidae, Homoptera)


99

Leptus (L.) elminus Haitlinger (larval) Leptus elminus Haitlinger, 2006g: 110

Type locality: Elmina Distribution: Ghana

Leptus (L.) errabundus Gabryś (ad.) Leptus villosus: Mihelčič 1964: 296 (non Achorolophus villosus Berlese, 1910) Leptus errabundus nom.nov.: Gabryś, 2000c: 65

Type locality: “St. Veit im Jauntal, Kärnten” Distribution: Austria

Leptus eslamizadehi Saboori (larval) Leptus eslamizadehi Saboori, 2002a: 550

Type locality: Uromieh, 1312,5 m Distribution: Iran Host: Unknown

Leptus (L.) esmailii Saboori et Ostovan (larval) Leptus esmailii: Saboori & Ostovan, 2000: 143

Type locality: Kermanshah Distribution: Iran Host: Eurygaster integriceps (Scutelleridae, Heteroptera)

Leptus (L.) eupodes (Berlese) (ad.) Abrolophus nemorum var. eupodes: Berlese, 1910: 352

Type locality: Risignano (Etruria) Distribution: Italy

Leptus (L.) faini Southcott (larval) Leptus faini: Southcott, 1993: 1531; 1999: 247

Type locality: New South Wales Distribution: Australia (NSW, S. Australia) Hosts: Psapharus infaustus (Col., Curculionidae); Isopeda sp., Holconia insignis (Sparassidae, Araneae)

Leptus (L.) fathipeuri Haitlinger et Saboori (larval) Leptus fathipeuri: Haitlinger & Saboori, 1996: 123

Type locality: Tabriz Distribution: Iran Host: from plants

100


Leptus (L.) filipinae Haitlinger (larval) Leptus filipinae Haitlinger, 2000a: 132

Type locality: Orosi (Costa Rica), Agua (Mexico), Belize Distribution: Costa Rica, Mexico, Belize Host: Lampyridae (Col.), undeterimined Heteroptera and from plants

Leptus (L.) fisheri Southcott (larval) Leptus fisheri: Southcott, 1999: 213

Type locality: Arkaba Creek, Flinders Range, South Australia Distribution: Australia (S. Australia) Host: Stenocotis depressa (Cicadellidae, Homoptera)

Leptus (L.) flindersi Southcott (larval) Leptus flindersi: Southcott, 1999: 144

Type locality: Glen Osmond, South Australia Distribution: Australia (S. Australia) Host: Calolampra irrorata (Blaberidae, Blattodea)

Leptus (L.) floridanus (Banks)(ad.) Rhyncolophus floridanus: Banks, 1896: 74 Rhyncholophus floridanus: Banks, 1907: 599 Leptus floridanus (Banks): Welbourn & Jennings, 1991: 568

Type locality: Punta Gorda, Florida Distribution: U.S.A. (Florida)

Leptus (L.) foliatus Southcott (larval) Leptus foliatus: Southcott, 1999: 181

Type locality: Alice Springs, Australia Distribution: Australia (Northern Territory) Host: locust (Acrididae, Orthoptera)

Leptus (L.) fortei Southcott (larval) Leptus fortei: Southcott, 1991: 755

Type locality: Mt Clarence, Albany (Western Australia) Distribution: Australia Host: Asilidae (Diptera)

Leptus (L.) fozicus Haitlinger (larval) Leptus fozicus: Haitlinger, 2004a: 152



101

Leptus (L.) francesi Southcott (larval) Leptus francesi Southcott, 1991: 719

Type locality: Brown Range, Inarlinga, Queensland Distribution: Australia (Queensland) Host: Aedes vigilax (Culicidae, Diptera), “tabanid marchfly” (Diptera)

Leptus (L.) furibundus Gabryś (ad.) Leptus diversus Mihelčič, 1958b: 46 (non Leptus diversus Mihelčič, 1958a) Leptus furibundus: Gabryś, 2000c: 65 (nom. nov. pro L. diversus Mihelčič, 1958b)

Type locality: Sierra Nevada, Veleta, 2760 m Distribution: Spain

Leptus (L.) gagrellae (Oudemans) (larval) Achorolophus gagrellae: Oudemans, 1910: 73; 1912: 178 Leptus gagrellae (Oudemans): Vitzthum, 1926: 159; Buitendijk, 1945: 347; Beron, 1975: 69

Type locality: Tjibodas, Java Distribution: Indonesia (Java) Host: Gagrella sp. (Phalangiidae, Opiliones)

Leptus (L.) gagzoi (Oudemans) (larval) Achorolophus gagzoi Oudemans, 1910e: 108; 1912: 161 Leptus gagzoi (Oudemans): Buitendijk, 1945: 347; Turk, 1948: 92; Feider, 1956: 150; Beron, 1975: 46

Type locality: Colon, Panama Distribution: Panama, Trinidad & Tobago Host: Orthoptera, Tettigoniidae

Leptus (L.) galerucae Feider (larval) Leptus galerucae Feider, 1967: 41; Southcott, 1992: 24

Type locality: Jassy (Romania) Distribution: Romania Host: Galeruca tanaceti (Chrysomelidae, Coleoptera)

Leptus (L.) gauphalus Haitlinger (larval) Leptus gauphalus: Haitlinger, 1990: 48; Southcott, 1993: 1531

Type localiy: Australia Distribution: Australia Host: Hypocilibe impunctata [=Nyctozoilus impunctatus] (Tenebrionidae, Coleoptera)

Leptus (L.) gennadicus Haitlinger (larval) Leptus gennadicus: Haitlinger, 2003a: 140

Type locality: Rhodes, Gennadi

102


Distribution: Greece (Rhodes) Host: Coleoptera, Buprestidae indet.

Leptus (L.) georgeae Southcott (larval) Leptus georgeae: Southcott, 1993: 1510

Type locality: South Australia Distribution: Australia (S. Australia) Host: Geometridae (Lep.)

Leptus (L.) ghiradellae Southcott (larval and deutonymph) Leptus ghiradellae: Southcott, 1992a: 105

Type locality: Tyringham, Massachusetts Distribution: USA (Massachusetts) Host: Leucania commoides, L. pseudargyria (Noctuidae, Lepidoptera)

Leptus (L.) gifuensis Kawashima (larval) Leptus gifuensis Kawashima, 1958: 199; Shiba, 1969: 103

Type locality: Kinkazan, Gifu City, Gifu Pref., Japan Distribution: Japan Host: Lycosa sp. (Lycosidae, Araneae)

Leptus (L.) giganticus Khot (ad.) Leptus giganticus: Khot, 1964a: 682

Type locality: Rishikesh, Uttar Pradesh Distribution: India (Uttar Pradesh)

Leptus (L.) glossinarum Fain et Elsen (larval) Leptus glossinarum: Fain & Elsen, 1972: 64

Type locality: Benzali Distribution: D.R. Congo (Zaire) Host: Glossina fuscipes quanzensis (Glossinidae, Diptera)

Leptus (L.) gracilipes (Berlese) (ad.) Abrolophus gracilipes Berlese, 1910: 352; 1913: 137 Leptus gracilipes (Berlese)

Loc. typ.: Valparaiso Distribution: Chile

Leptus (L.) groenlandicus (Trägårdh)(larval) Erythraeus groenlandicus Trägårdh, 1904b: 58 Leptus groenlandicus Southcott, 1992: 145


Type locality: Egedesminde Distribution: Greenland Host: ?

Leptus (L.) grossi Southcott (larval) Leptus grossi: Southcott, 1999: 162

Type locality: Hindmarsh Falls, South Australia Distribution: Australia (S. Australia) Host: extracted from moss

Leptus (L.) guilinicus Haitlinger (larval) Leptus guilinicus: Haitlinger, 2006a: 93

Type locality: Yangshuo n. Guilin Distribution: China Host: from herbaceous plants

Leptus (L.) guus Haitlinger (larval) Leptus guus: Haitlinger, 1990b: 51

Type locality: Turkmenia Distribution: Turkmenia Host: Trigonoscelis sp. (Coleoptera, Tenebrionidae)

Leptus (L.) gyas Fain et D’Amico (larval) Leptus gyas: Fain & D’Amico, 1997: 40

Type locality: Gabas, French Pyrenees Distribution: France Host: Gyas titanus (Phalangiidae, Opiliones)

Leptus (L.) halli Southcott (larval) Leptus halli: Southcott, 1993: 1537

Type locality: Australia Distribution: Australia Host: Cubicorhynchus maculatus (Col., Curculionidae)

Leptus (L.) hammameticus Haitlinger (larval) Leptus hammameticus: Haitlinger, 1998: 102

Type locality: Bir Boa, El Margaza n. Hammamet, North Tunisia, Tamersa, Sahara Distribution: Tunisia Host: Tenebrionidae (Col.)

103

104


Leptus (L.) heleus Southcott (larval) Leptus heleus: Southcott, 1993: 1525

Type locality: Northern Territory Distribution: Australia (N. Territory) Host: Helea squamosus (Coleoptera: Tenebrionidae)

Leptus (L.) hidakai Kawashima (larval) Leptus hidakai: Kawashima, 1958: 202; Baker & Selden, 1997: 183

Type locality: Kasaya-gun, Fukuoka Pref., Japan Distribution: Japan, Singapore Hosts: Opilio pentaspinulatus (Phalangiidae, Opiliones), Chiracanthium sp. (Clubionidae, Araneae), Miagrammopes singaporensis (Uloboridae, Araneae)

Leptus (L.) hirtipes (Berlese)(Ny) Abrolophus hirtipes: Berlese, 1910: 352; 1913: 131

Leptus hirtipes (Berlese): Vitzthum, 1926: 164 Type locality: Depok, Java Distribution: Java, “Prinzen-Insel” (Indonesia)

Leptus (L.) hitchcocki Southcott (larval) Leptus hitchcocki: Southcott, 1999: 265

Type locality: Finke River, Glen Helen, 20 mi. N of Hermannsburg Distribution: Australia (Northern Territory) Host: Urodacus sp. (Scorpionidae, Scorpiones)

Leptus (L.) holgeri Haitlinger (larval) Leptus holgeri Haitlinger, 1999b: 57

Type locality: Vangvieng Distribution: Laos Host: undetermined Orthoptera

Leptus (L.) horiacus Haitlinger (larval) Leptus horiacus: Haitlinger, 1994c: 140

Type locality: Syria Distribution: Syria Host: Adesmia carinata (Tenebrionidae, Coleoptera)

Leptus (L.) hospeticus Haitlinger (larval) Leptus hospeticus: Haitlinger, 2002a: 177

Type locality: Hospet, South India Distribution: India Host: Oedaleus abruptus (Acrididae, Orthoptera)


105

Leptus (L.) hozumii Shiba (larval) Leptus hozumii Shiba, 1976: 180


Leptus (L.) hringuri Haitlinger (larval) Leptus hringuri Haitlinger, 2000c: 154

Type locality: Machu Picchu, 2500 m Distribution: Peru Host: Naupotus sp. (Coleoptera: Curculionidae: Brachyderinae)

Leptus (L.) hupingshanicus Zheng (larval) Leptus hupingshanicus: Zheng, 1996b: 229

Type locality: Mt. Hupingshan, Shimen Co., Hunan Prov. Distribution: China (Hunan Prov.) Host: Insecta indet.

Leptus (L.) ignotus (Oudemans)(larval) Erythraeus ignotus: Oudemans, 1903: 92; 1904: 94; Dudich et al., 1940: 55 Achorolophus ignotus (Oudemans): Oudemans, 1905c: 9; 1911: 19; 1912: 156; Bruyant, 1911: 258; Sellnick, 1940: 37; Starkoff & Starkoff, 1950: 96; Schweizer, 1951: 164; Lindroth et al., 1973: 131 Leptus ignotus (Oudemans): Sellnick, 1958: 43; Haitlinger, 1987d: 340; 2002e: 149; 2007b: 62; Gabrys & Mąkol, 1991: 709; Fain & Baugnée, 1996: 22 ? Erythraeus groenlandicus Trägårdh, 1904b: 58 Leptus zbelutkaicus Haitlinger, 1987d: 342

Type locality of Erythraeus ignotus Oudemans: Borkum, Holland Type locality of Erythraeus groenlandicus Trägårdh: Egedesminde, Greenland Type locality of Leptus zbelutkaicus Haitlinger: Nowa Zbelutka, Kielce prov., Poland Distribution: Belgium, France, Great Britain, ?Greenland, Holland, Hungary, Iceland, Poland, Sweden, Switzerland, U.S.A. Hosts: Lepidoptera, Opiliones

Leptus (L.) ignotus danelli Southcott (larval) Leptus ignotus danelli: Southcott, 1992a: 37

Type locality: Tyringham, Massachusetts Distribution: U.S.A. (Massachusetts) Host: Hyppa xylinoides, Simyra henrici, Leucania linda (Noctuidae, Lepidoptera)

106


Leptus (L.) iguacuicus Haitlinger (larval) Leptus iguacuicus: Haitlinger, 2004a: 147


Leptus (L.) ilzae Haitlinger (larval) Leptus ilzae Haitlinger, 1999b: 52

Type locality: Vangvieng Distribution: Laos Host: Hypomeces squamosus Herbst (Curculionidae, Tanymecinae, Col.)

Leptus (L.) incertus Gabryś (ad.) Leptus calvatus Mihelčič, 1958b:46 (non Leptus calvatus Willmann, 1951) Leptus incertus Gabryś, 2000c: 65

Type locality of Leptus calvatus Mihelčič, 1958: Veleta, Sierra Nevada, 2860 m Distribution: Spain

Leptus (L.) indianensis Fain, Gummer et Whitaker (larval) Leptus indianensis: Fain, Gummer & Whitaker, 1987: 139; Southcott, 1992: 23; Cokendolpher, 1993: 134; McAloon & Durden, 2000: 561

Type locality: Crawfordsville, Montgomery Co., Indiana Distribution: U.S.A. (Indiana, Tennessee, Illinois, Virginia) Hosts: Leiobunum ventricosum, L. nigripes, L. speciosum, L. calcar, L. longipes and other Leiobunum spp. (Phalangiidae, Opiliones)

Leptus (L.) indicus Khot (ad.) Leptus indicus: Khot, 1964a: 682


Leptus (L.) intermedius Meyer et Ryke (ad.) Leptus intermedius: Meyer & Ryke, 1959: 317; Meyer & Ueckermann, 1989: 20

Type locality: Bathurst Distribution: South Africa

Leptus (L.) japonicus Kawashima (larval) Leptus japonicus: Kawashima, 1958: 205

Type locality: Cape Sata, Kimotsuki-gun, Kagoshima Pref., Japan Distribution: Japan Hosts: Coleoptera, Mecoptera, Hymenoptera, Homoptera, Orthoptera, Diptera


107

Leptus (L.) jenseni Southcott (larval) Leptus jenseni: Southcott, 1993: 1514

Type locality: Queensland Distribution: Australia (Queensland) Host: Phalidura sp. (Coleoptera: Curculionidae, Amycterinae)

Leptus (L.) jocquei Fain et Elsen (larval) Leptus jocquei: Fain & Elsen, 1987: 113; Cokendolpher, 1993: 134

Type locality: Dembo, [2200 m], Nyika Plateau, Malawi Distribution: Malawi Host: Opiliones (Cristina lettowi, Phalangiidae)

Leptus (L.) josifovi Beron (larval) Leptus josifovi: Beron, 1975: 49; Southcott, 1992: 18; Haitlinger, 2004d: 145; 2006b: 108; 2006e: 106; 2007a: 74; 2007b: 62; 2007d: 530 Leptus (Leptus) josifovi Beron: Haitlinger, 2006c: ; 2006f: 38

Type locality: Ropotamo Valley (Bulgaria) Distribution: Bosnia and Herzegovina, Bulgaria, Croatia, France (Corsica), Greece (Samos), Italy (Sardinia), Portugal, Switzerland Host: Phyllomorpha laciniata (Coreidae, Heteroptera); Oedipoda caerulescens, Oedaleus decorus, Calliptamus italicus (Acrididae) and other Orthoptera

Leptus (L.) kalaallus Southcott (larval) Leptus kalaallus: Southcott, 1992: 82

Type locality: Ikia, Greenland Distribution: Greenland Host: Mitopus morio (Phalangiidae, Opiliones)

Leptus (L.) kamalii Karimi Iravanlou et Saboori (larval) Leptus kamalii: Karimi Iravanlou & Saboori, 2001: 165

Type locality: Varamin Distribution: Iran Host: Dociostaurus hauensteini (Acrididae, Orthoptera)

Leptus (L.) karachiensis Anwarullah et Ahsan (larval) Leptus karachiensis Anwarullah & Ahsan, 1970: 405

Type locality: Karachi Distribution: Pakistan

108


Leptus (L.) killingtoni Turk (larval) Leptus (Achorolophus) killingtoni: Turk, 1945a: 138; Killington & Bathe, 1946: 257 Leptus killingtoni: Turk, 1953: 26; Southcott, 1992: 54; Haitlinger, 2006c Leptus (Leptus) killingtoni: Haitlinger, 2006e: 106

Type locality: Alderney Heath on the northern outskirts of Parkstone, Dorset. Distribution: Azores, Great Britain, Spain Host: Odonata, Coenagrionidae (Pyrrhosoma nymphula, Enallagma cyathigerum, Coenagrion puella, etc.), Lepidoptera (Pseudaletia unipunctata, Noctuidae)

Leptus (L.) korematus Southcott (larval) Leptus korematus: Southcott, 1999: 233

Type locality: Pilgangoora Well, Port Hedland, Western Australia Distribution: Australia (W. Australia, S. Australia, Northern Territory) Host: Urodacus varians, U. hartmeyeri (Scorpionidae, Scorpiones), Lychas sp. (Buthidae, Scorpiones)

Leptus (L.) kuroshimaensis Kato et Kitahara (larval) Leptus kuroshimaensis: Kato & Kitahara, 1958: 171

Type locality: Kuroshima Distribution: Japan (Kuroshima) Host: Tabanidae (Diptera)

Leptus (L.) kyushuensis Ishii (larval) Leptus kyushuensis: Ishii, 1953: 160

Type locality: Kyushu Distribution: Japan (Kyushu) Hosts: Cicadidae (Homoptera) and Tabanidae (Diptera)

Leptus (L.) laplandicus Southcott (larval) Erythraeus phalangioides var. gracilipes: Trägårdh, 1910: 485; non Erythraeus phalangoides (de Geer, 1778: 134); non Erythraeus gracilipes (Kramer, 1897: 77); non Achorolophus gracilipes Oudemans, 1910: 48; 1912: 168 (= Leptus oudemansi Southcott, 1992a: 88, as nom. nov.) Achorolophus ignotus Oudemans, 1912: 158 (part) Leptus laplandicus Southcott, 1992a: 88

Type locality: Lapland Distribution: Sweden Host: under stones and in moss

Leptus (L.) laviniacus Haitlinger (larval) Leptus laviniacus: Haitlinger, 2002a: 180; 2004b: 158

Type locality: Mount Lavinia (Sri Lanka) and other places in Sri Lanka and India Distribution: India, Sri Lanka, Cambodia Host: Catantops innotabilis, Oxya nitidula (Catantopinae, Orthoptera; Acrididae indet.)


109

Leptus (L.) leleupi Fain et Elsen (larval) Leptus leleupi: Fain & Elsen, 1987: 115

Type locality: Uluguru, 1300 m, Tanzania Distribution: Tanzania Host: Oreoparnus sp. (Staphylinidae, Pselaphinae, Coleoptera)

Leptus (L.) lighti Southcott (larval) Leptus lighti: Southcott, 1999: 190

Type locality: Glen Osmond, South Australia Distribution: Australia (South Australia) Host: Chortoicetes terminifera (Acrididae, Orthoptera)

Leptus (L.) lomani (Oudemans) (larval) Erythraeus lomani: Oudemans, 1902b: 39; 1903: 141 Achorolophus lomani (Oudemans): Oudemans, 1905: 19; 1911: 126; 1912: 175 Leptus lomani (Oudemans): Feider, 1956: 150; Beron, 1975: 69; Cokendolpher, 1993: 134

Type locality: Chile, t.l. restr. is Corral, 39° 53’ S, 73° 25’ W, Valdivia (Cokendolpher, 1993) Distribution: Chile Host: Discocyrtus funestus (Gonyleptidae, Opiliones)

Leptus (L.) longipes Willmann (ad.) Leptus longipes: Willmann, 1951c: 152; Franz, 1954: 382

Type locality: Zurndorf, Austria Distribution: Austria

Leptus (L.) longipilis (Berlese)(ad.) Abrolophus longipilis Berlese, 1910: 352; 1913: 131 Leptus longipilis (Berlese): Willmann, 1951c: 152; Franz, 1954: 382; Gabrys & Mąkol, 1994: 128; 1995: 383; 1996: 488; Haitlinger, 2002e: 148

Type locality: Kissingen, Germany and Valombrosa, Italy Distribution: Austria, Germany, Italy, Poland

Leptus (L.) longispinosus (Berlese) Leptus longispinosus (Berlese): Willmann, 1954: 242

Type locality: Czech Rep. Distribution: Czech Rep.

Leptus (L.) lorarius Southcott (larval) Leptus lorarius: Southcott, 1999: 120

Type locality: Luain, Papua New Guinea Distribution: Papua New Guinea Host: “caught free”

110


Leptus (L.) lovaniensis Fain et Elsen (larval) Leptus lovaniensis: Fain & Elsen, 1987: 114

Type locality: Kimuenza near Kinshasa, D.R. Congo Distribution: D.R. Congo (Zaire) Host: Scarabaeidae (Coleoptera)

Leptus (L.) machilidis Southcott (larval) Leptus machilidis: Southcott, 1992a: 121; Haitlinger, 2002e: 149

Type locality: Kletno nr. Stronie Slaskie (voiv. Walbrzych), res. “Niedzwiedzia Cave – Rock of Pulina” Distribution: Poland Host: Machilis sp. (Machilidae, Thysanura)

Leptus (L.) maculatus (Banks) (ad.) Rhyncholophus maculatus: [Banks, 1894: 216] ? Banks, 1904a: 143 Erythraeus (Atomus) maculatus Banks: Banks, 1904: 29 Atomus maculatus: Banks, 1915: 40; ?Augustson, 1939: 193 (Erythraeinae) Leptus maculatus (Banks): Southcott, 1961a: 503

Type locality: U.S.A. (California) Distribution: U.S.A.

Leptus (L.) madagascariensis André (larval) Leptus madagascariensis André, 1941: 190

Type locality: Madagascar Distribution: Madagascar Host: Stenopirates longipes (Henicocephalidae, Heteroptera)

Leptus (L.) maldonadoicus Haitlinger (larval) Leptus maldonadoicus Haitlinger, 2000c: 151

Type locality: “30 km east from Puerto Maldonado, at river Madre de Dios” Distribution: Peru Host: Crotonotus sp. (Pseudophyllidae, Orthoptera)

Leptus (L.) managarus Haitlinger (larval) Leptus managarus: Haitlinger, 1990b: 53

Type locality: Java Distribution: Indonesia (Java) Host: Huechys chtyselectra (Homoptera, Cicadina, Tibicinidae)


111

Leptus (L.) maranaensis Haitlinger (larval) Leptus maranaensis: Haitlinger, 1987g: 703

Type locality: Marana n. Fianarantsoa Distribution: Madagascar Host: Orthoptera indet.

Leptus (L.) mariae Haitlinger (larval) Leptus mariae: Haitlinger, 1987d: 344; 1992: 502; 1998: 107; 2000b: 190; 2002e: 148; 2006c; 2006e: 106; 2007a: 75; 2007b: 62; Southcott, 1992: 502; Gabryś, 1991: 105; Fain, Baugnée & Hidvegi, 1992: 336; Beenen, 2001: 201; Mąkol & Gabryś, 1999: 30 Leptus gabrysi Southcott, 1992: 137; Gabryś & Mąkol, 1996: 488, Fain & Baugnée, 1996: 22

Type locality of L. mariae Haitlinger: Sulistrowiczkin. Sobotka, Wroclaw Prov., Poland Type locality of L. gabrysi Southcott: Res. “Bielinek on Odra” near Cedynia (voiv. Szczecin), Poland Distribution: Belgium, Bulgaria, Holland, Hungary, Italy (incl. Capri), Norway, Poland, Slovenia, Spain, Switzerland Hosts: Phyllobius urticae (Curculionidae, Coleoptera), Lochmaea caprae (Col., Chrysomelidae), Col.(Lagriidae), Cicadetta montana (Cicadidae, Homoptera), Tenebrionidae indet.; Forficula auricularia (Forficulidae, Dermaptera) (for L. gabrysi)

Leptus (L.) mariani Haitlinger (larval) Leptus mariani: Haitlinger, 1991: 266

Type locality: Porto Alegre, Rio Grande do Sul Distribution: Brazil Host: Stolas festiva (Col., Chrysomelidae, Cassidinae)

Leptus (L.) maringensis Fain et Elsen (larval) Leptus maringensis: Fain & Elsen, 1972: 64

Type locality: Baringa, D.R.Congo Distribution: D.R.Congo (Zaire) Host: Glossina fuscipes fuscipes (Glossinidae, Diptera)

Leptus (L.) masaimaraicus Haitlinger (larval) Leptus masaimaraicus Haitlinger, 2001b: 477

Type locality: Masai Mara (S 1o08’, E 35o05’) Distribution: Kenya Host: “from plants in meadows”

Leptus (L.) meloidarum Beron (larval) Leptus meloidarum: Beron, 1975: 56; Southcott, 1992: 24; Haitlinger, 1998: 106

Type locality: Vratsa, Bulgaria

112


Distribution: Bulgaria, Italy, Kirghizstan Host: Meloidae, Cicindelinae (Carabidae), Tenebrionidae (Coleoptera), Orthoptera, Opiliones

Leptus (L.) millipedius Southcott (larval) Leptus millipedius: Southcott, 1992a: 25

Type locality: Madeira Distribution: Madeira Host: Cylindroiulus hirticauda, Ommatoiulus moreleti (Julidae, Diplopoda)

Leptus (L.) minno Southcott (larval) Leptus minno: Southcott, 1999: 269

Type locality: Wattle Flat, South Australia Distribution: Australia (S. Australia) Host: Isopeda frenchi (Sparassidae, Araneae)

Leptus (L.) mirenae Haitlinger (larval) Leptus mirenae: Haitlinger, 1994b: 165

Type locality: South Spain Distribution: Spain Host: Julodis pilosa (Buprestidae, Coleoptera)

Leptus (L.) miromiri Haitlinger (larval) Leptus miromiri: Haitlinger, 1992: 505

Type locality: Distr. Lublin and Biala Podlaska, Poland Distribution: Poland Host: “from plants”

Leptus (L.) mogadoranus Haitlinger (larval) Leptus mogadoranus: Haitlinger, 1990c: 46

Type locality: Es – Saouira (Mogador) Distribution: Morocco Host: Ecphoroma hemisphaerica (Tenebrionidae, Coleoptera)

Leptus (L.) molochinus (C.L. Koch) (ad.) Rhyncholophus molochinus C.L. Koch, 1837: 16, 18; 1841: 430; 1842: 50 Balaustium molochinus (Koch): Buitendijk, 1945: 349 Leptus molochinus (C.L. Koch): Oudemans, 1913a: 14; 1913b:10; 1937: 1926; Schweizer, 1951: 140; Willmann, 1951a: 157, 1951c: 151; Franz, 1954: 382; Schmölzer, 1993: 61; Mihelčič, 1958a: 280; 1958b: 45; Schweizer & Bader, 1963: 306; Gabryś & Mąkol, 1991: 709; 1994: 128; 1995: 383; 1996: 488


113

Type locality: Regensburg, Germany Distribution: Austria, Germany, Holland, Hungary, Norway, Poland, Spain, Switzerland

Leptus (L.) monolithosicus Haitlinger (larval) Leptus monolithosicus Haitlinger, 2003b: 134

Type locality: Rhodes (Monolithos) Distribution: Greece (Rhodes) Host: Oedipoda sp. (Acrididae, Orthoptera)

Leptus (L.) monteithi Southcott (larval) Leptus monteithi: Southcott, 1993: 1512

Type locality: Tasmania Distribution: Tasmania Host: Lepioproctis sp. (Colletidae, Hymenoptera)

Leptus (L.) murteri Schweizer (ad.) Leptus murteri Schweizer, 1951: 143; Cooreman, 1955c: 14; Schweizer & Bader, 1963: 307

Type locality: Buffalora, Murter, Switzerland Distribution: Belgium, Italy, Switzerland

Leptus (L.) nearcticus Fain, Gummer et Whitaker (larval) Leptus nearcticus Fain, Gummer & Whitaker, 1987: 137; Southcott, 1992: 121

Type locality: Brazil, Clay Co., Indiana Distribution: U.S.A. (Indiana) Hosts: Leiobunum longipes, L. nigripes, L. vittatum (Phalangiidae, Opiliones)

Leptus (L.) nemorum (C.L. Koch) (ad.) Rhyncholophus nemorum: C.L.Koch, 1836: fasc. 1; 1841: 430; 1842: 50; Berlese, 1885: 137; Canestrini, 1897: 461; Kramer, 1898: 422; Halbert, 1915: 114; Jablonowski, 1918: 3 Rhyncolophus nemorum (C.L. Koch): Lucas, 1840: 474; 1850: 474 Achorolophus nemorum: Oudemans, 1906: 108 Abrolophus nemorum: Berlese, 1913: 131 Ritteria nemorum Koch: Hull, 1918: 25 (and var. vertex Kramer); George, 1913: 109 Erythraeus nemorum (C.L. Koch): Karppinen, 1958: 44 Leptus nemorum (C.L. Koch): Oudemans, 1913b:16; 1937: 1927; Schweizer, 1951: 136; Willmann, 1951c: 151; 1954: 241; Franz, 1954: 382; Bottazzi, 1956: 43; Schmölzer, 1993: 61; Gabryś & Mąkol, 1994: 128; 1995: 383; 1996: 488; Mąkol & Gabryś, 1999: 30

Type locality: Regensburg, Germany Distribution: Austria, Czech Rep., Denmark, England, Finland, Germany, Holland, Hungary, Ireland, Italy, Poland, Switzerland; ? New Guinea, New Britain

114


Leptus (L.) nikanori Haitlinger (larval) Leptus nikanori Haitlinger, 2000a: 132

Type locality: Irazu, ca. 3000 m Distribution: Costa Rica Host: from plants

Leptus (L.) norrisi Southcott (larval) Leptus norrisi: Southcott, 1999: 157

Type locality: Merimbula, New South Wales Distribution: Australia (N.S.W.) Host: Allomachilis sp. (Archaeognatha, Meinertellidae)

Leptus (L.) ochroniger Oudemans (ad., N) Achorolophus opilionis Oudemans, 1913a: 16 Leptus ochroniger Oudemans, 1913b: 15; Irk, 1939b: 168; Buitendijk, 1945: 347

Type locality: Hoensbroek (Limburg, Holland) Distribution: Austria, Hungary, Holland

Leptus (L.) ogazulacus Haitlinger (larval) Leptus ogazulacus: Haitlinger, 1990a: 29

Type locality: Mozambique Distribution: Mozambique Host: Dichtha cubica (Tenebrionidae, Coleoptera)

Leptus (L.) olafi Haitlinger (larval) Leptus olafi Haitlinger, 1991: 268

Type locality: Colombia Distribution: Colombia Host: Ataitula hoffini (Carabidae, Cicindellinae, Col.)

Leptus (L.) olamukijacus Haitlinger (larval) Leptus olamukijacus Haitlinger, 2001b: 476

Type locality: Olamukija, Masai Mara (S 1o08’, E 35o05’) Distribution: Kenya Host: Orthoptera indet.

Leptus (L.) onnae Haitlinger (larval) Leptus onnae Haitlinger, 2000a: 137; 2004a: 154

Type locality: Agua n. Palenque


115

Distribution: Mexico, Brazil Host: from plants

Leptus (L.) ornatus Womersley (ad.) Leptus ornatus Womersley, 1934: 224; 1936: 118

Type locality: Rottnest Island, West Australia Distribution: Australia (W.A., N.S.W.)

Leptus (L.) orthopterarum Beron (larval) Leptus orthopterarum Beron, 1975: 52; Haitlinger, 2006c; 2006e: 106

Type locality: Primorsko (Bulgaria) Distribution: Bulgaria, Spain Host: Bucephaloptera bucephala (Tettigoniidae, Orthoptera), unknown Orthoptera

Leptus (L.) orthrius Southcott (larval) Leptus orthrius: Southcott, 1993: 1506

Type locality: Northern Territory Distribution: Australia (N. Territory) Host: Orthrius sepulchralis (Col., Cleridae)

Leptus (L.) oudemansi (Karppinen) (larval) Achorolophus gracilipes: Oudemans, 1910: 48; 1912: 168 Leptus gracilipes (Oudemans): Buitendijk, 1945: 347 Achorolophus oudemansi: Karppinen, 1958: 43 (pro Achorolophus gracilipes Oudemans, 1910, nec Kramer, 1897) Leptus oudemansi (Karp.): Southcott, 1992

Type locality: “Saramaca – Expeditie” Distribution: Surinam Host: Cynorta sp. (Cosmetidae, Opiliones)

Leptus (L.) ovatus Khot (ad.) Leptus ovatus: Khot, 1964a: 686


Leptus (L.) oxyae Vishnupriya et Mohanasundaram (larval) Leptus oxyae Vishnupriya & Mohanasundaram, 1988: 255

Type locality: Tamil Nadu Distribution: India Host: Oxya nitidula (Acrididae, Orthoptera)

116


Leptus (L.) parvulus Mihelčič (ad) Leptus parvulus Mihelčič, 1958b: 45


Leptus (L.) pasopaicus Haitlinger (larval) Leptus pasopaicus Haitlinger, 1990a: 30

Type locality: Namibia Distribution: Namibia Host: Tenebrionidae indet.

Leptus (L.) pelebinus Haitlinger (larval) Leptus pelebinus Haitlinger, 2006g: 110

Type locality: Pelebina, ca. 150 km from Natitingou Distribution: Benin Host: undetermined Orthoptera

Leptus (L.) phalangii (de Geer) (larval) Acarus phalangii: de Geer, 1778; 1783: 204 Leptus phalangii: Irk, 1939b: 168; Dudich et al., 1940: 55; Buitendijk, 1945: 348; Franz, 1954: 382; Willmann, 1951c: 154; 1955: 211

Type locality: Sweden Distribution: Austria, Germany, Hungary, Sweden Host: Opiliones

Leptus (L.) phuketicus Southcott (larval) Leptus phuketicus: Southcott, 1994: 166

Type locality: Phuket Is., Thailand Distribution: Thailand Hosts: Gagrellula niveata (Phalangiidae, Opiliones)

Leptus (L.) phyllotretae Feider (larval) Leptus phyllotretae Feider, 1956: 148; Southcott, 1992: 42

Type locality: La Feliceni, Distr. Odorhei, and Iaşi, Romania Distribution: Romania Hosts: Phyllotreta spp. (Chrysomelidae, Coleoptera)

Leptus (L.) pincheni Southcott (larval) Leptus pincheni: Southcott, 1999: 209

Type locality: Philip I., Victoria Distribution: Australia (Victoria) Host: grasshopper Apotrips sp. (Acrididae, Orthoptera)


Leptus (L.) pistoris Southcott (larval) Leptus pistoris: Southcott, 1999: 255

Type locality: Engelbrook Reserve, South Australia Distribution: Australia (S. Australia) Host: Urodacus manicatus (Scorpionidae, Scorpiones)

Leptus (L.) polythrix Fain et Elsen (larval) Leptus polythrix Fain & Elsen, 1987: 112; Cokendolpher, 1993: 134

Type locality: Dembo, 2200 m, Nyika Plateau, Malawi Distribution: Malawi Host: Opiliones (Cristina lettowi, Phalangiidae)

Leptus (L.) poonaensis Khot (ad.) Leptus poonaensis: Khot, 1964a: 682

Type locality: Poona, Maharashtra Distribution: India

Leptus (L.) pozzoicus Haitlinger (larval) Leptus pozzoicus Haitlinger, 2007d: 532

Type locality: Porto Pozzo (Sardinia) Distribution: Sardinia Host: undetermined Orthoptera

Leptus (L.) puniceus Southcott (larval) Leptus puniceus: Southcott, 1999: 132

Type locality: Proserpine, Queensland Distribution: Australia (Queensland) Host: “caught free-living”

Leptus (L.) puylaerti Fain et Elsen (larval) Leptus puylaerti Fain & Elsen, 1987: 111; Cokendolpher, 1993: 134

Type locality: Chowo Rocks, 2100 m, Nyika Plateau, Malawi Distribution: Malawi Host: Opiliones (Cristina lettowi, Phalangiidae)

Leptus (L.) pyrenaeus André (larval) Leptus pyrenaeus: André, 1953: 150; Southcott, 1992: 17

Type locality: Banyuls-sur-Mer, France Distribution: France Host: Mesobuthus occitanus (Buthidae, Scorpiones)

117

118


Leptus (L.) repens Shiba (ad.) Leptus repens: Shiba, 1969: 103

Type locality: Shiba Heights IBP Area Distribution: Japan

Leptus (L.) rosellae Haitlinger (larval) Leptus rosellae Haitlinger, 1999a: 63

Type locality: Kusadası Distribution: Turkey Host: Orthoptera (? Acrididae)

Leptus (L.) rubricatus (C.L.Koch)(ad.) Rhyncholophus rubricatus: C.L.Koch, 1837: 16, 13; 1841: 430; 1842: 50 Achorolophus rubricatus (C. L. Koch): Oudemans, 1913a: 15 Leptus rubricatus (C.L. Koch): Oudemans, 1913b: 12; 1937: 1928; Dudich et al., 1940: 5; Buitendijk, 1945: 348; Schweizer, 1951: 140; Franz, 1954: 382; Mihelčič, 1958a: 280; 1958b: 45; Gabrys & Mąkol, 1991: 709; Schmölzer, 1993: 61

Type locality: Regensburg, Germany Distribution: Austria, Czech Rep., Germany, Holland, Hungary, Norway, Poland, Spain, Switzerland

Leptus (L.) ruginus Southcott (larval) Leptus ruginus: Southcott, 1992: 78

Type locality: Tyringham, Massachusetts Distribution: USA (Massachusetts) Host: Leucania pseudargyria (Noctuidae, Lepidoptera)

Leptus (L.) rwandae Fain et Jocqué (larval) Leptus rwandae Fain & Jocqué, 1996: 102

Type locality: Rwanda Distribution: Rwanda Hosts: Systenoplacis sp. (Zodariidae, Araneae)

Leptus (L.) saigusai Kawashima (larval) Leptus saigusai Kawashima, 1958: 197; Treat, 1975: 218

Type locality: Aomori Pref., Japan Distribution: Japan Hosts: Noctuidae, Pyralidae, Notodontidae, Geometridae (Lepidoptera)


119

Leptus (L.) samsingensis Gupta Leptus samsingensis Gupta, 1992: 123

Type locality: West Bengal, Distr. Darjiling, Samsing Distribution: India

Leptus (L.) sayi Southcott (larval) Leptus sayi Southcott, 1992a: 103

Type locality: Tyringham, Massachusetts Distribution: USA (Massachusetts) Host: Amphipoea americana (Noctuidae, Lepidoptera)

Leptus (L.) schedingi (Oudemans) (larval) Achorolophus schedingi: Oudemans, 1911a:125; 1912: 165 Leptus schedingi (Oudemans): Buitendijk, 1945: 348; Feider, 1956: 150; Beron, 1975: 68; Treat, 1975: 221

Type locality: Corral, S. Chile Distribution: Chile Host: Lepidoptera, Rhopalocera

Leptus (L.) scutellatus Southcott (larval) Leptus scutellatus: Southcott, 1999: 152

Type locality: Babiang (PNG) Distribution: Papua New Guinea Host: “free-living”

Leptus (L.) shimenensis Zheng (larval) Leptus shimenensis Zheng, 1996b: 233

Type locality: Mt. Hupingshan, Shimen Co., Hunan Prov. Distribution: China, Hunan Prov. Host: Insecta indet.

Leptus (L.) siculiformis (Trägårdh)(ad.) Rhyncholophus siculiformis Trägårdh, 1904: 64; 1908: 31

Leptus siculiformis (Trägårdh): Type locality: Heluan, Cairo (Egypt) Distribution: Egypt, Tanzania

Leptus (L.) siculus (Canestrini) (ad.) Rhyncholophus siculus Canestrini, 1884: 698; 1885: 150; Berlese, 1894: f. LXXI, 1; Kramer, 1898: 422 Belaustium siculum (Can.): Oudemans, 1906b: 108 Leptus siculus (Canestrini): Cooreman, 1946: 4 Achorolophus siculus (Can.): Starkoff & Starkoff, 1950: 96

120


Balaustium siculum (G. Can.): Bottazzi, 1956: 44

Type locality: Sciacca (Sicily) Distribution: Cyprus, Italy (incl. Sicily), Morocco, ? New Britain

Leptus (L.) siemsseni Oudemans (larval) Leptus siemsseni Oudemans, 1910e: 109; 1912: 158; Buitendijk, 1945: 348; Feider, 1956: 150; Zhang, 2001b: 23

Type locality: “Futschau, Fokien”, China Distribution: China (Fujian) Host: unknown

Leptus (L.) sieversi (Oudemans)(larval) Achorolophus sieversi Oudemans, 1911: 124; 1912: 170 Leptus sieversi : Buitendijk, 1945: 348; Feider, 1956: 150; Beron, 1975: 46; Fain et al., 1987: 135

Type locality: Puerto Cabello Distribution: Venezuela

Leptus (L.) sigthori (Oudemans)(ad.) Achorolophus sigthori Oudemans, 1913a: 17 Leptus sigthori (Oudemans): Oudemans, 1913b: 20; Sellnick, 1940: 36; 1949: 129; Buitendijk, 1945: 348

Type locality: Sittard (Limburg, Holland) Distribution: Holland, Iceland, Sweden

Leptus (L.) similis Fain et Elsen (larval) Leptus similis Fain & Elsen, 1987: 113

Type locality: Boga, D.R.Congo Distribution: D.R. Congo (Zaire) Host: Chlorion ciliatum var. maxillae (Sphecidae, Hymenoptera)

Leptus (L.) simonettae Haitlinger (larval) Leptus simonettae Haitlinger, 2000a: 137

Type locality: Tikal (Guatemala), Puerto Gringo (Honduras) Distribution: Guatemala, Honduras Host: small undetermined Acrididae (Orthoptera)

Leptus (L.) singhi Saboori et Arbabi (larval) Leptus singhi Saboori & Arbabi, 2003: 175

Type locality: Uttar-Pradesh Distribution: India (Uttar-Pradesh) Host: unknown


121

Leptus (L.) slivovi Beron (larval) Leptus slivovi Beron, 1975: 54; Haitlinger, 1987d: 342; 1987f: 107; 1992: 500; 2007b: 61; Southcott, 1992a: 18; 1992: 500; Gabryś, 1988: 23; Gabryś & Mąkol, 1994: 128

Type locality: Ropotamo Valley (Bulgaria) Distribution: Bulgaria, Liechtenstein, Norway, Poland Host: Laelia coenosa (Limantriidae), Arctia caja (Arctiidae), Zygaena sp. (Zygaenidae), Agapetes galathea (Satyridae) (Lepidoptera)

Leptus (L.) smithi Southcott (larval) Leptus smithi: Southcott, 1999: 247

Type locality: Canberra, A.C.T. Distribution: Australia Host: Urodacus manicatus (Scorpionidae, Scorpiones)

Leptus (L.) soddagus Haitlinger (larval) Leptus soddagus Haitlinger, 1990a: 31

Type locality: Usambara, Tanzania Distribution: Tanzania Host: Elateridae indet. (Col.)

Leptus (L.) southcotti Beron (larval) Leptus southcotti Beron, 1975: 47; Southcott, 1992: 16

Type locality: Potop (Bulgaria) Distribution: Bulgaria Host: Collembola

Leptus (L.) spinalatus Southcott (larval) Leptus spinalatus: Southcott, 1993: 1476

Type locality: Northern Territory (Australia) Distribution: Australia (N. Territory) Host: Glenodon sp. (Myrmeleontidae, Neuroptera)

Leptus (L.) stefani Haitlinger (larval) Leptus stefani Haitlinger, 1991: 266

Type locality: Colombia Distribution: Colombia Host: Ptychoderes speciosus (Col., Anthribidae)

Leptus (L.) stieglmayri (Oudemans)(larval) Erythraeus stieglmayri: Oudemans, 1905b: 237 Achorolophus stieglmayri (Oudemans): Oudemans, 1905c: 9; 1912: 163

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Leptus stieglmayri: Buitendijk, 1945: 348; Feider, 1956: 150; Beron, 1975: 68; Fain, 1987e: 135

Type locality: Santa Cruz, Rio Grande do Sul, Brazil Distribution: Brazil Host: Opiliones

Leptus (L.) stolae Haitlinger (larval) Leptus stolae: Haitlinger, 1987e: 357; 1991: 272

Type locality: Espirito – Santo Distribution: Brazil Hosts: Stolas nudicollis, S. aenea, S. chalybea, S. oblita (Chrysomelidae, Cassidinae, Col.)

Leptus (L.) sudanensis (Oudemans)(larval) Achorolophus sudanensis Oudemans, 1911b: 138; 1912: 173 Leptus sudanensis (Oudemans): Buitendijk, 1945: 348; Feider, 1956: 150; Beron, 1975: 69

Type locality: “Khartoem”, Sudan Distribution: Sudan Host: Adesmia dilatata (Tenebrionidae, Coleoptera)

Leptus (L.) sulciscutus Zheng (larval) Leptus sulciscutus Zheng, 1996 b: 239

Type locality: M. Hupingshan, Shimen Co., Hunan Prov. Distribution: China (Hunan Prov.) Host: Insecta indet.

Leptus (L.) swani Southcott (larval) Leptus swani: Southcott, 1991: 731

Type locality: Glenelg, South Australia Distribution: Australia Host: Tabanidae (Diptera)

Leptus (L.) tammuzi Haitlinger (larval) Leptus tammuzi: Haitlinger, 1994c: 142

Type locality: Syria Distribution:Israel, Saudi Arabia, Syria Host: Pimelia mittrei, Vieta millingeni (Col., Tenebrionidae)

Leptus (L.) tarranus Southcott (larval) Leptus tarranus: Southcott, 1993: 1482

Type locality: Victoria Distribution: Australia (Victoria) Host: Tenebrionidae (Col.)


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Leptus (L.) terebrans Vitzthum (larval) Leptus terebrans Vitzthum, 1926: 160

Type locality: Tandjoeng Karang, Lampong, S. Sumatra Distribution: Indonesia (Sumatra)

Leptus (L.) tetrigius Southcott (larval) Leptus tetrigius Southcott, 1988c: 153

Type locality: Deerwood, Kuruwita (slopes of Botiyagala Mt.), > 300 m Distribution: Sri Lanka Host: Scelimena gavialis (Tetrigidae, Orthoptera)

Leptus texanus (Banks) (ad.) Rhyncholophus texanus: Banks, 1894: 217 Leptus texanus (Banks): Welbourn & Jennings, 1991: 568

Type locality: Texas Distribution: U.S.A. (Texas)

Leptus (L.) tindalei Southcott (larval) Leptus tindalei Southcott, 1999: 202

Type locality: Belair, S. Australia Distribution: Australia (South Australia) Host: grasshopper (Orthoptera)

Leptus (L.) tiranicus Haitlinger (larval) Leptus tiranicus Haitlinger, 2006d: 349

Type locality: Margerita, Playa Parguito n. El Tirano Distribution: Venezuela Host: undetermined Asilidae (Diptera) and legs of undetermined Orthoptera

Leptus (L.) titinius Southcott (larval) Leptus titinius: Southcott, 1993: 1496

Type locality: Western Australia Distribution: Australia (W. Australia) Host: Titinia bicolor (Col., Curculionidae)

Leptus (L.) torresianus Southcott (larval) Leptus torresianus: Southcott, 1988: 104; 1993: 1481

Type locality: Mt. Hemmant, N. of Daintree, Queensland Distribution: Australia (Queensland) Host: Cicadas – Venustria superba, Tanasa doddi (Homoptera, Auchenorhyncha, Cicadidae), Coleoptera (Curculionidae, Cryptorhynchinae)

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Leptus (L.) treati Welbourn (larval and adult) Leptus treati Welbourn, in Welbourn & Jennings, 1991: 569

Type locality: Piscataquis Co., ME Distribution: U.S.A. Host: Choristoneura fumiferana (Tortricidae, Lepidoptera)

Leptus (L.) triacanthus Southcott (larval) Leptus triacanthus: Southcott, 1999: 125

Type locality: Babiang Distribution: Papua New Guinea Host: “caught free-living”

Leptus (L.) trimaculatus (Rossi) (ad.) Trombidium trimaculatum Rossi, 1794: 5; Hermann, 1804: 27; Latreille, 1817: 118; 1829: 284; Dugès, 1834: 16; 1836: 302; Hahn, 1835: 64 Rhyncholophus trimaculatus: C.L. Koch, 1836: fa.1; 1841: 430; 1842: 50; Canestrini & Fanzago, 1877: 69; Canestrini, 1885: 149; Jablonowski, 1918: 3; Schweizer, 1922: 83 Rhyncolophus trimaculatus: Lucas, 1840: 473; 1850: 473 Belaustium trimaculatum: Oudemans, 1897: 120 Rhyncholophus (Ritteria) trimaculata: Thor, 1900: 9 Abrolophus trimaculatus: Berlese, 1913: 131 Ritteria trimaculata: Thor, 1900: 9; Hull, 1918: 25 Rhyncholophus trimaculatum (Rossi): Oudemans, 1937: 1929 (Leptus) Erythraeus trimaculatus: Karppinen, 1958: 45 Leptus trimaculatum: Turk, 1953: 26 Leptus trimaculatus: Oudemans, 1913b: 13; Sellnick, 1931: 696; Irk, 1939b: 168; Franke, 1940: 153; Cooreman, 1943; Buitendijk, 1945: 348; Feider, 1948: 208; Starkoff & Starkoff, 1950: 96; Schweizer, 1951: 135; Willmann, 1951c:151; 1954: 241; 1955: 211; Franz, 1954: 383; Bottazzi, 1956: 43; Mihelčič, 1958: 281; Schweizer & Bader, 1963: 303; Gabryś, 1988: 23; Wendt et al., 1992: 58; Gabryś & Mąkol, 1991: 709; 1994: 128; 1995: 383; Beron, 1995: 6; Haitlinger, 2002e: 148 Leptus echinopus: Beron, 1975: 60 (larval); Haitlinger, 1987d: 340; 1992: 500; 2000b: 191; 2002d: 555; 2003d: 4; 2003f: 8; 2007b: 61; Gabryś, 1988: 23; Southcott, 1992: 500

Type locality: Italy Type locality of Leptus echinopus Beron, 1975: Bulgaria Distribution: Austria, Belgium, Bulgaria, Croatia, Czech Rep., Denmark, Finland, France, Germany, Great Britain (England and Scotland), Greece, Holland, Hungary, Italy, Japan, Latvia, Poland, Romania, Slovak Rep., Slovenia, Spain, Switzerland; Wendt et al. (1992) and Haitlinger (2007c) concluded, that L. echinopus is a synonyme, being the larval form of L. trimaculatus Host (of the larve, described as Leptus echinopus Beron: Collembola, Boletophagus reticulatus (Tenebrionidae, Coleoptera), Erythraeus sp. (Erythraeidae, Acariformes)


Leptus (L.) trucidatus (Hull)(ad.) Achorolophus trucidatus: Hull, 1923 Leptus trucidatus: Southcott, 1993: 1494

Type locality: Western Australia Distribution: Australia

Leptus (L.) truncatus Southcott (larval) Leptus truncatus: Southcott, 1993: 1544

Type locality: Victoria Distribution: Australia (Victoria) Host: Pedilophorus venustus (Col., Byrrhidae)

Leptus (L.) ubudicus Haitlinger (larval) Leptus ubudicus: Haitlinger, 2006d: 225

Type locality: Ubud (Bali) Distribution: Indonesia (Bali) Host: from herbaceous plants

Leptus (L.) urodaci Southcott (larval) Leptus urodaci: Southcott, 1999: 187

Type locality: Engelbrook Res., Mt. Lofty Ranges, Kangaroo I. (S. Australia) Distribution: Australia (S. Australia) Host: Urodacus manicatus (Scorpionidae, Scorpiones)

Leptus (L.) ursyni Haitlinger (larval) Leptus ursyni Haitlinger, 1991: 265

Type locality: Chile Distribution: Chile Host: Scotobius planatus (Tenebrionidae, Coleoptera)

Leptus (L.) utheri Southcott (larval) Leptus utheri: Southcott, 1993: 1533

Type locality: New South Wales Distribution: Australia (NSW) Host: Parahyborrhynchus convexiusculus (Col., Curculionidae)

Leptus (L.) variatus Mihelčič (ad.) Leptus diversus var. variatus Mihelčič, 1958b: 46 Leptus variatus: Gabryś, 2000c: 66 (stat. nov.)


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126


Leptus (L.) vertex (Kramer) (ad.) Rhyncholophus vertex Kramer, 1886: 263; Schweizer, 1922: 83; Trägårdh, 1910: 488 Rhyncholophus nemorum var. vertex: Berlese, 1887: fasc. XL, 3 Rhyncholophus (Ritteria) vertex Kramer: Sig Thor, 1900: 57 Erythraeus vertex (Kramer): Trägårdh, 1904b: 57 Ritteria nemorum var. vertex Kramer: Hull, 1918: 25 Leptus vertex (Kramer): Oudemans, 1913b: 18; Irk, 1939b: 168; Sellnick, 1940: 36; Schweizer, 1951: 139; Lindroth et al., 1973: 131; ; Gabryś, 1990 (Thesis); Gabryś & Mąkol, 1991: 709; 1994: 129; 1995: 383; 1996: 488; Mąkol & Gabryś, 1999: 30; Haitlinger, 2002e: 149

Type locality: Germany Distribution: Austria, Germany, Great Britain, Greenland, Hungary, Iceland, Italy, Norway, Poland, Switzerland

Leptus (L.) verticiformis Trägårdh (ad.) Leptus verticiformis Trägårdh, 1971: 57

Type locality: Faroe Is. Distribution: Faroe Is.

Leptus (L.) villosus (Berlese) (ad.) Achorolophus villosus Berlese, 1910: 348 Leptus villosus (Berlese): Khot, 1964a: 686 Leptus villosus (Kramer)

Type locality: Corfu, Greece Distribution: Greece, India (Himachal Pradesh, 13500 ft.),

Leptus (L.) waldockae Fain (larval) Leptus waldockae Fain, 1991: 69

Type locality: Learmonth, West Australia Distribution: Australia (West Australia) Host: Lychas alexandrinus Hirst (Buthidae, Scorpiones)

Leptus (L.) warregense (Hirst) (ad.) Belaustium [sic] warregense Hirst, 1928: Leptus warregense Womersley, 1934: 223; Halliday, 1998: 59

Type locality: Distribution: Australia

Leptus (L.) welbourni Southcott (larval) Leptus welbourni Southcott, 1992a: 115

Type locality: Tyringham, Massachusetts Distribution: U.S.A. (Massachusetts) Host: Leucania commoides (Noctuidae, Lepidoptera)


127

Leptus (L.) yoshimii Shiba (ad.) Leptus yoshimii Shiba, 1976: 175

Type locality: Pasoh Forest, Malaya Distribution: Malaysia Host: Leucania commoides (Noctuidae, Lepidoptera)

Leptus (L.) zhangi Saboori et Atamehr (larval) Leptus zhangi Saboori & Atamehr, 1999: 159

Type locality: Uromieh Distribution: Iran Host: Hyponomeuta malinella (Yponomeutidae, Lepidoptera)

Leptus (L.) zhejiangensis Zheng (larval) Leptus zhejiangensis Zheng, 2003: 56

Type locality: Zhejiang Distribution: China Host: Siobla ferox (Tenthredinidae, Hymenoptera)

Leptus (L.) zhutingensis Zheng (larval) Leptus zhutingensis Zheng, 1996a: 62

Type locality: Zhuting, Zhuzhou Co., Hunan Prov. Distribution: China (Hunan Prov.) Host: Colaspoides opaca (Coleoptera, Chrysomelidae); Homoptera (Cicadidae)

Leptus sp. Leptus sp.: De Armas & Trueba, 2003: 133

Distrinution: Dominican Republic Host: Phrynus kennidae (Phrynidae, Amblypygi)

Subgenus Amaroptus Haitlinger (larval) Amaroptus Haitlinger, 2000c: 156 (in the same paper also as “Amaroptes” and “Amaropus”, err. typ.)

Type species: Leptus (Amaroptus) vuki Haitlinger, 2000

Leptus (Amaroptus) vuki Haitlinger (larval) Leptus (Amaroptus) vuki Haitlinger, 2000c: 157

Type locality: 30 km east from Puerto Maldonado Distribution: Peru Hosts: unidentified Catantopinae (Acrididae, Orthoptera)

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Subfamily Callidosomatinae Southcott Sphaerolophinae Kishida, 1939: 48 Callidosominae Southcott, 1957: 97 Callidosomatinae Southcott, 1961a: 521

Type genus: Callidosoma Womersley, 1936

Tribe Callidosomatini Southcott Callidosomini Southcott, 1957: 97 Callidosomatini Southcott, 1972: 1

Type genus: Callidosoma Womersley, 1936

Genus Caeculisoma Berlese (ad. and larv.) Caeculisoma Berlese, 1888: 186

Type species: Caeculisoma tuberculatum Berlese, 1888

Subgenus Caeculisoma s.str. Berlese Caeculisoma Berlese, 1888: 186

Type species: Caeculisoma tuberculatum Berlese, 1888

Caeculisoma (C.) afrum Cooreman (ad.) Caeculisoma afrum Cooreman, 1958: 47

Type locality: Kihemba, Masarcha, 2200 m, Kivu Distribution: D.R. Congo (Kivu)

Caeculisoma (C.) argus argus Vitzthum (ad.) Caeculisoma argus Vitzthum, 1926: 169; Womersley, 1936; 120; Cooreman, 1958: 45

Type locality: near Lake Toba, Sumatra Distribution: Sumatra (Indonesia), Kangaroo Is., S. Australia (Australia)

Caeculisoma (C.) argus io Southcott (ad.) Caeculisoma argus io: Southcott, 1961b: 174

Type locality: Glen Osmond (South Australia) Distribution: Australia

Caeculisoma (C.) carmenae Haitlinger Caeculisoma carmenae Haitlinger, in print

Type locality: South Africa Distribution: South Africa


129

Caeculisoma (C.) claviger Canestrini (ad.) Caeculisoma claviger Canestrini, 1897: 462 Coeculisoma claviger Canestrini: Oudemans, 1906b: 108 Caeculisoma clavigerum: Cooreman, 1958: 44

Type locality: “Friedrich Wilhelm Hafen”, New Guinea Distribution: Papua New Guinea

Caeculisoma (C.) cooremani Southcott (larval) Caeculisoma cooremani Southcott, 1972: 26; Key, 1990: 11

Type locality: 4 miles S. of Nilemah H.S., Hamelin Pool area, West Australia Distribution: Australia (W. Australia) Host: “from the grasshopper Urnisiella sp. (Acrididae, Orthoptera)”

Caeculisoma (C.) cordipes Vitzthum (ad.) Caeculisoma cordipes Vitzthum, 1935: 91; Cooreman, 1958: 47

Type locality: Uahuka, 2900 feet (885 m) alt. Distribution: Marquesan Islands (Uahuka)

Caeculisoma (C.) darwiniense Southcott (larval) Caeculisoma darwiniense Southcott, 1961b: 164; 1972: 28; Key, 1990: 11; Beron: in prep.

Type locality: Coomalie Creek, Northern Territory, Australia Distribution: Australia (Northern Territory, Queensland, New South Wales, South Australia), New Guinea Host: locusts (Goniaea sp. aff. hyalina, Macrazelota cervina, Peakesia vitripennis, Austroicetes tricolor, Austracris guttulosa) (Acrididae, Orthoptera), Macrotoma sp. (Coleoptera, Cerambycidae)

Caeculisoma (C.) haussa Beron (ad.) Caeculisoma haussa Beron, 2000: 66

Type locality: Maiduguri, N. Nigeria Distribution: Nigeria

Caeculisoma (C.) hunanica Zheng (ad.) Caeculisoma hunanica Zheng, 2002b: 61

Type locality: Hunan, Distribution: China (Hunan)

Caeculisoma (C.) huxleyi Southcott (larval) Caeculisoma huxleyi Southcott, 1972: 32

Type locality: Nelson Distribution: New Zealand Host: “ex moth, Xanthorrhoe sp.” (Lepidoptera)

130


Caeculisoma (C.) infernale Vitzthum (ad.) Caeculisoma infernale Vitzthum, 1926: 173-177; Cooreman, 1958: 46

Type locality: “Pedada-Bai, Lampong” (South Sumatra) Distribution: Indonesia (Sumatra)

Caeculisoma (C.) johnstoni Womersley (ad.) Caeculisoma johnstoni Womersley, 1934: 241; Cooreman, 1958: 46

Type locality: South Australia Distribution: Australia (South Australia)

Caeculisoma (C.) montana (Rainbow) (ad.) Rhyncholophus montanus Rainbow, 1906: 156 Caeculisoma montana (Rainbow): Womersley, 1934: 237 Caeculisoma montanum (Rainbow): Cooreman, 1958: 44

Type locality: Mount Kosciusko, F.C.T., Australia Distribution: Australia

Caeculisoma (C.) mouldsi Southcott (larval) Caeculisoma mouldsi Southcott, 1988: 107

Type locality: Mt. Hemmant, N. Daintree, Queensland Distribution: Australia (Queensland) Hosts: Cicadas – Venustria superba, Tamasa doddi, Psaltoda fumipennis (Cicadidae), Mardalana suffusa (Tibicinidae) (Auchenorhyncha, Homoptera)

Caeculisoma (C.) nasutum Hirst (ad.) Caeculiosoma [lapsus typ.] nasutum Hirst, 1928: 568 Caeculisoma nasutum Hirst: Womersley, 1934: 236; Cooreman, 1958: 46

Type locality: Cairns Distr. (Queensland) Distribution: Australia (Queensland)

Caeculisoma (C.) nestori Haitlinger (larval) Caeculisoma nestori Haitlinger, 2004f: 437

Type locality: Foz do Iguazu Distribution: Brazil Host: from plants

Caeculisoma (C.) plantacionis Beron (ad.) Caeculisoma (C.) plantacionis Beron, in prep.

Type locality: Belik Plantation, New Ireland Distribution: New Ireland (Papua New Guinea)


Caeculisoma (C.) septentrionale Beron (ad.) Caeculisoma (C.) septentrionale Beron, in prep.

Type locality: Chimakothi, 1900-2300 m Distribution: Bhutan

Caeculisoma (C.) sparnoni Southcott (larval) Caeculisoma sparnoni Southcott, 1972: 31

Type locality: Panorama, Adelaide, South Australia Distribution: Australia (South Australia) Host: free living

Caeculisoma (C.) sulcatum (Canestrini)(ad.) Rhyncholophus sulcatus Canestrini, 1898a: 481; 1898b: 397 Belaustium sulcatum (Canestrini): Oudemans, 1906b: 108; Vitzthum, 1924: 357-359 Caeculisoma sulcatum (Canestrini): Vitzthum, 1926: 168 Caeculisoma sulcatum (Vitzthum): Cooreman, 1958: 45

Type locality: Erima, New Guinea Distribution: New Guinea, Krakatau (Indonesia)

Caeculisoma (C.) tuberculatum Berlese (ad.) Caeculisoma tuberculatum Berlese, 1888: 186; Oudemans, 1906b: 107; Cooreman, 1958: 44

Type locality: Buenos Aires, Asuncion Distribution: Argentina, Paraguay; ? New Guinea

Subgenus Papuacaeculisoma Beron (ad.) Papuacaeculisoma Beron: in prep.

Type species: Caeculisoma (Papuacaeculisoma) chapmani Beron, in prep.

Caeculisoma (Papuacaeculisoma) chapmani Beron (ad.) Caeculisoma (Papuacaeculisoma) chapmani Beron: in prep.

Type locality: Kolonoboi (New Ireland), Lae, Morobe Prov. (New Guinea) Distribution: Papua New Guinea

Caeculisoma (Papuacaeculisoma) novaeirlandiae Beron (ad.) Caeculisoma (Papuacaeculisoma) novaeirlandiae Beron: in prep.

Type locality: Kaluan 2, rain forest Distribution: New Ireland

Genus Callidosoma Womersley (larval and ad.) Callidosoma Womersley, 1936: 120

Type species: Caeculisoma ripicola Womersley, 1934 (ad.)

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132


Callidosoma apollo Southcott (larval and deutonymph) Callidosoma apollo Southcott, 1972: 11; Treat, 1985: 118

Type locality: Yavapai Co., Arizona Distribution: Arizona, Massachusetts (U.S.A.) Hosts: Protorthodes alfkeni, Leucania linda (Noctuidae) and other Lepidoptera

Callidosoma dasypodiae Womersley (larval) Erythraeus dasypodiae Womersley, 1934: 245 Callidosoma dasypodiae (Womersley): Southcott, 1966: 815; 1972: 17; Treat, 1975: 201

Type locality: Canberra Distribution: Australia (A.C.T., Queensland) Host: Lepidoptera (“on peacock moth” – Womersley, 1934; “moth Sericia spectans Guenée” – Southcott, 1972)

Callidosoma daviesi Treat (larval) Callidosoma daviesi Treat, 1985: 101

Type locality: Maricopa Co., Arizona Distribution: U.S.A. (Arizona) Host: Microlepidoptera indet.

Callidosoma droozi Treat (larval) Callidosoma droozi Treat, 1985: 104

Type locality: North Carolina, U.S.A. Distribution: U.S.A. (North Carolina) Host: Melanolophia canadaria choctawae (Geometridae, Lepidoptera)

Callidosoma galtoni Southcott (larval) Callidosoma galtoni Southcott, 1972: 6

Type locality: Beetaloo Valley, South Australia Distribution: U.S.A, Australia Host: grasshopper (Orthoptera)

Callidosoma guatemalensis Treat (larval) Callidosoma guatemalensis Treat, 1985: 105

Type locality: La Avellana, Guatemala Distribution: Guatemala Host: mosquito, Mansonia titillans (Culicidae, Diptera)

Callidosoma harlani Treat (larval) Callidosoma harlani Treat, 1985: 94

Type locality: Fort Gulick, Panama


133

Distribution: Panama Hosts: Lepidoptera (Noctuidae and Pyralidae)

Callidosoma heatherae Southcott (larval) Callidosoma heatherae Southcott, 1972: 8

Type locality: Grovely, near Brisbane Distribution: Australia (Queensland)

Callidosoma huachuca Treat (larval) Callidosoma huachuca Treat, 1985: 108

Type locality: Cochise County, Carr Canyon, Huachuca Mountains, 5000 ft., Arizona Distribution: U.S.A. (Arizona) Host: Rejectaria albisinuata (Lepidoptera, Noctuidae)

Callidosoma insulare Treat (larval) Callidosoma insulare Treat, 1985: 115

Type locality: North Carolina, Dare Co., Buxton, Hatteras Island Distribution: U.S.A. (North Carolina) Host: Bleptina medialis (Lepidoptera, Noctuidae)

Callidosoma leodegari Haitlinger (larval) Callidosoma leodegari Haitlinger, 2004e: 251

Type locality: Brazil Distribution: Brazil, Argentina Host: unknown

Callidosoma matsumuratettix Tseng, Yang et Pan Callidosoma matsumura tettix Tseng, Yang & Pan, 1976

Type locality: Taiwan Distribution: Taiwan

Callidosoma metzi Sharma, Drooz et Treat (larval and adult) Callidosoma metzi Sharma, Drooz & Treat, 1983: 120; Treat, 1985: 120

Type locality: Research Triangle Park, N. C. Distribution: U.S.A. (North Carolina, Arkansas, Louisiana, Mississippi, New Jersey, W. Virginia), Panama Host: Anacamptodes vellivolata and other moths (Lepidoptera: Geometridae, Cossidae, Tortricidae, Lycaenidae, Pyralidae)

134


Callidosoma quinteri Treat (larval) Callidosoma quinteri Treat, 1985: 108

Type locality: Lancaster Co., Adamstown, Pennsylvania Distribution: U.S.A. (Pennsylvania) Host: Catocala serena (Noctuidae, Lepidoptera)

Callidosoma ripicolum (Womersley) (ad.) Caeculisoma ripicola Womersley, 1934: 238 Callidosoma ripicola (Womersley): Womersley, 1936: 120

Type locality: South Australia Distribution: Australia (S. Australia, Victoria)

Callidosoma rostratum Southcott (larval) Callidosoma rostratum Southcott, 1972: 10

Type locality: Babiang, New Guinea Distribution: Papua New Guinea Host: in leaf litter

Callidosoma sorkini Treat (larval) Callidosoma sorkini Treat, 1985: 109

Type locality: Arizona, Maricopa Co., Wickenberg Distribution: U.S.A. (Arizona) Host: Microlepidoptera undet.

Callidosoma tiki Southcott (larval) Callidosoma tiki Southcott, 1972: 21; Treat, 1975: 202

Type locality: Titirangi, Auckland Distribution: New Zealand Host: “fly” (Diptera), Lepidoptera (Crambus apicellus, Scoparia halopis, S. trivirgata)

Callidosoma tindalei Southcott (larval) Callidosoma tindalei Southcott, 1972: 23; Treat, 1975: 205

Type locality: “at 5600 ft. near Mt. Kosciusko” Distribution: Australia Host: “at base of wing of Oreixenica correae (Nymphalidae, Lepidoptera)”

Callidosoma treati Southcott (larval) Callidosoma treati Southcott, 1972: 15; 1985: 122; Treat, 1975: 205

Type locality: College Park, Maryland Distribution:U.S.A. (Maryland, Mississippi, Massachusetts, Florida, Louisiana, N. Carolina) Hosts: Lepidoptera (Pseudaletia unipuncta and others, Geometridae, Tineidae, Noctuidae, Notodontidae)


Callidosoma wallaceanum Southcott (larval) Callidosoma wallaceanum Southcott, 1972: 13

Type locality: Pelotas, Brazil Distribution: Brazil Host: Lepidoptera, Noctuidae (Cirphis adultera)

Callidosoma welbourni Treat (larval) Callidosoma welbourni Treat, 1985: 111; Haitlinger, 2004g: 130

Type locality: Sao Paulo, Piracicaba Distribution: Brazil, Dominican Republic Host: “Halisidota” annulosa (Lepidoptera)

Callidosoma womersleyi Southcott (ad.) Callidosoma womersleyi Southcott, 1946: 43; 1972: 4


Genus Carastrum Southcott (larval) Carastrum Southcott, 1988: 195

Type species: Carastrum ferrari Southcott, 1988

Carastrum ferrari Southcott (larval) Carastrum ferrari Southcott, 1988: 195

Type locality: Durras, N.S.W., Australia Distribution: Australia (N.S.Wales) Hosts: Pyrellia tasmaniae and Helina sp. (Muscidae, Diptera)

Carastrum sanurensis Haitlinger (larval) Carastrum sanurensis Haitlinger, 2006d: 220

Type locality: Sanur, Bali Distribution: Bali (Indonesia) Hosts: from herbaceous plants

Genus Dambullaeus Haitlinger (larval) Dambullaeus Haitlinger, 2001a: 53

Type species: Dambullaeus piae Haitlinger, 2001

Dambullaeus piae Haitlinger (larval) Dambullaeus piae Haitlinger, 2001a: 53

Type locality: Dambulla Distribution: Sri Lanka Hosts: from plants

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136


Genus Harpagella Southcott (larval) Harpagella Southcott, 1996: 257

Type species: Harpagella moxonae Southcott, 1996

Harpagella moxonae Southcott (larval) Harpagella moxonae Southcott, 1996: 257

Type locality: Kinglake Nat. Park Distribution: Australia (Victoria) Host: Drosophila (Scaptodrosophila) sp., Drosophila inornata (Drosophilidae, Diptera)

Genus Momorangia Southcott (larval) Momorangia Southcott, 1972: 34

Type species: Momorangia jacksoni Southcott, 1972

Subgenus Momorangia Southcott (larval) Momorangia: Southcott, 1972: 34

Type species: Momorangia jacksoni Southcott, 1972

Momorangia (Momorangia) gabini Haitlinger (larval) Momorangia gabini Haitlinger, 2004c: 179

Type locality: Masai Mara Distribution: Kenya Host: from plants

Momorangia (M.) jacksoni Southcott (larval) Momorangia jacksoni Southcott, 1972: 34

Type locality: Mt. Williams, Arthur’s Pass, 5000 ft. Distribution: New Zealand Host: cicada Melampsalta oromelaena (Cicadidae, Homoptera)

Momorangia (M.) vallata Southcott (larval) Momorangia vallata Southcott, 1972: 36

Type locality: South Ridge, Low Peak, Mt. Luxmore, S. side Te Anau at 4150 ft. Distribution: New Zealand Hosts: cicada (Melampsalta oromelaena – Cicadidae, Homoptera), moth (Agrotis ypsilon, Noctuidae, Lepidoptera)

Subgenus Neomomorangia Fain et Santiago-Blay (larval) Neomomorangia Fain & Santiago-Blay, 1993: 340

Type species: Momorangia (Neomomorangia) asphaerae Fain et Santiago-Blay, 1993


137

Momorangia (Neomomorangia) asphaerae Fain et Santiago-Blay (larval) Momorangia (Neomomorangia) asphaerae Fain & Santiago-Blay, 1993: 340

Type locality: Minas Gerais Distribution: Brazil Host: Asphaera sp. (Chrysomelidae: Alticinae, Coleoptera)

Tribe Charletoniini Southcott Charletoniini Southcott, 1961a: 522; 1988a: 203

Type genus: Charletonia Oudemans, 1.5.1910 – larval (syn. Sphaerolophus Berlese, 1910 – adult)

Genus Abrolophus Berlese (larval and ad.) Abrolophus Berlese, 1891, A.M.S. 59 (1), 2 (as subgenus of Rhyncholophus; non Abrolophus Berlese, 1893 = Leptus; nec Berlese, 1916 = Leptus) Discolophus Kishida, 1939: 48

Type species of Abrolophus Berlese: Abrolophus quisquiliarum (Hermann) Berlese, 1891 Type species of Discolophus Kishida: Discolophus tokunagai Kishida, 1939

Abrolophus aequalipes (Schweizer et Bader)(comb. nov.)(ad.) Balaustium aequalipes Schweizer & Bader, 1963: 301

Type locality: National park, Val da la Föglia, 2000 m Distribution: Switzerland

Abrolophus aitapensis (Southcott)(larval) Hauptmannia aitapensis Southcott, 1948: 256; Haitlinger, 1987b: 915; 1987g: 721; 2002d: 118; 2006a: 84; 2006d: 219 Abrolophus aitapensis: Haitlinger, 2006a: 84

Type locality: Deia Creek in the Aitape region, New Guinea Distribution: Macao, Madagascar, New Guinea, Vietnam, Indonesia (Bali)

Abrolophus albidoinfumatus Berlese (ad.)(Inc. sed.) Abrolophus albidoinfumatus Berlese, 1916b: 127

Type locality: East Africa Distribution: East Africa

Abrolophus anomalus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium anomalus Schweizer & Bader, 1963: 289

Type locality: “Nordostschweiz, Hoher Kasten (Säntisgebiet), 1950 m” Distribution: Switzerland

138


Abrolophus artemisiae (Schrank)(ad.) Acarus artemisiae Schrank, 1803 Smaridia artemisiae (Schrank) Balaustium artemisiae (Schrank): Buitendijk, 1945: 348 Balaustium lapidarium Willmann: 1950: 191; 1951: 148; 1956: 248; Franz, 1954: 383 (as “n. spec. i.l.”) Abrolophus lapidarius (Willmann): Gabryś, 1988: 24 Abrolophus artemisiae (Schrank): Gabryś & Mąkol, 1990: 709; 1994: 129; 1995: 383

Type locality: Austria Distribution: Austria, France, Germany, Holland, Hungary, Poland

Abrolophus arvensis (Banks)(ad.) Erythraeus arvensis Banks, 1912: 96 Legneria arvensis (Banks): Oudemans, 1941: 179 Abrolophus arvensis (Banks): Southcott, 1961a: 536

Type locality: Utah, Salt Lake City Distribution: USA (Utah)

Abrolophus baderi Gabryś (ad.) Balaustium longipes Schweizer & Bader, 1963: 289 Abrolophus baderi: Gabryś, 2000c: 66 (nom. nov. pro Balaustium longipes Schweizer et Bader non Balaustium longipes Willmann, both Abrolophus)

Type locality: Saillon (Wallis) Distribution: Switzerland

Abrolophus basiliensis (Schweizer et Bader)(comb. nov.)(ad.) Balaustium basiliense Schweizer & Bader, 1963: 300

Type locality: Dinkelberg: Bettingen, 350 m Distribution: Switzerland

Abrolophus basumtwiensis Haitlinger (larval) Abrolophus basumtwiensis Haitlinger, 2006g: 113

Type locality: Basumtwi, Kumasi (Ghana) Distribution: Ghana Host: “from herbaceous plants”

Abrolophus batotensis Khot (ad.) Abrolophus batotensis Khot, 1965: 74

Type locality: Batot (Kashmir State) Distribution: India (Kashmir)


139

Abrolophus benoni (Haitlinger) (larval) Hauptmannia benoni Haitlinger, 2002d: 115; 2004h: 215 Abrolophus benoni: Haitlinger, 2007b: 59; 2007d: 538

Type locality: Santana, Madeira Distribution: Canary Is. (La Palma), Madeira Host: from plants

Abrolophus bohdani (Haitlinger) (larval) Hauptmannia bohdani Haitlinger, 2003d: 603; 2003f: 67 Abrolophus bohdani: Haitlinger, 2007b: 59

Type locality: Strawczyn n. Kielce Distribution: Poland Host: from plants

Abrolophus brevipalpis (Schweizer et Bader)(comb. nov.)(ad.) Balaustium brevipalpe Schweizer & Bader, 1963: 287

Type locality: National park, Alp Murter, 2500 m Distribution: Switzerland

Abrolophus brevus (Womersley)(ad.) Belaustium brevum Womersley, 1934: 227 Balaustium brevum Wom.: Halliday, 1998: 56 Abrolophus brevus (Womersley): Halliday, 2001: 327

Type locality: Riverton, South Australia Distribution: Australia (S.A.)

Abrolophus complanatus (Berlese)(ad.) Achorolophus complanatus Berlese, 1910: 348 Abrolophus complanatus (Berlese): Bernini et al., 1994-96: 40

Type locality: Germany Distribution: Germany, Italy

Abrolophus crassitarsus (Schweizer)(ad.) Balaustium crassitarsum Schweizer, 1951: 155 Abrolophus crassitarsus (Schweizer): Southcott, 1961a: 536; Gabryś & Mąkol, 1996: 489; Haitlinger, 2002e: 147

Type locality: La Schera, 2550 m, Swiss Alps Distribution: Hungary, Poland, Switzerland

Abrolophus crimensis Haitlinger (larval) Abrolophus crimensis: Haitlinger, 2007c

Type locality: Crimea, 7 km east from Alushta (44°45’ N, 34°29’ E)

140


Distribution: Ukraine Host: from herbaceous plants

Abrolophus curtipapillus (Schweizer)(ad.) Balaustium curtipapillum Schweizer, 1951: 155; Franz, 1954: 383; Cooreman, 1955c: 13; Schweizer & Bader, 1963: 282 Abrolophus curtipapillum (Schweizer): Bernini et al., 1994-96: 40

Type locality: Swiss Alps, Buffalora, Val Ftur, 2000 m Distribution: Austria, Italy, Switzerland

Abrolophus cuspidatus Shiba (ad.) Abrolophus cuspidatus: Shiba, 1969: 104

Type locality: Shiga Heights IBP Area Distribution: Japan

Abrolophus delhiensis Khot (ad.) Abrolophus delhiensis Khot, 1965: 71


Abrolophus densipapillus (Schweizer)(ad.) Balaustium densipapillum Schweizer, 1951: 157; Sellnick, 1958: 42; Schweizer & Bader, 1963: 293 Abrolophus densipapillum Schw.: Schmölzer, 1993: 61

Type locality: Swiss Alps, National park, Murtaröl, Val dal Botsch, 2000-2500 m Distribution: Sweden, Switzerland

Abrolophus engadinensis (Schweizer et Bader)(comb. nov.)(ad.) Balaustium engadinense Schweizer & Bader, 1963: 297

Type locality: Nationalpark, S-chanf, 1650 m Distribution: Switzerland

Abrolophus epigaeus (Berlese)(ad.) epigaeus Berlese, 1918 Abrolophus epigaeus (Berlese): Bernini et al., 1994-96: 40

Type locality: Italy Distribution: Italy

Abrolophus evansi (Schweizer et Bader)(comb. nov.)(ad.) Balaustium evansi Schweizer & Bader, 1963: 294

Type locality: Stabenchod, 2600 m Distribution: Switzerland


141

?Abrolophus falconeri (Hull)(ad.) Achorolophus falconerii Hull, 1918: 29

Type locality: Yorkshire Distribution: England

Abrolophus frisius Southcott (ad.) Balaustium insulare Willmann, 1952: 168 Abrolophus frisius Southcott, 1961a: 534 (nom. nov. pro Balaustium insulare Willmann, 1952, non Abrolophus insularis Womersley, 1934)

Type locality: Wangerooge I. Distribution: Germany

Abrolophus glauerti (Womersley) (ad.) Belaustium glauerti Womersley, 1934: 226; Halliday, 1998: 56 Balaustium glauerti Womersley: Halliday, 1998: 56 Abrolophus glauerti (Womersley): Halliday, 2001: 327

Type locality: Perth (W.A.) Distribution: Australia (W.A.)

Abrolophus halberti (Cooreman)(ad.) Rhyncholophus Passerinii Halbert, 1920, nec Rh. passerinii Berlese, 1904 Balaustium halberti: Cooreman, 1946: 7; Turk, 1953: 26 Abrolophus halberti: Gabryś, 1992b: 233

Type locality: Malahide, Ireland Distribution: Cyprus, Ireland

Abrolophus handschini (Schweizer et Bader)(comb. nov.)(ad.) Balaustium handschini Schweizer & Bader, 1963: 286

Type locality: Konkordia, 2800 and 2850 m Distribution: Switzerland

Abrolophus humberti (Haitlinger) (larval) Hauptmannia humberti Haitlinger, 1996: 444; 2003g: 67 Abrolophus humberti: Haitlinger, 2007b: 59

Type locality: Tatra Mountains, Tomanowa Valley, alt. 1450 m Distribution: Poland

Abrolophus iacobi Feider (ad.) Abrolophus iacobi Feider, [1980: 3], 1981: 68

Type locality: Romania Distribution: Romania

142


Abrolophus incanescens Berlese (ad.) Abrolophus incanescens Berlese, 1916b: 128

Type locality: La Plata Distribution: La Plata (? Argentina)

Abrolophus insularum (Womersley)(ad.) Belaustium insularum Womersley, 1934: 229 Abrolophus insularis (Womersley): Southcott: 1961a: 534; Halliday, 1998: 56

Type locality: Rottnest Island, West Australia Distribution: Australia (West Australia)

Abrolophus iraninejadi Saboori et Hajiqanbar (larval) Abrolophus iraninejadi: Saboori & Hajiqanbar, 2005: 149

Type locality: Miandoab (Iran) Distribution: Iran Host: free-living

Abrolophus jurassicus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium jurassicum Schweizer & Bader, 1963: 299

Type locality: Jura, Schauenburgerfluh, 660 m Distribution: Switzerland

Abrolophus khanjanii (Haitlinger et Saboori) (larval) Hauptmannia khanjanii Haitlinger & Saboori, 1996:123; Haitlinger, 2002d: 118 Abrolophus khanjanii: Haitlinger, 2007b: 59

Type locality: Toiserkan Distribution: Iran Host: from plants (Medicago sativa – alfalfa)

Abrolophus laevicristatus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium laevicristatum Schweizer & Bader, 1963: 302

Type locality: Jura, Schauenburgerfluh, 660 m Distribution: Switzerland

Abrolophus littorale (Womersley)(ad.) Belaustium littorale Womersley, 1934: 226 Abrolophus littorale (Womersley): Halliday, 2001: 327

Type locality: Poin Perron, West Australia, under seaweed Distribution: Australia (W.A.)


143

Abrolophus longicollis (Oudemans)(larval) Achorolophus longicollis Oudemans, 1910a: 31 Hauptmannia longicollis (Oudemans): Oudemans, 1910b: 49; 1912: 181; Buitendijk, 1945: 347; Sellnick, 1958: 43; Lindroth et al., 1973: 131; Haitlinger, 1987h: 351; 2002d: 118; 2002e: 148; 2003f: 7; 2003g: 66; Gabryś, 1988: 24 Abrolophus longicollis (Oud.): Haitlinger, 2007b: 59

Type locality: Sneek, Holland Distribution: Holland, Czech Rep., Iceland, Poland, Sweden

Abrolophus longipapillus (Schweizer)(comb. nov.)(ad.) Balaustium longipapillum Schweizer, 1951: 154; Schweizer & Bader, 1963: 283

Type locality: Murter, Mt. Mezdi (S-charl), 2500 m Distribution: Switzerland

Abrolophus longipes (Willmann) (ad.) Balaustium longipes Willmann, 1951c: 150 Abrolophus longipes: Gabryś, 2000c: 66

Type locality: Zurndorf, Austria Distribution: Austria

Abrolophus longulus (Willmann)(comb. nov.)(ad.) Balaustium longulum Willmann, 1951c: 149; Franz, 1954: 383

Type locality: Marz-Rohrbach, 386 m (Austria) Distribution: Austria

Abrolophus longus (Schweizer)(ad.) Balaustium longum Schweizer, 1951: 159 Abrolophus longus (Schweizer): Southcott, 1961a: 536; Mihelčič, 1971: 145

Type locality: Swiss Alps, Val Sassa, Furcla Trupchum, 2070-2786 m Distribution: Switzerland

Abrolophus marinensis Haitlinger (larval) Abrolophus marinensis Haitlinger, 2007d: 538

Type locality: Marine de Fatimore (Corsica) Distribution: France (Corsica) Host: “from herbaceous plants”

Abrolophus miniatus (Hermann)(ad.) Trombidium miniatum Hermann, 1804: 28 Trombidium hyperboreum Thorell, 1872: 162 Smaridia miniata (Hermann): Dugès, 1834: 12 (with”?”)

144


Rhyncholophus mineatus (Hermann): Halbert, 1915: 115 Rhyncholophus miniatus (Herm.): Trägårdh, 1910: 488; Jablonowski, 1918: 3; Schweizer, 1922: 81 Rhyncholophus albicomus: L. Koch, 1879: 125 Rhyncholophus hyperboreus (Thorell): Trägårdh, 1901: 60 Belaustium miniatum (Hermann): Hull, 1918: 26 Balaustium miniatum (Hermann): Irk,1939b: 168; Schweizer, 1951: 152; Turk, 1953: 26; Franz, 1954: 383; Bottazzi, 1956: 43; Sellnick, 1958: 42; Mihelčič, 1958b: 47; Schweizer & Bader, 1963: 290; Lindroth et al., 1973: 130 Erythraeus miniatus (Herm.): Trägårdh, 1904: 55-56 Abrolophus miniatus (Hermann): Schmölzer, 1993: 61; Gabryś & Mąkol, 1990: 709; 1994: 129; 1995: 383; 1996: 489; Gabryś, 1997: 226

Type locality: France Distribution: Algeria, Austria, France, Germany, Great Britain, Greenland, Hungary, Ireland, Italy, Poland, Russia, Switzerland

Abrolophus mirabelae Haitlinger (larval) Abrolophus mirabelae Haitlinger, 2007b: 57

Type locality: many, holotype from Sankt Möritz, 1850 m, Switzerland Distribution: France, Switzerland Host: from herbaceous plants

Abrolophus mortenseni (Southcott) (larval) Hauptmannia mortenseni Southcott, 1994: 173; Haitlinger, 2002d: 118 Abrolophus mortenseni: Haitlinger, 2007b: 59

Type locality: Koh Chang, Thailand Distribution: Thailand Host: Psocoptera

Abrolophus neobrevicollis Zhang et Goldarazena (larval) Abrolophus neobrevicollis Zhang & Goldarazena, 1996: 127; Goldarazena, Jordana & Zhang, 1999: 31; Haitlinger, 2002d: 115

Type locality: Pitillas, Navarra (Northern Spain) Distribution: Madeira, Spain Host: Chirothrips aculeatus (Thripidae, Thysanoptera)

Abrolophus newmani (Womersley) (ad.) Belaustium newmani Womersley, 1934: 228 Balaustium newmani Womersley: Halliday, 1998: 56 Abrolophus newmani (Womersley): Halliday, 2001: 327

Type locality: Perth (W.A.) Distribution: Australia (W.A.)


145

Abrolophus nivalis (Schmölzer) (larval) Hauptmannia nivalis Schmölzer, 1956: 560 Abrolophus nivalis: Haitlinger, 2007c

Type locality: Refuge Temple-Ecrins etc., Dauphiné, 2100-2740 m Distribution: France Host: in grass

Abrolophus norvegicus (Thor)(ad.) Rhyncholophus (Ritteria) norvegica Thor, 1900: 7 Rhyncholophus norvegicus (Thor): Halbert, 1915: 115 Achorolophus norvegicus Thor: Hull, 1918: 28 Leptus norvegicus (Thor): Turk, 1953: 26 Abrolophus norvegicus (Thor): Gabryś & Mąkol, 1991: 709; 1994: 129; 1995: 383; Gabryś, 1997: 226; Mąkol & Gabryś, 1999: 30; Haitlinger, 2002e: 148

Type locality: Norway Distribution: Great Britain, Hungary, Ireland, Norway, Poland

Abrolophus novaeguinensis (Gunther) (comb. nov.)(ad.) Belaustium novaeguinensis Gunther, 1941: 156

Type locality: Watut valley, Territory of New Guinea, “entagled in the scrotal hairs of a wallaby (Macropus sp.)” Distribution: Papua New Guinea

Abrolophus passerinii (Berlese)(ad.) Erythraeus Passerinii Berlese, 1904: 16 Abrolophus passerini (Berlese): Gabryś, 1992c: 233; 1997: 226; Gabryś & Mąkol, 1994: 130

Type locality: Meloria, Italy Distribution: Ireland, Italy, Poland

Abrolophus penelopae Haitlinger (larval) Abrolophus penelopae Haitlinger, 2006h: 81

Type locality: Jinka, South Ethiopia Distribution: Ethiopia Host: from plants

Abrolophus perifi (Schweizer et Bader)(comb. nov.)(ad.) Balaustium perifi Schweizer & Bader, 1963: 288

Type locality: National park, “Weg nach Punt Perif”, 1830 m Distribution: Switzerland

146


Abrolophus perlongus (Schweizer)(ad.) Balaustium perlongum Schweizer, 1951: 160 Abrolophus perlongum (Schweizer): Southcott, 1961a: 536

Type locality: Swiss National park, Stabel-chod, 2200 m Distribution: Switzerland

Abrolophus pseudolongicollis (Haitlinger) (larval) Hauptmannia pseudolongicollis Haitlinger, 1987h: 351; 1996; 2002d: 118; 2003d: 3; 2003g: 67; 2004d: 144; Gabryś, 1988: 24 Abrolophus pseudolongicollis: Haitlinger, 2007b: 57

Type locality: Lubiaz, Poland Distribution: Croatia, Italy, Liechtenstein, Poland, Slovenia

Abrolophus pseudolongicollis kiejstuti Haitlinger (larval) Abrolophus pseudolongicollis kiejstuti Haitlinger, 2006f: 36; 2007a: 71; 2007c

Type locality: 12 km E from Zarica (Bosnia and Herzegovina), Zagrade n. Ivangrad (Montenegro) Distribution: Bosnia and Herzegovina, Croatia, Greece, Moldova, Montenegro, Slovenia, Ukraine Host: from herbaceous plants

Abrolophus pulcher (G. Canestrini)(ad.) Rhyncholophus pulcher G. Canestrini, 1885: 148 Abrolophus pulcher (G. Canestrini): Bernini et al., 1994-96: 40

Type locality: Genovesato, Italy Distribution: Italy

Abrolophus quadrisetus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium quadrisetum Schweizer & Bader, 1963: 284

Type locality: Berner Alpen, Désorhorn, 2500 m Distribution: Switzerland

Abrolophus quinquesetus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium quinquesetum Schweizer & Bader, 1963: 281

Type locality: Berner Alpen, Finsteraarhorn, 3287 m Distribution: Switzerland

Abrolophus quisquiliarus (Hermann)(ad.) Trombidium quisquiliarum Hermann, 1804: 32 Erythraeus quisquiliarum: Latreille, 1817: 48; 1823: 168; 1829: 284 Balaustium quisquiliarum (Hermann): Oudemans, 1937: 1948; Willmann, 1939: 129; 1949: 129; 1954: 244; 1956: 248; Buitendijk, 1945; Schweizer, 1951: 154; Schmölzer, 1956: 560; Turk, 1953: 26; Franz, 1954: 383; Cooreman, 1955c: 14; Sellnick, 1958: 42; Schweizer & Bader, 1963: 295


147

Belaustium quisquiliarum (Herm.): Oudemans, 1910: 13; 1913a: 13; 1913b: 6; Hull, 1918: 27 Smaridia quisquiliarum ?: Dugès, 1834: 12 Smaridia quisquiliarum: Dugès, 1836: 493 Erythraeus nivosus Contarini, 1847: 162 (nom. nudum) Rhyncholophus quisquiliarum (Hermann): C.L. Koch, 1842: 50; Canestrini, 1885: 150; Berlese, 1900: 163 Abrolophus quisquiliarum (Hermann): Starkoff & Starkoff, 1950: 96; Gabryś, 1988: 24; Schmölzer, 1993: 61; Bernini et al., 1994-96: 40; Haitlinger, 2002e: 148 Abrolophus quisquiliarus (Hermann): Gabryś, 1988: 24; Gabryś & Mąkol, 1991: 709; 1994: 130

Type locality: Strassburg (France) Distribution: Austria, Belgium, England, France, Germany, Holand, Hungary, Italy, Poland, Russia (Kaliningrad), Spain, Switzerland

Abrolophus quisquiliarus sigma (Mihelčič) (ad.) Balaustium quisquiliarum sigma Mihelčič, 1964: 298

Type locality: “Zetterfeld bei Lienz”, ca.1800 m Distribution: Austria

Abrolophus raripapillus (Schweizer)(ad.) Balaustium raripapillum Schweizer, 1951: 159; Schmölzer, 1956: 560 Abrolophus raripapillus Mihelčič, 1971: 145

Type locality: Cluozza, Alp Casana, 1800-2200 m, Swiss Alps Distribution: France, Switzerland

Abrolophus rhopalicus (C.L.Koch)(ad.) Rhyncholophus rhopalicus C.L. Koch, 1837: 16; 1842: 50 Belaustium rhopalicus (C.L. Koch): Oudemans, 1913a: 13; 1913b: 3; Sellnick, 1931: 696; Willmann, 1937: 160 Balaustium rhopalicus (C.L. Koch): Buitendijk, 1945: 349 Balaustium rhopalicum (C.L. Koch): Sellnick, 1940: 37; Willmann, 1951c: 148; 1956: 248; Schweizer, 1951: 156; Lindroth et al., 1973: 130 Abrolophus rhopalicum (C.L. Koch): Schmölzer, 1993: 61 Abrolophus rhopalicus (C.L. Koch): Southcott, 1961a: 536; Mihelčič, 1971: 145; Gabryś, 1988: 24; 1997: 226

Type locality: Regensburg, Germany Distribution: Austria, Germany, Greece, Holland, Iceland, Poland, Switzerland

Abrolophus ripicola (Womersley) (ad.) Belaustium ripicola Womersley, 1934: 228 Balaustium ripicola Wom.: Halliday, 1998: 56 Abrolophus ripicola Womersley: Khot, 1965: 71; Halliday, 2001: 327

Type locality: Hindmarsh River, Victor Harbour, South Australia Distribution: Australia, India

148


Abrolophus rubipes (Berlese et Trouessart) (ad.) Rhyncholophus miniatus Hermann, var. rubipes Berlese & Trouessart, 1889: 11 Rhyncholophus rubripes: Halbert, 1915: 115; 1920: 145 Abrolophus rubipes: Witte & Storch, 1973: 441; Wendt & Witte, 1985: 685 Belaustium rubripes: Hull, 1918: 26; Haarløv, 1942: 33 Balaustium rubripes: Sellnick, 1949: 130; Evans & Browning, 1953: 418; Turk, 1953: 26 Ritteria hirsutus: George, 1910: 182 (= Rhyncholophus hirsutus George, see Hull, 1918: 26)

Type locality: Wimereux (Pas-de-Calais) Distribution: England, Faroe Is., France, Greenland, Ireland, Sweden

Abrolophus sabulosus (Halbert) (ad.) Rhyncholophus sabulosus Halbert, 1915: 116 Belaustium sabulosum (Halbert): Hull, 1918: 27 Balaustium sabulosum (Halbert): Schweizer, 1951: 158; Turk, 1953: 26; Mihelčič 1958b: 47 Balaustium sabulosum (Halbert) var. majus Mihelčič, 1958b: 47 Abrolophus sabulosum (Halbert): Schmölzer, 1993: 61 Abrolophus sabulosus (Halbert): Mihelčič, 1971: 145

Type locality: Mulranny sandhills, County Mayo, Ireland Distribution: Austria, England, Ireland, Spain, Switzerland

Abrolophus schweizeri (Evans) (ad.) Balaustium sabulosum Schweizer, 1951: 158 (nec B. sabulosum Halbert) Balaustium schweizeri Evans, 1952: 670 Abrolophus schweizeri (Evans): Gabryś, 2001: 227

Type locality: Reading, England Distribution: England, Switzerland

Abrolophus scopularius (Hull) (comb. nov.) (ad.) Belaustium scopularium Hull, 1918: 27

Type locality: St. Anne’s-on-the-Sea, Lancashire Distribution: England

Abrolophus setipapillus Berlese (ad.) Abrolophus setipapillus Berlese, 1916b: 129

Type locality: La Plata Distribution: La Plata (? Argentina)

Abrolophus simplex Berlese (ad.) Abrolophus simplex Berlese, 1916b: 127

Type locality: South Ethiopia, Kounhi Distribution: Ethiopia


149

Abrolophus spectabilis Berlese (ad.) Abrolophus spectabilis Berlese, 1916b: 126

Type locality: East Africa Distribution: East Africa

Abrolophus spiculosus Ueckermann (ad.) Abrolophus spiculosus Ueckermann, in Meyer & Ueckermann, 1989: 20

Type locality: Rooiputs, Kalahari Gemsbok Nat. Park, Cape Provimce Distribution: South Africa

Abrolophus strojnyi Gabryś (ad.) Abrolophus strojnyi Gabryś, 1992b: 233; Gabryś & Mąkol, 1994: 130; Haitlinger, 2002e: 147

Type locality: Plazowska Gora n. Narol, Przemysl province (Poland) Distribution: Hungary, Poland

Abrolophus subquisquiliarus (Schweizer et Bader)(comb. nov.) (ad.) Balaustium subquisquiliarum Schweizer & Bader, 1963: 295

Type locality: Val d’Acqua, 2000 m Distribution: Switzerland

Abrolophus tardus (Halbert)(ad.) Rhyncholophus tardus Halbert, 1915: 116; 1920: 145 Belaustium tardum (Halbert): Hull, 1918: 27 Balaustium tardum (Halbert): Willmann, 1941: 64; 1950: 191; 1951c: 148; 1954: 244; Turk, 1953: 26 Abrolophus tardus (Halbert): Gabryś, 1988: 24; 1997: 226

Type locality: Bellacragher Bay, near Mulranny, Ireland Distribution: Austria, Bosnia and Herzegovina, Czech Rep., England, Ireland, Poland

Abrolophus tonsor (Southcott) (larval) Hauptmannia tonsor Southcott, 1996: 274 Abrolophus tonsor: Haitlinger, 2007b: 59; 2007d: 538

Type locality: New South Wales Distribution: Australia (NSW)

Abrolophus trinotatus Berlese (ad.) Abrolophus trinotatus Berlese, 1916b: 128

Type locality: La Plata Distribution: La Plata [Argentina]

150


Abrolophus trisetus (Schweizer et Bader)(comb. nov.)(ad.) Balaustium trisetum Schweizer & Bader, 1963: 286

Type locality: Mt. Tavrü, 2500 m Distribution: Switzerland

Abrolophus unimiri Haitlinger (larval) Abrolophus unimiri Haitlinger, 2006a: 84

Type locality: Yangshuo n. Guilin Distribution: China

Abrolophus unisetus (Schweizer et Bader) (comb. nov.) (ad.) Balaustium unisetum Schweizer & Bader, 1963: 298

Type locality: Säntis, 2500 m Distribution: Greenland, Switzerland

Abrolophus vittatus (Hull) (comb. nov.) (ad.) Belaustium vittatum Hull, 1918: 28

Type locality: Durham Distribution: Great Britain

Abrolophus welbourni Yao Hongren, Snider et Snider (larval) Abrolophus welbourni Yao Hongren, Snider R.J. & Snider R.M., 2000: 150; Haitlinger, 2007d: 538

Type locality: Michigan, Upper Peninsula, Dickinson County Distribution: U.S.A.

Abrolophus zelandicus Luxton (ad.) Abrolophus zelandicus Luxton, 1989: 416

Type locality: New Zealand Distribution: New Zealand

Genus Andrevella Southcott (larval) Andrévella Southcott, 1961a: 538

Type species: Erythraeus parkeri André, 1930

Andrevella parkeri (André) (larval) Erythraeus parkeri André, 1930: 545 Andrevella parkeri (André): Southcott, 1961a: 539

Type locality: Hyères, Var, France Distribution: France Host: larva of Cicindelidae (Coleoptera)


Andrevella persiangulfia Saboori, Hajiqanbar et Kamali (larval) Andrevella persiangulfia Saboori, Hajiqanbar & Kamali, 2007: 47

Type locality: The Persian Gulf beach Distribution: Iran Host: free living on soil

Genus Cecidopus Karsch (ad.) Cecidopus Karsch, 1879: 562 Sphaerolophus (Eupodolophus) Berlese, 1914: 146-7 Fessoniella Hirst, 1924: 1075

Type species of Cecidopus Karsch: Cecidopus diversipes Karsch, 1879 Type species of Fessoniella Hirst: F. mitchelli Hirst, 1924

Cecidopus chubbi (Berlese) (ad.) Sphaerolophus (Eupodolophus) chubbi Berlese, 1914: 147; Southcott, 1961a: 530 Cecidopus chubbi (Berlese): Cooreman, 1953: 3

Type locality: Clairmont near Durban Distribution: South Africa

Cecidopus diversipes Karsch (ad.) Cecidopus diversipes Karsch, 1879: 562; Cooreman, 1953: 2

Type locality: Ceylon (Sri Lanka) Distribution: Sri Lanka

Cecidopus mitchelli (Hirst) (ad.) Fessoniella mitchelli Hirst, 1924: 1075; Southcott, 1961a: 530 (? syn. of C. chubbi) Cecidopus mitchelli (Hirst): Cooreman, 1953: 3

Type locality: Table Mountain, Cape of Good Hope Distribution: South Africa

Cecidopus nigeriae Beron (ad.) Cecidopus nigeriae Beron, 2000: 70

Type locality: Wase Rock Game Res., Plateau State Distribution: Nigeria

Cecidopus qadrii Ahsan et Anwarullah (ad.) Cecidopus qadrii Ahsan & Anwarullah, 1970: 47

Type locality: Karachi Distribution: Pakistan

151

152


Cecidopus shyamae Khot (ad.) Cecidopus shyamae Khot, 1965: 69


Cecidopus straeleni Cooreman (ad.) Cecidopus straeleni Cooreman, 1953: 3

Type locality: Kalungwe, 1740 m, Nat. Park Upemba (DR Congo) Distribution: DR Congo

Genus Charletonia Oudemans Charletonia Oudemans, 1910c: 73 (larval)

Type species: Erythraeus singularis Oudemans, 1910 Sphaerolophus Berlese, 1910: 349 (adult)

Type species: Rhyncholophus globiger Berlese, 1885 Cavannea Berlese, 1910: 350 (ad.) (subgenus ?)

Type species: Rhyncholophus cavannae Berlese, 1885 Hehlenia Oudemans, 1941: 181

Type species: Rhyncholophus papillosus C.L. Koch, 1837 non Trombidium papillosum Hermann, 1804

Charletonia agatae Haitlinger (larval) Charletonia agatae Haitlinger, 1987g: 719

Type locality: Antananarivo, Isorana n. Fianarantsoa Distribution: Madagascar Host: from plants

Charletonia ahwazensis Haitlinger et Saboori (larval) Charletonia ahwazensis Haitlinger & Saboori, 2007: 65

Type locality: Ahwaz, Karaj (1312 m), Iran Distribution: Iran Host: undetermined Orthoptera

Charletonia alarobiaensis Haitlinger (larval) Charletonia alarobiaensis Haitlinger, 1987g: 716

Type locality: Alarobia – Vahiposa, middle Madagascar Distribution: Madagascar Host: Orthoptera indet. and plants

Charletonia alvedae Haitlinger (larval) Charletonia alvedae Haitlinger, 2000d: 384

Type locality: Puerto Maldonado, by the river Madre de Dios (Peru)


153

Distribution: Peru Host: ? Crotonotus sp. (Orthoptera, Pseudophyllidae)

Charletonia aranea Southcott (larval) Charletonia aranea Southcott, 1966: 801; 1991a: 79

Type locality: Srinagar, Kashmir Distribution: Kashmir (India) Host: Araneae

Charletonia arborum (Vitzthum) (comb. nov.) (ad.) Sphaerolophus arborum Vitzthum, 1926: 152

Type locality: Depok, Java Distribution: Indonesia (Java)

Charletonia areolata (Trägårdh) (larval) Erythraeus areolatus Trägårdh, 1908: 33; Oudemans, 1910e: 107; 1912: 139; Kishida, 1929: 98; Lawrence, 1940: 403; Buitendijk, 1945: 349; Evans, 1953: 280; Feider, 1954: 1021 Charletonia areolata (Trägårdh): Southcott, 1961a: 527; 1966: 788; 1991a: 100

Type locality: Kibonoto, Kilimanjaro, 1300-1600 m (Tanzania) Distribution: Kenya, Mozambique, Tanzania, Zululand (South Africa) Hosts: Orthoptera (Acrididae), Heteroptera (Miridae, Lygaeidae), Coleoptera (Tenebrionidae, Melyridae), Hymenoptera (Braconidae), Diptera (Dolichopodidae), Araneae (Cyrtophora sp.)

Charletonia arlettae Haitlinger (larval) Charletonia arlettae Haitlinger, 1987g: 708

Type locality: Andoharonomaitso n. Fianarantsoa Distribution: Madagascar Host: Neuroptera indet.

Charletonia austisensis Haitlinger (larval) Charletonia austisensis Haitlinger, 2007d: 533

Type locality: Austis, Sardinia Distribution: Italy (Sardinia) Host: undetermined Orthoptera

Charletonia banksi Southcott (larval) Charletonia banksi Southcott, 1966: 796; 1991a: 68; Welbourn, 1983: 130

Type locality: Capella, Queensland Distribution: Australia (Queensland, Northern Territory) Host: Oedaleus australis (Orthoptera)

154


Charletonia barbata Southcott (larval) Charletonia barbata Southcott, 1966: 768; 1991a: 68; Welbourn, 1983: 131 Charletonia grandjeani Southcott, 1966: 792; 1983: 131; 1991: 68; Welbourn, 1983: 131

Type locality: Bowen, Queensland Distribution: Australia (Queensland, Northern Territory) Host: Valanga irregularis (Orthoptera: Acrididae)

Charletonia beninensis Haitlinger (larval) Charletonia beninensis Haitlinger, 2006g: 119

Type locality: Grand Popo (Benin) Distribution: Benin, Ghana Host: undetermined Orthoptera and herbaceous plants

Charletonia berlesiana (Paoli)(larval) Balaustium berlesianum Paoli, 1937: 187 Charletonia berlesiana (Paoli): Southcott, 1961a: 528; 1991a: 65; 1966: 709 Charletonia (Ch.) berlesiana: Bernini et al., 1994-96: 40

Type locality: Italy Distribution: Italy Hosts: Orthoptera (Dociostaurus maroccanus, Prionotropis hystrix appula, Decticus albifrons, D. londoni)

Charletonia blascoi Southcott (larval) Charletonia blascoi Southcott, 1993b: 51

Type locality: Spain Distribution: Spain Hosts: Solenoxyphus sauledai (Miridae, Heteroptera)

Charletonia braunsi (Oudemans) (larval) Erythraeus braunsi Oudemans, 1910e: 106; 1912: 147; Kishida, 1929: 99; Lawrence, 1940: 403; Buitendijk, 1945: 349; Feider, 1954: 1021 Charletonia braunsi (Oudemans): Eyndhoven, 1944: 52; Buitendijk, 1945: 349; Southcott, 1961a: 527; 1966: 738; Haitlinger, 2006h: 87; 2006g: 120

Type locality: “Loss – eiland” near Konakry, Guinea Distribution: Ethiopia, Ghana, Guinea Host: Truxalis sp. (Acrididae, Orthoptera)

Charletonia brunni (Oudemans)(larval) Erythraeus brunni Oudemans, 1910e: 108; 1912: 142; Kishida, 1929: 99; Feider, 1954: 1021 Charletonia brunni (Oudemans): Eyndhoven, 1944: 52; Southcott, 1961a: 527; 1966: 703; Haitlinger, 2006h: 87; 2006g: 119


155

Balaustium brunni (Oudemans): Buitendijk, 1945: 348

Type locality: Wari, Forcados River (Nigeria) Distribution: Benin, Ghana, Ethiopia, Nigeria Host: Pyrgomorphidae (Orthoptera)

Charletonia bucephalia Beron (larval) Charletonia bucephalia Beron, 1975: 73; Southcott, 1991a: 65; Haitlinger, 2000c: 386; 2004d: 149 Charletonia tamarae Haitlinger, 1984: 385

Type locality of Ch. bucephalia: Primorsko, distr. Burgas Type locality of Ch. tamarae: Agi Apostoli n. Kalamos, Greece, on Orthoptera indet. Distribution: Bulgaria, Croatia, Greece Host: Bucephaloptera bucephala (Orthoptera, Tettigoniidae)

Charletonia buforania (Womersley) (larval) Erythraeus buforanius Womersley, 1934: 245; Feider, 1954: 1019 Charletonia buforania (Womersley): Southcott, 1961a: 528; 1966: 724; 1991: 68

Type locality: Mullewa, Western Australia Distribution: Australia (Western Australia, South Australia) Host: Buforania sp. (Orthoptera, Acrididae)

Charletonia canadensis (Smiley)(comb. nov.)(ad.) Sphaerolophus canadensis Smiley, 1968: 15

Type locality: Vineland Station, Ontario Distribution: Canada (Ontario)

Charletonia cardinalis (Pallas, sensu C.L. Koch)(larval, adult) Acarus cardinalis Pallas, 1772: 44 Rhyncholophus cardinalis C.L.Koch, 1837: fa.16, t.10; Canestrini & Fanzago, 1877: 73 Trombidium (Otonyssus) sudeticum Oudemans, 1897: 118 (non Müller) Erythraeus singularis: Oudemans, 1910a: 31; 1912: 130 (larva) Charletonia singularis: Oudemans, 1910d: 73; Kawashima, 1961: 17; Southcott, 1961a: 525; 1966: 806; 1991a: 99; Shiba, 1976: 187; Haitlinger, 1985: 433; 1987d: 347 Hehlenia singularis (C.L. Koch): Oudemans, 1941: 181 Smaris singularis (C.L. Koch): Buitendijk, 1945: 350 Sphaerolophus cardinalis (C.L. Koch) : Franke, 1940: 154; Willmann, 1951c: 151; Franz, 1954: 382; Witte, 1977: 74; 1978: 160; Treat, 1980: 205 (adult) Charletonia cardinalis (C.L. Koch): Southcott, 1991a: 99; Gabryś & Mąkol, 1994: 129; 1995: 383; Haitlinger, 2000c: 384; 2002e: 148; 2003f: 7

Type locality of Rhyncholophus cardinalis C.L. Koch, 1837, ad.: Germany Type locality of Erythraeus singularis Oudemans, 1910, larval: Wageningen, Holland

156


Distribution: Austria, Czech Rep., Germany, Holland, Poland, Russia (Kaliningrad), Sweden; Malaya, Japan, Turkey, Azerbaydjan; Australia Hosts: Liburnia pellucida, Aphrophora sp. (Homoptera), Heteroptera, Phyllotreta undulata (Chrysomelidae, Coleoptera), Lepidoptera, Mammalia

Charletonia cavannae (Berlese)(comb. nov.)(ad.) Rhyncholophus Cavannae Berlese, 1885a: 137; Jablonowski, 1918: 3 Sphaerolophus (Cavannea) cavannae var. corcyraeus: Berlese, 1910: 350; Sellnick, 1931: 694 Cavannea cavannae (Berlese): Turk, 1953: 26 Sphaerolophus (Cavannea) cavannae (Berl.): Feider & Chioreanu, 1977: 270

Type locality: Piedimonte di Alife (“Fioretino, Campania ed Calabria”), Italy Distribution: England, Greece (Kerkira, “var. corcyreus Berl.”), Hungary, Italy

Charletonia cilissa (Cooreman) (comb. nov.) (ad.) Cavannea cilissa Cooreman, 1955a: 128 Sphaerolophus (Cavannea) cilissa (Cooreman)

Type locality: “Au pied de l’Amanus, à l’est de Beylan”, Asia Minor Distribution: Turkey

Charletonia cooremani (Feider et Chioreanu) (comb. nov.) (larval) Sphaerolophus (Cavannea) cooremani Feider & Chioreanu, 1977: 271

Type locality: Prosperous Bay, 184-500 m (Sainte-Helene) Distribution: Sainte-Helene

Charletonia cuglierensis Haitlinger (larval) Charletonia cuglierensis Haitlinger, 2007d: 535

Type locality: Cuglieri, Sardinia Distribution: Italy (Sardinia) Host: undetermined Orthoptera

Charletonia cursor (Vitzthum) (comb. nov.) (ad.) Sphaerolophus cursor Vitzthum, 1926: 156

Type locality: Djakarta (“Hoorn in der Bucht von Batavia”) Distribution: Indonesia (Java)

Charletonia dalegori Haitlinger (larval) Charletonia dalegori Haitlinger, 2003b: 50; 2004d: 149; 2006f: 37; 2007d: 533

Type locality: Kalavarda, Rhodes Distribution: Croatia, France (Corsica), Greece (incl. Rhodes) Host: Caliptamus italicus (Tettigoniidae), Acrididae, undet. Orthoptera


Charletonia damavandica Karimi Iravanlou, Kamali et Talebi (larval) Charletonia damavandica Karimi Iravanlou, Kamali & Talebi, 2002: 131

Type locality: Varamin Distribution: Iran Host: Orthoptera, Acrididae (Acrotylus insubricus, Mioscirus wagneri)

Charletonia danangensis Haitlinger (larval) Charletonia danangensis Haitlinger, 1986: 643

Type locality: Danang Distribution: Vietnam Host: Noctuidae (Lepidoptera)

Charletonia delhiensis (Khot) (comb. nov.) (ad.) Sphaerolophus delhiensis Khot, 1965: 66


Charletonia domawiti Haitlinger (larval) Charletonia domawiti Haitlinger, 2004f: 436


Charletonia dorotae Haitlinger (larval) Charletonia dorotae Haitlinger, 1987g: 711

Type locality: Ambila – Levaitso, East Madagascar Distribution: Madagascar Host: undet. Orthoptera

Charletonia edytae Haitlinger (larval) Charletonia edytae Haitlinger, 1987g: 705

Type locality:Mananjary. East Madagascar Distribution: Madagascar Host: Odonata indet.

Charletonia enghoffi Southcott (larval) Charletonia enghoffi Southcott, 1991a: 103

Type locality: Tenerife, Barranco de las Cuevas, 200-280 m Distribution: Canary Islands (Tenerife) Host: Bunochelis canariana (Phalangiidae, Opiliones)

157

158


Charletonia falcata Southcott (larval) Charletonia falcata Southcott, 1966: 699; 1991a: 79

Type locality: Nusite, Aitape Region, New Guinea Distribution: New Guinea Host: Curculionidae (Col.)

Charletonia feideri Southcott (larval) Charletonia feideri Southcott, 1966: 752; 1991: 69

Type locality: many places in Queensland, Victoria, ACT, Northern Territory, Western Australia and Tasmania Distribution: Australia, Tasmania Host: Gastrimargus musicus (Acrididae, Orthoptera)

Charletonia froggatti (Oudemans) (larval) Erythraeus froggatti Oudemans, 1910a: 31 (nec Erythraeus froggatti: Berlese, 1920 = Morieria novae – hollandiae Oudemans, 1941); 1912: 144; Buitendijk, 1945: 348 (frogatti) Charletonia froggatti (Oudemans): Oudemans, 1910c: 73; Southcott, 1961a: 527; 1966: 715; 1991a: 65

Type locality: Conjola, New South Wales Distribution: Australia (New South Wales) Host: Podacanthus wilkinsoni (Phasmodea) and Trombidiidae s.l. gen. sp.

Charletonia ghanensis Haitlinger (larval) Charletonia ghanensis Haitlinger, 2006g: 116

Type locality: Elmina Distribution: Ghana Host: “from herbaceous plants”

Charletonia gigas (Khot) (comb. nov.) (ad.) Sphaerolophus (Cavannea) gigas Khot, 1965: 64


Charletonia glifadaensis Haitlinger (larval) Charletonia glifadaensis Haitlinger, 2003b: 50; 2007d: 532

Type locality: A. Glifada, Rhodes Distribution: France (Corsica), Greece (Rhodes) Host: Oedipoda sp. (Orthoptera: Acrididae: Oedipodinae)

Charletonia globigera (Berlese)(ad.) Rhyncholophus globiger Berlese, 1885a: 139; Schweizer, 1922: 83 Belaustium globigerum (Berlese): Oudemans, 1905c: 9; 1913b: 1; 1914: 2


159

Sphaerolophus globiger (Berlese): Berlese, 1910: 349; Cooreman, 1943: 14; Willmann, 1949: 128; Turk, 1953: 25; Weidner, 1954: 41; Franz, 1954: 382; Sellnick, 1958: 42; Karppinen, 1958: 45; Schweizer & Bader, 1963: 310; Gabryś, 1988: 24 Sphaerolophus globigerum (Berlese): Buitendijk, 1945: 349 Achorolophus globiger Berlese: Hull, 1918: 28 Rhyncholophus (Ritteria) globigera Berlese: Thor, 1900: 7 Charletonia globigera (Berlese): Gabryś & Mąkol, 1994: 129; 1995: 383 Charletonia (Ch.) globigera: Bernini et al., 1994-96: 40

Type locality: Pratiglione (Ivrea, Italy) Distribution: Austria, Belgium, England, Finland, France, Germany, Holland, Italy, Norway, Poland, Sweden, Switzerland; ? Guatemala

Charletonia goliatha (Berlese)(ad.) Sphaerolophus goliathus Berlese, 1910: 349 Charletonia goliatha (Berlese): Southcott, 1991: 65

Type locality: Sidney, Australia Distribution: Australia

Charletonia grandpopensis Haitlinger (larval) Charletonia grandpopensis Haitlinger, 2006g: 116

Type locality: Grand Popo (Benin) Distribution: Benin Host: “from herbaceous plants”

Charletonia hermanni Southcott (larval) Charletonia hermanni Southcott, 1966: 749; 1991a: 67

Type locality: Scott’s Creek, Victoria Distribution: Australia (Victoria) Host: Phaulacridium vittatum (Acrididae, Orthoptera)

Charletonia huensis Haitlinger (larval) Charletonia huensis Haitlinger, 1986a: 641

Type locality: Hue Distribution: Vietnam

Charletonia hunanensis Zheng (larval) Charletonia hunanensis Zheng, 1996a: 65

Type locality: Zhuzhou, Hunan Prov. Distribution: China (Hunan Prov.) Hosts: Dragonfly (Odonata: Libellulidae), damselfly (Odonata: Megapodagriidae)

160


Charletonia ishiii Southcott (larval) Sphaerolophus sp.: Ishii, 1954: 65 Charletonia ishiii Southcott, 1991a: 79

Type locality: Fukuoka (Kyushu) Distribution: Japan Host: Mecopoda (Mecopodidae, Orthoptera)

Charletonia iwonae Haitlinger (larval) Charletonia iwonae Haitlinger, 1987g: 707

Type locality: Andoharonomaito n. Fianarantsoa, Antananarivo Distribution: Madagascar Host: from Lepidoptera and Neuroptera indet. and from plants

Charletonia jaegerskioeldi (Trägårdh)(larval) Erythraeus jaegerskioeldi Trägårdh, 1904a: 67; Oudemans, 1910e: 106; Kishida, 1929: 99 Charletonia jaegerskioeldi (Trägårdh): Oudemans, 1910d: 88; Southcott, 1961a: 526; 1966: 695; 1991a: 79 Balaustium jaegerskioeldi (Trägårdh): Buitendijk, 1945: 348

Type locality: Sinai Distribution: Egypt (Sinai) Host: grasshopper (Orthoptera)

Charletonia jolantae Haitlinger (larval) Charletonia jolantae Haitlinger, 1986a: 645

Type locality: Danang, Vietnam Distribution: Vietnam Host: Orthoptera

Charletonia justynae Haitlinger (larval) Charletonia justynae Haitlinger, 1987g: 712

Type locality: Antananarivo Distribution: Madagascar Host: undet. Orthoptera

Charletonia kaliksti Haitlinger (larval) Charletonia kaliksti Haitlinger, 2003b: 52; 2006e: 106; 2007b: 61

Type locality: Gennadi, Rhodes Distribution: France, Greece (Rhodes), Spain Host: Aiolopus sp. (Orthoptera: Acrididae: Oedipodinae)


161

Charletonia kalithensis Haitlinger (larval) Charletonia kalithensis Haitlinger, 2006b: 108; 2006f: 38

Type locality: Kalithea, Samos Distribution: Greece (Cont. and Samos) Host: from plants

Charletonia keralicus Ramaraju et Mohanasundaram (larval) Charletonia keralicus Ramaraju & Mohanasundaram, 1998: 208

Type locality: Kerala, Vellayani Distribution: India (Kerala) Host: undet. Acrididae (Orthoptera)

Charletonia keyi Southcott (larval) Charletonia keyi Southcott, 1983: 123

Type locality: Mt. White, 3 km S. of Coen (Queensland) Distribution: Australia (Queensland) Host: grasshopper Greyacris profundesulcata (Orthoptera, Acridodea, Pyrgomorphidae)

Charletonia kibonotensis (Trägårdh) (larval) Erythraeus kibonotensis Trägårdh, 1908: 34; Oudemans, 1910e: 106; 1912: 149; Kishida, 1929: 99; Buitendijk, 1945: 350 Erythraeus kibonatensis : Feider, 1954: 1021 Charletonia kibonotensis : Southcott, 1961a: 526; 1966: 742; 1991a: 100; Haitlinger, 1987g: 715

Type locality: Kibonoto, Kilimanjaro, Tanzania Distribution: Madagascar, Tanzania, Nigeria Host: Orthoptera (Acrididae), Lepidoptera and from plants

Charletonia krendowskyi (Feider) (larval) Erythraeus krendowskyi Feider, 1954: 1018 Charletonia krendowskyi (Feider): Southcott, 1961a: 528; 1966: 780; 1991a: 67; Beron, 1975: 70; Haitlinger, 2003b: 53; 2004d: 149; 2007a: 75; Hailinger & Saboori, 2007: 68

Type locality: Agigea (Romania) Distribution: Bulgaria, Croatia (Pag), Greece (incl. Rhodes and Corfu), Iran, Moldova, Romania, Ukraine Host: Mantis religiosa (Mantodea, Mantidae), unidentified Orthoptera

Charletonia lankensis Southcott (larval) Charletonia lankensis Southcott, 1988c: 158

Type locality: Labugama, 24 miles ESE Colombo Distribution: Sri Lanka Host: Eucriotettix tricarinatus (Tetrigidae, Orthoptera)

162


Charletonia lawrencei Southcott (larval) Charletonia lawrencei Southcott, 1966: 746; 1991a: 100

Type locality: Durban, Natal Distribution: South Africa Host: Orthoptera (Acrididae)

Charletonia lombokensis Haitlinger (larval) Charletonia lombokensis Haitlinger, 2006d: 222

Type locality: Lombok, Kuta Distribution: Indonesia (Lombok) Host: large Orthoptera

Charletonia mehranensis Haitlinger et Saboori (larval) Charletonia mehranensis Haitlinger & Saboori, 2007: 61

Type locality: Mehran, Varamin (Iran) Distribution: Iran Host: Anacridium aegyptium (Orthoptera, Acrididae)

Charletonia milloti (André) (ad.) Erythraeus milloti André, 1946: 268 Sphaerolophus milloti (André): Southcott, 1961a: 531 Charletonia milloti (André):

Type locality: Tananarive Distribution: Madagascar

Charletonia minuta (Khot) (comb. nov.) (ad.) Sphaerolophus minutus Khot, 1965: 66


Charletonia miyazakii (Kawashima)(larval) Callidosoma miyazakii Kawashima, 1958: 194 Charletonia miyazakii (Kawashima): Kawashima, 1961a:17; Southcott, 1966: 772; 1991a: 79

Type locality: Kinkazan, Gifu Sity, Gifu Pref., Japan Distribution: Japan Host: Theridion sp. (Theridiidae, Araneae) and other spiders

Charletonia mollicomus (Berlese) (ad.) Sphaerolophus (Cavannea) mollicomus Berlese, 1910: 351 Charletonia (Cavannea) mollicomus: Bernini et al., 1994-96: 40

Type locality: Cerchio, Aquila, Italy Distribution: Italy


Charletonia naivashae Southcott (larval) Charletonia naivashae Southcott, 1991a: 108

Type locality: Kinja at Lake Naivasha, 1950 m Distribution: Kenya Host: Erythraeinae (Erythraeidae, Acariformes, ? Erythrites sp.)

Charletonia nazeleae Karimi Iravanlou, Kamali et Talebi (larval) Charletonia nazeleae Karimi Iravanlou, Kamali & Talebi, 2002: 136

Type locality: Karaj Distribution: Iran Hosts: Heliopteryx humeralis, Palasiella turcomana (Orthoptera, Acrididae)

Charletonia nishidai Southcott (larval) Charletonia nishidai Southcott, 1991a: 112

Type locality: California, Smith Ck., Mt Hamilton, Santa Clara Co. Distribution: U.S.A. (California) Host: Camnula pellucida (Orthoptera, Acrididae)

Charletonia nova (Oudemans)(ad.) Sphaerolophus novus Oudemans, 1927: 230; 1928: 91 Charletonia nova (Oudemans)

Type locality: Buru Distribution: Buru, Indonesia

Charletonia ojimai (Kishida)(larval) Erythraeus ojimai Kishida, 1929: 100; 1930: 16 Callidosoma ojimai (Kishida): Kawashima, 1958: 195 Charletonia ojimai: Kawashima, 1961a: 18; Southcott, 1961a: 527; 1966: 814; Treat, 1975: 210

Type locality: Oita-shi, Kyushyu, Japan Distribution: Kyushyu (Japan) Host: “Rice moth” – Chilo simplex (Crambidae, Lepidoptera)

Charletonia okinawaensis (Kitahara et Takara)(larval) Callidosoma okinawaensis Kitahara & Takara, 1964: 63 Charletonia okinawaensis (Kitahara et Takara): Southcott, 1991a: 89

Type locality: Okinawa Distribution: Japan (Okinawa) Hosts: Mantodea, Orthoptera

163

164


Charletonia oudemansi Southcott (larval, Pn, Dn) Charletonia oudemansi Southcott, 1966: 710; 1991a: 65; Treat, 1975: 212; Welbourn, 1983: 131

Type locality: N.S.Wales, 6 miles N. of Wombeyan Caves Distribution: Australia (N.S.Wales, Victoria, S. Australia, Tasmania) Host: Phasmodea, Lepidoptera

Charletonia paolii Southcott (larval) Charletonia paolii Southcott, 1966: 785

Type locality: 38 miles NE. of Clermont, Queensland Distribution: Australia Host: Orthoptera (Acrididae, Catantopinae)

Charletonia perthensis (Womersley)(larval) Erythraeus perthense Womersley, 1934: 244 Charletonia perthensis (Womersley): Southcott, 1961a: 528; 1966: 695; 1991a: 73

Type locality: King’s Park, Perth, Western Australia Distribution: Australia (Western Australia) Host: ?

Charletonia philometra Southcott (larval) Charletonia philometra Southcott, 1991a: 119

Type locality: Idaho, N.E. Fruitland Distribution: U.S.A. (Idaho) Host: Philodromidae (Araneae)

Charletonia politricha (Berlese) (comb. nov.) (ad.) Sphaerolophus politrichus Berlese, 1910: 349

Type locality: “Caraca”[Caracas] Distribution: [Venezuela]

Charletonia porcinus (Hull) (comb. nov.) (ad.) Achorolophus porcinus: Hull, 1918: 28 Leptus porcinus (Hull): Turk & Turk, 1952: 500 Sphaerolophus porcinus Hull: Southcott, 1961a: 531

Type locality: Newham bog, Northumberland Distribution: England

Charletonia rageaui Southcott (larval) Charletonia rageaui Southcott, 1966: 758; 1991a: 79

Type locality: Noumea Distribution: New Caledonia Host: Cydistomyia sp.(Tabanidae, Diptera) and Libellulidae (Odonata)


165

Charletonia rocciai Treat et Flechtmann (larval) Charletonia rocciai Treat & Flechtmann, 1979: 117; Southcott, 1991a: 66

Type locality: Jau, Sao Paulo, Brazil Distribution: Brazil Host: Metagonistylum minense (Diptera, Tachinidae)

Charletonia saboorii Karimi Iravanlou, Kamali et Talebi (larval) Charletonia saboorii Karimi Iravanlou, Kamali & Talebi, 2002: 142

Type locality: Varamin Distribution: Iran Host: Heteracris littoralis similis (Acrididae, Orthoptera)

Charletonia salti (Evans) (comb. nov.) (ad.) Sphaerolophus salti: Evans, 1953: 277

Type locality: Kilimanjaro, Tanzania, 3695 m Distribution: Tanzania

Charletonia samosensis Haitlinger (larval) Charletonia samosensis Haitlinger, 2006b: 110

Type locality: Votsalakia (Samos) Distribution: Greece (Samos) Host: from plants

Charletonia sanctaehelenae (Feider et Chioreanu) (comb. nov.) (ad.) Sphaerolophus (Cavannea) sanctaehelenae Feider & Chioreanu, 1977: 279

Type locality: High Central Ridge, 700-800 m (Sainte-Helene) Distribution: Sainte-Helene

Charletonia shiroyama Yaita, Kato et Toriyama (larval) Charletonia shiroyama: Yaita, Kato & Toriyamama, 1961: 168; Kawashima, 1961b: 235; Southcott, 1991a: 95; 2004b: 158; Haitlinger, 2004; 2006d: 220 Charletonia domrowi: Southcott, 1966: 691; Welbourn, 1983: 131

Type locality: Okinawa Distribution: Japan (Okinawa), Malaysia, Australia (Queensland, Northern Territory, Western Australia), Cambodia, Indonesia (Lingga Is., Bali, Lombok) Host: Xanterriaria mediocris (Acrididae, Orthoptera), Orthoptera indet.

Charletonia similis (Vitzthum) (comb. nov.) (ad.) Sphaerolophus similis: Vitzthum, 1926: 155

Type locality: Wailima, Sumatra Distribution: Sumatra (Indonesia)

166


Charletonia southcotti Kawashima (larval) Charletonia southcotti: Kawashima, 1961a:16; Southcott, 1966: 811; 1991a: 79

Type locality: Fukuoka-Pref., Kyushu Distribution: Japan (Kyushu) Host: Gagrella japonica (Opiliones, Phalangiidae)

Charletonia spinosa (Berlese) (comb. nov.) (ad.) Sphaerolophus spinosus Berlese, 1910: 350

Type locality: Sierra Leone Distribution: Sierra Leone

Charletonia striaticeps Southcott (larval) Charletonia striaticeps Southcott, 1991a: 73

Type locality: 10 km NE of Carnegie H.S., Western Australia Distribution: Australia (Western Australia, S. Australia, Northern Territory, Queensland) Host: Pycnostictus seriatus (Acrididae, Orthoptera)

Charletonia subnuda (Berlese) (comb. nov.) (ad.) Sphaerolophus subnudus Berlese, 1910: 350

Type locality: Matusinos Distribution: South America

Charletonia sureshi (Ramaraju et Mohanasundaram) (comb. nov.) (larval) Sphaerolophus sureshi Ramaraju & Mohanasundaram, 1998: 210

Type locality: Madurai, Tamil Nadu Distribution: India (Tamil Nadu) Host: Attractomorpha crenulata (Acrididae, Orthoptera)

Charletonia swaziana (Lawrence) (larval) Erythraeus swazianus Lawrence, 1940: 401 Charletonia swaziana (Lawrence): Southcott, 1961a: 528; 1966: 732; 1991a: 100

Type locality: Piet Retief Distribution: Swaziland Hosts: Acorypha mosambicus, Gastromargus virescens (Acrididae, Orthoptera)

Charletonia taiwanensis Tsai et Chow (larval, Dn) Charletonia taiwanensis Tsai & Chow, 1988: 171; Southcott, 1991a: 98; Chow, 1996: 359

Type locality: Nangkang Distribution: Taiwan Host: Condracris rosea (Acrididae, Orthoptera)


167

Charletonia tatianae Haitlinger (larval) Charletonia tatianae Haitlinger, 1987g: 704

Type locality: Marana n. Fianarantsoa Distribution: Madagascar Host: “collected from plants”

Charletonia tragardhi Southcott (larval) Charletonia tragardhi Southcott, 1966: 721; 1991a: 65

Type locality: 30 miles N of Wauchope, Northern Territory Distribution: Australia (W. Australia, N. Territory) Host: Macrotoma sp. (Coleoptera, Cerambycidae)

Charletonia venus Southcott (larval) Erythraeus phalangioides [sic]: Oudemans, 1903a: 142 (larva); Oudemans, 1912: 135 (larva) (non Erythraeus phalangoides (de Geer, 1778)(ad.) Charletonia phalangioides : Kawashima, 1961a: 17 Charletonia venus Southcott, 1961a: 527; 1966: 704; 1991a: 99; Bernini et al., 1996-94: 40

Type locality: Maderno, Lombardy, Italy Distribution: Greece, France, Italy Hosts: Orthoptera, Acrididae (Stenobothrus, Oedipoda coerulescens); Mantodea (Mantis religiosa, Mantidae)

Charletonia vitzthumi Southcott (larval) Charletonia vitzthumi Southcott, 1966: 728; 1991a: 79

Type locality: 8 miles WSW of Thorntonia, Queensland Distribution: Australia (Queensland, Northern Territory, Western Australia) Host: Acrididae (Orthoptera)

Charletonia volzi (Oudemans) (larval) Erythraeus volzi: Oudemans, 1910e: 107; 1912: 152; Vitzthum, 1926: 159; Kishida, 1929: 99; Buitendijk, 1945: 350 Charletonia volzi (Oudemans): Eyndhoven, 1944: 52; Southcott, 1961a: 527; 1966: 762; 1991a: 79; Haitlinger, 2004b: 158; 2006d: 220

Type locality: Kwalu (NE. Sumatra), “Buitenzorg”(= Bogor)(W. Java) Distribution: New Guinea, Indonesia (Java, Sumatra, Lingga Is.), Cambodia, Burma Host: Orthoptera (Mecopoda elongata, Mecopodidae)

Charletonia westraliensis (Womersley)(ad.) Sphaerolophus westraliense Womersley, 1934: 231 Charletonia westraliensis (Womersley): Southcott, 1991a: 65

Type locality: Bridgetown (W.Australia) Distribution: Australia

168


Charletonia womersleyi Southcott (larval) Charletonia womersleyi Southcott, 1966: 776;1991a: 66; Fain & Baugnée, 1996: 22

Type locality: Farnham Royal (England) Distribution: Belgium, England Host: “Psyllid” (Homoptera, Psyllodea); Ribautodelphax pungens (Delphacidae), Stenocranus minutus (Delphacidae), Adarrus multinotatus (Euscelidae)(all Homoptera)

Charletonia wrighti Southcott (larval) Charletonia wrighti Southcott, 1991a: 120

Type locality: New Jersey, Burlington Co., Duke’s Bridge Bog Distribution: U.S.A. (New Jersey) Host: Lycaena (Tharsalea) epixanthe (Lepidoptera, Lycaenidae)

Charletonia zorani Haitlinger (larval) Charletonia zorani Haitlinger, 2004d: 150

Type locality: Krk Is., Sv. Duh n. Omišalj Distribution: Croatia (Krk) Host: Orthoptera indet.

Genus Grandjeanella Southcott (larval) Grandjeanella Southcott, 1961a: 539

Type species: Hauptmannia westraliense Womersley, 1934

Grandjeanella ainae Haitlinger (larval) Grandjeanella ainae Haitlinger, 2002f: 194

Type locality: Can Picafort, Mallorca Distribution: Mallorca (Balear Is., Spain) Host: “from plants”

Grandjeanella bella Zhang (larval) Grandjeanella bella Zhang, in Zhang & Goldarazena, 1996: 138

Type locality: Bojnurd (Iran) Distribution: Iran Host: Sericothrips sp. (Thripidae, Thysanoptera)

Grandjeanella charadrae Southcott (larval) Grandjeanella charadrae Southcott, 1996: 270

Type locality: Workanda Creek NP, Belair, S. Australia Distribution: Australia (SA) Host: “sweeping vegetation”


169

Grandjeanella haitlingeri Goldarazena et Zhang (larval) Grandjeanella haitlingeri Goldarazena & Zhang, 1997: 231; Goldarazena, Jordana & Zhang, 1999b: 26

Type locality: Navarra – Nafarroa, Tafalla (Spain) Distribution: Spain Hosts: Heteroptera indet., Pentatoma sp. (Pentatomidae), Orius sp. (Anthocoridae)

Grandjeanella kamalii Saboori et Atamehr (larval) Grandjeanella kamalii Saboori & Atamehr, 2000: 2

Type locality: Uromieh Distribution: Iran Hosts: free living on Phaseulus vulgaris (Fabaceae)

Grandjeanella multisetosa Zhang et Goldarazena (larval) Grandjeanella multisetosa Zhang & Goldarazena, 1996: 133; Goldarazena & Zhang, 1998; Goldarazena, Jordana & Zhang, 1999a: 31; Goldarazena, Zhang & Jordana, 2000: 79; Haitlinger, 2004d: 144; 2006b: 108; 2006e: 106; 2006f: 36

Type locality: Navarra-Nafarroa, Tafalla (Spain) Distribution: Croatia, Greece (incl. Samos), Spain, Turkey, Ukraine Hosts: Thrips tabaci and other Thysanoptera (Thripidae, Aeolothripidae), also from plants

Grandjeanella pangela Southcott (larval) Grandjeanella pangela Southcott, 1996: 261

Type locality: Bullace’s Glen, near Hall’s Ga, Grampians Distribution: Australia (Victoria) Hosts: from moss, etc.

Grandjeanella shepherdae Southcott (larval) Grandjeanella shepherdae Southcott, 1996: 266

Type locality: Barkley, near Avoca Distribution: Australia (Victoria) Hosts: “ex stomach of rabbit (o.c.)”

Grandjeanella westraliensis (Womersley) (larval) Hauptmannia westraliense Womersley, 1934: 249; Southcott, 1948: 260 Grandjeanella westraliensis (Womersley): Southcott, 1961a: 539

Type locality: Chittering, W. Australia Distribution: Australia Host: “under stones”

170


Genus Hauptmannia Oudemans (larval) Achorolophus Oudemans, 1910a: 31 (partim) Hauptmannia Oudemans, 1910b: 48 (larva of Abrolophus ?) Balaustioides Southcott, 1989c: 173 (synonymised by Haitlinger, 1996: 443)

Type species of Hauptmannia Oudemans: Achorolophus longicollis Oudemans, 1910 Type species of Balaustioides Southcott: B. tuxeni Southcott, 1989

Hauptmannia baardi Haitlinger (larval) Hauptmannia baardi Haitlinger, 2004h: 215

Type locality: El Pilar, La Palma, ca. 1300 m, Canary Is. Distribution: Canary Is. (La Palma) Host: from herbaceous plants

Hauptmannia brevicollis Oudemans (larval) Hauptmannia brevicollis Oudemans, 1910e: 109; Buitendijk, 1945: 347; Sellnick, 1958: 43; Willmann, 1949: 129; Lindroth et al., 1973: 131; Somermaa, 1973; Gabryś, 1988: 24; Haitlinger, 1986b: 181; 1987h: 160; 1995: 263; 2002e: 148; 2003f: 7; 2003d: 2; 2003g: 62; 2005a: 74; 2007b: 56; 2007c; Mąkol & Gabryś, 1999: 30

Type locality: Arnhem, Holland Distribution: Andorra, Austria, Czech Rep., Finland, Germany, Holland, Iceland, Lithuania, Moldova, Norway, Poland, Russia, Slovakia, Slovenia, Sweden, Ukraine Host: Thysanoptera, Homoptera (Javesella pellucida, Delphacidae)

Hauptmannia fossor Southcott (larval) Hauptmannia fossor Southcott, 1996: 274

Type locality: Homebush, New South Wales Distribution: Australia (N.S.W.) Host: “from soil debris”

Hauptmannia gracilenta Willmann (larval) Hauptmannia gracilenta Willmann, 1937: 162

Type locality: Neuwerk Distribution: Germany

Hauptmannia iranica Haitlinger et Saboori (larval) Hauptmannia iranica Haitlinger & Saboori, 1996:120 Type locality: Noshahr Distribution: Iran Host: from plants


171

Hauptmannia kazimierae Haitlinger (larval) Hauptmannia kazimierae Haitlinger, 1986b: 184; 1987i: 162; 2002e: 148; 2003d: 2; 2003g: 63; 2005a: 74; 2007b: 57; Southcott, 1994: 171; Gabrys, 1988: 25; Gabrys & Mąkol, 1994: 130

Type locality: Poland (Walbrrzych, Wroclaw and Bielsko – Biala voivodeships) Distribution: Andorra, Poland, Slovakia, Slovenia, Switzerland

Hauptmannia kotorensis Haitlinger (larval) Hauptmannia kotorensis Haitlinger, 2007a: 70

Type locality: Radanovici n. Kotor (Montenegro), Slovenia, Croatia (incl. Krk I.) Distribution: Croatia, Montenegro, Slovenia Host: from herbaceous plants

Hauptmannia miyatakei Kawashima (larval) Hauptmannia miyatakei Kawashima, 1958: 208

Type locality: Hirao, Fukuoka City, Fukuoka Pref., Japan Distribution: Japan Hosts: Homotoma radiatum, Trioza sp. (Psyllidae, Homoptera)

Hauptmannia ostovani Haitlinger et Saboori (larval) Hauptmannia ostovani Haitlinger & Saboori, 1996: 118

Type locality: Kazeron Distribution: Iran Host: Aphididae (Homoptera)

Hauptmannia parvum Schweizer et Bader (larval) Hauptmannia parvum Schweizer & Bader, 1963: 313

Type locality: Blockhaus Purcher, 2000 m, Switzerland Distribution: Switzerland

Hauptmannia podorasensis Haitlinger (larval) Hauptmannia podorasensis Haitlinger, 2006f: 36; 2007a: 67

Type locality: Podorasac (Bosnia and Herzegovina) Distribution: Bosnia and Herzegovina, Croatia, Greece Host: “from herbaceous plants”

Hauptmannia sardiniensis Haitlinger (larval) Hauptmannia sardiniensis Haitlinger, 2007d: 540

Type locality: Porto Pozzo, Sardinia Distribution: Italy (Sardinia) Host: “from herbaceous plants”

172


Hauptmannia silesiacus Haitlinger (larval) Hauptmannia silesiacus Haitlinger, 1986b: 189; 1987i: 163; 2000c: 388; 2002d: 554; 2002e: 148; 2003d: 3; 2003g: 65; 2004d: 144; 2007b: 57; Gabryś, 1988: 25

Type locality: Wroclaw – Swojec, Poland Distribution: France, Greece (Samos), Poland, Slovakia Host: from plants

Hauptmannia stanislavae Haitlinger (larval) Hauptmannia stanislavae Haitlinger, 1986b: 188; 2000c: 388; 2002e: 148; 2002g: 554; 2003g: 66; 2004d: 144; Gabryś, 1988: 25

Type locality: Wroclaw voivodeship, Poland Distribution: France, Poland, Slovakia Host: unknown (from herbaceous plants)

Hauptmannia trifarius Shiba (larval) Hauptmannia trifarius Shiba, 1976: 185; Haitlinger, 2002d: 118

Type locality: Pasoh Forest Distribution: Malaysia Host: unknown

Hauptmannia viticola Fain et Cobanoglu (larval) Hauptmannia viticola Fain & Cobanoglu, 1998: 63

Type locality: Salubakal Village, Bozcaada Distribution: Turkey Host: unknown insect

Hauptmannia willmanni Schweizer (larval) Hauptmannia willmanni Schweizer, 1951: 164; Schweizer & Bader, 1963: 312

Type locality: Stabel – chod, 2600 m, Swiss Alps Distribution: Switzerland

Hauptmannia wratislaviensis Haitlinger (larval) Hauptmannia wratislaviensis Haitlinger, 1986b: 188; 1987i: 161;1994a: 408; 2000c: 388; 2002e: 148; 2003f: 7; 2007c; Gabryś, 1988: 25; Southcott, 1989: Balaustioides tuxeni Southcott, 1989c: 173 (synonymised by Haitlinger, 1996: 443)

Type locality: Wroclaw voiwodeship, Poland Distribution: Czech Rep., Holland, Iceland, Lithuania, Moldova, Norway, Poland, Russia, GB (Scotland), Slovenia, Switzerland, Ukraine


Hauptmannia yanlingica Zheng (larval) Hauptmannia yanlingica Zheng, 2002: 89

Type locality: Hunan Distribution: China (Hunan) Host: Diptera Brachycera

Hauptmannia sp. Distribution: India (Maharashtra) Host: Stenchaetothrips minutus (Thripidae, Thysanoptera)

Genus Iguatonia Haitlinger (larval) Iguatonia Haitlinger, 2004f: 438

Type species: Iguatonia barbillae Haitlinger, 2004

Iguatonia barbillae Haitlinger (larval) Iguatonia barbillae Haitlinger, 2004f: 436

Type locality: Foz do Iguacu Distribution: Brazil Host: from plants and from Homoptera ad. indet.

Genus Neoabrolophus Khot (ad.) Neoabrolophus Khot, 1965: 74

Type species: Neoabrolophus brevisetorum Khot, 1965

Neoabrolophus bisetis (Evans)(ad.) Balaustium bisetis Evans, 1953: 274 Neoabrolophus bisetis (Evans): Shiba, 1976: 185

Type locality: Kilimanjaro, 12 450 ft.(3800 m) Distribution: Tanzania

Neoabrolophus brevisetorum Khot (ad.) Neoabrolophus brevisetorum Khot, 1965: 74

Type locality: Kalingpong (Bengal) Distribution: India

Neoabrolophus hirsutus Shiba (ad.) Neoabrolophus hirsutus Shiba, 1976: 183


173

174


Neoabrolophus longisetorum Khot (ad.) Neoabrolophus longisetorum Khot, 1965: 77

Type locality: Rishikesh Distribution: India (Uttar Pradesh)

Genus Pussardia Southcott (larval) Pussardia Southcott, 1961a: 538

Type species: Hauptmannia mullewaensis Womersley, 1934

Pussardia mullewaensis (Womersley)(larval) Hauptmannia mullewaensis Womersley, 1934: 250; Southcott, 1948: 261 Pussardia mullewaensis (Womersley): Southcott, 1961a: 538

Type locality: Mullewa, W. Australia Distribution: Australia (W.A.)

Genus Rudaemannia Haitlinger Rudaemannia Haitlinger, 2000d: 386

Type species: Hauptmannia rudaensis Haitlinger, 1986

Rudaemannia rudaensis (Haitlinger) (larval) Hauptmannia rudaensis Haitlinger, 1986b: 182; Gabryś, 1988: 24 Rudaemannia rudaensis (Haitlinger): Haitlinger, 2000d: 386; 2003f: 68; 2007b: 61; 2007c Hauptmannia viburnicola Fain & Cobanoglu, 1998: 67; Haitlinger, 2000d: 386

Type locality: Ruda Milicka near Wroclaw, Poland Type locality of Hauptmannia viburnicola Fain & Cobanoglu: Karagol, Ankara, from Viburnum lantana Distribution: Greece (Rhodes), Moldova, Poland, Switzerland, Turkey Host: Thysanoptera

Subfamily Phanolophinae Southcott Phanolophinae Southcott, 1946: 173

Type genus: Phanolophus André, 1927

Genus Phanolophus André Phanolophus André, 1927: 169 ?Platylophus Berlese, 1910: 351 (nom. dubium)(non Swainson, 1832 – Aves)

Type species: Phanolophus nasica André, 1927

Phanolophus oedipodarum (Frauenfeld)(ad. et larv.) Rhyncholophus oedipodarum Frauenfeld, 1868: 889 Erythraeus oedipodarum (von Frauenfeld, 1868): Oudemans, 1939: 80 Phanolophus nasica André, 1927: 170; Daniel & Samšinak, 1955: 1242 Smaris nasica: Vitzthum, 1931: 148; André, 1932a: 885


175

Smaridia magnifica Berlese, 1918: 181 (type from Dociostaurus maroccanus, Orthoptera, Apulia, Italy) Smaris magnifica Berlese: André, 1932a: 884 Phanolophus oedipodarum (André): Southcott, 1961a: 451; Haitlinger, 1993: 85; 2004d: 152; 2006f: 37

Type locality: “la Reghaia (près d’Alger)”, Algeria Distribution: Algeria, Austria, Azerbaydjan, Croatia, Czech Rep., Cyprus, Greece, Italy Hosts: Calliptamus italicus, C. barbarus, Rhacocleis germanica, Euchortippus declivus, Dociostaurus maroccanus, Oedipoda coerulescens, Chortippus vagans (Acrididae), Acrometopa macropoda, Decticus albifrons (Tettigoniidae), Ephippiger cavannai (Bradyporidae)(Orthoptera)

Subfamily Balaustiinae Southcott Balaustiidae Grandjean, 1947a: 12 Balaustiinae Southcott, 1957: 98 (in part) Balaustiinae Southcott, 1961a: 540 (restricted)

Type genus: Balaustium von Heyden, 1826

Genus Balaustium von Heyden (ad. and larval) Balaustium von Heyden, 1826: 309 Belaustium: Gervais, 1844: 164

Type species: Trombidium murorum Hermann, 1804

Balaustium affine Willmann (ad.) Balaustium affine Willmann, 1954: 244

Type locality: Moravia Distribution: Czech Rep.

Balaustium afghanicum Cooreman (ad.) Balaustium afghanicum Cooreman, 1960: 15

Type locality: Chamchir ghar, Kandahar, Panjvai, Afghanistan Distribution: Afghanistan

Balaustium angustum Evans (ad.) Balaustium angustum Evans, 1953: 276

Type locality: Kilimanjaro, 3810 m Distribution: Tanzania

Balaustium antarcticum (Trägårdh)(ad.) Erythraeus antarcticus Trägårdh, 1907: 25 Belaustium [sic] antarcticum (Trägårdh): Trägårdh, 1931: 625 Balaustium or Abrolophus ? – Southcott, 1961a: 546 Balaustium antarcticum (Trägårdh): Feider & Chioreanu, 1977: 270

Type locality: Falkland Is. Distribution: Falkland Is.

176


Balaustium araneipes Cooreman (ad.) Balaustium araneipes Cooreman, 1956: 5 as nom. nov. pro Rhyncholophus araneoides (Berlese) sensu Halbert, 1920, nec Achorolophus araneoides Berlese, 1910

Type locality: Malahide, Ireland Distribution: Algeria, Ireland

Balaustium araneoides (Berlese)(ad.) Achorolophus araneoides Berlese, 1910: 349 Balaustium araneoides (Berlese): Willmann, 1951a: 353; 1951c: 148; Evans & Browning, 1953: 418 (araneipes?); Franz, 1954: 383; Bernini et al., 1996-94: 40

Type locality: Palermo, Sicily Distribution: Austria, England, Ireland, Sicily

Balaustium barloventensis Haitlinger (larval) Balaustium barloventensis Haitlinger, 2004h: 216

Type locality: Barlovento, La Palma Distribution: Canary Is. (La Palma) Host: from herbaceous plants

Balaustium bipilum Meyer et Ryke (ad.) Balaustium bipilum Meyer & Ryke, 1959: 312

Type locality: Potchefstroom Distribution: South Africa

Balaustium bolivianum Beron (ad.) Balaustium bolivianum Beron, in prep.

Type locality: above La Paz, 4480 m Distribution: Bolivia

Balaustium brunoni Haitlinger (larval) Balaustium brunoni Haitlinger, 2005a: 514

Type locality: Argentina Distribution: Argentina Host: from plants

Balaustium bulgariense Oudemans (ad.) Balaustium bulgariense Oudemans, 1926: 150; Irk, 1939b: 168; Willmann, 1954: 244

Type locality: Bulgaria Distribution: Austria, Bulgaria, Czech Rep.


177

Balaustium dowelli Smiley (ad.) Balaustium dowelli Smiley, 1964: 110; 1966: 25

Type locality: Conway County, Arkansas Distribution: U.S.A.

Balaustium florale Grandjean (larval) Balaustium florale Grandjean, 1947b: 327; Haitlinger, 2002f: 196

Type locality: Corsica Distribution: Corsica, Mallorca Host: flowers

Balaustium graminum Meyer et Ryke (ad.) Balaustium graminum Meyer & Ryke, 1959: 316

Type locality: Grabouw Distribution: South Africa

Balaustium innocentae Haitlinger (ad.) Balaustium innocentae Haitlinger, 2006a: 95

Type locality: Lingkou n. Xian Distribution: China

Balaustium kacperi Haitlinger (larval) Balaustium kacperi Haitlinger, 1996: 450

Type locality: Studzienna n. Wambierzyce (voi. Walbrzych), ca. 500 m. Distribution: Poland

Balaustium kendalli Welbourn (ad.) Balaustium kendalli Wellbourn, in Welbourn & Jennings, 1991: 567

Type locality: Piscataquis Co., ME Distribution: U.S.A. (ME)

Balaustium madeirense Willmann (ad.) Balaustium madeirense Willmann, 1939a: 18; Cooreman, 1955a: 126; 1955c: 14; Bernini et al., 1996-94: 40

Type locality: Madeira Distribution: Belgium, Greece, Italy, Madeira, Turkey

Balaustium malpaisesensis Haitlinger (larval) Balaustium malpaisesensis Haitlinger, 2004h: 218

Type locality: Malpaises, Tigalate, Fuencaliente (La Palma) Distribution: Canary Is. (La Palma) Host: from herbaceous plants

178


Balaustium medardi Haitlinger (larval) Balaustium medardi Haitlinger, 2000c: 388

Type locality: Tiahuanacu (Bolivia), ca. 4000 m; Peru (2900-3500 m) Distribution: Bolivia, Peru Host: from plants

Balaustium medicagoense Meyer et Ryke (ad.) Balaustium medicagoense Meyer & Ryke, 1959: 310; Halliday, 2001: 327

Type locality: Caledon (South Africa) Distribution: South Africa, Australia

Balaustium minodorae Haitlinger (larval) Balaustium minodorae Haitlinger, 2000e: 73

Type locality: Xochimilko Distribution: Mexico Host: from plants

Balaustium murorum (Hermann)(ad.) Trombidium murorum Hermann, 1804: 28 Erythraeus murorum: Latreille, 1817: 423; 1823: 168; Audouin, 1824: 291; Dugès, 1836: 302 Rhyncholophus murorum (Hermann): C.L. Koch, 1837: fa.16, t.12; 1842: 50 Belaustium murorum (Hermann): Oudemans, 1906: 87; Sellnick, 1931: 696; Vitzthum, 1935; Karppinen, 1958: 45 Balaustium murorum (Hermann): Irk, 1939: 168; Buitendijk, 1945: 348; Willmann, 1949: 128; 1951c: 151; 1954: 244; Feider, 1948: 209; Sellnick, 1949: 130;1958: 42; Schweizer, 1951: 147; Turk, 1953: 26; Franz, 1954: 383; Cooreman, 1955c: 13; Schweizer & Bader, 1963: 278; Rack, 1983: 157; Gabryś, 1988: 25; 2000: 55; Gabryś & Mąkol, 1994: 130; 1996: 489; Bernini et al., 1996-94: 40; Haitlinger, 2002e: 149

Type locality: France Distribution: Austria, Belgium, Czech Rep., England, Finland, Germany, Greece, Holland, Hungary, Italy (incl. Sicily), Poland, Romania, Sweden, Switzerland, Tunisia

Balaustium neominiatum Mihelčič (ad.) Balaustium neominiatum Mihelčič, 1958a: 283

Type locality: Vallecas, Spain Distribution: Spain

Balaustium neomurorum Schweizer (ad.) Balaustium neomurorum Schweizer, 1951: 151; Franz, 1954: 383; Schweizer & Bader, 1963: 280 Abrolophus neomurorum Schw.: Schmölzer, 1993: 61

Type locality: Swiss Alps, Scanfs, Scarl etc., 1750-2100 m Distribution: Austria, Switzerland


Balaustium nikae Haitlinger (larval) Balaustium nikae Haitlinger, 1996: 453; 2003f: 7; 2004d: 156; 2006f: 38

Type locality: Rogalinek (voi. Poznan) Distribution: Croatia, Czech Rep., Greece, Poland, Slovakia, Ukraine

Balaustium obtusum Trägårdh (ad.) Belaustium [sic] obtusum Trägårdh, 1931a: 624 Balaustium obtusum: Southcott, 1961a: 546; Feider & Chioreanu, 1977: 270

Type locality: Masatierra, Juan Fernandez Distribution: Juan Fernandez

Balaustium papillatum Mihelčič (ad.) Balaustium papillatum Mihelčič, 1958b: 47


Balaustium putmani Smiley (ad.) Balaustium putmani Smiley, 1968: 13; Putman, 1970: 76

Type locality: Ontario, Canada Distribution: Canada (Ontario)

Balaustium rajmundi Haitlinger (larval) Balaustium rajmundi Haitlinger, 1996: 451

Type locality: Pieniny Mts., Kroscienko (voi. Nowy Sacz), ca. 460 m Distribution: Poland

Balaustium southcotti Feider et Chioreanu (ad.) Balaustium southcotti Feider & Chioreanu, 1977: 283

Type locality: High Peak and other points, 450-760 m, Sainte-Helene Distribution: Sainte-Helene

Balaustium soydani Haitlinger (larval) Balaustium soydani Haitlinger, 2000e: 70

Type locality: Panajechal Distribution: Guatemala Host: from plants

Balaustium submurorum Schweizer et Bader (ad.) Balaustium submurorum Schweizer & Bader, 1963: 280

Type locality: Birsfelden, 260 m Distribution: Switzerland

179

180


Balaustium unidentatum (Trägårdh)(ad.) Erythraeus unidentatus Trägårdh, 1904b: 56 Rhyncholophus unidentatus Tr.: Schweizer, 1922: 82 Balaustium unidentatus (Trägårdh): Gabryś & Haitlinger, 1986: 472 Balaustium unidentatum (Trägårdh): Schweizer & Bader, 1963: 278; Gabryś, 1988: 625, 2000: 55

Type locality: Greenland Distribution: Greenland, Poland, Switzerland

Balaustium veletense Mihelčič (ad.) Balaustium veletense Mihelčič, 1958b: 47


Balaustium vignae Meyer et Ryke (ad.) Balaustium vignae Meyer & Ryke, 1959: 310

Type locality: Kaap Muiden Distribution: South Africa

Balaustium wratislaviensis Haitlinger (larval) Balaustium wratislaviensis Haitlinger, 1996: 454

Type locality: Wroclaw; paratype from Germany, Osterode – Harz Distribution: Germany, Poland Hosts: Pipistrellus pipistrellus (Vespertilionidae, Chiroptera) in Germany, Parus major (Paridae, Aves) in Poland

Balaustium xerothermicum Gabryś (ad.) Balaustium xerothermicum: Gabryś, 2000a: 47

Type locality: Sutno on Bug River at Siemiatycze (Podlaskie Province) Distribution: Poland

Balaustium zhangi Saboori (larval, deut. and ad.) Balaustium zhangi Saboori, 2001: 171

Type locality: Nowshahr Distribution: Iran Host: on soil and Hydrangea paniculata

Balaustium sp. Balaustium sp.: Haitlinger, 2002f: 196

Type locality: Picafort, Mallorca Host: Tarentola mauritanica (Gekkonidae, Sauria, Reptilia)


Balaustium sp. Balaustium sp.: Haitlinger & Walter, 1997: 106

Locality: Osterode/Harz (Germany) Host: Pipistrellus pipistrellus (Mammalia, Chiroptera, Vespertilionidae)

Genus Bursaustium Haitlinger (larval) Bursaustium Haitlinger, 2000c: 390

Type species: Bursaustium gaspari Haitlinger, 2000

Bursaustium gaspari Haitlinger (larval) Bursaustium gaspari Haitlinger, 2000c: 390; 2006f: 37

Type locality: Bursa, Turkey Distribution: Turkey

Bursaustium norbakhshi Saboori (larval) Bursaustium norbakhshi Saboori, 2002b: 375

Type locality: Shahrkord, H. Norbakhsh Distribution: Iran Host: “free living on wheat”

Genus Fozustium Haitlinger (larval) Fozustium Haitlinger, 2005a: 517

Type species: Fozustium paranensis Haitlinger, 2005

Fozustium paranensis Haitlinger (larval) Fozustium paranensis Haitlinger, 2005a: 517


Genus Guatustium Haitlinger (larval) Guatustium Haitlinger, 2000e: 74

Type species: Guatustium biljanae Haitlinger, 2000

Guatustium biljanae Haitlinger (larval) Guatustium biljanae Haitlinger, 2000e: 74

Type locality: Panajechal Distribution: Guatemala

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182


Genus Italustium Haitlinger (larval) Italustium Haitlinger, 2000e: 79

Type species: Italustium eframi Haitlinger, 2000

Italustium eframi Haitlinger (larval) Italustium eframi Haitlinger, 2000e: 79

Type locality: near Móggio (Alps), ca. 900 m Distribution: Italy

Genus Lomeustium Haitlinger (larval) Lomeustium Haitlinger, 2006g: 120

Type species: Lomeustium togoensis Haitlinger, 2006

Lomeustium togoensis Haitlinger (larval) Lomeustium togoensis Haitlinger, 2006g: 121

Type locality: Lome (Togo), Winneba, Kumasi (Ghana), at lake Bosumtwi, Grand Popo (Benin) Distribution: Togo, Benin, Ghana Host: “from herbaceous plants”

Genus Moldoustium Haitlinger (larval) Moldoustium Haitlinger, 2007c

Type species: Moldoustium baltiensis Haitlinger, 2007

Moldoustium baltiensis Haitlinger (larval) Moldoustium baltiensis Haitlinger, 2007c

Type locality: Balti (Moldova) Distribution: Moldova, Ukraine Host: herbaceous plants

Genus Microsmarialla Khot (ad.) Microsmarialla Khot, 1963: 557

Type species: Microsmarialla coniferana Khot, 1963

Microsmarialla coniferana Khot (ad.) Microsmarialla coniferana Khot, 1963: 560

Type locality: Gwalior (Madhya Pradesh), India Distribution: India (Madhya Pradesh)

Genus Microsmaris Hirst (ad.) Microsmaris Hirst, 1926: 613

Type species: Microsmaris mirandus Hirst, 1926


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Microsmaris hirsti Womersley (ad.) Microsmaris hirsti Womersley, 1934: 233

Type locality: Freshwater Bay, Swan River, Perth, West Australia Distribution: Australia (West Australia)

Microsmaris joannae Hirst (ad.) Microsmaris goannae Hirst, 1928: 570; Womersley, 1934: 232 Microsmaris joannae Southcott, 1961a: 550

Type locality: Fulerton, Adelaide, South Australia Distribution: Australia (South Australia)

Microsmaris minutus Womersley (ad.) Microsmaris minutus Womersley, 1934: 234

Type locality: Mount Lofty, South Australia Distribution: Australia (South Australia)

Microsmaris mirandus Hirst (ad.) Microsmaris mirandus Hirst, 1926: 613; Vitzthum, 1931: 148; Womersley, 1934: 232; Oudemans, 1941: 182; Southcott, 1946: 46; 1961a: 548; Grandjean, 1947b: 332

Type locality: Christchurch, New Zealand Distribution: Australia (West Australia), New Zealand

Microsmaris pinnata Khot (ad.) Microsmaris pinnata Khot, 1963: 557

Type locality: Bikaner (Rajastan) and Delhi Distribution: India

Microsmaris sparsepinnata Khot (ad.) Microsmaris sparsepinnata Khot, 1963: 557

Type locality: Bikaner, Rajastan Distribution: India

Genus Mypongia Southcott (larval) Mypongia Southcott, 1961a: 554 Myopongia Haitlinger, 2000e: 69; 2006g: 121 (lapsus typ.)

Type species (original designation): Mypongia brevipes Southcott, 1961

Mypongia brevipes Southcott (larval) Mypongia brevipes Southcott, 1961a: 554 Myopongia brevipes Southcott: Haitlinger, 2000e: 69 (lapsus typ.)

Type locality: South Australia Distribution: Australia (South Australia)

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Genus Neobalaustium Willmann (ad.) Neobalaustium Wilmann, 1951: 349

Type species: Neobalaustium caspersi Willmann, 1951

Neobalaustium borceanum Feider et Suciu (ad.) Neobalaustium borceanum Feider & Suciu, 1959: 439; 1963: 51

Type locality: Agigea, reg. Constantsa, Romanian Dobroudja Distribution: Romania

Neobalaustium caspersi Willmann (ad.) Neobalaustium caspersi Willmann, 1951: 349; Southcott, 1961a: 548

Type locality: Pomorie, E. Bulgaria Distribution: Bulgaria

Genus Palenqustium Haitlinger (larval) Palenqustium Haitlinger, 2000e: 80

Type species: Palenqustium leanderi Haitlinger, 2000

Palenqustium leanderi Haitlinger (larval) Palenqustium leanderi Haitlinger, 2000e: 80

Type locality: Palenque Distribution: Mexico

Genus Pollux Southcott (larval) Pollux Southcott, 1961a: 558

Type species: Pollux workandae Southcott, 1961

Pollux cristatus (Womersley) (larval) Belaustium cristatum Womersley, 1934: 251 Microsmaris cristatus: Southcott, 1946: 46 Pollux cristatus (Womersley): Haitlinger, 2002c: 173

Type locality: Glen Osmond, South Australia Distribution: Australia (S.A.) Host: free and attached to Jassids (Jassidae, Homoptera) and other insects.

Pollux kovalamicus Haitlinger (larval) Pollux kovalamicus Haitlinger, 2002c: 173

Type locality: Kovalam Distribution: India Host: from plants


Pollux workandae Southcott (larval) Pollux workandae Southcott, 1961a: 563

Type locality: South Australia Distribution: Australia (South Australia) Host: “from soil and vegetation”

Genus Wartookia Southcott (ad.) Wartookia Southcott, 1961a: 550

Type species: Wartookia rebeccae Southcott, 1961

Wartookia rebeccae Southcott (ad.) Wartookia rebeccae Southcott, 1961a: 550

Type locality: Rosebrook, Wartook, Victoria Distribution: Australia (Victoria)

Subfamily Myrmicotrombiinae Southcott Myrmicotrombiinae Southcott, 1957: 98 (non Myrmicotrombiinae Feider, 1955: 75)

Type genus: Myrmicotrombium Womersley, 1934

Genus Myrmicotrombium Womersley (ad.) Myrmicotrombium Womersley, 1934: 186

Type species: Myrmicotrombium brevicristatum Womersley, 1934

Subgenus Myrmicotrombium (Myrmicotrombium) Womersley Myrmicotrombium Womersley, 1934: 186 Myrmicothrombium: Thor & Willmann, 1947: 227

Type species: Myrmicotrombium brevicristatum Womersley, 1934

Myrmicotrombium (M.) brevicristatum Womersley (ad.) Myrmicotrombium brevicristatum Womersley, 1934: 186; Southcott, 1957: 92; 1961a: 514


Subgenus Graecotrombium Beron (ad.) Myrmicotrombium (Graecotrombium) Beron, 1990: 67

Type species: Myrmicotrombium (Graecotrombium) mirum Beron, 1990

Myrmicotrombium (Graecotrombium) mirum Beron (ad.) Myrmicotrombium (Graecotrombium) mirum Beron, 1990: 68

Type locality: Keratea, Attica, Greece Distribution: Greece

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186


Species incertae sedes: “Erythraeus” pachypus (Hull)(ad.) Rhyncholophus pachypus Hull, 1918: 25 Erythraeus pachypus (Hull): Turk, 1953: 26

Type locality: West Allendale, Cleveland (England) Distribution: England

“Erythraeus” penninus (Hull) (ad.) Rhyncholophus penninus Hull, 1918: 24 Erythraeus penninus (Hull): Turk, 1953: 26

Type locality: West Allendale, England Distribution: England

Species inquirendae: Atomus longilinea Ewing, 1917: 155 – USA (Ohio) – “It resembles Fessonia sp.” (Miller, 1925: 110) Atomus niger Ewing, 1917: 154 – USA (Illinois) Atomus quadrirubipes Ewing – USA (Ohio, Miller, 1925: 110) Smaris spegazzinii Berlese, 1916 – “La Plata” Smaris granatae Di Caporiacco, 1938 Smaris intermedia Lombardini, 1949 Smaris australis Banks, 1896 – USA (Florida, Punta Gorda, Banks, 1896: 74, also Banks, 1904a: 143) – “six eyes” ?! Smaris longilinealis Ewing, 1909a – USA (Illinois) (“a double pair of eyes”); Fessonia longilinealis (Ewing), acc. to Miller, 1925: 112, Ohio Smaris longilinealis Ewing, 1909b – USA (Illinois) (“small single pair of eyes”) Smaridia pulcherrima Berlese, 1917 – East Africa Smaridia hungarica Karpelles, 1893 – Hungary Smaridia intermedia Lombardini, 1949 – Italy, on Musca domestica (certainly not an Erythraeoidea) Smaridia heteropoda Berlese, 1910 – South Africa (Capetown, comp. to H. ampulligera) Erythraeus spinatus Banks, 1894 – USA Erythraeus carolinus Banks, 1947 – USA, North Carolina Erythraeus munsteri Meyer et Ryke, 1959 – South Africa Erythraeus tolteka Vitzthum, 1933 – Mexico (Guerero) – no spines on palpe! Erythraeus alluaudi Berlese, 1920 – East Africa Erythraeus italicus Berlese, 1920 – Italy (Pisano) Erythraeus ursinus Berlese, 1920 – Chili, 3000 m Erythraeus posticatus Banks, 1916 – USA, California Erythraeus tonsus (L. Koch, 1879) – St. Paul Island, Pribilov Isl., Alaska (= Rhyncholophus tonsus from Russia) Erythraeus eucomus Berlese, 1920 – Germany


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Erythraeus decalvatus Berlese, 1920 – Germany Erythraeus mucronatus Trägårdh, 1908 – Tanzania Erythraeus meruensis Trägårdh, 1908 – Tanzania Erythraeus arvensis Banks, 1912 – USA (Utah) Erythraeus phalangiolus Ewing, 1917 – USA (Ohio); see also Miller, 1925: 111 Erythraeus arenicola Hall, 1912 – USA Erythraeus quadratus Ewing, 1917: 153 – USA (Illinois)(two pairs of eyes) Achorolophus brachiosus Berlese, 1916 – La Plata Achorolophus complanatus Berlese, 1910 – Germany Achorolophus humeratus Berlese, 1916 – East Africa (“Kounhi; Onotchocha”) Achorolophus boccettii Lombardini, 1958 (Leptus ?) Rhyncholophus angustipes Banks, 1907, nom. nov. pro R. gracilipes Banks, 1905 Rhyncholophus arenicola Andersén, 1863 – North Europe Rhyncholophus arenicola Hall, 1912 – USA (Erythraeus arenicola in Augustson, 1940: 193) Rhyncholophus attolus Banks, 1916 – Australia (NSW) Rhyncholophus brevitarsus Ewing, 1909 – USA Rhyncholophus canonicalis Pavesi, 1876: 448 – Italy, Capraja [Erythraeus ?] Rhyncholophus cavernarum Packard, 1888 – USA (Rhagidia ? – see Southcott, 1961a: 412) Rhyncholophus cinctipes Banks, 1894 – USA [Erythraeus cinctipes – Banks, 1915: 41] Rhyncholophus hispidus Brady, 1877 – England Rhyncholophus dugesii Lucas, 1846 – USA [Erythraeus dugesii (Lucas), acc. to Southcott, 1961a: 512] Rhyncolophus echinatus Banks, 1913 – Mexico (San José del Cabo)[Erythraeinae!] Rhyncholophus elongatus Banks, 1899 – USA Rhyncholophus erinaceus Stoll, 1886-1893 – Guatemala Rhyncholophus erythreus Ewing, 1909a: 58 – USA (Illinois)(“Eyes large, single and situated laterally”) Rhyncolophus [Rhincholophus] filipes Berlese, 1888 – Brasil, Mato Grosso [Erythraeinae] Rhyncolophus floridanus Banks, 1896 – USA (Florida, Punta Gorda, Banks, 1896: 74, also Banks, 1904a: 143) Rhyncholophus gloriosus Berlese, 1888 – Paraguay, Argentina Rhyncholophus gracilipes Banks, 1904 – Santa Rosa Island, California; British Columbia Rhyncholophus gracilipes Banks, 1904, Canada, British Columbia; USA, Illinois (Ewing, 1909: “single pair of eyes”); California, Santa Rosa I. (Banks, 1904a: 365) Rhyncholophus [Rhincholophus] impectus Berlese, 1888 – Argentina, Paraguay (4 eyes Erythraeus ?) Rhyncholophus lentiginosus C.L. Koch, 1837: 16, 22; 1842: 51 Rhyncholophus leprosus Ewing, 1909a: 60 – USA (Illinois) (“single pair of eyes”) Rhyncholophus longipes Banks, 1894 – USA (Florida, Indiana) Rhyncholophus longitarsus Ewing, 1909 – USA (“Columbia, Mo.”, a single pair of large eyes) (p. 58)

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Rhyncholophus maculatus Banks, 1894 – USA (Colorado, Florida, Arizona - Banks, 1902b: 220) Rhyncholophus miniaceus var. pulchellus Leonardi, 1901 Rhyncholophus moestus Banks, 1914 – “one eye-spot each side” (p. 56) Rhyncholophus montanus Banks, 1894 – USA Rhyncholophus pallidipes Karsch (Algeria, Tripolitania); Lucas, 1845-47: 312 (Tunis) sub R. pallipes (laps. orthogr.) Rhyncholophus parvipollicus Ewing, 1909 – USA (= Atomus parvipollicus Ewing, Illinois, Ohio) Rhyncholophus parvisetosus Ewing, 1909a: 57 – USA (Illinois)[= Erythraeus parvisetosus (Ewing, 1909) – Banks, 1915: 41; Miller, 1925: 111] Rhyncholophus parvus Banks, 1894 – USA; Illinois (Ewing, 1909: “single pair of eyes”); Colorado (Banks, 1895) Rhyncolophus pedatus Banks, 1913 – Mexico (Hermosillo)[Erythraeinae!] Rhyncholophus pedestris Berlese, 1888 – Argentina; also Berlese & Leonardi, 1901: 16 Rhyncholophus phloginus C.L. Koch, 1837 – (renamed Fessonia phloginus by Oudemans, 1941: 181) Rhyncholophus pilosus Banks, 1894 – USA (= Erythraeus pilosus (Banks) – Miller, 1925: 111, Ohio) Rhyncholophus pollicaris Ewing, 1909 – USA (Illinois), “a single pair of eyes” Rhyncholophus punctatus Banks, 1904 – USA (Florida, Banks, 1904a: 143) Rhyncholophus quadrioculus Ewing, 1909a: 59 – USA (Illinois)(“Two pairs of eyes”) Rhyncholophus quinquemaculatus (Stoll, 1886-1893) (Trombidium q.) – Argentina; also Berlese & Leonardi, 1901: 16 Rhyncholophus retentus Banks, 1916 – Australia, Victoria Rhyncolophus robustus Banks, 1895 – USA (Fort Collins, Colorado); Arizona (Banks, 1902b: 220) Rhyncholophus roseus Banks, 1894 – USA (in Grognieria – see Oudemans, 1941: 179) Rhyncholophus sagittatus Canestini, 1898 – Erima (New Guinea). Compared with R. globiger (now Charletonia), but with two kinds of settae. Rhyncholophus stolli Banks, 1913 – Costa Rica, San Jose (p. 685), “one eye each side” Rhyncholophus texanus Banks, 1894 – USA Rhyncholophus tonsus L. Koch, 1879 – Russia Rhyncholophus vernalis L. Koch, 1881 – Balears Rhyncholophus (Ritteria) plumosa Thor, 1900 – Norway Rhyncholophus (Apectolophus) simplex Leonardi, 1901 – Argentina Rhyncholophus simplex Banks – (Florida, also Banks, 1904a: 143) Rhyncholophus (Achorolophus) cursitans Leonardi, 1901 – Chile (Temuco, compared with “Rh.” cardinalis) Rhyncholophus (Abrolophus) striglipes Leonardi, 1901 Ritteria mantonensis George, 1907 – (= Rhyncholophus mantonensis in Hull, 1918) England (p. 357) Belaustium [sic] harrisonii Hull, 1918 – England (p. 27)


189

Doubtful or excluded names (see the revision of Southcott, 1961a) Acarus sambuci Schranck, 1781 (type of genus Smaris Latreille, 1796, later replaced as type by Trombidium squamatum Hermann, 1804, because Acarus sambuci Schranck was considered belonging to Tetranychidae) Cloquetia Oudemans, 1941 – proposed pro Rhyncholophus medioareolatus Kramer, 1898 from South America – both are nomina dubia Corerythrolophus Radford, 1950, not Coerythrolophus, as in Southcott, 1961a (lapsus calami) – replacement name for Erythrolophus Berlese, 1920 (praeoccup.) Ctenerythraeus Berlese, 1918 – a trombidiid mite Discolopheae, Caeculiosomateae [sic], Sphaerolopheae – used as “tribal names” by Kishida (1939), as well as “Sphaerolophinae”, which was not used by subsequent authors Dupreia Oudemans, 1941 – proposed for Smaridia pileifera Karpelles, 1891 – a trombidiid mite Euplatylophus Radford, 1950 – replacement name for Platylophus Berlese Erythraeus (Ptilolophus) claviger Berlese, 1916 – from “La Plata”, South America – nomen dubium Erythrombium Berlese, 1910 – nomen dubium Erythrombium eusisyrum Berlese, 1910 from Palermo (Sicily) – nomen dubium Subgenus Erythrolophus Berlese, 1920 – (nom. praeoccup. from Erythrolophus Swinhoe, 1892); the two described species in this “subgenus” – Erythraeus (Erythrolophus) froggatti Berlese, 1910 = novaehollanidae Oudemans, 1941 from Australia, or Africa and Erythraeus (Erythrolophus) mollis Berlese, 1923 from New Caledonia – may belong to Erythraeinae (4 eyes) or even to genus Erythraeus, but their identity and nomenclatorial status remain doubtful until redescription. Fallopia Oudemans, 1941 and Rhyncholophus poriferus Kramer, 1998 from New Britain – nomina dubia Grognieria Oudemans, 1941 and Rhyncholophus agilis Canestrini, 1884 – nomina dubia Hehlenia hermanni Oudemans, 1941 – unrecognizable Hehlenia hermanni Oudemans, 1941 – as nom. nov. pro Smaridia papillosa Berlese, 1884 non Trombidium papillosum Herm., 1804, nec Rhyncholophus papillosus C.L. Koch, 1837 – unrecognizable Hehlenia kochi Oudemans, 1941 – as nom. nov. pro Rhyncholophus papillosus C.L. Koch, 1837 – unrecognizable Platylophus Berlese, 1910 (praeoccup. from Platylophus Swainson, 1832) – homonym Platylophus argasoides Berlese, 1910 – from “Mechieria”(Africa bor.)(Algeria ? – unrecognizable, possibly synonym of Phanolophus oedipodarum Frauenfeld, 1868 = Ph. nasica André, 1927) Subgenus Ptilolophus Berlese, 1916 (not “Ptilophus”, as in Southcott, 1961a – lapsus calami) – nomen dubium Legneria Oudemans, 1941 – ? syn. of Abrolophus

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Leuchsia Oudemans, 1941 – junior synonym of Smaris Latr. Niemannia Oudemans, 1941 – syn. of Hirstiosoma Womersley (type Smaridia ampulligera Berlese, 1887) Rhyncholophus tridentifer Ewing, 1909 – member of Neotrombidium Leonardi, 1901 (see Southcott, 1961a: 412) Samouellea Oudemans, 1941 – ? syn. of Smaris Latreille Samouellea depilata (Berlese, 1988) for Smaridia depilata Berlese from South America – may be a good species as Smaris depilata (Berl.) – needs redescription Smaridia vestita Gervais, 1849 – unrecognizable; renamed Leuchsia vestita (Gerv.) by Oudemans, 1941: 186 Smaridia villosa Dugès, 1834 – renamed Hehlenia villosa by Oudemans, 1941: 181 Smaris mamillatus Say (ad.): Oudemans, 1937: 952 for Erythraeus mamillatus Say, 1821: 70 (ad.); E. mamillatus (Say): Banks, 1907: 598; Labidostomma mamillatus (Say): Jacot, 1938: 125; Banks, 1947: 129; Southcott, 1961a: 574. Note: Southcott (loc.cit.) expressed the opinion that it would be better to accept the view of Banks (1947) and to leave the species of Say in the fam. Labidostommatidae and that the “Smaris mamillatus”, figured by Baker and Wharton (1952) is a different species, really belonging to the genus Smaris. Type locality: Georgia and East Florida. Distribution: U.S.A. (Georgia, Florida) Veithia Oudemans, 1941 – nomen dubium, unrecognizable. The species Veithia assmuthi Oudemans, 1941 was redescribed by Zhang (2003) as Fessonia assmuthi (Oud.)

Other names, published in Erythraeoidea Acarus bicolor Schrank, 1803 Acarus hexapus Schrank, 1803 Astoma latirostre: Latreille, 1829 Erythraeus assimilis (Herm.): Oudemans, 1937: 1938 Leptus araneii Say, 1821 (see Oudemans, 1937: 1919) Leptus insectorum Latreille, 1817 Leptus latirostris: Latreille, 1817 Leptus oribatus Johnston, 1848 Leptus phalangiorum Brebisson, 1827 Ocypete Say, 1821 Rhyncholophus Hermann Dugès, 1834 Rhyncholophus humeralis Karsch, 1881 (Japan) Rhyncholophus paludicola C.L. Koch, 1837 Trombidium assimile Latreille, 1817 Trombidium insectorum Oken, 1835 (Allg. Natugesch., 8: 662) Trombidium latirostre Hermann, 1804 Trombidium saccatum C.L. Koch, 1849 (nom. nud.)(Balaustium, acc. to Oudemans, 1937: 1953)


Fossil Erythraeoidea Genus Proterythraeus Vercammen-Grandjean (larval, fossil) Proterythraeus Vercammen-Grandjean, 1972: 329

Type species: Proterythraeus southcotti Vercammen-Grandjean, 1972

Proterythraeus southcotti Vercammen-Grandjean (larval, fossil) Proterythraeus southcotti Vercammen-Grandjean, 1972: 329

Distribution: Canadian Cretaceous amber

Genus Erythraeus Latreille (larval, fossil and rec.) Erythraeus foveolatus (C.L. Koch) Rhyncholophus foveolatus C.L. Koch in Koch & Berendt, 1854 Erythraeus foveolatus (Koch): Oudemans, 1937: 1940

Distribution: Baltic amber

Erythraeus hirsutissimus (C.L. Koch) Rhyncholophus hirsutissimus C.L. Koch in Koch & Berendt, 1854 Erythraeus hirsutissimus (Koch et Berendt)


Erythraeus lagopus (Koch et Berendt) Rhyncholophus lagopus C.L. Koch in Koch & Berendt, 1854 Erythraeus lagopus (C.L. Koch)


Erythraeus longipes (C.L. Koch) Rhyncholophus longipes C. L. Koch in Koch & Berendt, 1854 Erythraeus longipes (C.L. Koch): Oudemans, 1937: 1979


Erythraeus proavus (C.L. Koch) Rhyncholophus proavus C.L. Koch in Koch & Berendt, 1854 Erythraeus proavus (C.L. Koch)


Erythraeus raripilus (C.L. Koch) Rhyncholophus raripilus C.L. Koch in Koch & Berendt, 1854 Erythraeus raripilus (C.L. Koch)


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192


Genus Balaustium von Heyden Balaustium illustris (C.L. Koch) Rhyncholophus illustris C.L. Koch in Koch & Berendt, 1854 Balaustium illustris (C.L. Koch): Oudemans, 1937: 1940


Genus Leptus Latreille (larval, fossil et rec.) ? Leptus incertus (C.L. Koch) Rhyncholophus incertus C.L. Koch in Koch & Berendt, 1854 Leptus incertus (C.L. Koch): Oudemans, 1937 : 1926

Distribution: Baltic amberGenus Arythaena Menge (ad., fossil) Arythaena Menge, 1854

Type species: Arythaena troguloides Menge, 1854

Arythaena troguloides Menge (ad.) Arythaena troguloides Menge, 1854

Distribution: Baltic amber (Olig.) Notes of Zacharda & Krivoluckij (1985, p. 150-151): “The material of the Paleogene Baltic amber, as far as it exists at all, should be re-examined from the modern systematic viewpoint”. One well preserved specimen of the larva of Erythraeoidea is recorded in this paper from amber from Jantardach, Taymir peninsula, Russia (tentatively placed in the family Erythraeidae).


193

Taxonomical changes in this Catalogue Hirstiosoma glacialis (Schmölzer, 1956) – comb. nov. pro Smaris glacialis Schmölzer Erythraeus (E.) hirsti Beron – nom. nov. pro Erythraeus pilosus (Hirst, 1928), nec Erythraeus pilosus Banks, 1894 Erythraeus (E.) plumatus Beron, nom. nov. pro Erythraeus plumosus Tseng, Yang & Pan, 1976 (nomen praeoccupatum from E. plumosus Khot, 1963) Erythraeus (E.) chinensis (Zheng, 2002) – comb. nov. pro Bochartia chinensis Zheng Genus Helladerythraeus Beron, 1988 – stat. nov. from Erythraeus (Helladerythraeus) Beron Abrolophus aequalipes (Schweizer et Bader, 1963) – comb. nov. pro Balaustium aequalipes Schweizer et Bader Abrolophus anomalus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium anomalus Schweizer et Bader Abrolophus basiliensis (Schweizer et Bader, 1963) – comb. nov. pro Balaustium basiliense Schweizer et Bader Abrolophus brevipalpis (Schweizer et Bader, 1963) – comb. nov. pro Balaustium brevipalpe Schweizer et Bader Abrolophus engadinensis (Schweizer et Bader, 1963) – comb. nov. pro Balaustium engadinense Schweizer et Bader Abrolophus evansi (Schweizer et Bader, 1963) – comb. nov. pro Balaustium evansi Schweizer et Bader Abrolophus handschini (Schweizer et Bader, 1963) – comb. nov. pro Balaustium handschini Schweizer et Bader Abrolophus jurassicus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium jurassicum Schweizer et Bader Abrolophus longipapillus (Schweizer, 1951) – comb. nov. pro Balaustium longipapillum Schweizer Abrolophus novaeguinensis (Gunther, 1941) – comb. nov. pro Belaustium [sic] novaeguinensis Gunther Abrolophus perifi (Schweizer et Bader, 1963) – comb. nov. pro Balaustium perifi Schweizer et Bader Abrolophus quadrisetus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium quadrisetum Schweizer et Bader Abrolophus quinquesetus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium quinquesetum Schweizer et Bader Abrolophus scopularius (Hull, 1918) – comb. nov. pro Belaustium [sic] scopularium Hull Abrolophus subquisquiliarus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium subquisquiliarum Schweizer et Bader Abrolophus trisetus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium trisetum Schweizer et Bader

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Abrolophus unisetus (Schweizer et Bader, 1963) – comb. nov. pro Balaustium unisetum Schweizer et Bader Charletonia arborum (Vitzthum, 1926) – comb. nov. pro Sphaerolophus arborum Vitzthum Charletonia canadensis (Smiley, 1968) – comb. nov. pro Sphaerolophus canadensis Smiley Charletonia cilissa (Cooreman, 1955) – comb. nov. pro Cavannea cilissa Cooreman Charletonia cursor (Vitzthum, 1926) – comb. nov. pro Sphaerolophus cursor Vitzthum Charletonia delhiensis (Khot, 1965) – comb. nov. pro Sphaerolophus delhiensis Khot Charletonia gigas (Khot, 1965) – comb. nov. pro Sphaerolophus (Cavannea) gigas Khot Charletonia mollicomus (Berlese, 1910) – comb. nov. pro Sphaerolophus (Cavannea) mollicomus Berlese Charletonia politricha (Berlese, 1910) – comb. nov. pro Sphaerolophus politrichus Berlese Charletonia porcinus (Hull, 1918) – comb. nov. pro Achorolophus porcinus Hull Charletonia salti (Evans, 1953) – comb. nov. pro Sphaerolophus salti Evans Charletonia spinosa (Berlese, 1910) – comb. nov. pro Sphaerolophus spinosus Berlese Charletonia subnuda (Berlese, 1910) – comb. nov. pro Sphaerolophus subnudus Berlese Charletonia sureshi (Ramaraju et Mohanasundaram, 1998) – comb. nov. pro Sphaerolophus sureshi Ramaraju et Mohanasundaram


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Distribution of Erythraeoidea EUROPE Albania – Erythraeidae: Erythraeus regalis Andorra (Haitlinger, 2005b) – Erythraeidae: Hauptmannia brevicollis, H. kazimierae; Erythraeus malwinae Austria (Cokendolpher, 1993; Franz, 1954; Haitlinger, 1995; Irk, 1939a, 1939b; Mihelčič, 1964; 1971; Schmölzer, 1993; Turk, 1981; Turk & Schuster, 1983; Willmann, 1951b, 1951c, 1951d, 1953) – Smarididae: Hirstiosoma ampulligera; Erythraeidae: Erythraeus acis, E. adpendiculatus, E. glabrisetosus, E. kuyperi, E. nivalis, E. opilionoides, E. phalangoides (=E. dubiosus), E. regalis, E. rupestris (=E. imperialis), E. styriacus, Curteria curticristata; Leptus calvatus, L. curtipes, L. erabundus, L. longipes, L. longipilis, L. molochinus, L. nemorum, L. ochroniger, L. phalangii, L. rubricatus, L. trimaculatus, L. vertex, Charletonia cardinalis, Ch. globigera, Abrolophus artemisiae (= lapidarius), A. curtipapillus,A. longipes, A. longulus, A. longus, A. miniatus, A. quisquiliarus, A. quisquiliarus sigma, A. raripapillus, A. rhopalicus, A. sabulosus, A. tardus, Hauptmannia brevicollis, Balaustium araneoides, B. bulgariense, B. murorum, B. neomurorum Azerbaydjan (Gadjiev & Rahmatulina, Eds., 1996) – Smarididae: Fessonia sp.; Erythraeidae: Eatoniana plumipes forma transcaucasica, “Leptus ignotus”, Charletonia cardinalis (= singularis); Phanolophus oedipodarum (= nasica) Belgium (Cooreman, 1943, 1945; Fain, 1991a, 1992; Fain & Baugnée, 1996; Fain, Baugnée & Hidvegi, 1992; Willmann, 1935) – Erythraeidae: Erythraeus (= Bochartia) adrastus, E. regalis, Leptus beroni, L. charanyca, L. cooremani, L. curtipes, L. ignotus, L. mariae, L. murteri, L. trimaculatus, Charletonia globigera, Ch. womersleyi, Abrolophus quisquiliarus, Balaustium madeirense, B. murorum Bosnia and Herzegovina (Haitlinger, 2007a; Turk, 1981; Willmann, 1941) – Erythraeidae: Erythraeus (Zaracarus) budapestensis, Erythraeus (E.) styriacus, Leptus josifovi, Abrolophus pseudolongicollis kiejstuti, A. tardus, Hauptmannia podorasensis Bulgaria (Beron, 1975, 1982, 1995, 2005, in prep.; Haitlinger, 2007a; Oudemans, 1926; Willmann, 1951a) – Smarididae: Smaris squamata, S. philopempta; Hirstiosoma ampulligera; Erythraeidae: Erythraeus bulgaromontanus, E. kresnensis, E. regalis, E. rilensis, Leptus beroni, L. josifovi, L. mariae, L. meloidarum, L. orthopterarum, L. slivovi, L. southcotti, L. trimaculatus (= echinopus), Charletonia bucephalia, Ch. krendowskyi, Abrolophus sp., Hauptmannia sp.; Balaustium bulgariense, Neobalaustium caspersi Croatia (Jablonowski, 1918; Haitlinger, 2004d; 2007a) – Erythraeidae: Erythraeus budapestensis, E. crocatus, E. sibuljinicus, Leptus josifovi, L. trimaculatus, Charletonia bucephalia, Ch. dalegori, Ch. krendowskyi, Ch. zorani, Abrolophus longus, A. pseudolongicollis, Hauptmannia pogoracensis, Grandjeanella multisetosa, Balaustium nikae, Phanolophus oedipodarum Czech Rep. (Daniel, 1971; Daniel & Samšinak, 1955; Haitlinger, 2003f; Willmann, 1954) – Erythraeidae: Erythraeus kuyperi, Leptus absoloni, Leptus longispinosus, L. nemorum, L.

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rubricatus, L. trimaculatus (= echinopus), Abrolophus longicollis, A. tardus, Charletonia cardinalis, Hauptmannia brevicollis, H. kotorensis, H. wratislaviensis, Phanolophus oedipodarum, Balaustium affine, B. bulgariense, B. murorum, B. nikae Denmark (Bornebusch, 1930; Haarløv, 1957, 1960; Hallas, 1978; Southcott, 1961a) – Erythraeidae: Erythreus phalangoides, E. (= Bochartia) adrastus, Leptus nemorum, L. echinopus Greenland (Haarløv, 1942; Kramer, 1897, Trägårdh, 1904b) – Erythraeidae: Erythraeus phalangoides, Leptus groenlandicus, L. ignotus (= ? groenlandicus Tr.), L. vertex, L. kalaallus, Abrolophus miniatus, Balaustium unidentatum Faroe Is. (Trägårdh, 1931b, 1971) – Erythraeidae: Abrolophus rubipes, Leptus verticiformis Finland (Haitlinger, 2000b, Karppinen, 1958) – Erythraeidae: Erythraeus adpendiculatus, E. monikae, E. phalangoides, E. regalis, E. rupestris, Leptus nemorum, L. oudemansi, L. trimaculatus, Hauptmannia brevicollis, Charletonia globigera, Balaustium murorum France (continent.)(André, 1929, 1930, 1933, 1953; Brébisson, 1827; Cooreman, 1955b; Dugès, 1834, 1836; Fain & D’Amico, 1997; Grandjean, 1947a, 1947b; Hermann, 1804; Latreille, 1796, 1802, 1804, 1806, 1810, 1817, 1829; Schmölzer, 1956; Southcott, 1961a; Theodoridès, 1955) – Smarididae: Smaris squamata, Hirstiosoma latreillei, H. ampulligera, H. glacialis, Fessonia papillosa, Clipeosoma jupiter; Erythraeidae: Erythraeus adpendiculatus, E. cinereus, E. kuyperi, E. phalangoides, E. pseudonivalis, E. regalis, Forania mentonensis, Leptus beroni, L. gyas, L. pyrenaeus, L. trimaculatus, Abrolophus raripapillus, Charletonia globigera, Ch. venus, Abrolophus artemisiae, A. miniatus, A. nivalis, A. quisquiliarus, A. rubipes, Andrevella parkeri, Balaustium florale Corsica (Beron, in prep.; Grandjean, 1947b, Haitlinger, 2006d) – Smarididae: Hirstiosoma ampulligera, Erythraeidae: Leptus josifovi, Charletonia dalegori, C. glifadaensis, Abrolophus marinensis, Balaustium florale Germany (Berlese, 1910; Franke, 1940; Haitlinger, 1995, 1996, 1997b; Haitlinger & Walter, 1997; Hahn, 1835; Haller, 1882; Halliday, 2001; C.L. Koch, 1836-1841; Maixner et al, 1998; Munchberg, 1935, 1936, 1956; Oudemans, 1903; Pfeifer 1956; Rack, 1976; Southcott, 1961a; Weidner, 1954; Wendt, 1996; Wendt et al., 1992; Willmann, 1952, 1955; Wohltmann, 1995; Wohltmann et al., 2007) – Smarididae: Smaris squamata, Fessonia papillosa; Erythraeidae: Erythraeus adpendiculatus, E. electoralis, E. germanicus, E. malwinae, E. monikae, E. nivalis, E. opilionoides, E. phalangoides, E. principalis, E. regalis; Curteria episcopalis; Leptus beroni, L. ignotus, L. longipilis, L. molochinus, L. nemorum, L. phalangii, L. rubricatus, L. trimaculatus, L. vertex, Charletonia cardinalis, Ch. globigera, Abrolophus artemisiae, A. complanatus, A. frisius, A. miniatus, A. quisquiliarus, A. raripapillus, A. rhopalicus, Hauptmannia brevicollis, H. gracilenta, H. nivalis, Balaustium murorum, B. wratislaviensis; Inc. sedes: “Erythraeus eucomus, Erythraeus decalvatus” Great Britain (Cambridge, 1875, Evans, 1952; Evans & Browning, 1953; Fain & Greenwood, 1991; George, 1896, 1907, 1909, 1910, 1911, 1912, 1913; Haitlinger, 2000c; Halliday, 2001; Hull, 1918; Johnston, 1848; Killington & Bathe, 1946; Parker, 1962; Rack, 1974; Southcott, 1961a; Sulaiman, 1983; Turk, 1945a, 1945b, 1953) – Smarididae: Hirstiosoma ampulligera; Erythraeidae: Erythraeus germanicus, E. johnstoni, E. monikae, E. opilionoides, “E.” pachy-


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pus, “E”. penninus, E. phalangoides, E. regalis, Leptus ignotus, L. killingtoni, L. nemorum, L. trimaculatus, L. vertex, Charletonia globigera, Ch. womersleyi, Abrolophus falconeri, A. miniatus, A. norvegicus, A. quisquiliarus, A. rubipes, A. sabulosus, A. schweizeri, A. tardus, A. vittatus, Hauptmannia wratislaviensis, Balaustium araneoides, “B.” harrisoni, B. murorum Jersey (George, 1896; Long, 1980) – Erythraeidae: Eatoniana sp. (? plumipes) Greece (cont.)(Berlese, 1916; Beron, 1988, 1990, in prep.; Cooreman, 1955a; Haitlinger, 1984, 1993, 2006f ) – Smarididae: Smaris squamata, Fessonia papillosa; Erythraeidae: Helladerythraeus pachytibialis, Erythraeus (Zaracarus) preciosus, E. (Z.) budapestensis, Eatoniana sp. (? plumipes), ? Curteria curticristata, Leptus josifovi, L. trimaculatus, Charletonia bucephalia (= Ch. tamarae), Ch. krendowskyi, Ch. venus, Phanolophus oedipodarum, Abrolophus rhopalicus, A. pseudolongicollis kiestuti, Hauptmannia podorasensis, Grandjeanella multisetosa, Bursaustium gaspari, Balaustium madeirense, B. murorum, B. nikae, Myrmicotrombium mirum Rhodes (Beron, 1988, Beron, in prep.; Haitlinger, 2003a, 2003b) – Smarididae: Smaris squamata; Erythraeidae: Erythraeus rutgeri, Curteria graeca, Leptus andae, L. gennadicus, L. monolithosicus, Charletonia dalegori, Ch. glifadaensis, Ch. kaliksti, Ch. krendowskyi, Rudaemannia rudaensis Crete (Beron, in prep.) – Smarididae: Smaris squamata; Erythraeidae: Curteria graeca Samos (Haitlinger, 2006b) – Erythraeidae: Erythraeus (Zaracarus) kastaniensis, E. (Z.) passidonicus, Hauptmannia silesiacus, Grandjeanella multisetosa, Leptus josifovi, Charletonia kalithensis, Ch. samosensis Corfu (Berlese, 1890; Sellnick, 1931) – Erythraeidae: Leptus trimaculatus, L. villosus Kephallonia (Sellnick, 1931) – Erythraeidae: Balaustium murorum Levkas (Sellnick, 1931) – Smarididae: Smaris squamata; Erythraeidae: Abrolophus rhopalicus Kithnos (Beron, 1988) – Erythraeidae: Curteria graeca Holland (The Netherlands) (Oudemans, 1903, 1905a, 1906, 1910a, 1910b, 1914; Southcott, 1992) – Erythraeidae: Curteria episcopalis; Erythraeus adpendiculatus, E. cinereus, E. kuyperi, E. opilionoides, E. phalangoides, E. regalis; Leptus beroni, L. ignotus, L. mariae, L. molochinus, L. ochroniger, L. rubricatus, L. sigthori, L. trimaculatus; Charletonia cardinalis, Ch. globigera; Abrolophus artemisiae, A. longicollis, A. quisquiliarus; Hauptmannia brevicollis; Balaustium murorum Hungary (Dudich, Kolosvary & Szalay, 1940; Fain & Ripka, 1998; Gabryś & Mąkol, 1991, 1996; Jablonowski, 1918; Karpelles, 1893; Willmann, 1938) – Smarididae: Fessonia papillosa; Hirstiosoma latreillei; Clipeosoma jupiter; Erythraeidae: Erythraeus acis, E. adpendiculatus, E. budapestensis, E. cinereus, E. electoralis, E. jowitae, E. opilionoides, E. phalagoides, E. regalis, Curteria southcotti, Leptus beroni, L. clethrionomydis, L. ignotus, L. longipilis, L. mariae, L. molochinus, L. nemorum, L. ochroniger, L. phalangii, L. rubricatus, L. trimaculatus (= echinopus), L. vertex, Abrolophus artemisiae, A. crassitarsus, A. miniatus, A. norvegicus, A. quisquiliarus, A. strojnyi, Charletonia cavannae; Balaustium murorum. Inc. sedes: “Smaridia hungarica”

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Iceland (Lindroth et al., 1973; Sellnick, 1940; Southcott, 1989c) – Erythraeidae: Erythraeus regalis, E. phalangoides; Leptus beroni, L. ignotus, L. sigthori, L. vertex, Abrolophus longicollis, A. rhopalicus; Hauptmannia brevicollis, H. wratislaviensis (as Balaustioides tuxeni) Ireland (Halbert, 1915, 1920) – Erythraeidae: Erythraeus regalis, Leptus beroni, L. nemorum, Abrolophus halberti, A. miniatus, A. passerinii, A. rubipes, A. sabulosus, A. tardus, Balaustium araneipes, B. araneoides Italy (continent.)(Berlese, 1885, 1910, 1913, 1918, ; Bernini et al., 1994-1996; Bottazzi, 1951, 1956; Canestrini G., 1884, 1885; Canestrini G. & R. Canestrini. 1882; Canestrini & Fanzago, 1877; di Caporiacco, 1938; Contarini, 1843, 1847; Cooreman, 1955c; Haitlinger, 1996, 1998, 2000d, 2003d; Lombardini, 1949, 1958; Oudemans, 1901, 1941; Paoli, 1937; Rossi, 1794; Starkoff & Starkoff, 1950; Zhang, 2003) – Smarididae: Smaris granatae, S. squamata, Fessonia papillosa; Hirstiosoma ampulligera; Veithia schneideri; Erythraeidae: Erythraeus acis, E. adpendiculatus, E. cinereus, E. crocatus, E. electoralis, E. hibernans, E. italicus, E. kuyperi, E. opilionoides, E. phalangoides, E. regalis, E. similis; Leptus agilis, L. beroni (= holmiae), L. boccettii, L. curtipes, L. dubius, L. eupodes, L. ignotus, L. longipilis, L. mariae, L. meloidarum, L. murteri, L. nemorum, L. pulchellus, L. siculus, L. trimaculatus, L. vertex; Charletonia berlesiana, Ch. cardinalis, Ch. cavannae, Ch. globigera, Ch. mollicomus, Ch. venus; Abrolophus complanatus, A. curtipapillus, A. epigaeus, A. longipilis, A. miniatus, A. passerinii, A. pseudolongicollis, A. pulcher, A. quisquiliarus; Phanolophus oedipodarum; Balaustium madeirense, B. murorum, Italustium eframi Sicily (Berlese, 1882, 1888, 1910; Canestrini, 1884) – Smarididae: Smaris squamata; Erythraeidae: Erythraeus acis, E. cinereus; Leptus araneoides, L. siculus; Balaustium araneoides, B. murorum; “Erythrombium eusisyrum” – nomen dubium Sardinia (Beron, in print, Haitlinger, 2006d, Lombardini, 1958) – Smarididae: Hirstiosoma ampulligera; Smaris squamata; Fessonia papillosa; Erythraeidae: Erythraeus cinereus, E. electoralis, E. regalis, Leptus josifovi, L. pozzoicus, Charletonia austisensis, C. cuglierensis, Hauptmannia sardiniensis, “Achorolophus boccetii” Latvia (Haitlinger, 2000b) – Erythraeidae: Leptus trimaculatus (=echinopus) Liechtenstein (Haitlinger, 2007b) – Erythraeidae: Leptus slivovi, Abrolophus pseudolongicollis, Hauptmannia brevicollis Lithuania (Haitlinger, 2000b; 2003d) – Erythraeidae: Abrolophus brevicollis; Hauptmannia wratislaviensis Macedonia (Turk, 1981) – Erythraeidae: Erythraeus styriacus Moldova (Haitlinger, 2007c) – Erythraeidae: Rudaemannia rudaensis, Hauptmannia brevicollis, H. wratislaviensis, Moldoustium baltiensis Montenegro (Haitlinger 2007a) – Erythraeidae: Erythraeus opilionides, E. (Zaracarus) budapestensis, Abrolophus pseudolongicollis kiejstuti, Hauptmannia kotorensis Norway (Åbro, 1991; Buitendijk, 1945; Haitlinger, 2000b; Mąkol & Gulvik, 2002; Thor, 1900a, 1900b; Trägårdh,1904b) – Smarididae: Hirstiosoma ampulligera; Erythraeidae: Erythraeus gertrudae, E. opilionoides, E. phalangoides, E. regalis; Leptus beroni,


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L. mariae, L. molochinus, L. rubricatus, L. slivovi, L. vertex; Charletonia globigera, Abrolophus norvegicus; Hauptmannia brevicollis, H. wratislaviensis; Inc. sedis: “Rhyncholophus (Ritteria) plumosa” Poland (Frenzel, 1936; Gabryś, 1988, 1991a, 1991b, 1992a, 1992c, 1997; Gabryś & Haitlinger, 1986; Gabryś & Mąkol, 1994, 1995, 1997, 2000; Gabryś, Tomaszewska, Mąkol & Chorbinski, 1995; Haitlinger, 1985, 1986b, 1987a, 1987d, 1987h, 1987i, 1992, 1996, 2000c, 2002b, 2002e, 2003f; Halliday, 2001; Ignatowicz, 1974, 1975; Kielczewski & Kashyna, 1965; Mąkol & Gabryś, 1999; Oudemans, 1912; Southcott, 1992; Vitzthum, 1935b; Willmann, 1949, 1950, 1956; Winiarska, 1998; Witalinski, 1985) – Smarididae: Hirstiosoma latreillei; Clipeosoma jupiter; Erythraeidae: Erythraeus acis, E. adpendiculatus, E. cinereus, E. eleonorae, E. elwirae, E. gertrudae, E. jowitae, E. kuyperi, E. malwinae, E. monikae, E. opilionoides, E. phalangoides (= E. dubiosus), E. regalis, E. rupestris (= E. imperialis); Curteria episcopalis, C. southcotti; Leptus beroni (= L. holmiae), L. clethrionomydis, L. gabrysi, L. ignotus (= L. zbelutkaicus), L. longipilis, L. machilidis, L. mariae, L. miromiri, L. molochinus, L. nemorum, L. rubricatus, L. slivovi, L. trimaculatus (= echinopus), L. vertex; Charletonia cardinalis (= Ch. singularis), Ch. globigera; Abrolophus artemisiae, A. bohdani, A. crassitarsus, A. humberti, A. lapidarius, A. longicollis, A. miniatus, A. norvegicus, A. passerinii, A. pseudolongicollis, A. quisquiliarus, A. rhopalicus, A. strojnyi, A. tardus; Hauptmannia brevicollis, H. kazimierae, H. silesiaca, H. stanislavae, H. wratislaviensis; Rudaemannia rudaensis; Balaustium kacperi, B. murorum, B. nikae, B. rajmundi, B. unidentatum, B. wratislaviensis, B. xerothermicum Portugal (continental)(Haitlinger, 2006e) – Erythraeidae: Leptus josifovi Madeira (Haitlinger, 2002d; Southcott, 1992, 1976; Willmann, 1939a) – Erythraeidae: Leptus millipedius; Abrolophus benoni, A. neobrevicollis; Balaustium madeirense Azores (McNeill & Treat, 1992) – Erythraeidae: Leptus killingtoni Romania (Jablonowski, 1918; Cooreman, 1951; Feider, 1948, 1950, 1954, 1955, 1956, 1967, 1980, 1981; Feider & Suciu, 1959, 1963; Szalay, 1931) – Smarididae: Smaris squamata; Erythraeidae: Erythraeus electoralis, E. phalangoides, E. regalis, Leptus galerucae, L. phyllotretae, L. trimaculatus, Abrolophus iacobi; Charletonia krendowskyi; Balaustium murorum; Neobalaustium borceanum Russia (all, continent.)(Buitendijk, 1945; Dwigubsky, 1802; Haitlinger, 2003d; Koch L., 1879) – Erythraeidae: Erythraeus cinereus, E. rupestris (=imperialis), Leptus beroni, Charletonia cardinalis [Kaliningrad], Abrolophus quisquiliarus [Kaliningrad], Hauptmannia brevicollis, H. wratislaviensis, “Rhyncholophus tonsus” Kuril Is. (Thor & Ushida, 1933) – Erythraeidae: Erythraeus phalangoides Bering I. (Banks, 1899) – Erythraeidae: “Erythraeus elongatus Banks” (Inc. sedis – ? Trombidiidae s.l.) Slovakia (Haitlinger, 2002d; Jablonowski, 1918; Stasiov, 2003) – Erythraeidae: Erythraeus gertrudae, E. monikae, Leptus beroni, L. trimaculatus (=echinopus), Hauptmannia brevicollis, H. kazimierae, H. stanislavae, H. silesiaca, Balaustium nikae

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Slovenia (Haitlinger, 2003d, 2007a; Southcott, 1992; Turk, 1981) – Erythraeidae: Erythraeus gertrudae, E. monikae, E. styriacus, Leptus beroni, L. mariae, L. trimaculatus (= echinopus), Abrolophus pseudolongicollis, Hauptmannia brevicollis, H. kazimierae, H. kotorensis, H. wratislaviensis Spain (continent.) (Gabryś, 2000c; Zhang & Goldarazena, 1996; Goldarazena & Zhang, 1997, 1998a, 1998b; Goldarazena, Jornada & Z.-Q. Zhang, 1999a, 1999b; Goldarazena, Loomis & Jordana, 1999; Haitlinger, 1990c, 1994b, 2006e; Mayoral & Barranco, 2004, 2006; Mihelčič, 1958a, 1958b; Southcott, 1993b, 1995b) – Smarididae: Smaris bipartita, S. squamata, Hirstiosoma ampulligera; Erythraeidae: Abalakeus lorestanicus, Erythraeus acis, E. acutipilis, E. campestri, E. imperfectus, E. lancifer, E. phalangoides, E. preciosus, E. regalis, E. southcotti; Eatoniana plumipes; Leptus akkus, L. curtipes, L. diversus, L. furibundus, L. incertus, L. killingtoni, L. mariae, L. mirenae, L. molochinus, L. orthopterarum, L. parvulus, L. rubricatus, L. trimaculatus, L. variatus; Grandjeanella haitlingeri, G. multisetosa; Charletonia blascoi, Ch. kaliksti; Abrolophus neobrevicollis, A. quisquiliarus, A. veletensis; Balaustium neominiatum Balear Is. (L. Koch, 1882, Haitlinger, 2002f ) – Erythraeidae: Erythraeus picaforticus, E. southcotti, E. preciosus; Grandjeanella ainae; Balaustium florale; Inc. sedis: “Rhyncholophus vernalis” Canary Is. (Haitlinger, 2004g; Haitlinger & Saboori, 1996; Southcott, 1991a) – Erythraeidae: Erythraeus fabiolae [Tenerife], Curteria (=Pedroerythraeus) ernesti [La Palma], Leptus tinnae, Charletonia enghoffi [Tenerife], Hauptmannia baardi [La Palma], Abrolophus benoni [La Palma], Balaustium barloventensis [La Palma], B. malpaisesensis [La Palma] Sweden (Danell, 1987; Sellnick, 1949, 1958; Somermaa, 1973; Southcott, 1992a; Willmann, 1943) – Erythraeidae: Erythraeus kuyperi, E. phalangoides, Leptus beroni, L. ignotus, L. laplandicus, L. phalangii, L. sigthori, L. vertex, Charletonia cardinalis, Ch. globigera, Abrolophus densipapillus, A. longicollis, A. rubipes, Hauptmannia brevicollis, Balaustium murorum Switzerland (Gabryś, 2000c; Haitlinger, 1994a; 2007b; Haller, 1885; Schweizer, 1922, 1951, 195 ; Schweizer & Bader, 1963) – Smarididae: Hirstiosoma ampulligera; Erythraeidae: Erythraeus acis, E. dubiosus, E. kuyperi, E. mariolae, E. nivalis, E. phalangoides, E. regalis, Eatoniana plumipes, Curteria curticristata, Leptus beroni, L. curtipes, L. ignotus, L. josifovi, L. mariae, L. molochinus, L. murteri, L. rubricatus, L. trimaculatus (= echinopus), L. vertex, Charletonia globigera, Abrolophus aequalipes, A. anomalus, A. baderi, A. basiliensis, A. brevipalpis, A. crassitarsus, A. densipapillus, A. engadinensis, A. evansi, A. handschini, A. jurassicus, A. laevicristatus, A. longipapillus, A. longus, A. miniatus, A. perifi, A. perlongus, A. quadrisetus, A. quinquesetus, A. quisquiliarus, A. raripapillus, A. rhopalicus, A. sabulosus, A. schweizeri, A. subquisquiliarus, A. trisetus, A. unisetus, Hauptmannia parvum, H. kazimierae, H. willmanni, H. wratislaviensis, Rudaemannia rudaensis, Balaustium murorum, B. neomurorum, B. submurorum, B. unidentatum


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Turkey (all)(Cooreman, 1955a; Fain & Cobanoglu, 1998; Haitlinger, 1999a; 2000d; Goldarazena, Zhang & Jordana, 2000; Saboori, Cakmak & Nouri-Gonbalani, 2004; Saboori, Cobanoglu & Bayram, 2004; Bayram & Cobanoglu, 2005) – Erythraeidae: Erythraeus ankaraicus, E. kresnensis, E. sifi; E. (Zaracarus) didonae, E. (Zaracarus) aydinicus; Leptus rosellae, Charletonia cardinalis, Ch. cilissa, Hauptmannia viburnicola, Rudaemannia rudaensis (= Hauptmannia viticola), Grandjeanella multisetosa, Balaustium madeirense, Bursaustium gaspari Ukraine (Haitlinger, 2007c) – Erythraeidae: Erythraeus (Zaracarus) preciosus, E. (E.) kuyperi, Leptus trimaculatus, Grandjeanella multisetosa, Hauptmannia brevicollis, H. wratislaviensis, Abrolophus pseudolongicollis kiejstuti, A. crimensis, Moldoustium baltiensis, Balaustium nikae AFRICA Algeria (André, 1927a; Berlese, 1910; Cooreman, 1956; Lucas, 1846, 1864) – Erythraeidae: Erythraeus regalis, E. tragardhi, Curteria fageli, Eatoniana plumipes, Abrolophus miniatus, Phanolophus oedipodarum (= nasica), Balaustium araneipes; Inc. sedis: “Rhyncholophus pallipes” Benin (Haitlinger, 2006g) – Erythraeidae: Leptus pelebinus, Charletonia beninensis, Ch. brunni, Ch. grandpopensis, Lomeustium togoensis Congo – Kinshassa (D.R. Congo, former Zaire)(Cooreman, 1958; Fain & Elsen, 1987) – Erythraeidae: Leptus benzaliensis, L. carpenteri, L. glossinarum, L. lovaniensis, L. maringensis, L. similis; Caeculisoma afrum, Caecidopus straeleni Egypt (Trägårdh, 1904a) – Erythraeidae: Erythraeus tragardhi; Leptus siculiformis; Charletonia jaegerskioeldi [Sinai]; Inc. sedis: “Lucasiella plumipes” Ethiopia (Berlese, 1916b, Haitlinger, 2006h, Lombardini, 1941) – Erythraeidae: Erythraeus (Z.) jinkaensis, Erythraeus sp.; Leptus dalicus, L. dinekaicus, Abrolophus simplex, A. penelopae, Charletonia braunsi, Ch. brunni Ghana (Haitlinger, 1990, 2006g) – Erythraeidae: Leptus bertoldi, L. elminus, L. abrafoicus, Abrolophus basumtwiensis, Charletonia beninensis, Ch. braunsi, Ch. brunni, Ch. ghanensis, Lomeustium togoensis Guinea (Oudemans, 1910) – Erythraeidae: Charletonia braunsi Ivory Coast (Fain & Elsen, 1972) – Erythraeidae: Leptus benzaliensis Kenya (Haitlinger, 1990a, 2001b, 2004c; Southcott, 1991a) – Erythraeidae: Leptus aggoratus, L. benzaliensis, L. bogoriacus, L. olamukijacus, L. masaimaraicus; Charletonia areolata, Ch. naivashae; Momorangia gabini Libya (Beron, in prep.; Pavesi, 1884) – Erythraeidae: “Rhyncholophus pallidipes” (Inc. sedis) Madagascar (André, 1941, 1946; Haitlinger, 1987g) – Erythraeidae: Leptus aldonae, L. madagascariensis, L. maranaensis; Charletonia agatae, Ch. alarobiaensis, Ch. arlettae, Ch. dorotae, Ch. edytae, Ch. iwonae, Ch. justynae, Ch. kibonotensis, Ch. milloti, Ch. tatianae; Abrolophus aitapensis

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Malawi (Fain & Elsen, 1987; Cokendolpher, 1993) – Erythraeidae: Leptus bicristatus, L. jocquei, L. polythrix, L. puylaerti Morocco (Haitlinger, 1990c) – Erythraeidae: Leptus mogadoranus, L. siculus Mozambique (Haitlinger, 1990a) – Erythraeidae: Leptus ogazulacus, Charletonia areolata, Neoabrolophus sp. Namibia (Haitlinger, 1990a) – Erythraeidae: Leptus pasopaicus Niger (Southcott, 1994b) – Erythraeidae: Abalakeus chekei Nigeria (Beron, 2000, Beron, in prep.; Buitendijk, 1945; Oudemans, 1910) – Smarididae: Fessonia nigeriae; Erythraeidae: Leptus sp., Caeculisoma haussa, Cecidopus nigeriae, Charletonia brunni, Ch. kibonotensis; Neoabrolophus sp. Rwanda (Fain, 1996; Fain & Elsen, 1987; Fain & Jocqué, 1996) – Erythraeidae:Opserythraeus hoffmannae, Leptus aureliani, L. cavernicola, L. rwandae Sainte-Helene (Feider & Chioreanu, 1977) – Erythraeidae: Charletonia cooremani, Ch. sanctaehelenae, Balaustium southcotti Sierra Leone (Berlese, 1910) – Erythraeidae: Charletonia spinosa South Africa (Berlese, 1910; Halliday, 2001; Haitlinger [in print], Lawrence, 1937, 1940, 1944; Meyer & Ryke, 1959; Meyer et al., 1973; Meyer & Ueckermann, 1989) – Smarididae: Kraussiana brevicristata; Erythraeidae: Kakamasia pilosa, K. polygona, Eatoniana namaquaensis, Leptus atticolus, L. intermedius, L. assagasicus, Abrolophus spiculosus, Charletonia areolata, Ch. lawrencei, Cecidopus chubbi, C. mitchelli, Caeculisoma carmenae, Balaustium bipilum, B. graminum, B. medicagoense, B. vignae; Genera, recorded by Meyer et al., 1973 as living in South Africa, but without precise data: Andrevella, Erythraeus (= Bochartia and Parerythraeus), Hauptmannia, Sphaerolophus (= Charletonia), Erythrites, Mypongia, Grandjeanella, Clavismaris, Clipeosoma, Smaris; Inc. sedes: “Erythraeus munsteri”, “Smaridia heteropoda“ Sudan (André, 1962; Oudemans, 1911b) – Erythraeidae: Erythraeus nigritiensis, Leptus sudanensis Swaziland (Lawrence, 1940) – Erythraeidae: Charletonia swaziana Tanzania (Beron, in prep.; Evans, 1953; Fain & Elsen, 1987; Haitlinger, 1990a; Trägårdh, 1908) – Erythraeidae: Erythraeus nasalis, Leptus leleupi, L. siculiformis, L. soddagus, Charletonia areolata, Ch. kibonotensis, Ch. salti, Neoabrolophus bisetis, Balaustium angustum; Inc. sedes: “Erythraeus mucronatus, Erythraeus meruensis” Togo (Haitlinger, 2006g) – Erythraeidae: Lomeustium togoensis Tunisia (Haitlinger, 1998; Pavesi, 1884; Trägårdh, 1904a) – Erythraeidae: Leptus hammameticus, Balaustium murorum; Inc. sedis: “Rhyncholophus ? plumipes”, “Rhyncholophus pallidipes” Zambia (Haitlinger, 1990a) – Erythraeidae: Leptus aggoratus Zimbabwe (Welbourn, 1983) – Erythraeidae: Leptus sp. “East Africa” (Berlese, 1916) – Erythraeidae: Inc. sedes: “ Erythraeus alluaudi, Leptus humeratus, Abrolophus albidoinfumatus, A. spectabilis, Smaridia pulcherrima”


203

ASIA Afghanistan (Cooreman, 1960) – Erythraeidae: Balaustium afghanicum Bhutan (Beron, in prep.) – Smarididae: Kraussiana sp.; Erythraeidae: Caeculisoma septentrionale Burma (Myanmar)(Haitlinger, 2004b; Southcott, 1957, 1963) – Smarididae: Fessonia australiensis, Erythraeidae: Leptus astrubali, Myrmicotrombium sp., Charletonia volzi Cambodia (Haitlinger, 2004b) – Erythraeidae: Erythraeus kacperi, Leptus laviniacus, L. astrubali, Charletonia shiroyama, Ch. volzi China (Goosmann, 1925; Kishida, 1939; Trägårdh, 1937; Zhang, 2001a, 2001b; Zhang, Li & Xian, 1997; Zhang, Zhang & Lin, 2000; Zheng Bo-yi, 1996a, 1996b, 2001, 2002a, 2002b, 2002c, 2003; Haitlinger, 2006a; Beron, in prep.) – Smarididae: Fessonia australiensis; Erythraeidae: Abalakeus bambusae, Erythraeus chinensis, E. jacoti, E. yangshounicus, E. zhangi, Claverythraeus mongolicus, Podosmaridia longipes, Leptus brachypodos, L. coloanensis, L. dolichopodos, L. guilinicus, L. hupingshanicus, L. siemsseni, L. shimensis, L. sulciscutus, L. zhejiangensis, L. zhutingensis, Abrolophus aitapensis [Macao], A. unimiri, Caeculisoma hunanica, Charletonia hunanensis, Neophanolophus chinensis, N. liui, Hauptmannia yanlingica, Balaustium innocentae Philippines (Cokendolpher, 1993) – Erythraeidae: Leptus sp. Taiwan (Kishida, 1935; Tsai & Chow, 1988; Tseng, Yang & Pan, 1976; Yen, Tsai & Chow, 1991; Chow Y.S. 1996) – Erythraeidae: Erythraeus plumatus, Callidosoma matsumuratettix, Charletonia taiwanensis Cyprus (Cooreman, 1946; Haitlinger, 1993; Zhang, 1995a) – Smarididae: Smaris pinus; Erythraeidae: Leptus siculus, Abrolophus halberti, Phanolophus oedipodarum India (Agarwal & Dhiman, 1989; Beron, in prep.; Chinniah & Mohanasundaram, 1999; Dhiman, 1987; Dhiman & Sharma, 1995; Dhiman & Singh, 1988; Gupta, 1985, 1992; Haitlinger, 1998, 2002c; Jadhava & Varma, 1987; Khot, 1963a, 1963b, 1964, 1965; Mohanasundaram & Parameswaran, 1989; Oudemans, 1941; Saboori & Arbabi, 2003; Southcott, 1966; Zhang, 1995b, 2003; Ramaraju & Mohanasundaram, 1998; Vishnupriya & Mohanasundaram, 1988) – Smarididae: Fessonia assmuthi, F. australiensis; Neophanolophus shibai; Erythraeidae: Erythraeus delhiensis, E. elegans, E. indicus, E. orientalis, E. plumosus, E. serratociliatus, Leptus aeolopae, L. alkmenae, L. chrotogonae, L. giganticus, L. hospeticus, L. indicus, L. laviniacus, L. ovatus, L. oxyae, L. poonaensis, L. singhi, L. villosus, Neoabrolophus brevisetorum, N. longisetorum, Cecidopus shyamae, Charletonia aranea, Ch. delhiensis, Ch. gigas, Ch. keralicus, Ch. minuta, Ch. sureshi, Abrolophus batotensis, A. delhiensis, A. ripicola, Hauptmannis sp.; Pollux kovalamicus, Microsmarialla coniferana, Microsmaris pinnata, M. sparsepinnata Indonesia (Berlese, 1904, 1910; Haitlinger, 1990, 1998, 2006c; Oudemans, 1910, 1928; Vitzthum, 1926) – Erythraeidae: Leptus admeti [Sumatra], L. balicus [Bali], L. gagrellae [Java], L. hirtipes [Java], L. managarus [Java], L. terebrans [Sumatra], L. ubudicus [Bali], Caeculisoma argus [Sumatra], C. infernale [Sumatra], C. sulcatum [Krakatau], Charletonia arborum [Java], Ch. cursor [Java], Ch. lombokensis [Lombok], Ch. nova

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[Buru], Ch. shiroyama [Lingga, Bali, Lombok], Ch. similis [Sumatra], Ch. volzi [Sumatra, Java, Lingga, Bali], Carastrum sanurensis [Bali]; Abrolophus aitapensis [Bali] Iraq (Shamsuddin & Mohammad, 1979) – Leptus sp. Iran (Babolmorad & Saboori, 2000; Gabryś, 1989; Zhang & Goldarazena (1996), Haitlinger & Saboori, 1996, 2007; Barimani, Saboori & Ghasemi, 2002; Karimi Iravanlou & Saboori, 2000, 2001; Karimi Iravanlou, Kamali & Talebi, 2002; Khanjani & Ueckermann, 2005; Saboori, 2000a, 2000b, 2002a, 2002b; Saboori & Akrami, 2001; Saboori & Atamehr, 1999, 2000; Saboori & Babolmorad, 2000; Saboori, Goldarazena & Khajeali, 2004; Saboori & Nowzari, 2001; Saboori & Ostovan, 2000, Saboori & Lachinani, 2003; Saboori, Nowzari & Bagheri-Zenouz, 2004; Saboori & Hajiqanbar, 2005; Saboori, Hajiqanbar & Kamali, 2007) – Erythraeidae: Abalakeus lorestanicus; Erythraeus akbariani, E. garmsaricus, E. hypertrichotus, E. mirjavehi, E. sabrinae, E. shojaii, E. tehranicus, E. (Zaracarus) iranicus, E. (Z.) kharrazii, E. (Z.) kurdistanensis, E. (Z.) longipedus, E. (Z.) rajabii, E. (Z.) ueckermanni; Leptus fathipeuri, L. esmailii, L. eslamizadehi, L. kamalii, L. zhangi; Charletonia ahwazensis, Ch. damavandica, Ch. krendowskyi, Ch. mehranensis, Ch. nazeleae, Ch. saboorii, Abrolophus iraninejadi, A. khanjanii, Andrevella persiangulfia, Hauptmannia iranica, H. ostovani; Grandjeanella kamalii, G. bella, Balaustium zhangi, Bursaustium norbakhshi Israel (André, 1951; Shulov, 1957; Haitlinger, 1994c) – Smarididae: Hirstiosoma sp.; Erythraeidae: Erythraeus phalangoides, E. [Bochartia] kuyperi, Leptus tammuzi, Charletonia (=Sphaerolophus) sp. Japan (Baker & Selden, 1997; Ishii, 1953, 1954; Hara & Hanada, 1962; Kawashima, 1958, 1961a, 1961b; Karsch, 1881; Kato & Kitahara, 1958; Kishida, 1929a, 1929b, 1930; Kitahara & Takara, 1964; Morikawa, 1963; Shiba, 1969, 1981; Yaita, Kato & Toriyama, 1961; Yano & Ehara, 1982; Zheng & Takafuji, 1998) – Erythraeidae: Erythraeus phalangoides nipponicus; Leptus asahinai, L. gifuensis, L. hidakai, L. japonicus, L. kuroshimaensis, L. kyushuensis, L. repens, L. saigusai, L. trimaculatus; Charletonia ishiii, Ch. miyazakii, Ch. ojimai, Ch. okinawaensis [Okinawa], Ch. shiroyama [Okinawa], Ch. southcotti; Hauptmannia miyatakei; Abrolophus cuspidatus [inc. sed.: Rhyncholophus humeralis Karsch] Kazakhstan (Haitlinger, 1998) – Erythraeidae: Leptus auliacus Kirghizstan (Haitlinger, 1998) – Erythraeidae: Leptus meloidarum Korea (North)(Beron, in prep.) – Erythraeidae: Erythraeus sp. Laos (Haitlinger, 1999) – Erythraeidae: Leptus augusti, L. holgeri, L. ilzae Lebanon – Erythraeidae: Abrolophus halberti Malaysia (Haitlinger, 1999b, Shiba, 1976) – Smarididae: Kraussiana mitsukoae, Neophanolophus morii, Clipeosoma furtadoi; Erythraeidae: Leptus acuminatus, L. agenori, L. calidus, L. cameronensis, L. chibai, L. hozumii, L. yoshimii; Charletonia shiroyama; Neoabrolophus hirsutus; Hauptmannia trifarius Mongolia (Haitlinger, 1998) – Erythraeidae: Leptus batoricus Nepal (Beron, in prep.) – Smarididae: Kraussiana sp.; Erythraeidae: Erythraeus sp., Leptus sp., Charletonia sp.


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New Guinea (Papua New Guinea and West New Guinea)(Beron, 2002, in prep.; Canestrini G., 1897, 1898; Gunther, 1941; Haitlinger, 1990b; Kramer, 1898; Oudemans, 1905a, 1905c, 1906b; Southcott, 1948, 1972, 1984, 1993, 1999) – Smarididae: Trichosmaris papuana, Clipeosoma copiolarum; Erythraeidae: Leptus boggohoranus, L. debeauforti, L. draco, L. calcar, L. cheesmanae, L. lorarius, L. scutellatus, L. triacanthus, Callidosoma rostratum, Caeculisoma chapmani, C. claviger, C. darwini, C. mouldsi, C.novaeirlandiae, C. plantacionis,C. sulcatum, ? C. tuberculatum, Charletonia falcata, Ch. volzi, Abrolophus aitapensis, A. novaeguinensis [“Rhyncholophus poriferus” – nom. dubium] Pakistan (Ahsan & Anwarullah, 1970; Anwarullah & Ahsan, 1970) – Erythraeidae: Cecidopus qadrii, Leptus karachiensis Saudi Arabia (Haitlinger, 1994c) – Erythraeidae: Leptus tammuzi Singapore (Baker & Selden, 1997) – Erythraeidae: Leptus hidakai Sri Lanka (Haitlinger, 2001a, 2002a; Karsch, 1879; Southcott, 1988c) – Erythraeidae: Leptus laviniacus, L. tetrigius, Charletonia lankensis, Cecidopus diversipes, Dambullaeus piae Syria (Haitlinger, 1994c) – Erythraeidae: Leptus horiacus, L. tammuzi Thailand (Haitlinger, 1999a; Southcott, 1994) – Erythraeidae: Leptus addari, L. agenori, L. astrubali, L. phuketicus, Abrolophus mortenseni Turkmenia (Haitlinger, 1990b) – Erythraeidae: Leptus guus; Inc. sedis: “Eatoniana plumifer forma transcaucasica” Vietnam (Haitlinger, 1986, 1987b) – Erythraeidae: Charletonia danangensis, Ch. jolantae, Ch. huensis, Abrolophus aitapensis NORTH AMERICA Bermuda Is. (Banks, 1903) – Erythraeidae: “Rhyncholophus sp.” Canada (Banks, 1916; Cadogan & Laing, 1977, 1981; Smiley, 1968; Southcott, 1992a) – Erythraeidae: Leptus albertensis, Charletonia canadensis, Balaustium putmani; Inc. sedis:”Rhyncholophus gracilipes” Mexico (Banks, 1898, 1913; Beron, 1996, in prep.; Cokendolpher, 1993; Crespo & Soto, 1979; Haitlinger, 2000e; Hoffmann & Mendez, 1973; Southcott, 1963; Vitzthum, 1933; Wharton, 1938) – Smarididae: Calorema azteka, Smaris boneti, S. grandjeani christensoni, S. lanceolata, Fessonia australiensis, F. lacrimosa, F. lappacea, F. scobina, Clavismaris conifera, Hirstiosoma bolivari, Trichosmaris d. dispar; Erythraeidae: Erythraeus sp., Augustsonella southcotti, Erythraxus bisetosus, Tepoztlana sandovali, Paraphanolophus halffteri, Leptus filipinae, L. onnae; Balaustium minodorae, Palenqustium leanderi; Inc. sedes: “Erythraeus tolteka, Rhyncolophus pedatus, Rh. echinatus” U.S.A. (Adler & Brown, 1986; Augustson, 1940; Banks, 1894, 1895, 1896, 1901, 1902a, 1902b, 1904a, 1904b, 1907, 1912, 1916a, 1947; Beron, in prep.; Childers & Rock, 1980; Cockerell, 1907; Cokendolpher, 1993; Cokendolpher, Horner & Jennings, 1979; Ebeling, 1934; Ewing, 1909, 1910, 1917; Fain, Gummer & Whitaker, 1987; Grogan, 1982; Hall, 1912; Hartzell, 1918; McAloon & Durden, 2000; McDaniel & Bolen, 1981; MacKay, Grimsley & Cokendolpher, 1992; Miller, 1925; Reillo, 1989; Resh & Haag, 1974; Say, 1821; Sharma,

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Drooz & Treat, 1983; Sharma, Farrier & Drooz, 1983; Sorkin, 1982; Southcott, 1963, 1966, 1972, 1989b, 1991a, 1992a; Treat, 1975, 1978, 1979, 1985, Welbourn & Jennings. 1991; Welbourn & Young, 1987; Yao Hongren, Snider & Snider, 2000; Young & Welbourn, 1987, 1988) – Smarididae: Smaris mamillatus, S. grandjeani christensoni, Fessonia lappacea, Clavismaris cybaea, Hirstiosoma bolivari, Trichosmaris d. dispar, T. jacoti; Erythraeidae: Erythraeus exilipes, E. simplex, Augustsonella tuberculata, Lasioerythraeus johnstoni, L. shirleyanneae, L. whitcombi, Leptus bakeri, L. californicus, L. calix, L. cercopius, L. clarki, L. coreophilus, L. droozi, L. floridanus, L. ghiradellae,L. hitchcocki, L. ignotus, L. ignotus danelli, L. indianensis, L. maculatus, L. nearcticus, L. ruginus, L. sayi, L. texanus, L. treati, L. welbourni, Callidosoma apollo, C. daviesi, C. droozi, C. galtoni, C. huachuca, C. insulare, C. metzi, C. quinteri, C. sorkini, C. treati, Charletonia buforania, Ch. nishidai, Ch. paolii, Ch. perthensis, Ch. wrighti, Abrolophus welbourni, Balaustium kendalli, B. dowelli; Inc. sedes: “Smaris australis, Smaris longilinealis; Erythraeus spinatus, E. carolinus, E. posticatus, E. tonsus, E. arvensis, Rhyncholophus angustipes Banks, 1907, nom. nov. pro Rh. gracilipes Banks, 1905, Rh. brevitarsus, Rh. cavernarum, Rh. cinctipes, Rh. elongatus, Rhyncholophus erythreus, R. floridanus, Rh. gracilipes, Rh. leprosus, Rh. longipes, Rh. longitarsus, Rh. maculatus, Rh. moestus, Rh. montanus, Rh. parvipollicus, Rh. parvisetosus, Rh. parvus, Rh. pilosus, Rh. pollicaris, Rh. punctatus, Rh. quadrioculus, Rh. robustus, Rh. roseus, Rh. texanus, Rh. tridentifer” SOUTH AND CENTRAL AMERICA Argentina (Berlese, 1888, 1916b; Haitlinger, 2004e, 2005a) – Erythraeidae: Leptus calvescens, Caeculisoma tuberculatum, Callidosoma leodegari, Abrolophus incanescens, A. magnificus, A. setipapillus, A. trinotatus, Balaustium brunoni; Inc. sedes: “Rhyncholophus impectus, Rh. pedestris, Rh. quinquemaculatus, Rh. (Apectolophus) simplex “ Belize (Haitlinger, 2000a; Smiley, 1968) – Erythraeidae: Paraphanolophus metcalfei, Leptus filipinae Bolivia (Beron, in prep.; Haitlinger, 2000c) – Erythraeidae: Erythraeus bolivianus, Lasioerythraeus welbourni, Balaustium bolivianum, B. medardi Brazil (Berlese, 1888; Fain & Santiago-Blay, 1993; Haitlinger, 1987e, 1991, 2004a, 2004e, 2004d, 2005a; Oudemans, 1905b, Rosa & Flechtmann, 1980; Southcott, 1972; Treat, 1985; Treat & Flechtmann, 1979; Coracini & Samuels, 2002) – Erythraeidae: Leptus adaminae, L. alberti, L. calvescens, L. cyrili, L. fozicus, L. iguacuicus, L. mariani, L. onnae, L. stieglmayri, L. stolae, Callidosoma leodegari, C. wallaceanum, C. welbourni, Caeculisoma nestori, Charletonia rocciai, Ch. domawiti, Iguatonia barbillae, Momorangia asphaerae, Fozustium paranensis Chile (continent.)(Casaneuva & Angulo, 1995; Cokendolpher, 1993; Gervais, 1849; Leonardi, 1901; Oudemans, 1902, 1911, 1912; Welbourn & Young, 1988) – Smarididae: Smaris nicoleti; Erythraeidae: Leptus lomani, L. schedingi, L. ursyni, L. gracilipes; Inc. sedes: “Erythraeus ursinus, Rhyncholophus (Abrolophus) striglipes, Rhyncholophus (Achorolophus) cursitans, Rhyncholophus gloriosus”, “Rhyncholophus andinus“


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Juan Fernandez Is. (Trägårdh, 1931a) – Erythraeidae: Balaustium obtusum Colombia (Haitlinger, 1991, Losada, 1947) – Erythraeidae: Leptus olafi, L. stefani Costa Rica (Banks, 1913a; Haitlinger, 2000a; Southcott, 1962, 1995a) – Smarididae: Clavismaris cybaea, Surasmaris longirostris; Erythraeidae: Leptus filipinae, L. nikanori; Inc. sedis: “Rhyncholophus stolli” Cuba (Beron, in prep.; Prieto & Bauza, 1999) – Erythraeidae: Leptus sp. Dominican Rep. (Botosaneanu & Bolland, 1997, De Armas & Trueba, 2003; Haitlinger, 2004g) – Erythraeidae: Lasioerythraeus johnstoni; Leptus cabareticus, Leptus sp.; Callidosoma welbourni Ecuador (continent.) Galapagos Is. (Banks, 1924) – Erythraeidae: Erythraeus remotus Falkland Is. (Trägårdh, 1907) – Erythraeidae: Balaustium antarcticum Guatemala (Haitlinger, 2000e; Southcott, 1963, 1989a, 1992; Stoll, 1886-1893; Treat, 1985) – Smarididae: Smaris grandjeani christensoni, Fessonia serrata, Trichosmaris d. dispar, T. dispar dentella; Erythraeidae: Leptus ariel, L. simonettae, Callidosoma guatemalensis, ? Charletonia globigera, Balaustium soydani, Guatuscium biljanae; Inc. sedis: “Rhyncholophus erinaceus” Honduras (Haitlinger, 2000a) – Erythraeidae: Leptus simonettae Panama (Southcott, 1963; Treat, 1985; Zhang, 2001) – Smarididae: Calorema azteka, Smaris g. grandjeani grandjeani, S. lanceolata, S. zeteki; Erythraeidae: Leptus gagzoi, Callidosoma harlani Paraguay (Berlese, 1888) – Erythraeidae: Leptus calvescens, Caeculisoma tuberculatum, C. metzi Peru (Beron, in prep.; Flechtmann, 1980; Haitlinger, 2000c; Schuster & Turk, 1987) – Erythraeidae: Erythraeus ghilarovi, E. peruvianus, E. titikaka, Leptus annikae, L. ariel, L. hringuri, L. maldonadoicus, L. vuki, Charletonia alvedae, Balaustium medardi Surinam (Oudemans, 1910) – Erythraeidae: Leptus oudemansi Trinidad and Tobago (Turk, 1948) – Erythraeidae: Leptus gagzoi Venezuela (Berlese, 1910; Oudemans, 1911, 1912; Haitlinger, 2006d) – Erythraeidae: Leptus olafi, L. sieversi, L. tiranicus, Charletonia politricha OCEANIA Australia (continental)(Banks, 1916b; Berlese, 1910; Fain, 1991; Haitlinger, 1990b; Halliday, 1998, 2001; Greenslade, 1985; Greenslade & Southcott, 1980; Hirst, 1928; James, 1995, 2000, James, O’Malley & Rayner, 1995; Key, 1990, 1991; Koch L., 1865; Lee & Southcott, 1979; Rainbow, 1906; Southcott, 1946a, 1946b, 1948, 1957, 1960, 1961a, 1961b, 1961c, 1966, 1972, 1983, 1988b, 1991, 1993, 1996, 1997, 1999; Tribe, 2000; Womersley, 1934, 1936; Womersley & Southcott, 1941) – Smarididae: Smaris arenicola, S. prominens, Fessonia australiensis, F. taylori, Hirstiosoma scalare; Sphaerotarsus allmani, S. claviger, S. leptopilus, S. monticolus, S. ripicolus, S. womersleyi; Erythraeidae: Erythraeus gregoryi, E. guttatus, E. osmondensis, E. hirsti (= E. pilosus Hirst), E. stuarti, Erythrites reginae,

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E. guttatus, E. urrbrae, E. womersleyi, Erythrellus imbricatus, Erythroides clavatus, E. darwini, E. macdonnelli, E. neoserratus, E. serratus, Rainbowia celeripes, R. imperator (= Bochartia oudemansi), Leptus agrotis, L. alopecurus, L. alvimordax, L. anomalus, [L. antepodianus], L. bankensis, L. barmeedius, L. bathypogonus, L. batjallus, L. baudini, L. belicolus, L. calcar, L. carduus, L. cerambycius, L. charon, L. cheesmanae, L. chelonethus, L. clavatus, L. clelandi, L. comosus, L. cultellus, L. elderi, L. faini, L. fisheri, L. flindersi, L. foliatus, L. fortei, L. francesi, L. gauphalus, L. georgeae, L. grossi, L. halli, L. heleus, L. henseni, L. hitchcocki, L. korematus, L. lighti, L. minno, L. norrisi, L. ornatus, L. orthrius, L. pincheni, L. pistoris, L. puniceus, L. scutellatus, L. smithi, L. spinalatus, L. swani, L. tarranus, L. tindalei, L. titinius, L. torresianus, L. triacanthus, L. truncatus, L. urodaci, L. utheri, L. waldockae, L. warregense, Callidosoma dasypodiae, C. galtoni, C. heatherae, C. ripicolum, C. tindalei, C. womersleyi, Caeculisoma argus argus, C. argus io, C. cooremani, C. darwiniense, C. johnstoni, C. montana, C. mouldsi, C. nasutum, C. sparnoni, Carastrum ferrari, Harpagella moxonae, Charletonia banksi, Ch. barbata, Ch. buforania, Ch. domrowi, Ch. feideri, Ch. froggatti, Ch. goliatha, Ch. grandjeani, Ch. hermanni, Ch. keyi, Ch. oudemansi, Ch. paolii, Ch. perthensis, Ch. shiroyama, Ch. striaticeps, Ch. tragardhi, Ch. vitzthumi, Ch. westraliensis, Abrolophus brevus, A. glauerti, A. insularis, A. littorale, A. newmani, A. ripicola, A. tonsor, Hauptmannia fossor, Pussardia mullewaensis, Grandjeanella charadrae, G. westraliensis, G. pangela, G. shepherdae, Microsmaris hirsti, M. joannae, M. minutus, M. mirandus, Wartookia rebeccae, Mypongia brevipes, Pollux workandae, Balaustium medicagoense, Myrmicotrombium brevicristatum; Inc. sedes: “Rhyncholophus attolus, Rh. retentus” Tasmania (Halliday, 2001; Seeman & Miller, 2002; Southcott, 1993; Womersley & Southcott, 1941) – Smarididae: Hirstiosoma tasmaniensis; Erythraeidae: Leptus monteithi, L. charon, Charletonia feideri, Ch. oudemansi, Balaustium medicagoense Caroline Is. (Southcott, 1961a) – Smarididae: Kraussiana sp. Marquesan Is. (Vitzthum, 1935) – Erythraeidae: Caeculisoma cordipes New Zealand (Forster & Forster, 1973; Luxton, 1989; Southcott, 1972, 1988a; Womersley, 1936; Womersley & Southcott, 1941; Zhang, 2000; Clark, 2004) – Smarididae: Hirstiosoma novaehollandiae; Erythraeidae: Neosmaris novaezealandiae, Taranakia lambi, Erythrites jacksoni, Ramsayella rangitata, Abrolophus zelandicus, Callidosoma tiki, Caeculisoma huxleyi, Momorangia jacksoni, M. vallata, Microsmaris mirandus New Caledonia (Berlese, 1923; Southcott, 1966) – Erythraeidae: Charletonia rageaui; “Erythraeus (Erythrolophus) mollis”(Inc. sedis)


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Hosts of Erythraeoidea Arthropoda Class Arachnida Order Pseudoscorpiones – Leptus chelonethus Order Opiliones – Leptus ignotus, L. meloidarum, L. phalangii, L. stieglmayri Fam. Phalangiidae – Leptus beroni, L. bicristatus, L. gagrellae, L. gyas, L. indianensis, L. jocquei, L. kalaallus, L. nearcticus, L. phuketicus, L. polythrix, L. puylaerti, Charletonia enghoffi Fam. Opilionidae – Leptus hidakai Fam. Cosmetidae – Leptus oudemansi Fam. Gonyleptidae – Leptus lomani Order Scorpiones Fam. Buthidae – Leptus korematus, L. pyrenaeus, L. waldockae Fam. Scorpionidae – Leptus barmeedius, L. baudini, L. carduus, L. hitchcocki, L. korematus, L. pistoris, L. smithi, L. urodaci Fam. Bothriuridae – Leptus charon Order Amblypygi Fam. Phrynidae – Leptus sp. Order Araneae – Charletonia aranea Fam. Araneidae – Charletonia areolata Fam. Attidae – Leptus atticolus Fam. Lycosidae – Leptus gifuensis Fam. Clubionidae – Leptus hidakai Fam. Zodariidae – Leptus rwandae Fam. Theridiidae – Callidosoma miyazakii Fam. Uloboridae – Leptus hidakai Fam. Sparassidae – Leptus charon, L. faini, L. minno Order Acariformes Fam. Erythraeidae – Erythrites womersleyi, Erythroides serratus, Leptus trimaculatus Fam. Trombidiidae s.l. – Charletonia froggatti Class Diplopoda Order Julida Fam. Julidae – Leptus millipedius Class Entognatha Order Collembola – Leptus trimaculatus (= echinopus), L. southcotti Order Thysanura Fam. Machilidae – Erythraeus kresnensis, Leptus machilidis Fam. Meinertellidae – Leptus norrisi (from Allomachilis) Class Insecta – Leptus brachypodos, L. hupingshanicus, L. shimensis, L. sulciscutus

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Order Orthoptera – Leptus agenori, L. balicus, L. bankensis, L. bathypogonus, L. cheesmanae, L. draco, L. holgeri, L. japonicus, L. josifovi, L. maranaensis, L. meloidarum, L. pozzoicus, L. tindalei, L. tiranicus, Charletonia ahwazensis, Ch. alarobiaensis, Ch. austisensis, Ch. dorotae, Ch. krendowskyi, Ch. lombokensis, Ch. shiroyama Acridodea – Charletonia keyi Fam. Acrididae – Abalakeus chekei, Erythrites jacksoni, Ramsayella rangitata, Leptus calcar, L. californicus, L. cultellus, L. dubius, L. foliatus, L. josifovi, L. kamalii, L. laviniacus, L. lighti, L. oxyae, L. pincheni, ?L. rosellae, L. simonettae, Leptus vuki, Caeculisoma cooremani, C. darwiniense, Charletonia areolata, Ch. banksi, Ch. barbata, Ch. berlesiana, Ch. braunsi, Ch. buforania, Ch. damavandica, Ch. feideri, Ch. hermanni, Ch. mehranensis, Ch. nazeleae, Ch. saboorii, Ch. shiroyama, Ch. striaticeps, Ch. swaziana, Ch. venus, Phanolophus oedipodarum Fam. Pamphagidae – Charletonia berlesiana Fam. Pyrgomorphidae – Leptus chrotogonae, Charletonia brunni, Ch. keyi Fam. Pseudophyllidae – Leptus annikae, L. maldonadoicus, Charletonia alvedae Fam. Tetrigidae – Leptus bakeri, L. tetrigius Fam. Tettigoniidae – Leptus gagzoi, L. orthopterarum, Charletonia berlesiana, Ch. bucephalia, Ch. dalegari, Ch. feideri, Ch. barbata (= grandjeani), Abalakeus lorestanicus, Phanolophus oedipodarum Fam. Bradyporidae – Phanolophus oedipodarum Fam. Mecopodidae – Charletonia ishiii, Ch. volzi Order Mantodea Fam. Mantidae – Charletonia krendowskyi Order Phasmodea – Taranakia lambi, Leptus cheesmanae, Charletonia froggatti Fam. Phasmatidae – Leptus draco Order Blattodea Fam. Blaberidae – Leptus flindersi Order Dermaptera Fam. Forficulidae – Leptus mariae Order Odonata – Leptus draco, Charletonia edytae Fam. Libellulidae – Charletonia hunanensis Fam. Megapodagriidae – Charletonia hunanensis Fam. Coenagrionidae – Leptus killingtoni Order Heteroptera – Erythraeus hirsti, E. plumosus, Leptus filipinae, Charletonia cardinalis (= singularis) Fam. Miridae – Lasioerythraeus johnstoni, Charletonia areolata, Ch. blascoi Fam. Tingidae – Erythraeus shojaii Fam. Lygaeidae – Charletonia areolata Fam. Scutelleridae – Leptus esmailii Fam. Pentatomidae – Lasioerythraeus johnstoni, Grandjeanella haitlingeri Fam. Coreidae – Leptus coreophilus, L. josifovi


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Fam. Gelastocoridae – Leptus batjallus Fam. Anthocoridae – Grandjeanella haitlingeri ?Fam. Henicocephalidae – Leptus madagascariensis Order Homoptera – Leptus beroni, L. japonicus, Erythraeus gertrudae Cicadina (Auchenorhyncha) Fam. Delphacidae – Charletonia cardinalis, Paraphanolophus metcalfei, Hauptmannia brevicollis Fam. Cicadidae – Leptus zhutingensis, L. draco, L. kyushuensis, L. mariae, L. torresianus, Caeculisoma mouldsi, Momorangia vallata Fam. Cicadellidae – Neophanolophus shibai, Erythraeus adrastus, E. rajabii, E. berninensis, E. (Zaracarus) aydinicus, Leptus fisheri Fam. Fulgoridae – Leptus admeti Fam. Cercopidae – Leptus cercopius, Charletonia cardinalis Fam. Tibicinidae – Leptus managarus, Caeculisoma mouldsi “cicada” – Leptus clelandi Fam. Jassidae – Polux cristatus Aphidodea – Erythraeus iranicus, E. longipedus, E. southcotti Fam. Aphididae – Erythraeus akbariani, E. ankaraicus, E. kuyperi, E. sabrinae, E. southcotti, E. preciosus, Hauptmannia ostovani Fam. Aleyrodidae – Lasioerythraeus johnstoni Fam. Psyllidae – Leptus elderi Order Coleoptera – Leptus japonicus Fam. Belidae – Leptus belicolus Fam. Cicindelidae – Leptus boggohoranus, L. meloidarum, L. olafi, Andrevella parkeri Fam. Carabidae – Leptus dalicus Fam. Chrysomelidae – Leptus alberti, L. alkmenae, L. zhutingensis, L. ariel, L. galerucae, L. mariae, L. mariani, L. phyllotretae, L. stolae, L. zhutingensis, Charletonia cardinalis Fam. Cleridae – Leptus orthrius Fam. Tenebrionidae – Leptus aggoratus, L. akkus, L. anomalus, L. auliacus, L. batoricus, L. bertoldi, L. gauphalus, L. guus, L. hammameticus, L. heleus, L. horiaticus, L. meloidarum, L. mogadoranus, L. ogazulacus, L. pasopaicus, L. sudanensis, L. tammuzi, L. tarranus, L. trimaculatus (= echinopus), L. ursyni, Charletonia areolata Fam. Melyridae – Charletonia areolata Fam. Byrrhidae – Leptus truncatus Fam. Buprestidae – Leptus mirenae, L. gennadicus Fam. Elateridae – Leptus soddagus Fam. Curculionidae – Leptus augusti, L. faini, L. halli, L. hringuri, L. ilzae, L. jenseni, L. mariae, L. titinius, L. torresianus, L. utheri, Charletonia falcata Fam. Scarabaeidae – Leptus benzaliensis, L. lovaniensis Fam. Cerambycidae – Leptus cerambycius, Charletonia tragardhi, Caeculisoma darwiniense Fam. Lycidae – Leptus cyrili Fam. Lampyridae – Leptus filipinae

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Fam. Pselaphidae – Leptus leleupi Fam. Lagriidae – Leptus mariae Fam. Meloidae – Leptus meloidarum Fam. Anthribidae – Leptus stefani Order Neuroptera – Charletonia iwonae Fam. Myrmeleontidae – Leptus spinulatus Order Mecoptera – Leptus japonicus Order Thysanoptera – Erythraeus osmondensis, Lasioerythraeus johnstoni, Hauptmannia brevicollis, Rudaemania rudaensis Fam. Thripidae – Grandjeanella bella, Hauptmannia sp. Order Psocoptera Fam. Liposcelididae – Smaris prominens Fam. Myopsocidae – Erythraeus hirsti, Erythrites urrbrae Order Hymenoptera – Leptus japonicus Fam. Formicidae – Erythraeus adrastus, Leptus clarki, L. debeauforti Fam. Apidae – Leptus ariel Fam. Sphecidae – Leptus similis Fam. Braconidae – Charletonia areolata Fam. Colletidae – Leptus monteithi Order Diptera – Leptus beroni, L. japonicus, Callidosoma tiki Nematocera – Leptus cavernicolus Fam. Culicidae – Leptus francesi, Callidosoma guatemalensis Fam. Dolichopodidae – Erythraeus lancifer, Charletonia areolata Fam. Drosophilidae – Harpagella moxonae Fam. Tabanidae – Leptus alopecurus, L. alvimordax, L. francesi, L. kuroshimaensis, L. kyushuensis, L. swani Fam. Muscidae – Carastrum ferrari Fam. Asilidae – Leptus bathypogonus, L. fortei, L. tiranicus Fam. Calliphoridae – Leptus charon Fam. Stomoxydidae – Leptus comosus Fam. Ceratopogonidae – Erythraeidae Fam. Bombyliidae – Leptus dubius Fam. Simuliidae – Charletonia krendowskyi Fam. Glossinidae – Leptus benzaliensis, L. carpenteri, L. glossinarum, L. maringensis Order Trichoptera – Leptus sp. Order Lepidoptera – Leptus beroni, L. cooremani, L. ignotus, L. schedingi (from Rhopalocera), Callidosoma welbourni, Charletonia cardinalis, Ch. iwonae Microlepidoptera – Callidosoma daviesi, C. sorkini Fam. Anthelidae – Leptus charon Fam. Tortricidae – Leptus droozi, L. treati, Callidosoma metzi Fam. Tineidae – Callidosoma treati


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Fam. Pyralidae – Leptus saigusai, Callidosoma harlani, C. metzi, C. tiki Fam. Yponomeutidae – Leptus zhangi Fam. Geometridae – Leptus georgeae, L. saigusai, Callidosoma droozi, C. metzi, C. treati, Charletonia cardinalis, Taranakia lambi Fam. Cossidae –Callidosoma metzi Fam. Nymphalidae – Callidosoma tindalei Fam. Satyridae – Leptus slivovi Fam. Lymantriidae – Leptus asahinai, L. slivovi Fam. Lycaenidae – Callidosoma metzi Fam. Zygaenidae – Leptus slivovi Fam. Arctiidae – Leptus slivovi Fam. Notodontidae – Leptus saigusai, Callidosoma treati Fam. Noctuidae – Taranakia lambi, Leptus agrotis, L. albertensis, L. calix, L. charanyca, L. ghiradellae, L. ignotus danelli, L. ruginus, L. saigusai, L. sayi, L. welbourni, L. yoshimii, Callidosoma apollo, C. dasypodiae, C. harlani, C. huachuca, C. insulare, C. quinteri, C. treati, C. wallaceanum, Caeculisoma huxleyi, Momorangia vallata, Charletonia danangensis Chance Hosts Class Mammalia – Charletonia cardinalis Order Chiroptera – Balaustium sp. Order Rodentia – Leptus clethrionomydis

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Index of Erythraeoidea The names, accepted as valid in the present Catalogue, are in bold Abalakeus 59 abrafoicus, Leptus 85 Abrolophus 137 absoloni, Leptus 86 Acariformes 36 Acarus 70, 73, 137, 155 Achorolophus 85 acis, Erythraeus 63 acuminatus, Leptus 86 acutipilis, Erythraeus 63 adaminae, Leptus 86 addari, Leptus 86 admeti, Leptus 86 adpendiculatus, Acarus 63 adpendiculatus, Erythraeus 63 adpendiculatus, Rhyncholophus 63 adrastus, Bochartia 63 adrastus, Erythraeus 63 aeolopae, Leptus 86 aequalipes, Abrolophus 137 aequalipes, Balaustium 137 affine, Balaustium 175 afghanicum, Balaustium 175 afrum, Caeculisoma 128 agatae, Charletonia 152 agenori, Leptus 86 aggoratus, Leptus 87 agilis, Grognieria 87, 189 agilis, Leptus 87 agilis, Rhyncholophus 87, 189 agrotis, Leptus 87 ahwazensis, Charletonia 152 ainae, Grandjeanella 168 aitapensis, Abrolophus 137 aitapensis, Hauptmannia 137 akbariani, Erythraeus 63

akkus, Leptus 87 alarobiaensis, Charletonia 152 albertensis, Leptus 87 alberti, Leptus 87 albicomus, Rhyncholophus 144 albidoinfumatus, Abrolophus 137 aldonae, Leptus 87 alkmenae, Leptus 88 allmani, Sphaerotarsus 56, 57 alluaudi, Erythraeus 186 alopecurus, Leptus 88 alvedae, Charletonia 152 alvimordax, Leptus 88 Amaroptus 127 ampulligera, Hirstiosoma 55 ampulligera, Niemannia 55 ampulligera, Smaridia 55 ampulligera, Smaris 55 andae, Leptus 88 Andrevella 150 angustipes, Rhyncholophus 187 angustum, Balaustium 175 ankaraicus, Erythraeus 64 annikae, Leptus 88 anomalus, Abrolophus 137 anomalus, Balaustium 137 anomalus, Leptus 88 antarcticum, Balaustium 175 antarcticum, Belaustium 175 antarcticus, Erythraeus 175 antepodianus, Leptus 79 antipodianus, Leptus 79 aokii, Erythraeus 64 Apectolophus 62 Apobolostigmata 48 apollo, Callidosoma 132


appendiculatus, Erythraeus 63 aranea, Charletonia 153 araneii, Leptus 190 araneipes, Balaustium 176 araneoides, Achorolophus 176 araneoides, Balaustium 176 araneoides, Leptus 88 araneoides, Rhyncholophus 176 arborum, Charletonia 153 arborum, Sphaerolophus 153 arenicola, Erythraeus 187 arenicola, Rhyncholophus 187 arenicola, Smaris 52 areolata, Charletonia 153 areolatus, Erythraeus 153 argasoides, Platylophus 189 argus, Caeculisoma 128 ariel, Leptus 88 arlettae, Charletonia 153 artemisiae, Abrolophus 138 artemisiae, Acarus 138 artemisiae, Balaustium 138 artemisiae, Smaridia 138 arvensis, Abrolophus 138 arvensis, Erythraeus 138 arvensis, Legneria 138 Arythaena (fossil) 192 asahinai, Leptus 89 asphaerae, Momorangia 137 assagasicus, Leptus 89 assimile, Trombidium 190 assimilis, Erythraeus 190 assmuthi, Fessonia 48 assmuthi, Veithia 48 astrubali, Leptus 89 Atomus 85 atticolus, Leptus 89 attolus, Rhyncholophus 187 augusti, Leptus 89 Augustsonella 59 auliacus, Leptus 89

aureliani, Leptus 90 austisensis, Charletonia 153 australiensis, Fessonia 49 australis, Smaris 186 aydinicus, Erythraeus 76 azteka, Calorema 48 baardi, Hauptmannia 170 baderi, Abrolophus 138 bakeri, Leptus 90 Balaustiidae 175 Balaustiinae 175 Balaustioides 170 Balaustium 175 balicus, Leptus 90 baltiensis, Moldoustium 182 bambusae, Abalakeus 59 bankensis, Leptus 90 banksi, Charletonia 153 banksi, Erythraeus 187 barbata, Charletonia 154 barbillae, Iguatonia 173 barloventensis, Balaustium 176 barmeedius, Leptus 90 basiliense, Balaustium 138 basiliensis, Abrolophus 138 basumtwiensis, Abrolophus 138 bathypogonus, Leptus 90 batjallus, Leptus 90 batoricus, Leptus 91 batotensis, Abrolophus 138 baudini, Leptus 91 Belaustium 175 belicolus, Leptus 91 bella, Grandjeanella 168 beninensis, Charletonia 154 benoni, Abrolophus 139 benoni, Hauptmannia 139 benzaliensis, Leptus 91 berlesei, Leptus 91 berlesiana, Charletonia 154 berlesianum, Balaustium 154

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berninensis, Erythraeus 64 beroni, Leptus 91 bertoldi, Leptus 92 bicolor, Acarus 190 bicristatus, Leptus 92 biljanae, Guatustium 181 bipartita, Smaris 52 bipilum, Balaustium 176 biscutatus, Smaris 194 bisetis, Balaustium 173 bisetis, Neoabrolophus 173 bisetosa, Erythraeus 78 bisetosus, Erythraxus 78 blascoi, Charletonia 154 boccettii, Achorolophus 187 boccettii, ? Leptus 187 Bochartia 62 Bockartia 62 boggohoranus, Leptus 92 bogoriacus, Leptus 92 bohdani, Abrolophus 139 bohdani, Hauptmannia 139 bolivari, Hirstiosoma 56 bolivianum, Balaustium 176 bolivianus, Erythraeus 64 boneti, Smaris 52 borceanum, Neobalaustium 184 brachiosus, Achorolophus 187 brachypodos, Leptus 92 braunsi, Charletonia 154 braunsi, Erythraeus 154 brevicollis, Hauptmannia 170 brevicristata, Fessonia 50 brevicristata, Kraussiana 50 brevicristatum, Myrmicotrombium 185 brevipalpe, Balaustium 139 brevipalpis, Abrolophus 139 brevipes, Mypongia 183 brevisetorum, Neoabrolophus 173 brevitarsus, Rhyncholophus 187 brevum, Balaustium 139

brevum, Belaustium 139 brevus, Abrolophus 139 brunoni, Balaustium 176 brunni, Balaustium 155 brunni, Charletonia 154 brunni, Erythraeus 154 bucephalia, Charletonia 155 budapestensis, Erythraeus 76 buforania, Charletonia 155 buforanius, Erythraeus 155 bulgariense, Balaustium 176 bulgaromontanus, Erythraeus 64 Bursaustium 181 cabareticus, Leptus 92 Caeculiosomateae 189 Caeculiosoma 130 Caeculisoma 128 calcar, Leptus 93 calidus, Leptus 93 californicus, Leptus 93 calix, Leptus 93 Callidosoma 131 Callidosomatinae 128 Callidosomatini 128 Callidosominae 128 Callidosomini 128 callitricha, Oecosmaris 51 Calorema 48 calvatus, Leptus 93, 106 calvescens, Abrolophus 93 calvescens, Achorolophus 93 calvescens, Leptus 93 calvescens, Rhyncholophus 93 cameronensis, Leptus 93 campestri, Erythraeus 64 canadensis, Charletonia 155 canadensis, Sphaerolophus 155 canonicalis, Rhyncholophus 187 capillatus, Erythraeus 64 Carastrum 135 cardinalis, Acarus 155


cardinalis, Charletonia 155 cardinalis, Rhyncholophus 155 cardinalis, Sphaerolophus 155 carduus, Leptus 94 carmenae, Caeculisoma 128 carolinus, Erythraeus 186 carpenteri, Leptus 94 caspersi, Neobalaustium 184 cavannae, Cavannea 156 cavannae, Charletonia 156 cavannae, Rhyncholophus 156 cavannae, Sphaerolophus (Cavannea) 156 Cavannea 152 cavernarum, Rhyncholophus 187 cavernicola, Leptus 94 Cecidopus 151 celeripes, Erythraeus 84 celeripes, Rainbowia 84 celeripes, Rhyncholophus 84 cerambycius, Leptus 94 cercopius, Leptus 94 chapmani, Caeculisoma 131 charadrae, Grandjeanella 168 charanyca, Leptus 94 Charletonia 152 Charletoniini 137 charon, Leptus 94 cheesmanae, Leptus 95 chekei, Abalakeus 59 chelonchus, Leptus 95 chelonethus, Leptus 95 chibai, Leptus 95 chinensis, Bochartia 64 chinensis, Erythraeus 64 chinensis, Neophanolophus 50 christensoni, Smaris grandjeani 52 chrotogonae, Leptus 95 chubbi, Cecidopus 151 chubbi, Sphaerolophus (Eupodolophus) 151 cilissa, Cavannea 156 cilissa Charletonia 156

cilissa, Sphaerolophus (Cavannea) 156 cinctipes, Erythraeus 187 cinctipes, Rhincholophus 187 cinereum, Trombidium 65 cinereus, Erythraeus 65 cinereus, Rhyncholophus 65 cinereus, Rhyncolophus 65 clarki, Leptus 95 clavatus, Erythroides 80 clavatus, Leptus 95 Claverythraeus 59 claviger, Caeculisoma 129 claviger, Erythraeus (Ptilolophus) 62, 189 claviger, Sphaerotarsus 57 clavigerum, Caeculisoma 129 Clavismaris 54 clelandi, Leptus 96 clethrionomydis, Leptus 96 Clipeosoma 55 Cloquetia 189 coccineus, Leptus 96 coccineus, Pediculus 96 Coerythrolophus (lapsus calami) 189 coloanensis, Leptus 96 communis, Rhyncholophus 72 comosus, Leptus 96 complanatus, Achorolophus 139 complanatus, Abrolophus 139 conifera, Clavismaris 54 coniferana, Microsmarialla 182 cooperi, Smaris 52 cooremani, Caeculisoma 129 cooremani, Charletonia 156 cooremani, Leptus 96 cooremani, Sphaerolophus (Cavannea) 156 copiolarum, Clipeosoma 55 corcyreus, Sphaerolophus (Cavannea) 156 cordipes, Caeculisoma 129 coreophilus, Leptus 96 Corerythrolophus 189 crassitarsum, Balaustium 139

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crassitarsus, Abrolophus 139 crimensis, Abrolophus 139 cristatum, Balaustium 184 cristatum, Belaustium 184 cristatus, Microsmaris 184 cristatus, Pollux 184 crocatus, Balaustium 65 crocatus, Erythraeus 65 crocatus, Rhyncholophus 65 cuglierensis, Charletonia 156 cultellus, Leptus 97 cursor, Charletonia 156 cursor, Sphaerolophus 156 Curteria 60 curticristata, Curteria 60 curticristata, Morieria 60 curtipapillum, Abrolophus 140 curtipapillum, Balaustium 140 curtipapillus, Abrolophus 140 curtipes, Leptus 97 cuspidatus, Abrolophus 140 cybaea, Clavismaris 54 cyrili, Leptus 97 dalegori, Charletonia 156 dalicus, Leptus 97 damavandica, Charletonia 157 Dambullaeus 135 danangensis, Charletonia 157 danelli, Leptus ignotus 105 darwini, Erythroides 80 darwiniense, Caeculisoma 129 dasypodiae, Callidosoma 132 dasypodiae, Erythraeus 132 daviesi, Callidosoma 132 debeauforti, Achorolophus 97 debeauforti, Erythraeus 97 debeauforti, Leptus 97 decalvatus, Erythraeus 65 degeerii, Rhyncolophus 71 delhiensis, Abrolophus 140 delhiensis, Charletonia 157

delhiensis, Erythraeus 65 delhiensis, Paraerythraeus 65 delhiensis, Sphaerolophus 157 densipapillum, Abrolophus 140 densipapillum, Balaustium 140 densipapillus, Abrolophus 140 dentella, Trichosmaris dispar 58 depilata, Samouellea 190 depilata, Smaridia 52, 190 depilata, Smaris 190 didonae, Erythraeus 76 dinekaicus, Leptus 97 Discolopheae 189 Discolophus 137 dispar dentella, Trichosmaris 58 dispar dispar, Trichosmaris 57 diversipes, Cecidopus 151 diversus, Leptus 98, 101 dolichopodos, Leptus 98 domawiti, Charletonia 157 domrowi, Charletonia 165 dorotae, Charletonia 157 dowelli, Balaustium 177 draco, Leptus 98 droozi, Callidosoma 132 droozi, Leptus 98 dubiosus, Erythraeus 71 dubius, Achorolophus 98 dubius, Leptus 98 Dugesia 51 Dugesii, Erythraeus 75 dugesii, Rhyncholophus 75 Dupreia 189 Eatonia 61 Eatoniana 61 echinatus, Rhyncolophus 187 echinopus, Leptus 124 edytae, Charletonia 157 eframi, Italustium 182 elderi, Leptus 98 electoralis, Erythraeus 65


electoralis, Rhyncholophus 65 elegans, Erythraeus 65 elegans, Paraerythraeus 65 eleonorae, Erythraeus 76 elminus, Leptus 99 elongatus, Rhyncholophus 187 elwirae, Erythraeus 66 engadinense, Balaustium 140 engadinensis, Abrolophus 140 enghoffi, Charletonia 157 epigaeus, Achorolophus 140 epigaeus, Abrolophus 140 episcopale, Erythraeus 60 episcopalis, Balaustium 60 episcopalis, Curteria 60 episcopalis, Rhyncholophus 60 erinaceus, Rhyncholophus 187 ernesti, Curteria 60 ernesti, Pedroerythraeus 60 errabundus, Leptus 99 Erythraeidae 58 Erythraeinae 58 Erythraeoidea 48 Erythraeus 62 erythreus, Rhyncholophus 187 Erythraxus 78 Erythrellus 78 Erythrites 79 Erythroides 80 eslamizadehi, Leptus 99 esmailii, Leptus 99 eucomus, Erythraeus 186 Euplatylophus 189 eupodes, Abrolophus nemorum var. 99 eupodes, Leptus 99 Eupodolophus 151 eusisyrum, Erythrombium 189 evansi, Abrolophus 140 evansi, Balaustium 140 exilipes, Erythraeus 66 fabiolae, Erythraeus (Zaracarus) 76

fageli, Curteria 61 fageli, Morieria 61 faini, Leptus 99 falcata, Charletonia 158 falconeri, Abrolophus 141 falconerii, Achorolophus 141 Fallopia 189 fathipeuri, Leptus 99 feideri, Charletonia 158 ferrari, Carastrum 135 Fessonia 48 Fessoniella 151 Fessoniinae 48 filipes, Rhincholophus [sic] 187 filipinae, Leptus 100 fisheri, Leptus 100 flavopictus, Erythraeus 71 flavus, Erythraeus 60 flindersi, Leptus 100 florale, Balaustium 177 floridanus, Leptus 100 floridanus, Rhyncholophus 100 floridanus, Rhyncolophus 100 foliatus, Leptus 100 Forania 81 fortei, Leptus 100 fossor, Hauptmannia 170 foveolatus, Erythraeus (fossil) 191 foveolatus, Rhyncholophus 191 fozicus, Leptus 100 Fozustium 181 francesi, Leptus 101 frisius, Abrolophus 141 froggatti, Erythraeus 158 froggatti,Charletonia 158 furibundus, Leptus 101 furtadoi, Clipeosoma 55 gabini, Momorangia 136 gabrysi, Leptus 111 gagrellae, Achorolophus 101 gagrellae, Leptus 101

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gagzoi, Achorolophus 101 gagzoi, Leptus 101 galerucae, Leptus 101 galtoni, Callidosoma 132 garmsaricus, Erythraeus 66 gaspari, Bursaustium 181 gauphalus,Leptus 101 gennadicus, Leptus 101 georgeae, Leptus 102 georgei, Eatoniana 62 Gerites 79 germanicus, Erythraeus 66 gertrudae, Erythraeus 66 ghanensis, Charletonia 158 ghilarovi, Erythraeus 66 ghiradellae, Leptus 102 gifuensis,Leptus 102 giganticus, Leptus 102 gigas, Charletonia 158 gigas, Sphaerolophus (Cavannea) 158 glabrisetosus, Erythraeus 66 glacialis, Hirstiosoma 56 glacialis, Smaris 56 glauerti, Abrolophus 141 glauerti, Balaustium 141 glauerti, Belaustium 141 glifadaensis, Charletonia 158 globiger, Achorolophus 159 globiger, Rhyncholophus 158 globiger, Sphaerolophus 159 globigera, Charletonia 158 globigera, Rhyncholophus (Ritteria) 159 globigerum, Belaustium 158 globigerum, Sphaerolophus 159 gloriosus, Rhyncholophus 187 glossinarum, Leptus 102 goannae, Microsmaris 183 goliatha, Charletonia 159 goliathus, Sphaerolophus 159 gracilenta, Hauptmannia 170 gracilipes, Abrolophus 102

gracilipes, Achorolophus 108, 115 gracilipes, Leptus 102 gracilipes, Rhyncholophus 71 graeca, Curteria 61 Graecotrombium 185 graminum, Balaustium 177 granatae, Smaris 186 Grandjeanella 168 grandjeani christensoni, Smaris 52 grandjeani grandjeani, Smaris 52 grandjeani, Charletonia 154 grandjeani, Leuchsia 52 grandpopensis, Charletonia 159 gregoryi, Erythraeus 67 gregoryi, Parerythraeus 67 groenlandicus, Erythraeus 102 groenlandicus, Leptus 102 Grognieria 189 grossi, Leptus 103 guatemalensis, Callidosoma 132 guilinicus, Leptus 103 guttatus, Erythraeus 79 guttatus, Erythrites (Gerites) 79 guus, Leptus 103 gyas, Leptus 103 haitlingeri, Grandjeanella 169 halberti, Abrolophus 141 halberti, Balaustium 141 halffteri, Paraphanolophus 83 halleri, Rhyncholophus 62 halli, Leptus 103 hammameticus, Leptus 103 handschini, Abrolophus 141 handschini, Balaustium 141 harlani, Callidosoma 132 Harpagella 135 harrisonii, Belaustium 188 Hauptmannia 170 haussa, Caeculisoma 129 heatherae, Callidosoma 133 Hehlenia 152


heleus, Leptus 104 Helladerythraeus 81 Hermann, Rhyncholophus 195 hermanni, Charletonia 159 hermanni, Hehlenia 189 heteropoda, Smaridia 54, 186 heteropoda, (?) Smaris 54 hexapus, Acarus 190 hibernans, Erythraeus 67 hidakai, Leptus 104 hirsti, Erythraeus 67 hirsti, Microsmaris 183 Hirstiosoma 55 Hirstiosomatinae 54 Hirstiosominae 54 hirsutissimus, Erythraeus (fossil) 191 hirsutissimus, Rhyncholophus 191 hirsutus, Neoabrolophus 173 hirsutus, Ritteria 148 hirtipes, Abrolophus 104 hirtipes, Leptus 104 hitchcocki, Leptus 104 hoffmannae, Opserythraeus 83 holgeri, Leptus 104 holmiae, Leptus 91 horiacus, Leptus 104 hospeticus, Leptus 104 hozumii, Leptus 105 hringuri, Leptus 105 huachuca, Callidosoma 133 huensis, Charletonia 159 humberti, Abrolophus 141 humberti, Hauptmannia 141 humeralis, Rhyncholophus 190 humeratus, Achorolophus 187 hunanensis, Charletonia 159 hunanica, Caeculisoma 129 hungarica, Smaridia 186 hupingshanicus, Leptus 105 huxleyi, Caeculisoma 129 hyperboreum, Trombidium 143

hyperboreus, Rhyncholophus 144 hypertrichotus, Erythraeus 67 iacobi, Abrolophus 141 ignotus danelli, Leptus 105 ignotus, Achorolophus 91, 105, 108 ignotus, Erythraeus 105 ignotus, Leptus 105 iguacuicus, Leptus 106 Iguatonia 173 illustris, Balaustium (fossil) 192 illustris, Rhyncholophus 191 ilzae, Leptus 106 imbricatus, Erythrellus 78 impectus, Rhincholophus [sic] 187 imperator, Erythraeus 84 imperator, Leptus 84 imperator, Rainbowia 84 imperialis, Erythraeus 73 imperialis, Rhyncholophus 73 imperfectus, Erythraeus 67 incanescens, Abrolophus 142 incertus, Leptus 106 incertus, (?) Leptus (fossil) 192 incertus, Rhyncholophus 191 indianensis, Leptus 106 indicus, Erythraeus 67 indicus, Leptus 106 indicus, Paraerythraeus 67 infernale, Caeculisoma 130 innocentae, Balaustium 177 insectorum, Leptus 190 insectorum, Trombidium 190 insulare, Callidosoma 133 insularis, Abrolophus 142 insularum, Abrolophus 142 insularum, Belaustium 142 intermedia, Smaris 186 intermedius, Leptus 106 io, Caeculisoma argus 128 iranica, Hauptmannia 170 iranicus, Erythraeus 76

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iraninejadi, Abrolophus 142 ishii, Charletonia 160 italicus, Erythraeus 67 Italustium 182 iwonae, Charletonia 160 jacksoni, Erythrites (? Urrbrites) 80 jacksoni, Momorangia 136 jacoti, Erythraeus 67 jacoti, Hirstiosoma 58 jacoti, Trichosmaris 58 jaegerskioeldi, Balaustium 160 jaegerskioeldi, Charletonia 160 jaegerskioeldi, Erythraeus 160 japonicus, Leptus 106 jenseni, Leptus 107 jinkaensis, Erythraeus 76 joannae, Microsmaris 183 jocquei, Leptus 107 johnstoni, Caeculisoma 130 johnstoni, Erythraeus 67 johnstoni, Lasioerythraeus 82 jolantae, Charletonia 160 josifovi, Leptus 107 jowitae, Erythraeus 68 jupiter, Clipeosoma 55 jurassicum, Balaustium 142 jurassicus, Abrolophus 142 justynae, Charletonia 160 kacperi, Balaustium 177 kacperi, Erythraeus 68 Kakamasia 81 kalaallus, Leptus 107 kaliksti, Charletonia 160 kalithensis, Charletonia 161 kamalii, Grandjeanella 169 kamalii, Leptus 107 Kamertonia 82 karachiensis, Leptus 107 kastaniensis, Erythraeus 77 kazimierae, Hauptmannia 171 kendalli, Balaustium 177

keralicus, Charletonia 161 keyi, Charletonia 161 khanjanii, Abrolophus 142 khanjanii, Hauptmannia 142 kharrazii, Erythraeus 77 kibonotensis, Erythraeus 161 kibonotensis, Charletonia 161 kibonatensis, Erythraeus 161 kiejstuti, Abrolophus pseudolongicollis 146 killingtoni, Leptus 108 kochi, Balaustium 189 kochi, Hehlenia 189 korematus, Leptus 108 kotorensis, Hauptmannia 171 kovalamicus, Pollux 184 Kraussiana 50 krendowskyi, Charletonia 161 krendowskyi, Erythraeus 161 kresnensis, Erythraeus 68 kurdistaniensis, Erythraeus 77 kuroshimaensis, Leptus 108 kuyperi, Bochartia 68 kuyperi, Erythraeus 68 kyushuensis, Leptus 108 lacrimosa, Fessonia 49 laevicristatum, Balaustium 142 laevicristatus, Abrolophus 142 lagopus, Erythraeus (fossil) 191 lagopus, Rhyncholophus 191 lambi, Taranakia 85 lanceolata, Smaris 52 lancifer, Erythraeus 77 lankensis, Charletonia 161 lapidarium, Balaustium 138 lapidarius, Abrolophus 138 laplandicus, Leptus 108 lappacea, Fessonia 49 Lasioerythraeus 82 latirostre, Astoma 190 latirostre, Trombidium 190 latirostris, Leptus 190


latreillei, Hirstiosoma 56 latreillei, Smaris 55, 56 laviniacus, Leptus 108 lawrencei, Charletonia 162 leanderi, Palenqustium 184 Legneria 190 leleupi, Leptus 109 lentiginosus, Rhyncholophus 187 leodegari, Callidosoma 133 leprosus, Rhyncholophus 187 Leptidae 58 Leptides 58 Leptinae 85 Leptoidae 58 leptopilus, Sphaerotarsus 57 Leptus 85 Leuchsia 51, 190 lighti, Leptus 109 littorale, Abrolophus 142 littorale, Belaustium 142 liui, Neophanolophus 51 lomani, Achorolophus 109 lomani, Erythraeus 109 lomani, Leptus 109 lombokensis, Charletonia 162 Lomeustium 182 longicollis, Abrolophus 143 longicollis, Achorolophus 143 longicollis, Hauptmannia 143 longilinea, Atomus 186 longilinealis, Smaris 186 longipapillum, Balaustium 143 longipapillus Abrolophus 143 longipedus, Erythraeus 77 longipes, Abrolophus 143 longipes, Balaustium 143 longipes, Erythraeus (fossil) 191 longipes, Leptus 109 longipes, Podosmaridia 83 longipes, Rhyncolophus 187 longipilis, Abrolophus 109

longipilis, Leptus 109 longirostris, Surasmaris 54 longisetorum, Neoabrolophus 174 longisetosus, Erythraeus 68 longispinosus, Leptus 109 longitarsus, Rhyncholophus 187 longulum, Balaustium 143 longulus, Abrolophus 143 longum, Balaustium 143 longus, Abrolophus 143 lorarius, Leptus 109 lorestanicus, Abalakeus 59 lovaniensis, Leptus 109 Lucasiella 61 macdonnelli, Erythroides 80 machilidis, Leptus 110 Macropus [in Erythraeidae] 61 maculatus, Atomus 110 maculatus, Leptus 110 maculatus, Rhyncholophus 110 madagascariensis, Leptus 110 madeirense, Balaustium 177 magnifica, Smaridia 175 magnifica, Smaris 175 majus, Balaustium sabulosum var. 148 maldonadoicus, Leptus 110 malpaisesensis, Balaustium 177 malwinae, Erythraeus 69 mamillatus, Erythraeus 190 mamillatus, Smaris 190 managarus, Leptus 110 maranaensis, Leptus 111 mariae, Leptus 111 mariani, Leptus 111 marinensis, Abrolophus 143 maringensis, Leptus 111 mariolae, Erythraeus 69 marmoratus, Abrolophus masaimaraicus, Leptus 111 matsumuratettix, Callidosoma 133 medardi, Balaustium 178

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224


medicagoense, Balaustium 178 medioareolatus, Rhyncholophus 189 mehranensis, Charletonia 162 meloidarum, Leptus 111 mentonensis, Bochartia 81 mentonensis, Forania 81 meruensis, Erythraeus 187 metcalfei, Paraphanolophus 83 metzi, Callidosoma 133 Microsmarialla 182 Microsmaris 182 millipedius, Leptus 112 milloti, Charletonia 162 milloti, Erythraeus 162 milloti, Sphaerolophus 162 mineatus, Rhyncholophus 144 miniata, Smaridia 143 miniatum, Balaustium 144 miniatum, Belaustium 144 miniatum, Trombidium 143 miniatus var. rubipes, Rhyncholophus 148 miniatus, Abrolophus 143 miniatus, Erythraeus 144 miniatus, Rhyncholophus 144 minno, Leptus 112 minodorae, Balaustium 178 minuta, Charletonia 162 minutus, Microsmaris 183 minutus, Sphaerolophus 162 mirandus, Microsmaris 183 mirabelae, Abrolophus 144 mirenae, Leptus 112 mirjavehi, Erythraeus 69 miromiri, Leptus 112 mirum, Myrmicotrombium 185 mitchelli, Cecidopus 151 mitchelli, Fessoniella 151 mitsukoae, Kraussiana 50 miyatakei, Hauptmannia 171 miyazakii, Callidosoma 162 miyazakii, Charletonia 162

moeritzensis, Erythraeus 69 moestus, Rhyncholophus 188 mogadoranus, Leptus 112 Moldoustium 182 mollicomus, Charletonia 162 mollicomus, Charleonia (Cavannea) 162 mollicomus, Sphaerolophus (Cavannea) 162 mollis, Erythraeus (Erythrolophus) 189 molochinus, Balaustium 112 molochinus, Leptus 112 molochinus, Rhyncholophus 112 Momorangia 136 mongolicus, Claverythraeus 60 monikae, Erythraeus 69 monolithosicus, Leptus 113 montana, Caeculisoma 130 montanum, Caeculisoma 130 montanus, Rhyncholophus 130 monteithi, Leptus 113 monticolus, Sphaerotarsus 57 Morieria 60 morii, Neophanolophus 51 mortenseni, Abrolophus 144 mortenseni, Hauptmannia 144 mouldsi, Caeculisoma 130 moxonae, Harpagella 136 mucronatus, Erythraeus 187 mullewaensis, Hauptmannia 174 mullewaensis, Pussardia 174 multisetosa, Grandjeannella 169 munsteri, Erythraeus 186 murorum, Balaustium 178 murorum, Belaustium 178 murorum, Erythraeus 178 murorum, Rhyncholophus 178 murorum, Trombidium 178 murteri, Leptus 113 Mypongia 183 Myrmicotrombiinae 185 Myrmicothrombium 185 Myrmicotrombium 185


naivashae, Charletonia 163 namaquamensis, Eatoniana 62 namaquensis, Eatoniana 62 namaquensis, Erythraeus 62 nasalis, Erythraeus 69 nasica, Phanolophus 174 nasica, Smaris 174 nasutum, Caeculisoma 130 nazeleae, Charletonia 163 nearcticus, Leptus 113 nemorum, Abrolophus 113 nemorum, Achorolophus 113 nemorum, Erythraeus 113 nemorum, Leptus 113 nemorum, Rhyncholophus 113 nemorum, Ritteria 113 Neoabrolophus 173 Neobalaustium 184 neobrevicollis, Abrolophus 144 neominiatum, Balaustium 178 Neomomorangia 136 neomurorum, Abrolophus 178 neomurorum, Balaustium 178 Neophanolophus 50 neoserratus, Erythroides 80 Neosmaris 82 nestori, Caeculisoma 130 newmani, Abrolophus 144 newmani, Balaustium 144 newmani, Belaustium 144 nicoleti, Smaris 53 nicoletii, Smaridia 53 nicoletii, Smaris 53 Niemannia 55, 190 niger, Rhyncholophus 72 nigeriae, Cecidopus 151 nigeriae, Fessonia 49 nigritiensis, Erythraeus 69 nikae, Balaustium 179 nikanori, Leptus 114 nipponicus, Erythraeus 71

nipponicus, Erythraeus phalangioides 71 nishidai, Charletonia 163 nivalis, Abrolophus 145 nivalis, Erythraeus 70 nivalis, Hauptmannia 145 nivalis, Rhyncholophus 70 nivosus, Erythraeus 147 norbakhshi, Bursaustium 181 norrisi, Leptus 114 norvegica, Rhyncholophus (Ritteria) 145 norvegicus, Achorolophus 145 norvegicus, Abrolophus 145 norvegicus, Leptus 145 norvegicus, Rhyncholophus 145 nova, Charletonia 163 novaeguinensis, Abrolophus 145 novaeguinensis, Belaustium 145 novaehollandiae, Hirstiosoma 56 novae - hollandiae, Morieria 158 novaeirlandiae, Caeculisoma 131 novaezealandiae, Neosmaris 83 novus, Sphaerolophus 163 obtusum, Balaustium 179 obtusum, Belaustium 179 ochroniger, Leptus 114 Ocypete 190 Oecosmaris 48 oedipodarum, Erythraeus 174 oedipodarum, Phanolophus 174 oedipodarum, Rhyncholophus 174 ogazulacus, Leptus 114 ojimai, Callidosoma 163 ojimai, Charletonia 163 ojimai, Erythraeus 163 okinawaensis, Callidosoma 163 okinawaensis, Charletonia 163 olafi, Leptus 114 olamukijacus, Leptus 114 onnae, Leptus 114 opilionides, Erythraeus 70 opilionis, Achorolophus 114

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opilionoides, Erythraeus 70 opilionoides, Rhyncholophus 70 Opserythraeus 83 oribatus, Leptus 190 orientalis, Erythraeus 70 ornatus, Leptus 115 orthopterarum, Leptus 115 orthrius, Leptus 115 osmondensis, Erythraeus 70 ostovani, Hauptmannia 171 oudemansi, Achorolophus 115 oudemansi, Bochartia 84 oudemansi, Charletonia 164 oudemansi, Erythraeus 84 oudemansi, Leptus 115 ovatus, Leptus 115 oxyae, Leptus 115 pachypus, “Erythraeus” 186 pachypus, Rhyncholophus 186 pachytibialis, Erythraeus (Helladerythraeus) 81 pachytibialis, Helladerythraeus 81 Palenqustium 184 pallidipes, Rhyncholophus 188 pallipes, Rhyncholophus 188 paludicola, Rhyncholophus 190 pangela, Grandjeanella 169 paolii, Charletonia 164 papillatum, Balaustium 179 papillosa, Fessonia 49 papillosa, Smaridia 49 papillosum, Smaris 49 papillosum, Trombidium 49 papillosus, Erythraeus 49 papillosus, Rhyncholophus 49 papillosus, Smaris 49 papuana, Trichosmaris 58 Paraerythraeus 62 paranensis, Fozustium 181 Paraphanolophus 83 Parasitengona 34 parkeri, Andrevella 150

parkeri, Erythraeus 150 parvipollicus, Atomus 188 parvipollicus, Rhyncholophus 188 parvisetosus, Erythraeus 188 parvisetosus, Rhyncholophus 188 parvulus, Leptus 116 parvum, Hauptmannia 171 parvus, Rhyncholophus 188 pasopaicus, Leptus 116 passerinii, Abrolophus 145 passerinii, Erythraeus 145 passerinii, Rhyncholophus 141 passidonicus, Erythraeus 77 pedatus, Rhyncholophus 188 pedestris, Rhyncholophus 188 Pedroerythraeus 60 pelebinus, Leptus 116 penelopae, Abrolophus 146 penninus, “Erythraeus” 186 penninus, Rhyncholophus 186 perifi, Abrolophus 145 perifi, Balaustium 145 perlongum, Balaustium 146 perlongus, Abrolophus 146 persiangulfia, Andrevella 151 perthense, Erythraeus 164 perthensis, Charletonia 164 peruvianus, Erythraeus 70 phalangii, Acarus 116 phalangii, Leptus 116 phalangioides var. Acis, Rhyncholophus 63 phalangioides var. gracilipes, Erythraeus 71 phalangioides, Apectolophus 70 phalangioides, Charletonia 167 phalangioides, Erythraeus 167 phalangioides, Rhyncholophus 70 phalangioides, Trombidium 70 phalangiolus, Erythraeus 187 phalangiorum, Leptus 190 phalangoides nipponicus, Erythraeus 71 phalangoides, Acarus 70


phalangoides, Erythraeus 70 phalangoides, Rhyncholophus 71 Phanolophinae 174 Phanolophus 174 philometra, Charletonia 164 philopempta, Sclerosmaris 51, 53 philopempta, Smaris 53 phloginus, Rhyncholophus 188 phloginus, Fessonia 188 phuketicus, Leptus 116 phyllotretae, Leptus 116 piae, Dambullaeus 135 picaforticus, Erythraeus 71 pileifera, Dupreia 189 pileifera, Smaridia 189 pilosa, Kakamasia 81 Pilosoma 51 pilosus, Erythraeus 71, 188 pilosus, Rhyncholophus 188 pincheni, Leptus 116 pinnata, Microsmaris 183 pinus, Smaris 53 pistoris, Leptus 117 plantacionis, Caeculisoma 130 Platylophus 189 plumatus, Erythraeus 72 plumifer, Eatoniana 62 plumifer, Rhyncholophus (Macropus) 61 plumipes, Eatoniana 61 plumipes, Erythraeus 61 plumipes, Erythraeus (Lucasiella) 61 plumipes, Lucasiella 61 plumipes, Rhyncholophus 61 plumosa, Rhyncholophus (Ritteria) 188 plumosus, Erythraeus 72 plumulosa, Hehlenia 189 pluto, Pilosoma 51 Podosmaridia 83 podorasensis, Hauptmannia 171 politricha, Charletonia 164 politrichus, Sphaerolophus 164

227

pollicaris, Rhyncholophus 188 Pollux 184 polonica, Kamertonia 82 polygona, Kakamasia 81 polythrix, Leptus 117 poonaensis, Leptus 117 porcinus, Achorolophus 164 porcinus, Charletonia 164 porcinus, Leptus 164 porcinus, Sphaerolophus 164 porifera, Fallopia 189 poriferus, Rhyncholophus 189 posticatus, Erythraeus 186 pozzoicus, Leptus 117 preciosus, Erythraeus 77 principalis, Balaustium 72 principalis, Erythraeus 72 principalis, Rhyncholophus 72 proavus, Erythraeus (fossil) 191 proavus, Rhyncholophus 191 prominens, Calyptostoma 53 prominens, Fessonia 53 prominens, Smaris 53 Prostigmata 34 Proterythraeus (fossil) 191 pseudolongicollis, Abrolophus 146 pseudolongicollis, Hauptmannia 146 pseudonivalis, Erythraeus 72 Ptilolophus 189 Ptilophus 189 pulchellus, Leptus 189 pulchellus, Rhyncholophus miniaceus var. 188 pulcher, Abrolophus 146 pulcher, Rhyncholophus 146 pulcherrima, Smaridia 186 punctatus, Rhyncholophus 188 puniceus, Leptus 117 Pussardia 174 putmani, Balaustium 179 puylaerti, Leptus 117 pyrenaeus, Leptus 117

228


quadratus, Erythraeus 187 qadrii, Cecidopus 151 quadrioculus, Rhyncholophus 188 quadrirubripes, Atomus 186 quadrisetum, Balaustium 146 quadrisetus, Abrolophus 146 quinquemaculatus, Rhyncholophus 188 quinquesetum, Balaustium 146 quinquesetus, Abrolophus 146 quinteri, Callidosoma 134 quisquiliarum sigma, Balaustium 147 quisquiliarum, Abrolophus 146 quisquiliarum, Balaustium 146 quisquiliarum, Belaustium 147 quisquiliarum, Erythraeus 146 quisquiliarum, Rhyncholophus 147 quisquiliarum, Smaridia 147 quisquiliarum, Trombidium 146 quisquiliarus, Abrolophus 146 quisquiliarus sigma, Abrolophus 147 rageaui, Charletonia 164 rajabii, Erythraeus 78 rajmundi, Balaustium 179 Ramsayella 84 rangitata, Ramsayella 84 raripapillum, Abrolophus 147 raripapillum, Balaustium 147 raripilus, Erythraeus (fossil) 192 raripilus, Rhyncholophus (fossil) 192 rebeccae, Wartookia 185 regale, Belaustium 72 regalis, Erythraeus 72 regalis, Rhyncholophus 72 regalis, Rhyncolophus 72 reginae, Erythraeus 79 reginae, Erythrites (E.) 79 reginae, Leptus 79 remotus, Erythraeus 73 repens, Leptus 118 retentus, Rhyncholophus 188 rhopalicum, Abrolophus 147

rhopalicum, Balaustium 147 rhopalicus, Abrolophus 147 rhopalicus, Balaustium 147 rhopalicus, Belaustium 147 rhopalicus, Rhyncholophus 147 Rhyncholophidae 58 Rhyncholophinae 58 Rhyncholophus 62 Rhyncolophides 58 Rhyncholophini 58 Rhyncholophoidae 58 Rhyncolophus 62 rilensis, Erythraeus 73 rileyi, Leptus 194 ripicola, Abrolophus 147 ripicola, Balaustium 147 ripicola, Belaustium 147 ripicola, Caeculisoma 57, 131, 134 ripicola, Callidosoma 134 ripicolum, Callidosoma 134 ripicolus, Sphaerotarsus 57 Ritteria 85 robustus, Rhyncholophus 188 rocciai, Charletonia 165 rosellae, Leptus 118 roseus, Rhyncolophus 188 rostratum, Callidosoma 134 rotundicollis, Hauptmannia rubescens, Rhyncholophus 63 rubescens, Rhyncolophus 63 rubipes, Abrolophus 148 rubipes, Rhyncholophus miniatus var. 148 rubricatus, Achorolophus 118 rubricatus, Leptus 118 rubricatus, Rhyncholophus 118 rubripes, Balaustium 148 rubripes, Belaustium 148 rubripes, Rhyncholophus 148 Rudaemannia 174 rudaensis, Hauptmannia 174 rudaensis, Rudaemannia 174


ruginus, Leptus 118 rupestris,Acarus 73 rupestris, Erythraeus 73 rupestris, Gamasus 73 rutgeri, Erythraeus 73 rwandae, Leptus 118 saboorii, Charletonia 165 sabrinae, Erythraeus 73 sabulosum, Abrolophus 148 sabulosum, Balaustium 148 sabulosum, Belaustium 148 sabulosus, Abrolophus 148 sabulosus, Rhyncholophus 148 saccatum, Trombidium 190 sagittatus, Rhyncholophus 188 saigusai, Leptus 118 salti, Charletonia 165 salti, Sphaerolophus 165 sambuci, Acarus 189 sambuci, Smaris 189 samosensis, Charletonia 165 Samouellea 51, 190 samsingensis, Leptus 119 sanctaechelenae, Charletonia 165 sanctaechelenae, Sphaerolophus (Cavannea) 165 sandovali, Tepoztlana 85 sanurensis, Carastrum 135 sardiniensis, Hauptmannia 171 sayi, Leptus 119 scalare, Hirstiosoma 56 scalaris, Hirstiosoma 56 schedingi, Achorolophus 119 schedingi, Leptus 119 schneideri, Fessonia 49 schneideri, Veithia 49 schweizeri, Abrolophus 148 schweizeri, Balaustium 148 Sclerosmaris 51 scobina, Fessonia 50 scopularium, Belaustium 148 scopularius, Abrolophus 148

scopulifera, Eatonia 61, 62 scutellatus, Leptus 119 septentrionale, Caeculisoma 131 sericea, Hirstiosoma 58 sericea, Smaris 58 serrata, Fessonia 50 serratociliatus, Erythraeus 73 serratociliatus, Paraerythraeus 73 serratus, Erythraeus 81 serratus, Erythroides 81 setipapillus, Abrolophus 148 shibai, Neophanolophus 51 shimensis, Leptus 119 shirleyanneae, Bochartia 82 shirleyanneae, Lasioerythraeus 82 shiroyama, Charletonia 165 shojaii, Erythraeus 74 shyamae, Cecidopus 152 sibuljinicus, Erythraeus 78 siculiformis, Leptus 119 siculiformis, Rhyncholophus 119 siculum, Balaustium 120 siculum, Belaustium 119 siculus, Achorolophus 119 siculus, Leptus 119 siculus, Rhyncholophus 119 siemsseni, Leptus 120 sieversi, Achorolophus 120 sieversi, Leptus 120 sifi, Erythraeus 74 signatus, Rhyncholophus 72 sigthori, Achorolophus 120 sigthori, Leptus 120 silesiacus, Hauptmannia 172 similis, Charletonia 165 similis, Leptus 120 similis, Rhyncholophus 74 similis, Sphaerolophus 165 similis, Erythraeus 74 simonettae, Leptus 120 simplex, Abrolophus 148

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simplex, Erythraeus 74 simplex, Rhyncholophus (Apectolophus) 188 singhi, Leptus 120 singularis, Charletonia 155 singularis, Erythraeus 152, 155 singularis, Hehlenia 155 singularis, Smaris 155 slivovi, Leptus 121 Smaridia 51 Smarididae 48 Smaridiidae 48 Smaridinae 48 Smaris 51 Smarisidae 48 smithi, Leptus 121 soddagus, Leptus 121 sorkini, Callidosoma 134 southcotti, Augustsonella 59 southcotti, Balaustium 179 southcotti, Charletonia 166 southcotti, Curteria 61 southcotti, Erythraeus 74 southcotti, Leptus 121 southcotti, Proterythraeus (fossil) 191 soydani, Balaustium 179 sparnoni, Caeculisoma 131 sparsepinnata, Microsmaris 183 spatulatopilis, Erythraeus 74 spectabilis, Abrolophus 149 spegazzinii, Smaris 186 Sphaerolopheae 189 Sphaerolophinae 128, 189 Sphaerolophus (Eupodolophus) 151 Sphaerotarsus 56 spiculosus, Abrolophus 149 spinalatus, Leptus 121 spinatus, Erythraeus 186 spinosa, Charletonia 166 spinosus, Sphaerolophus 166 squamata, Dugesia 53 squamata, Smaris 53

squamatum, Smaris 53 squamatum, Trombidium 53 squamatus, Rhyncholophus 53 squamatus, Smaris 53 stanislavae, Hauptmannia 172 stefani, Leptus 121 stieglmayri, Achorolophus 121 stieglmayri, Erythraeus 121 stieglmayri, Leptus 121 stolae, Leptus 122 stolli, Rhyncholophus 188 straeleni, Cecidopus 152 striaticeps, Charletonia 166 striglipes, Rhyncholophus (Abrolophus) 188 strojnyi, Abrolophus 149 stuarti, Erythraeus 74 styriacus, Erythraeus 75 submurorum, Balaustium 179 subnuda, Charletonia 166 subnudus, Sphaerolophus 166 subquisquiliarum, Balaustium 149 subquisquiliarus, Abrolophus 149 sudanensis, Achorolophus 122 sudanensis, Leptus 122 sudeticum, Trombidium (Otonyssus) 155 sulcatum, Belaustium 131 sulcatum, Caeculisoma 131 sulcatus, Rhyncholophus 131 sulciscutus, Leptus 122 Surasmaris 54 sureshi, Charletonia 166 sureshi, Sphaerolophus 166 swani, Leptus 122 swaziana, Charletonia 166 swazianus, Erythraeus 166 taiwanensis, Charletonia 166 tamarae, Charletonia 155 tammuzi, Leptus 122 Taranakia 84 tardum, Balaustium 149 tardum, Belaustium 149


tardus, Abrolophus 149 tardus, Rhyncholophus 149 tarranus, Leptus 122 tasmaniensis, Hirstiosoma 56 tatianae, Charletonia 167 taylori, Fessonia 50 tehranicus, Erythraeus 78 Tepotztlana 85 terebrans, Leptus 123 Terebrites 80 tetrigius, Leptus 123 texanus, Leptus 123 texanus, Rhyncholophus 123 tiki, Callidosoma 134 tindalei, Callidosoma 134 tindalei, Leptus 123 tinnae, Erythraeus 75 tiranicus, Leptus 123 titikaka, Erythraeus 75 titinius, Leptus 123 togoensis, Lomeustium 182 tokunagai, Discolophus 137 tolteka, Erythraeus 186 tonsor, Abrolophus 149 tonsor, Hauptmannia 149 tonsus, Erythraeus 186 tonsus, Rhyncholophus 188 torresianus, Leptus 123 tragardhi, Charletonia 167 tragardhi, Erythraeus 75 tragardhi, Parerythraeus 75 transcaspica, Eatoniana plumifer f. 62 treati, Callidosoma 134 treati, Leptus 124 triacanthus, Leptus 124 Trichosmaris 57 tridentifer, Rhyncholophus 190 trifarius, Hauptmannia 172 trimaculata, Rhyncholophus (Ritteria) 124 trimaculata, Ritteria 124 trimaculatum, Belaustium 124

trimaculatum, Leptus 124 trimaculatum, Trombidium 124 trimaculatus, Abrolophus 124 trimaculatus, Erythraeus 124 trimaculatus, Leptus 124 trimaculatus, Rhyncholophus 124 trimaculatus, Rhyncolophus 124 trinotatus, Abrolophus 149 trisetum, Balaustium 150 trisetus, Abrolophus 150 troguloides, Arythaena (fossil) 192 Trombidium [in Erythraeoidea] 124 trucidatus, Achorolophus 125 trucidatus, Leptus 125 truncatus, Leptus 125 tuberculata, Augustsonella 59 tuberculatum, Caeculisoma 131 tuberculatus, Erythraeus 59 tuxeni, Balaustioides 172 ubudicus, Leptus 125 ueckermanni, Erythraeus 78 unidentatum, Balaustium 180 unidentatus, Balaustium 180 unidentatus, Erythraeus 180 unimiri, Abrolophus 150 unisetum, Balaustium 150 unisetus, Abrolophus 150 urodaci, Leptus 125 urrbrae, Erythraeus 79 urrbrae, Erythrites (Urrbrites) 79 Urrbrites 79 ursinus, Erythraeus 186 ursyni, Leptus 125 utheri, Leptus 125 vallata, Momorangia 136 variatus, Leptus 125 variatus, Leptus diversus var. 125 Veithia 48, 190 veletense, Balaustium 180 veletensis, Abrolophus 200 venus, Charletonia 167

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vernalis, Rhyncholophus 188 vertex, Achorolophus 91 vertex, Erythraeus 126 vertex, Leptus 126 vertex, Rhyncholophus 126 vertex, Rhyncholophus (Ritteria) 126 vertex, Rhyncholophus nemorum var. 91 vertex, Ritteria nemorum var. 126 verticiformis, Leptus 126 vestita, Leuchsia 190 vestita, Smaridia 190 viburnicola, Hauptmannia 174 vignae, Balaustium 180 villosa, Hehlenia 190 villosa, Smaridia 190 villosus, Achorolophus 126 villosus, Leptus 126 viticola, Hauptmannia 172 vittatum, Belaustium 150 vittatus, Abrolophus 150 vitzthumi, Charletonia 167 volzi, Charletonia 167 volzi, Erythraeus 167 vuki, Leptus (Amaroptus) 127 waldockae, Leptus 126 wallaceanum, Callidosoma 135 warregense, Belaustium 126 warregense, Leptus 126 Wartookia 185 welbourni, Abrolophus 150 welbourni, Callidosoma 135 welbourni, Lasioerythraeus 82

welbourni, Leptus 126 westraliense, Hauptmannia 169 westraliense, Sphaerolophus 167 westraliensis, Charletonia 167 westraliensis, Grandjeanella 169 whitcombi, Erythraeus 82 whitcombi, Lasioerythraeus 82 willmanni, Hauptmannia 172 womersleyi, Callidosoma 135 womersleyi, Charletonia 168 womersleyi, Erythraeus 80 womersleyi, Erythrites (Terebrites) 80 womersleyi, Sphaerotarsus 57 workandae, Pollux 185 wratislaviensis, Balaustium 180 wratislaviensis, Hauptmannia 172 wrighti, Charletonia 168 xerothermicum, Balaustium 180 yangshuonicus, Erythraeus 75 yanlingica, Hauptmannia 173 yoshimii, Leptus 127 Zaracarus 75 zbelutkaicus, Leptus 105 zelandicus, Abrolophus 150 zeteki, Smaris 54 zhangi, Balaustium 180 zhangi, Erythraeus 75 zhangi, Leptus 127 zhejiangensis, Leptus 127 zhutingensis, Leptus 127 zorani, Charletonia 168


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Haitlinger R. 1984. Charletonia tamarae n.sp. (Acari, Erythraeidae) an ectoparasite of the Orthoptera from Greece. – Polskie Pismo Ent., Wroclaw, 54: 385-390. Haitlinger R. 1985. Charletonia singularis (Oudemans) and other rare mites (Acarina) collected on small mammals in Poland. – Bull. Ent. de Pologne, 55: 433-436. Haitlinger R. 1986a. Three new Charletonia (Acari, Erythraeidae) from Vietnam. – Bull. Entomol. de Pologne, 56: 641-647. Haitlinger R. 1986b. The genus Hauptmannia Oudemans, 1910 (Acari, Prostigmata, Erythraeidae) in Poland. – Bull. Entomol. de Pologne, 56:181-191. Haitlinger R. 1987a. The genus Erythraeus Latreille, 1806 (Acari, Prostigmata, Erythraeidae) in Poland (larvae). – Bull. Ent. de Pologne, 57: 725-734. Haitlinger R. 1987b. Hauptmannia aitapensis Southcott, 1947 (Acari, Erythraeidae) a new species of mites to fauna of Viet-Nam. – Bull. Entom. de Pologne, 56: 915-916. Haitlinger R. 1987d. The genus Leptus Latreille, 1796 and Charletonia Oudemans, 1910 (Acari, Prostigmata, Erythraeidae) in Poland (larvae). – Pol. Pismo Ent., 57: 339-349. Haitlinger R. 1987e. Leptus stolae sp. n. (Acari, Prostigmata, Erythraeidae) from Stolas nudicollis (Boh.)(Coleoptera, Chrysomelidae, Cassidinae) from Brazil. – Bull. Ent. de Pologne, 57: 357-359. Haitlinger R. 1987f. Anomalies in growth and supernumerary setae in larvae of the genus Charletonia Oudemans, 1910 and Leptus Latreille, 1796 (Acari, Prostigmata, Erythraeidae). – Przeglad Zoologiczny, 31(1): 107-109. Haitlinger R. 1987g. Larval Erythraeidae (Acari, Prostigmata) from Madagascar. – Bull. Ent. de Pologne, 57: 701-723. Haitlinger R. 1987h. Hauptmannia pseudolongicollis n.sp. (Acari, Prostigmata, Erythraeidae) from Poland. – Bull. Ent. de Pologne, 57: 351-355. Haitlinger R. 1987i. Further information on distribution of species from the genus Hauptmannia Oudemans, 1910 (Acari, Prostigmata, Eythraeidae) in Poland. – Przeglad zoologiczny, 31(2): 159-164. Haitlinger R. 1990a. Four new species of Leptus (Acari, Prostigmata, Erythraeidae) from tropical Africa. – Angew. Parasitol., 31: 29-33. Haitlinger R. 1990b. Four new species of Leptus Latreille, 1796 (Acari, Prostigmata, Erythraeidae) from Insects of Australia, New Guinea and Asia. – Wiadomosci Parazytol., 36 (1-3): 47-53. Haitlinger R. 1990c. Two new species of Leptus Latreille 1976 (Acari, Prostigmata, Erythraeidae) from Tenebrionidae (Coleoptera) with a key to Europaean and North African species. – Pol. Pismo Ent., 60: 45-49. Haitlinger R. 1991. List of mites occuring on insects in Poland. – Wiadomości Parazytologiczne, 37(1): 85-90. Haitlinger R. 1992. New data on distribution of larvae from the genus Leptus Latreille, 1796 (Acari, Prostigmata, Erythraeidae) in Poland with the description of Leptus miromiri n.sp. – Wiadomosci parazytologiczne, 37(4), 1991 [1992]: 499-506.


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Haitlinger R. 1993. Acari (Arachnida) and Anoplura (Insecta) collected on small mammals, reptiles and insects in Greece and Cyprus. – Biologia Gallo-hellenica, 20(1): 83-88. Haitlinger R. 1994a. Two new larval mites (Acari: Trombidiidae, Erythraeidae) from Switzerland. – Mitt. Schweiz. Entomol. Ges., 67: 405-410. Haitlinger R. 1994b. A larval mite (Acari, Erythraeidae) parasitizing the buprestid beetle (Insecta, Coleoptera) in Spain. – Graellsia, 50: 165-166. Haitlinger R. 1994c. Two new species of Leptus Latreille, 1796 (Acari: Prostigmata: Erythraeidae) associated with Tenebrionidae (Insecta: Coleoptera). – Israel J. Ent., 28: 139-149. Haitlinger R. 1995. New mites (Acari, Astigmata: Canestriniidae; Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae) for the fauna of Austria, Germany and Hercegovina with descriptions of four new species. – Linzer biol. Beitr., 27(1): 259-272. Haitlinger R. 1996. Seven new larval species of mites (Acari, Prostigmata: Erythraeidae and Trombidiidae) from Poland. – Wiadomosci Parazytologiczne, 42(4): 443-460. Haitlinger R. 1997a. New larval mites (Acari, Prostigmata: Erythraeidae, Trombidiidae) from Canary Islands. – Zool. baetica, 8: 123-132. Haitlinger R. 1997b. Data relating to the disribution and host-specifity of bat-infesting mites (Acari, Mesostigmata, Prostigmata, Astigmata) in Germany. – Drosera, 2: 95-112. Haitlinger R. 1998. Five new species of Leptus Latreille, 1796 (Acari: Prostigmata, Erythraeidae) from Asia and Africa. – Bonner Zool. Beitr., 48(1): 97-110. Haitlinger R. 1999a. Three new larval mites (Acari: Prostigmata: Eutrombidiidae, Erythraeidae, Trombellidae) from Australia, Turkey and Thailand. – Zeszyty naukowe Akademii rolniczej we Wroclawiu, Zootechnika, 45, 362: 57-73. Haitlinger R. 1999b. Six new species of Leptus Latreille, 1796 (Acari, Prostigmata, Erythraeidae) from South-East Asia. – Miscellània Zoològica, 22(2): 51-68. Haitlinger R. 2000a. Four new species of Leptus Latreille, 1796 (Acari: Prostigmata: Erythraeidae) from Central America. – Systematic & Applied Acarology, 5: 131-142. Haitlinger R. 2000b. Mites (Acari: Prostigmata: Erythraeidae, Trombidiidae) new to the fauna of Norway, Finland, Russia, Latvia and Lithuania, with a description of Podothrombium roari n. sp. – Entomologica Fennica, 11: 187-193. Haitlinger R. 2000c. Four new species of Leptus Latreille, 1796 (Acari, Prostigmata, Erythraeidae) from Peru. – Boll. Mus. reg. Sci. nat Torino, 17 (1): 149-162. Haitlinger R. 2000d. New larval mites (Acari: Prostigmata: Erythraeidae, Microtrombidiidae, Trombidiidae) from Turkey, Peru and Poland. – Wiadomości Parazytologiczne, 46(3): 379-396. Haitlinger R. 2000e. Five new species of Balaustiinae (Acari: Prostigmata: Erythraeidae) from Guatemala, Mexico and Italy. – Zesz. Nauk. Acad. Roln. we Wrocławiu, 400: 69-84. Haitlinger R. 2001a. Dambullaeus piae gen. n., sp. n. (Acari: Prostigmata: Erythraeidae) from Sri Lanka. – Zesz. Nauk. Acad. Roln. We Wrocławiu, 429: 51-54. Haitlinger R. 2001b. Four new species of Leptus (Acari, Prostigmata, Erythraeidae) from Republic of South Africa and Kenya. – Biologia, Bratislava, 56: 473-481.

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Haitlinger R. 2002a. Two new species of Leptus Latreille, 1796 and the first record of Leptus astrubali Haitlinger, 1999 (Acari: Prostigmata: Erythraeidae) from India and Sri Lanka. – Syst. Appl. Acarol., 7: 177-184. Haitlinger R. 2002b. Erythraeus (Erythraeus) malwinae Haitlinger, 1995 (Acari: Prostigmata: Erythraeidae) nowy gatunek roztocza dla fauny Polski. – Przegląd Zoologiczny, 46(1-2): 65-66. Haitlinger R. 2002c. Pollux kovalamicus sp. nov. (Acari: Prostigmata: Erythraeidae) from India. – Syst. Appl. Acarol., 7: 173-175. Haitlinger R. 2002d. A new larval Hauptmannia Oudemans, 1910 and the first record of Abrolophus neobrevicollis Zhang & Goldarazena, 1996 (Acari, Prostigmata, Erythraeidae) from Madeira. – Syst. parasitol., 53: 115-119. Haitlinger R. 2002e. Rostocze (Acari: Prostigmata: Erythraeoidea, Trombidioidea, Tanaupodoidea, Trombiculoidea bez Trombiculidae) Sudetów. – Przyroda Sudetów Zach., 5: 147-152. Haitlinger R. 2002f. Erythraeidae and Trombidiidae (Allothrombiinae) (Acari: Prostigmata) from Mallorca (Balearic Islands), with description of two new species. – Bol. Soc. Hist. Nat. Balears, 45: 191-197. Haitlinger R. 2002g. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae) from Slovakia. – Biologia, 57: 554-556. Haitlinger R. 2003a. Four new larval Erythraeidae (Acari, Prostigmata) from Rhodes, Greece. – Biologia, Bratislava, 58(2): 133-146. Haitlinger R. 2003b. Three new larval Charletonia Oudemans, 1910 (Acari: Prostigmata: Erythraeidae) and the first record of Charletonia krendowskyi (Feider, 1954) from Rhodes, Greece. – Systematic Parasitology, 56: 49-55. Haitlinger R. 2003c. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae, Calyptostomatidae) from Slovenia, with a description of Campylothrombium nadbori spec. nov. – Scopolia, 51: 1-11. Haitlinger R. 2003d. Hauptmannia bohdani n.sp. from Poland (Acari: Prostigmata: Erythraeidae). – Genus, Wroclaw, 14 (4): 603-607. Haitlinger R. 2003e. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae) from the Czech Republic, with a description of Podothrombium karlovaicus n.sp. – J. Nat. Mus. Nat. Hist. Ser., 172 (1-4): 5-12. Haitlinger R. 2003f. Distribution of the species of the genera Hauptmannia, Oudemans, 1910 and Rudaemannia Haitlinger, 2000 (Acari: Prostigmata: Erythraeidae) in Poland. – Przegląd Przyrodniczy, 14(1-2): 61-69. Haitlinger R. 2004a. Three new species of Leptus Latreille, 1796 and the first record of Leptus onnae Haitlinger, 2000 (Acari: Prostigmata: Erythraeidae) from Brazil. – Systematic & Applied Acarology, 9: 147-156. Haitlinger R. 2004b. New records of mites (Acari: Prostigmata: Erythraeidae) from Cambodia and Myanmar, with a description of Erythraeus (Erythraeus) kacperi sp. nov. – Systematic & Applied Acarology, 9: 157-161.


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Haitlinger R. 2004c. A new larval Momorangia (Acari: Prostigmata: Erythraeidae) from Kenya. – Systematic & Applied Acarology, 9: 179-182. Haitlinger R. 2004d. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Eutrombidiidae) from Croatia, with descriptions of three new species. – Natura Croatica, Zagreb, 13(2): 143-160. Haitlinger R. 2004e. Callidosoma leodegnari n. sp. and new record of Callidosoma welbourni Treat, 1985 (Acari: Prostigmata: Erythraeidae) from Argentina and Brazil. – Int. J. Acarol., 30(3): 251-254. Haitlinger R. 2004f. Charletonia domawiti n. sp., Caeculisoma nestori n. sp., and Iguatonia barbillae n. gen. and n. sp. from Brazil (Acari: Prostigmata: Erythraeidae). – Genus, 15(3): 435-444. Haitlinger R. 2004g. Larval Erythraeid mites new to the fauna of Dominican Republic, with a description of Leptus cabareticus sp. n. (Acari: Prostigmata: Erythraeidae). – Zeszyty Naukowe Akademii Rolniczej we Wrocławiu, Zootechnika, 50(488): 125-132. Haitlinger R. 2004h. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae) from La Palma, Canary Islands, Spain, with descriptions of four new species and a new genus. – Aracnologia, 10: 215-223. Haitlinger R. 2005a. A new genus and four new species of mites from Argentina, Brazil and Nicaragua (Acari: Prostigmata: Erythraeidae, Eutrombidiidae). – Genus, Wroclaw, 16 (4): 513-525. Haitlinger R. 2005b. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Microtrombidiidae) from Andorra, with description of Campylothrombium soldeuensis sp.n. – Revista Ibérica de Aracnología, 12: 73-77. Haitlinger R. 2006a. Eight new species and new records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Johnstonianidae) from China including Macao. – Syst. & Applied Acarology, 11: 83-105. Haitlinger R. 2006b. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae) from Samos, Greece, with descriptions of six new species. – Syst. & Applied Acarology, 11: 107-123. Haitlinger R. 2006c. New species and new records of mites (Acari: Prostigmata: Erythraeoidea) from Bali, Lombok and Linnga Islands, Indonesia. – Syst. & Applied Acarology, 12: 219-230. Haitlinger R. 2006d. Dasitrombium margeritanum sp.n., Leptus tiranicus sp.n. and the first record of L. olafi Haitlinger (Acari: Prostigmata: Neothrombiidae, Erythraeidae) ectoparasitic on Orthoptera and Diptera (Insecta) from Margerita, Venezuela. – Polskie Pismo Entomologiczne, 75: 347-357. Haitlinger R. 2006e. New records of larval mites (Acari: Prostigmata: Erythraeidae, Eutrombidiidae, Trombidiidae) from Portugal and Spain. – Revista Iberica de Aracnologia, 14: 105-108. Haitlinger R. 2006f. New records of mites (Acari: Prostigmata: Erythraeidae, Eutrombidiidae, Trombidiidae, Chyzeriidae, Leeuwenhoekiidae) from Greece. – Zeszyty Naukowe Uniw. Przyr. we Wroclawiu, Biologia i Hodowla zwierząt, LIV, 548: 35-42.

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Haitlinger R. 2006g. A new genus and nine new larval species (Acari: Prostigmata: Erythraeidae, Eutrombidiidae) from Benin, Ghana and Togo. – Rev. Ibér. Aracnol., 14: 109-127. Haitlinger R. 2006h. Four new species of Erythraeidae (Acari: Prostigmata) and the first records of Charletonia braunsi (Oudemans, 1910) and C. brunni (Oudemans, 1910) from Ethiopia. – Revista Ibérica de Aracnología, 12: 79-90. Haitlinger R. 2007a. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Eutrombidiidae) from the Balkan Peninsula. – Biologia, Section Zoology, Bratislava, 62(1) 67-77. Haitlinger R. 2007b. New records of mites (Acari: Prostigmata: Erythraeidae, Trombidiidae, Eutrombidiidae) from France, Liechtenstein and Switzerland, with descriptions of three new species. – Syst. and Applied Acarology, 12: 55-72. Haitlinger R. 2007c. New species and records of mites (Acari: Prostigmata: Erythraeidae, Johnstonianidae, Microtrombidiidae, Trombidiidae) from Moldova and Ukraine. – Biologia, Section Zoology, Bratislava, Haitlinger R. 2007d. New records of mites from Corsica and Sardinia, with descriptions of five new species (Acari: Prostigmata: Erythraeidae, Trombidiidae, Eutrombidiidae). – Genus, 18(3): 529-543. Haitlinger R., A. Saboori. 1996. Seven new larval mites (Acari, Prostigmata, Erythraeidae) from Iran. – Miscellánea Zool., 19: 117-131. Haitlinger R., A. Saboori. 2007. Two new larval ectoparasitic Charletonia Oudemans (Acari: Prostigmata: Erythraeidae) found on Orthoptera (Insecta), and the first record of Charletonia krendowskyi (Feider) in Iran. – Polish J. of Entomology, 76: 61-71. Haitlinger R., G. Walter. 1997. Data relating to the distribution and host-specifity of batinfesting mites (Acari, Mesostigmata, Prostigmata, Astigmata) in Germany. – Drosera, 97(2): 95-112. Halbert J. N. 1915. Clare Island Survey. Part 39. Acarinida. Section II. Terrestrial and marine Acarina. – Proc. R. Irish Acad., 39(2): 45-136. Halbert J. N. 1920. The Acarina of the seashore. – Proc. R. Irish Acad., 35B(7): 106-152. Hall H.V.M. 1912. Some marine and terrestrial Acarina of Laguna Beach. – Rep. Laguna Mar. Lab., Pomona Coll., Claremont, Cal. No 1: 177-186. Halliday B. 1998. Mites of Australia. A Checklist and Bibliography. – Monographs on Invertebrate Taxonomy, 5: 317 pp. Halliday R.B. 2001. Systematics and biology of the Australian species of Balaustium von Heyden (Acari: Erythraeidae). – Australian J. of Entomology, 40(4): 326-330. Haller G. 1882. Beitrag zur Kenntnis der Milbenfauna Würtembergs. – Jh. Ver. Vaterl. Naturk. Würtemberg, 38: 293-325. Haller G. 1885. Beiträge zur Kenntnis der schweizerischen Milbenfauna. – V. Sch. naturf. Ges. Zurich, 30: 82. Hara J., M. Hanada. 1963. Description of a new species of Erythraeid mite (Acarina: Erythraeidae) from Japan. – Jap. J. Sanit. Zool., 13 (4): 268-270. Hartzell A. 1918. A preliminary list of the Acarina of Iowa. – Iowa Acad. Sci., 25: 205-212.


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