Annotated Catalogue of the Carabid Beetles of Albania (Coleoptera: Carabidae): v. 64 (Pensoft Series Faunistica)

T “ Without doubt, this excellent catalogue, compiled with diligence and knowledge, will be an important tool for anyo...

Author: B.V. Gueorguiev

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“

Without doubt, this excellent catalogue, compiled with diligence and knowledge, will be an important tool for anyone, not only specialists on Balkan Carabidae, and will improve our insights into the systematics and distribution of carabid beetles. The hope of the author to place Albania with this catalogue side by side to Bulgaria (certainly the best-known regional fauna of Carabidae in the Balkan Peninsula, catalogued in 1995 by the same author and his father) is surely fulfilled. David Wrase

”

Pensoft Series Faunistica 64

Annotated Catalogue of the Carabid Beetles of Albania (Coleoptera: Carabidae)

his catalogue summarizes all available literature records and a lot of new material revised or determined by the author in various European museums and private collections. At present the carabid fauna of Albania can be considered as well-studied. The catalogue includes references to or data for 543 species, 103 genera and 31 tribes of Carabidae. Several dozen taxa of specific or supraspecific rank are added for the first time to the fauna of either the Balkan Peninsula or Albania. A moderate taxonomic contribution based on a study of types, topotypes or series of previously problematic species or subspecies from Albania and the neighbouring countries is also presented; as a result, 38 various taxonomic changes (new synonyms, revalidations, new statuses etc.) are introduced. All species are listed with detailed localities and references. The book is addressed to specialists in coleopterology and entomology, biogeographers and nature conservationists, collectors and all ground beetle lovers.

ANNOTATED CATALOGUE of the

Carabid Beetles of Albania (Coleoptera: Carabidae) by B.V. Guéorguiev

Annotated Catalogue of the Carabid Beetles of Albania (Coleoptera: Carabidae) Borislav V. Guéorguiev

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ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA / BORISLAV V. GUÉORGUIEV

Annotated Catalogue of the Carabid Beetles of Albania (Coleoptera: Carabidae) BORISLAV V. GUÉORGUIEV

SOFIA–MOSCOW 2007

ANNOTATED CATALOGUE OF THE CARABID BEETLES OF ALBANIA (COLEOPTERA: CARABIDAE) Borislav V. Guéorguiev

Editor: Ivailo Stoyanov Scientific readers: Augusto Vigna Taglianti & David W. Wrase

First published 2007 ISBN 978-954-642-295-8 Pensoft Series Faunistica No 64 ISSN 1312-0174

© PENSOFT Publishers All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted in any form by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior written permission of the copyright owner.

Pensoft Publishers Geo Milev Str. 13a, Sofia 1111, Bulgaria [email protected] www.pensoft.net

Printed in Bulgaria, February 2007

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Contents Introduction 7 Background of the study 7 Aims of the study 8 Short description of the studied area 8 Treatment of the taxa 10 Material and depositories of the collections 10 Acknowledgments 11 Abbreviations 13 Topographic treatment of some localities 14 Systematic part 16 Results 187 Conclusions 188 Taxonomic structure of the Albanian Carabidae (Table 1) 189 Appendix 193 References 205 Index 224

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Introduction The ground beetle fauna of Albania is not very well known taking into account the fact that an overall investigation based upon studied material and critical survey on the literature has never been made. So far, our knowledge of this fauna is still largely due to Apfelbeck’s well-known monograph (Apfelbeck, 1904) and other works of the same author published in the period 1901-1930 (Apfelbeck, 1901a; 1901b; 1902a; 1903; 1904; 1905; 1906; 1907a; 1907b; 1909; 1911; 1916; 1917; 1918a; 1918b; 1918c; 1919a; 1919b; 1930). Later, important faunistic contributions were made also by Csiki (1940), Schatzmayr (1943b), and Hieke et al. (1986). Dealing with the Bulgarian ground beetles, Hieke & Wrase (1988) mentioned 296 species (Nebriinae – 14, Cicindelinae – 5, Carabinae – 10, Omophroninae – 1, Brachininae – 10, Siagoninae – 1, Scaritinae – 12, Trechinae – 58, Harpalinae – 185) for Albania without detailed records. For political reasons, in the period 1945-1990 not many faunistic records were published, and relatively few materials from the country were available in the collections. Since 1990 collecting ground beetles there became possible again. Recently, 363 species (Nebriinae – 10, Cicindelinae – 8, Carabinae – 18, Omophroninae – 1, Brachininae – 11, Siagoninae – 1, Scaritinae – 20, Apotominae – 1, Trechinae – 55, Harpalinae – 238) have been mentioned for Albania in the most recent catalogue of Palaearctic Coleoptera by Löbl & Smetana (2003). Examination of the available collections added information about 55 new species for the country plus 119 species omitted in the last work as well as many new faunistic data for other species. The subsequent critical survey of the literature data brought to the conclusion that 543 species of Carabidae live in Albania. Moreover, some taxonomic changes and a lot of comments for many taxa have been made.

Background of the study My interest in the Albanian Carabidae has appeared under the stimulus of my late father, the well-known Bulgarian coleopterologist Vassil Borissov Guéorguiev (*1935 – †1996), immediately after the printing of the catalogue of the Bulgarian Carabidae (Guéorguiev & Guéorguiev, 1995). The insufficient level of exploration of the carabid fauna of the country at this time (because of the strong and prolonged political isolation and lack of regional specialists), along with relatively diverse and fresh material collected by Bulgarian zoologists in Albania, were the principal reasons for starting this investigation. The complicated orography of this country, particularly the complex relief, determines a great diversity of zonal (seacoast, lowland, hilly, forest, subalpine, alpine) and intrazonal (riparian, stony, limestone) habitats, that could favor INTRODUCTION

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a high diversity of the focused animal groups – the ground beetles. Finally, it was necessary to sort out some interpretation of toponyms in the region, which were incorrectly treated in the past (for example, some authors have erroneously cited records or have referenced old collection material as being from Albania).

Aims of the study The present study pursues two objectives, and the first one is the accomplishment of faunistic inventory of the Carabidae from Albania. This process includes discovery of all faunistic publications for Albania and identification or partial revision of the collections (for unpublished and published specimens, respectively) and their registration. The result of these activities is the improvement of the faunistic knowledge (species diversity and distribution) about the ground beetles not only on the territory of Albania, but also on the Balkan Peninsula, and in Europe on a large scale. The second objective of the study is taxonomical – examination of type material and revision of the status of some doubtful taxa from Albania and its adjacent regions (Montenegro, the Kosovo Region of Serbia, Macedonia and Northwestern Greece), which will contribute to the systematics of the group.

Short description of the studied area Albania (indigenous name Shqipëri) is situated in the southwestern part of the Balkan Peninsula and is meridionally 300-400 km long. The country occupies about 28,748 km2, and its maximal width is about 150 km. Albania is bordered in northwest with Montenegro, in northeast and east with the Kosovo Region of Serbia and Macedonia, and in south with Greece. In the west, Ionic Sea divides Albania from the Apennine Peninsula where the strait Otranto is the close water area between both lands. One third of the country, mainly located in the western part, lies lower than 300 m a.s.l., the territory at altitude between 300 and 1000 m is almost 40 % of the country, while that above 1000 m a.s.l. is about 30 %. The mountains situated in the east of Albania are part of the Dinaric mountainous system and they represent relatively young and high massifs. The highest top is Korabi Peak (2764 m) located on the east state border. The main Albanian rivers, Drin, Shkumbin, Semani, Vijosë, flow in direction northwest - southeast. The situation of the country in the Mediterranean area and in a zone of subtropical climate determines the principal features of the flora and fauna. The summer is dry and warm with average temperatures in July between 24 and 25 C°, the autumn and winter are mild and rainy. In Albania falls one of the most abundant rainfalls in Europe – between 1000 and 1800 mm per year in the western lowlands, and to 2500 mm in the mountains. The seasonal distribution of the precipitations is very uneven – in the sum8


Map of Albania

MAP OF ALBANIA

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mer the rainfall is just about 1/10 from the annual sum, and most plains and hilly parts of the country are characterized with heavy summer droughts. The days with average temperature more than 10 C° are about 300 (in the seacoast plains) and favor the development of Mediterranean flora and fauna. The evergreen sclerophilous Mediterranean forests are developed in the western part of Albania up to altitude of 400-500 m and in some other regions – higher. These forests consist of maquises (mosaically distributed in the country) where Olea sp., Quercus coccifera, Q. ilex, other Quercus spp., Laurus nobilis, Phyllirea angustifolia, Pistacia lentiskus, Juniperus oxicedrus and other dominate. The following higher zone is occupied by xeric oak-hornbeam forests. The two zones are ones mostly suffering from human activities – in many places those forests are destroyed and, in are replaced by farmlands or secondary xeric formations including mostly Paliurus spinacristi. The belt of the mesophilous oak-hornbeam and beech forests is developed at altitude from 900-1000 up to 1600-1800 m a.s.l. This vegetation is more abundant in the northern parts of Albania, where the mesophilous forests form the upper forest border. Normally in Northern Albania the coniferous communities are above the beech forests, but elsewhere inversions in the distribution of the deciduous and coniferous forests, owing to local humidity conditions, may occur. The orophytic (subalpine and alpine) grass belt starts from 1900-2000 m a.s.l. It may be noted that the intrazonal riverside communities consist mainly of Platanus orientalis.

Treatment of taxa The arrangement of the taxa followed the order used in the recent Palaearctic Catalogue (Löbl & Smetana, 2003), with only two exceptions: genus Sinechostictus (Bembidiini) is regarded as separate genus, according to the recent studies on the phylogeny of Trechitae (Grebennikov, 1997; Grebennikov & Maddison, 2005), and the arrangement of Tachyini follows Sciaky & Vigna Taglianti (2003). Synonyms are included if they consider taxa from Albania.

Material and depositories of the collections So far, faunal records for Albania were found in 279 publications, not all of them including original or detailed data. The main part of the studied materials are kept in three large Central European museums, which preserve the most diverse collections of ground beetle species from the Balkan Peninsula, as well as type material from the region. Common feature of these collections is that the principal part of the Albanian material had been collected before 1942. Both the most diverse and most numerous materials collected from the country are these in the Museum für Naturkunde der Humboldt Universität in Berlin. There, the collection of A. Bischoff is the most outstanding one; other notable collections are these of E. v. Oertzen and H. Hopp, V. Apfelbeck 10


and A. Winneguth, A. Penther and H. Zerny (most part), H. Priesner (most part), L. Mader, C. Lona, J. Müller. F. Hieke, G. Müller-Motzfeld and L. Behne (Hieke et al., 1986), as well as the materials collected in the second half of XX century by H. Peks and S. Peks. The second important depository is that in the Naturhistorisches Museum Wien, Vienna, Austria. From aspect of the species diversity the materials from Albania there are comparable with these in Berlin. The most significant ground beetle collections in Vienna are these of V. Apfelbeck and his laboratory assistant A. Winneguth, E. Csiki, A. Penther and H. Zerny, H. Priesner, L. Mader (most part), A. Bischoff and A. Winkler (most part). Additionally, small parts of the collections of Matzenauer, C. Ravasini and C. Lona, J. Müller, L. Weirather and J. Meschnigg are also housed in Vienna. Another place of important carabid collection is the Hungarian Natural History Museum, Budapest. The most important collections here are these of V. Apfelbeck and A. Winneguth, E. Csiki (most part) and Xh. Murraj. Other small collections housed in Budapest are those of M. Grabowski, L. Mader, C. Ravasini and C. Lona, A. Winkler and A. Bischoff. Recently, Fehér et al. (2004) also collected material of Carabidae from Albania that is currently in process of preparation. An important place as depository of type material from the Balkans is the collection of the Deutsches Entomologisches Institut in Müncheberg, Germany; some of the types deposited there have been reviewed. Materials from one American and two Balkan museums have been examined too. The most substantial collection among three is that housed in the National Museum of Natural History in Sofia. It has been collected by Bulgarian expeditions to Albania after 1992 and at present it is containing about 160 species. The other two collections are: i/ the one in the Museum of Natural Sciences, Tiranë (Albania), which in part was identified and published by Murraj (1968), and ii/ that in the Carnegie Museum of Natural History, Pittsburgh (USA), consisting of recently collected material of 57 species.1

Acknowledgments The most important part of the material was examined in the course of the author’s work under the European Union-funded Integrated Infrastructure Initiative “Synthesys”, which is actually the prime cause for the completion of this work (Applications DE-TAF-725, AT-TAF-758 and HU-TAF-817). For the work with the collec1

Except the aforementioned collections, there are other four European museums that may house more or less large number of Carabidae from Albania. At the present time a significant part of the former collection of V. Apfelbeck is preserving in the Zemaljski muzej Bosne i Hercegovine in Sarajevo (Bosnia Herzegovina). The Museo Civico di Storia Naturale in Trieste (Italy) and the Museo Civico di Storia Naturale in Milano (Italy) house the most part of the materials collected by the Italian expeditions in Albania before the Second World War. The first one is remarkable with the collections of C. Ravasini, C. Lona and J. Müller, and the second one - with that of A. Schatzmayr. Most probably, the collection of the Narodni Museum in Prague (Czech Republic) also contains materials from Albania. ACKNOWLEDGEMENTS

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tions and the materials on loan I am indebted to Prof. V. Andoni (MNST), Dr. P. Beron (NMNHS), Dr. R. Booth (BMNH), Mr. R. Davidson (CMNH), Mr. B. Jaeger (ZMHU), Dr. A. Popov (NMNHS), Dr. H. Schönmann (NMW), Dr. Gy. Szel (HNHM), Dr. M. Uglih (ZMHU) and Dr. L. Zerche (DEI). With pleasure, I record my deep appreciation to the colleagues who contributed this study giving their unpublished data (marked with the paragraphs “Material included” or “Additional material included”): in the first place Prof. Dr. A. Vigna Taglianti (Università di Roma”La Sapienza”, Roma, Italy) as well as Mr. P. Bulirsch (Prague, Czech Republic), Dr. F. Hieke (ZMHU), Mr. S. Hristovski (Institute of Biology, Skopje, Republic of Macedonia), Mr. B. Jaeger (ZMHU), Prof. Xh. Murraj (Tiranë, Albania) and Mr. D.W. Wrase (Berlin, Germany). It is worth mentioning that, Prof. Dr. A. Vigna Taglianti told me that several years ago he began the preparation of a checklist of the Carabidae of Albania, being still not completed. Generously, he entrusted to me the draft of this work which included a moderate number of records collected from the Italian expeditions to Albania carried out during the last 15 years. Dr. F. Hieke made valuable comments and improvements in the part about genus Amara, as well as kindly provided me the data of both unavailable papers and papers in press. The following colleagues also revised and improved different parts of the catalogue: Dr. P. Bonavita (Bembidiini; Università di Roma”La Sapienza”, Roma, Italy), Mr. P. Bulirsch (Scaritinae), Mr. J. Gebert (Cicindelinae; Rohne, Germany), Dr. P.M. Giachino (Speluncarius, Tapinopterus; Settore Fitosanitario Regionale, Torino, Italy), Mr. B. Jaeger (Stenolophina), Mr. P. Neri (Bembidiini; Forlí, Italy), Dr. B. Kataev (Harpalini; ZISP), Mr. L. Picciau (Speluncarius, Tapinopterus; Museo Regionale de Scienze Naturali, Torino), Dr. L. Toledano (Bembidiini; Museo Civico di Storia Naturale, Verona, Italy), Prof. Dr. A. Vigna Taglianti (whole draft, and especially Bembidiini) and Mr. D.W. Wrase (whole draft, and especially Harpalini). For the determination of material, I thank Dr. D. Fedorenko (Dyschirius; Moscow Pedagogical State University, Moscow, Russia), Dr. F. Hieke (Amara), Dr. B. Kataev (Anisodactylus, Harpalus, Ophonus, Trichotichnus, Acupalpus, Bradycellus), both Mr. G. Ledoux and Mr. Ph. Roux (Nebria; Paris, France), Mr. P. Neri (Bembidion), Mr. M. Pavesi (Syntomus, Olisthopus; MSNM, Italy) and Mr. D.W. Wrase (Notiophilus, Harpalus, Ophonus, Parophonus, Microlestes). Special thanks are due to Prof. Dr. A. Vigna Taglianti, Mr. D.W. Wrase, and Mr. I. Stoyanov (Central Laboratory of General Ecology, Sofia, Bulgaria), who made careful reviews and proposed important changes of the earlier versions of the manuscript on which this paper is based. Willingly, again Mr. I. Stoyanov helped me with the revision of the English text. Prof. Dr. A. Casale (Università di Sassari, Sassari) kindly allowed me the reproduction of his drawing of Abanotrechus beroni, and Dr. P. Stoev (NMNHS) gave me the basis of the map. I owe deep gratitude to my late farther for his strong and enthusiastic support in the initial stages of the work, as well as to my mother and my wife for their tolerance.

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Abbreviations I. Abbreviations of collections studied upon the European Union-funded Integrated Infrastructure Initiative “Synthesys” by the author DEI Deutsches Entomologisches Institut, Müncheberg, Germany HNHM Hungarian Natural History Museum (Magyar Természettudományi Múzeum), Budapest, Hungary MNHUB Museum für Naturkunde der Humboldt Universität zu Berlin, Bereich Zoologisches Museum, Berlin, Germany NMW Naturhistorisches Museum Wien, Vienna, Austria II. Abbrevuiations of other collections studied by the author BMNH The Natural History Museum, London, United Kingdom CMNH Carnegie Museum of Natural History, Pittsburgh, USA MNST Museum of Natural Sciences, Tirana, Albania NMNHS National Museum of Natural History, Sofia, Bulgaria III. Abbreviations of collections studied by other colleagues MCG Museo Civico di Storia Naturale “Giacomo Doria”, Genoa, Italy MHNG Muséum d’Histoire Naturelle, Geneva, Switzerland MNHP Muséum National d’Histoire Naturelle, Paris, France MSNM Museo Civico di Storia Naturale di Milano, Milano, Italy MTKD Museum für Tierkunde, Dresden, Germany NMPC National Museum Prague (Národni muzeum v Praze), Prague, Czech Republic ZISP Zoological Institute, St. Petersburg, Russia ZSM Zoologische Staatssammlung, Munich, Germany cAVT private collection A. Vigna Taglianti, Roma, Italy cBJ private collection B. Jaeger, Berlin, Germany cFC private collection F. Cassola, Rome, Italy cLR private collection G. Ledoux & P. Roux, Paris, France cPB private collection P. Bulirsch, Prague, Czech Republic cPN private collection P. Neri, Forlí, Italy cSH private collection S. Hristovski, Skopje, Macedonia cWH private collection W. Heinz, Schwanfeld, Germany cWR private collection D.W. Wrase, Berlin, Germany cXM private collection Xh. Murraj, Tiranë, Albania

ABBREVIATIONS

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IV. Other abbreviations and symbols used in the systematic part of the catalogue ! taxa still not well defined ? taxa with doubtful or questionable occurrence in Albania [ ] taxa that most probably inhabit Albania; they were recorded from neighbouring region # taxa incorrectly indicated for Albania, and therefore excluded from the catalogue f. female specimen/s m. male specimens/s s. specimen/s

Topographic treatment of some localities (first are the correct treatments of toponyms, then their most frequent variants in the examined references and collections) Buenë River = Bojana River Çukall Mt. = Cukali; Cucale Dukati = Fush-Dukati; Fushes Dukati; Dukat-Fshat Durrës = Durazzo Gjalicë Mt. or Gjalicë e Lumes = Montes Gjallica Ljums; Djalica Ljums Gjirokastër = Argirokastro Gostilë = Köstil Ibë (? Ibë e Postmë; ? Ibë e Sipërme) = Iba Karbunara e Madhe = Karbunara Kukës = Kukësi Lezhë = Alessio Liqen i Shkodrës = Shkodër Lake Llogora = Logora; Lungara Lura Mt. = Kunora e Lurës Lurë e Vjetër = Lurja Lushnje = Lushnia Maja e Rrabës = “Raba Massiv” Mal i Dajtit Mt. = Dajti Mt.; Dajt Massif Mal i Shenjt Mt. = Mal i Shêit Maranajt Mt. = Maranaj Nangë = Nangat North Albanian Alps = Prokletija Peshtrik Mt. = Paštrik; Pashtrik Rapshë = Rapša; Rapsa Shengjin = San Giovanni di Medua; Medua Shëndelli Mt. = Shendeli; Shen Ili 14


Shkëlzen Mt. = Shkölsen; Skölsen Shkodër = Scutari Stan-Karbunarë = Star Karbunara Tejas = Radomir (village in East Albania, Korab Mt.) Tepelenë = Tepelene Tërpan = Terpan Theth = Thethi Vermosh = Vermosa Vijosë River = Vijosa River Vlorë = Valona; Avlona Zepë Mt. = Zebia; Zebja

TOPOGRAPHIC TREATMENT OF SOME LOCALITIESEDITORIAL

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SYSTEMATIC PART Family CARABIDAE Latreille, 1802 Subfamily NEBRIINAE Laporte, 1834 Tribe NEBRIINI Laporte, 1834 1. Genus Eurynebria Ganglbauer, 1891 Notes. New genus for the fauna of Albania. 1.1. Eurynebria complanata (Linnaeus, 1767) Material studied. “Apfelb. Albanien”, 4 s. (MNHUB); “Albania: Valona.” / “complanata orientalis Typus! Apfb.”, 1 s. (HNHM); “Valona. Albania” / “Typus! Apf.”, 1 s. (HNHM); “Valona” / “v. albanica Cotypus! Apfb”, 4 s. (HNHM); “Südalbanien Kopstein” / “Eurynebria complanata var. albanica Apf.”, 2 s. (NMW); “Alban. Wght. Valona” / “Eurynebria complanata var. albanica Apf.”, 8 s. (NMW); “Alban. Wght. Valona”, 4 s. (MNHUB); “Albanien Avlona 08” / “E. complanata albanica Coll. Apfelbeck”, 4 s. (HNHM); “Valona Albanien V.1908 H. Hopp”, 2 s. (MNHUB); “Ruskuli Alb., Mader” / “Eurynebria complanata var. albanica Apf.”, 2 s. (NMW); “Shëng… 25.X.1965 kol. Murraj”, 2 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Shën Pjeter, Durrës District, 29.V.2004, 1 f., A. Paparistos leg. (cAVT). Notes. Eurynebria complanata orientalis Apfelbeck, E. complanata albanica and E. complanata var. albanica Apfelbeck are nomen nudum. The species was recorded hitherto only for European Turkey on the Balkans (Farkač & Janata, 2003: 78). New species for the fauna of Albania.

2. Genus Leistus Frölich, 1799 Subgenus Leistus s.str. 2.1. Leistus fulvibarbis Dejean, 1826 2.1.1. Leistus fulvibarbis fulvibarbis Dejean, 1826 References. Leistus fulvibarbis: Apfelbeck, 1904: 50 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1943b: 94 (Stan-Karbunarë); Perrault, 1986: 58, fig. 3; Hieke & Wrase, 1988: 21; Perrault, 1991: 198, fig. 22. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Skutari Bojana”, 6 s. (NMW; MNHUB); “Apfelb. Alban. spt. IV.-V.’05”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Logara Juni”, 1 s. (MNHUB); “leg. Bischoff Albania 1934” / “Logara”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); Tiranë Park, 1-9.IX.1997, 7 s., R. Ward leg. (CMNH). Notes. Farkač & Janata (2003: 82) have omitted L. fulvibarbis for Albania. Here, the species is confirmed for the country. 16


2.2. Leistus nitidus (Duftschmid, 1812) References. Leistus nitidus: Perrault, 1986: 58, fig. 3; Perrault, 1991: 198, fig. 23. Material studied. “Albania leg. Bischoff 1936” / “Shkëlzen”, 1 s. (MNHUB). Notes. Perrault displayed the species range as including North Albania (Perrault, 1986; Perrault, 1991). In the second work he noted also a single locality on the map. First detailed record for Albania. Subgenus Pogonophorus Latreille, 1802 2.3. Leistus apfelbecki Ganglbauer, 1891 Material included (studied A. Vigna Taglianti). Valbona, Lugu i Cirkut Mt., 2000 m, 16.VII.1993, 1 m., L. Latella leg. (cAVT). Notes. New species for the fauna of Albania. 2.4. Leistus magnicollis Motschulsky, 1866 References. Leistus magnicollis: Reitter, 1884: 36; Apfelbeck, 1904: 50; Apfelbeck, 1907a: 498, 501 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 21; Perrault, 1992: 21, fig. 11. Material studied. “Oroši (Ptr.) Alban. spt.”, 2 s. (NMW); “Merdita Oroschi”, 2 s. (HNHM); “Alban. Exp.’18 Pashtrik 4-14.7.”, 1 s. (NMW); “Albania leg. Bischoff 1935” / “Koritnik”, 1 s. (MNHUB); “Albania leg. Bischoff 1935” / “Llogora”, 3 s. (MNHUB); “Giallica” / “Albania leg. Bischoff 1936”, 9 s. (MNHUB); “Albania leg. Bischoff 1936” / “Shkalla Haruse” / “Korab”, 1 s. (MNHUB); “Cafa e Sharit Iballe” / “Albania leg. Mueller 1939”, 7 s. (MNHUB); “Maja e Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 s., L. Penev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). Notes. Farkač & Janata (2003: 83) have omitted L. magnicollis for Albania. Here the species is confirmed for the country. 2.5. Leistus parvicollis Chaudoir, 1869 References. Leistus parvicollis: Perrault, 1992: 18-19, 22, fig. 12; Farkač & Janata, 2003: 83. Material studied. “Oroši (Ptr.) Alban. spt.”, 1 s. (NMW); “Alban. spt. Maranai”, 1 s. (HNHM); “Alban. Exped. Kula Ljums 15-20.VIII.’18”, 1 s. (NMW); “Albania leg. Bischoff 1935” / “Cukale”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 7 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 7 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Mal i Krujes”, 2 s. (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Mal i Dajtit”, 1 s. (MNHUB); “Mali i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); Gjirokastër, 6.V.1994, 1 s., P. Stoev leg. (NMNHS). Notes. For the present, the differentiation of L. parvicollis from L. spinibarbis s.l. bases only on the ratio of the maximal width of head and the width of pronotum base (Perrault, 1992: 19). First detailed records for Albania.

COLEOPTERA: CARABIDAE: NEBRIINAE

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2.6. Leistus rufomarginatus (Duftschmid, 1812) References. Leistus rufomarginatus: Apfelbeck, 1907a: 498 (Oroshi); Hieke & Wrase, 1988: 21; Farkač & Janata, 2003: 84. Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.-V.’05”, 5 s. (NMW); “Albania leg. Bischoff 1932” / “Mal i Dajit”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “TomorBuranj Juli 1932”, 1 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 1 s. (MNHUB). 2.7. Leistus spinibarbis (Fabricius, 1775) 2.7.1. Leistus spinibarbis rufipes Chaudoir, 1843 [= punctatus Reitter, 1885] References. Leistus spinibarbis rufipes: Apfelbeck, 1904: 49 (Vlorë; Berat); Perrault, 1992: 18, 22 (fig. 12); Farkač & Janata, 2003: 84. Leistus spinibarbis: Anonymous, 1907: 279; Hieke & Wrase, 1988: 21. Leistus spinibarbis punctatus: Apfelbeck, 1909: 297 (Vlorë). Material studied. “Latif Skutari”, 2 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt. IV.-V.’05”, 12 s. (NMW); “Merdita Munela”, 4 s. (NMW); “Alban. spt. Maranai”, 9 s. (HNHM; MNHUB); “Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 9 s. (NMW); “Albania leg. Bischoff 1932” / “IgeEzren”, 1 s. (MNHUB); “Albania leg. Bischoff 1932” / “Kjori”, 2 s. (MNHUB); “leg. Bischoff Albania 1934” / “Mal i Krues”, 10 s. (MNHUB); “Giallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 24 s. (MNHUB); “Mal i Dajit” / “Albania leg. Bischoff 1936”, 11 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 23 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “23.IV.1958 leg. Xh. Murraj”, 10 s. (HNHM); Mal i Dajtit Mt., 10-11.V.1965, 4 s. (MNST). Notes. Except the material studied above I have seen more specimens in MNHUB, which remained unregistered.

3. Genus Nebria Latreille, 1802 Subgenus Alpaeus Bonelli, 1810 Notes. The systematic of the Balkan species from this group is complex and still not well settled. Comparison carried out between specimens from all the subspecies of Nebria aetolica Apfelbeck, 1901 s.l. in HNHM revealed that some of them are more closely related each other than to others. The subspecies have vicarious distribution in the alpine zones of the high mountains of South Albania, Southwestern Macedonia, North and Central Greece. Details in the pronotum form manifested that N. aetolica galicica, N. aetolica macedonica, N. aetolica papingutica and N. aetolica rambouseki are more similar each other than any of them to the remaining subspecies. For the reason to display more natural relations in this group the following nomenclatorial acts are proposed: Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. of Nebria (Alpaeus) aetolica macedonica Mařan, 1938 Nebria (Alpaeus) macedonica galicica Horvatovich, 1974, new assessment of Nebria (Alpaeus) aetolica galicica Horvatovich, 1974 Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974, new assessment of Nebria (Alpaeus) aetolica papingutica Horvatovich, 1974 Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assessment of Nebria (Alpaeus) aetolica rambouseki Mařan, 1938 18


3.1. Nebria attemsi Apfelbeck, 1908 References. Nebria attemsi: Bänninger, 1925: 272; Mařan, 1938: 51 (Korab Mt.); Csiki, 1940: 210 (Korab Mt.); Ledoux & Roux, 1992: 258; Farkač & Janata, 2003: 85. Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 40 s. (HNHM); “Akad. Balk. Exp. Csiki 1918” / “Albania Mts. Korab”, more than 70 s. (HNHM); “Alban. Exped. Korab, 1918, 24-31.VII.”, more than 80 s. (NMW); “Albania leg. Bischoff 1932” / “Tomor”, 1 s. (MNHUB); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” / “Derzë”, 2 s. (MNHUB); “Albania leg. Bischoff 1935” / “Korab”, more than 50 s. (MNHUB). Notes. Apfelbeck published three descriptions of this species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. 3.2. Nebria bosnica Ganglbauer, 1889 3.2.1. Nebria bosnica jezercensis Horvatovich, 1974 References. Nebria bosnica jezercensis: Horvatovich, 1974b: 90-91 (Maja e Jezercë Mt., loc. typ.); Huber, 2001: 78 (Maja e Jezercë Mt.); Farkač & Janata, 2003: 85 (Maja e Jezercë Mt.). 3.2.2. Nebria bosnica kulti Mařan, 1939 References. Nebria apfelbecki kulti: Mařan, 1939a: 20-21 (Shkëlzen Mt., loc. typ.). Nebria bosnica: Csiki, 1940: 210 (Shkëlzen Mt.). Nebria bosnica kulti: Horvatovich, 1974b: 90 (Shkëlzen Mt.); Huber, 2001: 78 (Shkëlzen Mt.); Farkač & Janata, 2003: 85 (Shkëlzen Mt.). Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 5 s. (HNHM); “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 syntype (HNHM); “Albania leg. Bischoff 1936” / “Shkëlzën”, 15 s. (MNHUB). 3.2.3. Nebria bosnica prokletiensis Horvatovich, 1974 References. Nebria apfelbecki: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near Vermosh); Mařan, 1939a: 20 (North Albanian Alps: Grebeni near Vermosh). Nebria bosnica prokletiensis: Horvatovich, 1974b: 91 (North Albanian Alps, loc. typ., state attribution unspecified); Farkač & Janata, 2003: 85. Material studied. “Prokletija Gbt. Dr. Penther”, 1 paratype (HNHM). Notes. N. bosnica apfelbecki Ganglbauer, 1891 inhabits only mountains in Eastern Bosnia Herzegovina and Western Montenegro. 3.2.4. Nebria bosnica sturanyi Apfelbeck, 1906 References. Nebria sturanyi: Apfelbeck, 1906: 1662-1663 (Maranajt Mt., loc. typ.); Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1918c: 160 (Maranajt Mt.); Bänninger, 1925: 270; Mařan, 1939a: 20 (Maranajt Mt.); Pretner, 1978: 146, 166. Nebria bosnica sturanyi: Horvatovich, 1974b: 89 (Maranajt Mt.); Huber, 2001: 77-78 (Maranajt Mt.); Farkač & Janata, 2003: 85 (Maranajt Mt.). Material studied. “Latif Alban spt.”, 1 s. (HNHM); “Alban. spt. Maranai”, 4 s. (MNHUB); “Maranai Alban.-occ.” / “Coll. Apfelbeck”, 1 s. (HNHM); “Maranai G.” / “Apfelb. Alban. spt. 11.V.05”, 3 s. (HNHM); “Apfelb. Alban. spt. IV.-V.05” / “Maranai” / “sturanyi Apfelb.”, 15 s. (NMW); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 6 s., P. Beron & B. Petrov leg. (cLR; NMNHS). COLEOPTERA: CARABIDAE: NEBRIINAE

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? Nebria bosnica tresnjevikensis Horvatovich, 1974 Notes. This geographical race inhabits the Montenegrin part of the North Albanian Alps and may live in the northernmost alpine spot in Albania too. Huber (2001: 81, Abb. 3) showed map where the subspecies is shown to live in the country in spite of lack of exact locality. ? Nebria dahli (Duftschmid, 1812) ? Nebria dahli montenegrina Apfelbeck, 1904 References. Nebria dahli velebitica: Apfelbeck, 1904: 56 (“Sv. Troice bei Prisren”). Nebria dahli: Horion, 1941: 81. Notes. The locality “Sv. Troice bei Prisren” (Apfelbeck, 1904) is situated on the northern slopes of the Shar Planina Mountains, in the Kosovo Region of Serbia. Probably Horion (1941) followed the same record. N. dahli may live in Albania since it occurs in the massif of Durmitor (Apfelbeck, 1904 sub N. dahli montenegrina), and N. dahli velebitica Heyden, 1884 inhabits only in Croatia (Farkač & Janata, 2003: 85). 3.3. Nebria ganglbaueri Apfelbeck, 1905 3.3.1. Nebria ganglbaueri ganglbaueri Apfelbeck, 1905 References. Nebria ganglbaueri: Apfelbeck, 1905: 240 (Albania / Kosovo border: Koritnik Mt., alpine belt, loc. typ.); Apfelbeck, 1907b: 636 (Koritnik Mt., alpine belt, loc. typ.); Bänninger, 1925: 270. Nebria ganglbaueri ganglbaueri: Horvatovich, 1975: 272 (Koritnik Mt.); Farkač & Janata, 2003: 86 (Koritnik Mt.). Material studied. “Latif Koritnik” / “ganglbaueri Apfelb.”, 2 syntypes (NMW). Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b), the older one is correctly treated as valid. 3.3.2. Nebria ganglbaueri gyalicaensis Horvatovich, 1975 References. Nebria ganglbaueri: Csiki, 1940: 210 (Gjalicë Mt.). Nebria ganglbaueri gyalicaensis: Horvatovich, 1975: 275-276 (Gjalicë Mt., loc. typ.); Farkač & Janata, 2003: 86 (Gjalicë Mt.). Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype and 18 paratypes (HNHM); “Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 1 paratype (NMW); “Alban. Exp. 18. Gjalica Ljums 17-26.VI.”, 16 s. (NMW). 3.3.3. Nebria ganglbaueri korabensis Horvatovich, 1975 References. Nebria ganglbaueri: Csiki, 1940: 210 (Korab Mt.). Nebria ganglbaueri korabensis: Horvatovich, 1975: 274-275 (Korab Mt., loc. typ.); Farkač & Janata, 2003: 86. Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 holotype and 22 paratypes (HNHM); “Alban. Exped. Korab 1918 24-31.VII.”, 23 s. (NMW); “Albania leg. Bischoff 1935” / “Korab”, more than 140 s., incl. 27 paratypes (MNHUB). Notes. Farkač & Janata (2003: 86) noted this subspecies for ex-Yugoslavia (Serbia and Montenegro). In fact its type locality is the Korab Mt. lying on the border between Albania and Macedonia. 3.3.4. Nebria ganglbaueri merditana Apfelbeck, 1906 References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Munella Mt., loc. typ.); Apfelbeck, 1907a: 500, 503 (Munella Mt.); Bänninger, 1925: 270; Mandl, 1964: 33-36; Horvatovich, 1974a: 359. Nebria ganglbaueri merditana: Horvatovich, 1975: 272-273 (Munella Mt.); Farkač & Janata, 2003: 86 (Munella Mt.). 20


Material studied. “Albania”, 1 s. (DEI); “Albania Merdita”, 1 s. designated incorrectly as syntype of Nebria ganglbaueri ditana Apfelbeck (DEI); “Merdita Munela” / “Coll. Apfelbeck”, 5 s. (HNHM); “Merdita Munela”, 1 s. designated incorrectly as syntype of Nebria ganglbaueri ditana Apfelbeck (DEI); “Merdita Munela”, 9 s. (MNHUB); “Merdita Munela” / “Vgth. 1906 Merdita”, 14 s. (NMW). Notes. Apfelbeck, 1906 described this taxon from two closely situated mountains (Munella Mt. and Zepë Mt.) situated in the area of Merdita. Also, he gave no exact “loc. typ.”. Later on, Horvatovich (1975) attributed the specimens from Zepë Mt. to the ssp. zebiaensis, so that “loc. typ.” of the ssp. merditana should be Munella Mt. Nebria ganglbaueri ditana Apfelbeck is either nomen nudum or incorrect spelling of the name merditana. In the second case the two specimens in DEI may be syntypes. 3.3.5. Nebria ganglbaueri zebiaensis Horvatovich, 1975 References. Nebria merditana: Apfelbeck, 1906: 1661-1662 (Merdita: Zepë Mt.); Apfelbeck, 1907a: 500, 503 (Zepë Mt.). Nebria ganglbaueri zebiaensis: Horvatovich, 1975: 273-274 (Zepë Mt., loc. typ.); Farkač & Janata, 2003: 86 (Zepë Mt.). Material studied. “Merdita Zebia” / “Coll.Apfelbeck”, 1 holotype and 4 paratypes (HNHM); “Merdita Zebia” / “Vgth. 1906 Merdita”, 2 s. (DEI). Notes. Further study on the validity of this race (with respect to the previous one) is needed. 3.4. Nebria germarii Heer, 1837 ! Nebria germarii ssp. Material studied. “Albania leg. Bischoff 1935” / “Maja e Jezeroe” / “Nebria!! wahrscheinlich” / “N. germari ssp. n. det. Dr. Horvatovich”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. For the present the subspecies status of the local population is not defined, but it probably tends to ssp. durmitorensis Apfelbeck, 1904. 3.5. Nebria macedonica Mařan, 1938 3.5.1. Nebria macedonica galicica Horvatovich, 1974 [= korabica Horvatovich, 1974] References. Nebria aetolica galicica: Horvatovich, 1974a: 360-361 (loc. typ. in Macedonian part of Galičica Mt.); Ledoux & Roux, 1992: 257 (Korab Mt.); Farkač & Janata, 2003: 84. Nebria aetolica korabica: Horvatovich, 1974a: 361 (loc. typ. in Macedonian part of Korab Mt.). Material studied. “Korab”, 1 paratype of Nebria aetolica korabica (HNHM). 3.5.2. Nebria macedonica papingutica Horvatovich, 1974 [= tomoricensis Horvatovich, 1974] References. Nebria aetolica tomoricensis: Mařan, 1945: 56 (Tomor Mt.); Horvatovich, 1974a: 364-365 (Tomor Mt., loc. typ. of tomoricensis; Tomor Mt.: Abbas Ali). Nebria aetolica papingutica: Horvatovich, 1974a: 363-364 (Maja e Papingut Mt., loc. typ. of ssp. papingutica; Maja e Policanit Mt.); Ledoux & Roux, 1992: 257 (Maja e Papingut Mt.; Tomor Mt.); Farkač & Janata, 2003: 84. Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s., including 3 paratypes of Nebria aetolica tomoricensis (HNHM; NMW); “Tomor Monte Toik” / “Albania 1922 Ravasini, Lona”, 1 paratype of Nebria aetolica tomoricensis (HNHM); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 holotype and 4 paratypes of Nebria aetolica tomoricensis (HNHM); “Tomori Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 5 COLEOPTERA: CARABIDAE: NEBRIINAE

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paratypes of Nebria aetolica tomoricensis (NMW); “Tomori Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Albania leg. Bischoff 1932” / “Tojk Tomor”, 10 paratypes of Nebria aetolica tomoricensis (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e Papingut”, 1 holotype and 26 paratypes of Nebria aetolica papingutica (HNHM; MNHUB); “Albania leg. Bischoff 1937” / “Maja e Policanit”, 2 paratypes of Nebria aetolica papingutica (HNHM); “Albania leg. Bischoff 1937” / “Mal i That”, 2 s. (HNHM). [Nebria macedonica rambouseki Mařan, 1938] Notes. This geographical race was described from the Macedonian part of the Jablanitsa Mt. (Mařan, 1938: 57; Mařan, 1945: 56). So far there is no sure record from Albania. Subgenus Boreonebria Jeannel, 1937 3.6. Nebria rufescens (Strøm, 1768) 3.6.1. Nebria rufescens rufescens (Strøm, 1768) [= gyllenhali (Schönherr, 1806)] References. Nebria gyllenhali: Hieke & Wrase, 1988: 22. 12. Nebria rufescens rufescens: Farkač & Janata, 2003: 88. Material studied. “Albania leg. Bischoff 1935” / “Shkëlzen”, 4 s. (MNHUB). Notes. First detailed record for Albania. Subgenus Eunebria Jeannel, 1937 3.7. Nebria jockischi Sturm, 1815 3.7.1. Nebria jockischi jockischi Sturm, 1815 References. Nebria jockischi: Hieke & Wrase, 1988: 22. Material studied. “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania leg. Bischoff 1936” / “Korab”, 8 s. (MNHUB). Notes. Farkač & Janata (2003: 90) have omitted N. jockischi for Albania. The species is here confirmed for the country with first detailed records. Subgenus Nebria s.str. 3.8. Nebria brevicollis (Fabricius, 1792) References. Nebria brevicollis: Apfelbeck, 1904: 54 (Velipoja; Vlorë); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes; Lushnje; Stan-Karbunarë; Berat); Hieke & Wrase, 1988: 22; Farkač & Janata, 2003: 92. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë). Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s. (HNHM); “Albania Sauk” / “2.V.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Dajti” / “2.6.1959 leg. Xh. Murraj”, 1 s. (HNHM); Tiranë, 10.V.1961, 1 s. (MNST); Mal i Dajtit Mt., 10-11.V.1965, 3 s. (MNST); Tiranë Park, 8.III.1998, 54 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH; NMNHS); “Albania Tomor Mts. Kajala e Tomorit, 26.V.2004, K. Harmos & D. Muránya”, 1 s. (HNHM).

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Additional material included (studied A. Vigna Taglianti). Tiranë, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 3 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT); Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 m., 2 f., P. Crucitti & C. Cavalier leg. (cAVT). Notes. Except the material studied above I have seen additionally more than 20 s. from different localities in MNHUB. 3.9. Nebria kratteri Dejean, 1831 3.9.1. Nebria kratteri kratteri Dejean, 1831 [= valonensis Apfelbeck, 1904] References. Nebria kratteri: Apfelbeck, 1904: 52 (South Albania); Apfelbeck, 1907a: 498, 501, 503 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1928: 46; Holdhaus, 1956: 7 (Mal i Dajtit Mt.); Magistretti, 1965: 79; Ledoux & Roux, 1990: 94. Nebria kratteri valonensis: Apfelbeck, 1904: 55 (Vlorë); Bänninger, 1925: 274; Mařan, 1939a: 19 (Vlorë); Jeanne, 1974: 66. Nebria kratteri ssp.: Schatzmayr, 1943b: 94 (cave “Stere Gropa e Vasos” near Dukati). Nebria kratteri kratteri: Farkač & Janata, 2003: 92. Material studied. “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Mal-i-Dajtit Tiranë” / “Albania Bischoff 1931”, 2 s. (DEI); “Tomor: Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara juni” / “Albania Bischoff 1932”, 1 s. (DEI); “Tomor-Buranj juli 1932” / “Albania Bischoff 1932”, 1 s. (DEI). Notes. The record “Chani Driskos bei Jannina” (Apfelbeck, 1904: 55) refers to Northwestern Greece. The locality at Mal i Shenit Mt. (Apfelbeck, 1907a), representing the northernmost known spot in the species range, needs verification. In MNHUB, I have seen more than 100 specimens from different localities, which remained unregistered. # Nebria salina Fairmaire & Laboulbène, 1854 References. Nebria brevicollis var. salina: Apfelbeck, 1909: 297 (Vlorë). Notes. This Atlantic species does not inhabit the Balkan Peninsula, so that the record for it referred to N. brevicollis.

Tribe NOTIOPHILINI Motschulsky, 1850 4. Genus Notiophilus Dumeril, 1806 4.1. Notiophilus aestuans Dejean, 1826 [= pusillus C.R. Waterhouse, 1833] References. Notiophilus aestuans: Csiki, 1940: 212 (Gjalicë Mt.). Notiophilus pusillus: Hieke & Wrase, 1988: 24. Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania, Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 2 s. (NMW); “Mal i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB). Notes. Bousquet & Barševskis (2003: 96) have omitted N. aestuans for Albania. Here it is confirmed for the country. The record “Peristeri-Gebirge bei Jannina” (Apfelbeck, 1904: 61 sub N. pusillus) refers to Greece. COLEOPTERA: CARABIDAE: NEBRIINAE

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4.2. Notiophilus aquaticus (Linnaeus, 1758) References. Notiophilus aquaticus: Hieke & Wrase, 1988: 24. Material studied. “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Notes. Bousquet & Barševskis (2003: 96) have omitted the species for Albania. Here it is confirmed for the country with first detailed records. 4.3. Notiophilus biguttatus (Fabricius, 1779) References. Notiophilus biguttatus: Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 25. Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania leg. Bischoff 1935” / “Korab”, 3 s. (MNHUB); “Albania leg. Bischoff 1935” / “Maja e Jezercoë”, 3 s. (MNHUB); “Albania leg. Bischoff 1936” / “Gjallica”, 3 s. (MNHUB); “Albania leg. Bischoff 1936” / “Shkëlzën”, 3 s. (MNHUB); “Albania leg. Mueller 1939” / “Kunora Drunit Cucale”, 8 s. (MNHUB); “Albania leg. Mueller 1939” / “Maja e Shpatit Iballe”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Lushnje, Divjaka National Park, 10.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). Notes. Bousquet & Barševskis (2003: 96) have omitted the species for Albania. Here it is confirmed for the country. 4.4. Notiophilus danieli Reitter, 1897 References. Notiophilus danieli: Apfelbeck, 1904: 64 (Vlorë); Ganglbauer, 1904: 648; Apfelbeck, 1909: 297 (Vlorë); Magistretti, 1965: 84; Hieke & Wrase, 1988: 25. Material studied. Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is confirmed for the country. 4.5. Notiophilus germinyi Fauvel, 1863 [= hypocrita Putzeys, 1866] References. Notiophilus hypocrita: Hieke & Wrase, 1988: 24. Material studied. “Alban. Exp.’ 18 Pashtrik 4-14.7.”, 3 s. (NMW); “Koritnik” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB). Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is confirmed for the country with first detailed records. 4.6. Notiophilus palustris (Duftschmid, 1812) References. Notiophilus palustris: Csiki, 1940: 212 (Gjalicë Mt.). Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM). Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is confirmed for the country.

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4.7. Notiophilus rufipes Curtis, 1829 References. Notiophilus rufipes: Apfelbeck, 1904: 64 (Velipoja near Lezhë); Hieke & Wrase, 1988: 25. Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “TomorBuranj Juli 1932” / “Albania Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); “Maja e Shpatit Iballe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); Tiranë Park, 1-9.IX.1997, 2 s., R. Ward leg. (CMNH). Notes. Bousquet & Barševskis (2003: 97) have omitted the species for Albania. Here it is confirmed for the country. 4.8. Notiophilus substriatus C.R. Waterhouse, 1833 References. Notiophilus substriatus: Csiki, 1940: 212 (Kula Ljums; foot of Gjalicë Mt.); Hieke & Wrase, 1988: 25. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Logara Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murrsi” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Tiranë Park, 22-31.X.1997, 1 s., R. Ward leg. (CMNH); Tiranë Park, 1-9.IX.1997, 1 s., R. Ward leg. (CMNH); Tiranë Park, 1218.XII.1997, 1 s., R. Ward leg. (NMNHS). Notes. Bousquet & Barševskis (2003: 98) have omitted the species for Albania. Here it is confirmed for the country.

Subfamily CICINDELINAE Latreille, 1802 Tribe CICINDELINI Latreille, 1802 5. Genus Calomera Motschulsky, 1862 5.1. Calomera fischeri (M. F. Adams, 1817) 5.1.1. Calomera fischeri fischeri (M. F. Adams, 1817) References. Cicindela fischeri: Apfelbeck, 1907a: 496, 501 (Kiri River near Mesi); Csiki, 1940: 209 (Kula Ljums); Hieke & Wrase, 1988: 11. Material studied. “Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); Shëngjin, 16.V.1963, 1 m. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS). Notes. Puchkov & Matalin (2003: 100) have omitted the species for Albania. Here it is confirmed for the country.

COLEOPTERA: CARABIDAE: CICINDELINAE

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5.2. Calomera littoralis (Fabricius, 1787) 5.2.1. Calomera littoralis nemoralis (Olivier, 1790) References. Cicindela lunulata var. nemoralis: Apfelbeck, 1894: 546. Cicindela lunulata: Murraj, 1968: 33 (Golem; Suhagora; Theth; Tomor Mt.). Lophyridia nemoralis nemoralis: Mandl, 1981: 150; 154. Calomera littoralis nemoralis: Puchkov & Matalin, 2003: 100. Material studied. Golem, 19.V.1958, 2 m. (MNST); Golem, 24.III. 1961, 1 m., 1 f. (MNST); Mal i Dajtit Mt.: Shkall Priskë, 30.VI.1960, 1 m. (MNST); Shëngjik, 20.VII.1961, 1 m. (MNST); Shëngjik, 16.V.1963, 2 m., 2 f. (MNST); Laguna Patog, 18.IX.1993, 4 s., S. Beshkov leg. (NMNHS); Velipoja, 19.IX.1993, 5 s., S. Beshkov leg. (NMNHS). Notes. Part of the material cited by Murraj (1968) was found and revised. Calomera lunulata (Fabricius, 1781) is a valid species with primarily North African range, in Europe it lives only in Sicily (Puchkov & Matalin, 2003: 101). [genus Cephalota Dokhtouroff, 1883] [subgenus Taenidia Rivalier, 1950] [Cephalota circumdata (Dejean, 1822)] [Cephalota circumdata circumdata (Dejean, 1822)] Notes. Most probably, the species inhabits the southern seacoast of Albania (see also Gebert, 1999: 30, Abb. 1.2.).

6. Genus Cicindela Linnaeus, 1758 Subgenus Cicindela s.str. 6.1. Cicindela campestris Linnaeus, 1758 6.1.2. Cicindela campestris olivieria Brullé, 1832 References. Cicindela campestris: Apfelbeck, 1904: 6 (Velipoja near Lezhë; Vlorë; Berat; Durrës); Apfelbeck, 1907a: 496 (Kiri River near Mesi); Apfelbeck, 1909: 297 (Vlorë; Berat); Csiki, 1940: 209 (Gjalicë Mt.; Korab Mt.); Murraj, 1968: 33 (Tiranë; Lurë; Sauk; Pashaliman; Kulla e Lumës); Hieke & Wrase, 1988: 9. Cicindela campestris pontica: Schatzmayr, 1943b: 93 (Karbunara e Madhe; Lushnje; Tërpan; “Poggio Boschetto”). Cicindela campestris olivieria: Mandl, 1944: 7; 12; 13 (Tiranë; Korab Mt.); Puchkov & Matalin, 2003: 104. Cicindela campestris campestris: Puchkov & Matalin, 2003: 103. Material studied. Dukati, 7.V.1958, 1 m. (MNST); Golem, 19.V.1958, 1 f. (MNST); Golem, 8.VII.1965, 1 f. (MNST); Mali-me Gropa, 3.VII.1958, 1 f. (MNST); Tiranë, 4.V.1959, 1 m. (MNST); Tiranë, 10.IV.1965, 1 f. (MNST); Sauk, 9.VII.1961, 1 f. (MNST); Sauk, 11.IV.1964, 2 m. (MNST); Sauk, 30.10.1963, 1 f. (MNST); Lurë, 18.VII.1961, 1 m. (MNST); Ibë, 24.IV.1962, 1 f. (MNST); Mal i Dajtit Mt., 10.V.1965, 3 f. (MNST); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS); North Albanian Alps, near cave B 52, 1600 m, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS); Petrela 15 km NE Tiranë, 300 m, 9.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). 26


Additional material included (studied F. Cassola). Rapsh, 15.V.2002, 1 s., A. Paparistos leg. (cFC); Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cFC); Vasjar (Tepelenë District), 18.IV.2003, 1 m., P. Crucitti & C. Cavalieri leg. (cFC). Notes. By the lack of modern study, the intraspecific relations of C. campestris s.l. in the Balkan Peninsula are obscure. For this reason all the data about it from Albania are referred to ssp. olivieria provisionally. The ssp. pontica Fischer von Waldheim, 1828 inhabits only the Black Sea Coast, so that the record for this race is referred to ssp. olivieria. Except the material studied above I have also seen additional specimens from Gjallica Lums and the Korab Mt. in HNHM. # Cicindela hybrida Linnaeus, 1758 References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke & Wrase, 1988: 10. Notes. I follow the view of Gebert (1995) and refer the literature records for C. hybrida to C. monticola albanica. 6.2. Cicindela monticola Menetries, 1832 6.2.1. Cicindela monticola albanica Apfelbeck, 1909 References. Cicindela hybrida: Apfelbeck, 1904: 5 (Vlorë); Csiki, 1940: 209 (Kula Ljums); Hieke & Wrase, 1988: 10. Cicindela hybrida albanica: Apfelbeck, 1909: 297 (Vlorë, loc. typ.; Berat, loc. typ.); Apfelbeck, 1917: 219 (Kiri River near Mesi, loc. typ.; Merdita: Munella, loc. typ.; Vlorë, loc. typ.); Apfelbeck, 1918a: 75 (Shkodër, loc. typ.; Orish, loc. typ.; Vlorë, loc. typ.); Apfelbeck, 1930: 161; Murraj, 1968: 33 (Golem; Suhagora; Tomor Mt.; Shëngjin); Cassola, 1973: 26. Cicindela sahlbergi albanica: Gebert, 1995: 16 (Buenë River near Shkorër; Mesi; Vlorë; Merdita: Munella; Berat). Cicindela monticola albanica: Puchkov & Matalin, 2003: 106. Material studied. “Skutari Bojana” / “C. v. albanica Apfb. Coll. Apfelbeck”, 2 s. (HNHM); “Albania Kula Ljums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania coll. Lichtn.”, 1 s. (HNHM); Shkumbi - Kavajes, 7.V.1958, 1 f. (MNST); Golem, 24.III.1961, 2 m., 3 f. (MNST); Divjake, 18.VI.1964, 1 f. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 2 s., B. Guéorguiev leg. (NMNHS); Leskovac - Barmachi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Apfelbeck published four consecutive descriptions of the ssp. albanica (Apfelbeck, 1909; Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930: 161), the oldest one is correctly treated as valid. 6.3. Cicindela sylvatica Linnaeus, 1758 6.3.1. Cicindela sylvatica sylvatica Linnaeus, 1758 References. Cicindela sylvatica sylvatica: Puchkov & Matalin, 2003: 107. 6.4. Cicindela transversalis Dejean, 1822 6.4.1. Cicindela transversalis transversalis Dejean, 1822 [= riparia Dejean, 1822] References. Cicindela hybrida riparia: Apfelbeck, 1907a: 496 (Kiri River near Mesi). Notes. Puchkov & Matalin (2003: 108) have omitted the species for Albania, therefore its occurrence in Albania needs further confirmation. COLEOPTERA: CARABIDAE: CICINDELINAE

27

7. Genus Cylindera Westood, 1831 Subgenus Cylindera s.str. 7.1. Cylindera germanica (Linnaeus, 1758) 7.1.1. Cylindera germanica germanica (Linnaeus, 1758) References. Cicindela germanica: Hieke & Wrase, 1988: 10. Cylindera germanica germanica: Puchkov & Matalin, 2003: 109. 7.1.2. Cylindera germanica muelleri (Magistretti, 1966) References. Cylindera germanica muelleri: Puchkov & Matalin, 2003: 109. Subgenus Eugrapha Rivalier, 1950 7.2. Cylindera arenaria (Füessly, 1775) 7.2.1. Cylindera arenaria viennensis (Schranck, 1781) References. Cicindela arenaria: Hieke & Wrase, 1988: 11. Cylindera arenaria viennensis: Puchkov & Matalin, 2003: 110. 7.3 Cylindera trisignata (Dejean, 1822) 7.3.1. Cylindera trisignata trisignata (Dejean, 1822) References. Cicindela literata sic!: Murraj, 1968: 33 (Shëngjin). Cylindera trisignata: Cassola, 1973: 33, Fig. 2. Cylindera trisignata trisignata: Puchkov & Matalin, 2003: 111. Material studied. Shëngjin, 16.V.1963, 4 m., 4 f. (MNST); Radona, Shkiela, 21.VI.1994, 8 s., B. Guéorguiev leg. (NMNHS). Notes. Cicindela litterata Sulzer, 1776 is synonym of Cylindera arenaria. Revision of material published by Murraj (1968) revealed that it is identical with C. trisignata.

8. Genus Myriochila Motschulsky, 1858 Notes. New genus for the fauna of Albania. Subgenus Myriochila s.str. Notes. New subgenus for the fauna of Albania. 8.1. Myriochila melancholica (Fabricius, 1798) 8.1. Myriochila melancholica melancholica (Fabricius, 1798) Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 815.VII.2006, 1 s., M. Komnenov leg. (cSH). Notes. New species for the fauna of Albania.

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Subfamily CARABINAE Latreille, 1802 Tribe CARABINI Latreille, 1802 9. Genus Calosoma Weber, 1801 Subgenus Calosoma s.str. 9.1. Calosoma inquisitor (Linnaeus, 1758) 9.1.1. Calosoma inquisitor inquisitor (Linnaeus, 1758) References. Calosoma inquisitor inquisitor: Bousquet et al., 2003: 119. Material studied. “Albania Rausa Dr. Grabowski”, 2 s. (HNHM); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Mal i Dajtit Mt., 27.V.2000, 1 f., A. Paparistos leg. (cAVT). Notes. First detailed records for Albania. 9.2. Calosoma sycophanta (Linnaeus, 1758) References. Calosoma sycophanta: Murraj, 1968: 33 (Sauk; Voskopojë; Shkodër); Hieke et al., 1986: 183 (Borshi near Vlorë); Bousquet et al., 2003: 120. Notes. Due to misidentifications found in the collection of Xh. Murraj, his literature record for C. sycophanta (Murraj, 1968) is included here with minimal reservation because of the easily recognition of this species. Subgenus Microcallisthenes Apfelbeck, 1918 9.3. Calosoma pentheri Apfelbeck, 1918 References. Calosoma pentheri: Apfelbeck, 1918c: 160-163 (eastern part of the North Albanian Alps, 1700 m, loc. typ.); Bousquet et al., 2003: 121. Material included (studied D.W. Wrase). “Prokletia-Geb. Alban., leg Popp”, 1 m. (cWR). 9.4. Calosoma relictum Apfelbeck, 1918 [= pentheroides Roubal, 1932] References. Calosoma relictum var. pentheroides: Roubal, 1932: 19. Calosoma relictum: Csiki, 1940: 209 (Korab Mt., Tejas); Bousquet et al., 2003: 121. Material examined. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 53 s. (HNHM); “Korab-Gebirge Albanien coll. Winkler”, 2 s. (HNHM).

10. Genus Carabus Linnaeus, 1758 Notes. I accept Březina’s (1999) “well geographical and morphologically defined” subspecies when dealing with the subgenera Megodontus, Procerus and Procrustes. COLEOPTERA: CARABIDAE: CARABINAE

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Subgenus Archicarabus Seidlitz, 1887 10.1. Carabus montivagus Palliardi, 1825 10.1.1. Carabus montivagus vellepiticus Hampe, 1850 [= sutomorensis Reitter, 1885] References. Carabus montivagus m. sutomorensis: Breuning, 1933: 677 (Vlorë; Korab Mt.); Breuning, 1978: 194. Carabus montivagus: Hieke & Wrase, 1988: 18. Carabus montivagus vellepiticus: Březina, 1999: 25; Bousquet et al., 2003: 129; Turin et al., 2003: 198. Subgenus Carabus s.str. 10.2. Carabus granulatus Linnaeus, 1758 10.2.1. Carabus granulatus interstitialis Duftschmid, 1812 [= aetolicus Schaum, 1857; miriditus Apfelbeck, 1901] References. Carabus granulatus var. miridita: Apfelbeck, 1901b: 121-122 (Lezhë, loc. typ.). Carabus granulatus miridita: Apfelbeck, 1904: 33-34 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë River Outfall); Monzini & Angelini, 1997: 39. Carabus granulatus aetolicus: Apfelbeck, 1904: 34 (Durrës; Vlorë); Vigna Taglianti et al., 2001: 72-73 (Velipoja near Lezhë). Carabus granulatus granulatus n. interstitialis: Breuning, 1933: 533 (Vlorë; Durrës); Breuning, 1978: 183. Carabus granulatus granulatus m. merdita sic!: Breuning, 1933: 534 (Velipoja near Lezhë); Breuning, 1978: 184. Carabus granulatus interstitialis v. miridita: Arnold, 1939: 158; Březina, 1999: 8 (Velipoja near Lezhë). Carabus granulatus m. miridita: Magistretti, 1965: 26. Carabus granulatus: Hieke & Wrase, 1988: 17. Carabus granulatus interstitialis: Bousquet et al., 2003: 136. Material included (studied A. Vigna Taglianti). Vlorë, 10.IV.1999, 1 f., A. Paparistos leg. (cAVT). Subgenus Chaetocarabus C.G. Thomson, 1875 10.3. Carabus intricatus Linnaeus, 1761 10.3.1. Carabus intricatus intricatus Linnaeus, 1761 [= montenegrinus Kraatz, 1876; chionobatus Born, 1918; weiratheri Breuning, 1934] References. Carabus intricatus montenegrinus: Apfelbeck, 1904: 28 (Durrës). Carabus intricatus var.: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m). Carabus intricatus: Klaptocz, 1910: 399 (Theth); Horion, 1941: 42. Ruspoli, 1965: 8; Geisthardt, 1975: 7; Hieke & Wrase, 1988: 16; Turin et al., 2003: 243 (North Albania). Carabus intricatus var. montenegrinus: Apfelbeck, 1918c: 160, 163 (North Albanian Alps, 1300-2000 m). Carabus intricatus chinobatus: Buresch & Kantardjieva, 1928: 84. Carabus intricatus intricatus m. montenegrinus: Breuning, 1934: 1043, 1045 (Merdita; Tiranë; Durrës; Vlorë). Carabus intricatus intricatus n. weiratheri: Breuning, 1934: 1046 (Vlorë); Magistretti, 1965: 45 (Vlorë); Breuning, 1978: 234 (Vlorë). Carabus intricatus intricatus var. weiratheri: Březina, 1999: 53. Carabus intricatus intricatus: Bousquet et al., 2003: 138. Material studied. Bizë, 27.VII.1966, 1 f. (MNST); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Razma, 1560 m, 12.VI.1995, 1 m., V. Sakalian leg. (NMNHS). 30


Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 1900 m, 10.VII.1993, 1 m., L. Latella leg. (cAVT). # subgenus Eucarabus Géhin, 1876 # Carabus catenulatus Scopoli, 1763 # Carabus catenulatus catenulatus Scopoli, 1763 [= catenatus Panzer, 1803; albanensis Géhin, 1885] References. Carabus catenatus var. albanensis: Apfelbeck, 1904: 35. Notes. Probably, the original record for this species from Albania is that in the description of albanensis Géhin, 1885. Apfelbeck (1904: 35) has based indirectly on this record, indicating no exact locality for the county. I cannot find the original description of this variation, and believe the species does not inhabit Albania (see also Turin et al., 2003). [subgenus Hygrocarabus C.G. Thomson, 1875] [Carabus variolosus Fabricius, 1787] [Carabus variolosus nodulosus Creutzer, 1799] References. Carabus variolosus nodulosus: Breuning, 1926: 19; Breuning, 1928a: 113; Buresch & Kantardjieva, 1928: 88. Carabus nodulosus: Turin et al., 2003: 240. Notes. Actually, the record “Albanien: Ljuboten” (Breuning, 1926: 22) concerns the territory of Republic of Macedonia (see also Breuning, 1935: 1169). The following authors had followed this data mentioning the species for Albania. In spite of this, C. variolosus may inhabit the marginal northern or northeastern regions of the country. [subgenus Limnocarabus Géhin, 1876] [Carabus clathratus Linnaeus, 1761] [Carabus clathratus auraniensis J. Müller, 1903] Notes. I cannot find any exact record for this species from Albania even though Turin et al. (2003: 155) showed in the map it occurs there. This rare Carabus-species may inhabit the country as Purkyně (1928) described the variation epirensis Purkyně, 1928 (synonym of C. clathratus auraniensis) from the Greek Epirus. Subgenus Megodontus Solier, 1848 10.4. Carabus caelatus Fabricius, 1801 10.4.1. Carabus caelatus caelatus Fabricius, 1801 [= sarajevoensis Apfelbeck, 1890; malissorum Apfelbeck, 1919; metalkanus Apfelbeck, 1919] References. Carabus caelatus sarajevensis sic!: Apfelbeck, 1907a: 498, 502 (Mal i Shenjt Mt.: top region, 1500 m). Carabus caelatus: Apfelbeck, 1907a: 500; Apfelbeck, 1917: 220 (Merdita); Apfelbeck, 1930: 162 (Merdita); Horion, 1941: 41; Březina, 1999: 73. Carabus caelatus malissorum: Apfelbeck, 1919a: 226-267 (North Albanian Alps, Maja e Jezerzë, loc. typ.); Apfelbeck, 1919b: 45-46 (North Albanian Alps, Maja e Jezerzë, loc. typ.). Carabus caelatus caelatus n. metalkanus: Breuning, 1935: 1297 (Merdita; Gjalicë e Lumes; Sarisaltik near Kruja; “Male Dojtitz”; Mal e Krujes); Breuning, 1978: 258. Carabus caelatus caelatus m. malissorum: Breuning, 1935: 1298 (North Albanian Alps; Maja e Jezercë); Breuning, 1978: 258. Carabus caelatus metalkanus: Arnold, 1939: 189. Carabus caelatus metalkanus v. malissorum: Arnold, 1939: 189. Carabus caelatus caelatus: Březina, 1999: 73; Bousquet et al., 2003: 163. Carabus caelatus sarajevoensis: Turin et al., 2003: 253. COLEOPTERA: CARABIDAE: CARABINAE

31

Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 2 m., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 3 m., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 1 m., 1 f., P. Dioli leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 1800 m, 9.VII.1993, 1 f., F. Sirtori leg. (cAVT). Notes. Apfelbeck published two description of the form malissorum (Apfelbeck, 1919; Apfelbeck, 1920), the older one is correctly treated as valid. 10.4.2. Carabus caelatus dalmatinus Duftschmid, 1812 [= procerus Reitter, 1885] References. Carabus caelatus dalmatinus: Apfelbeck, 1904: 25 (Velipoja near Lezhë); Wohlberedt, 1909: 127 (near Liqen i Shkodrës); Březina, 1999: 74; Bousquet et al., 2003: 163; Turin et al., 2003: 253 (Velipoja; Lezhë). Carabus dalmatinus procerus: Apfelbeck, 1918c: 160 (North Albanian Alps). Carabus caelatus var. procerus: Apfelbeck, 1918c: 163 (North Albanian Alps, 1300-2000 m). Carabus caelatus dalmatinus n. procerus: Breuning, 1935: 1300 (Lezhë District); Breuning, 1978: 258. Material included (studied A. Vigna Taglianti). Durrës, 13,VI.1998, 1 f., A. Paparistos leg. (cAVT). Notes. The mentioning of the locality “Vilajet” (Turin et al., 2003: 253) is misunderstanding. Actually this Turkish word is an equivalent to the English one “district”. 10.5. Carabus croaticus Dejean, 1826 10.5.1. Carabus croaticus durmitorensis Apfelbeck, 1904 [= ljubetensis Apfelbeck, 1917; babinjensis Apfelbeck, 1919] References. Carabus croaticus babinjensis: Apfelbeck, 1919a: 226 (North Albanian Alps, loc. typ.); Apfelbeck, 1919b: 45 (North Albanian Alps, loc. typ.); Krätschmer & Drovenik, 1977: 215 (Abb.1); Turin et al., 2003: 254 (Babinje Planina Mt.). Carabus croaticus ljubetensis n. ljubetensis: Breuning, 1935: 1291 (North Albanian Alps; Babinje Planina Mt.); Breuning, 1978: 257. Carabus croaticus ljubetensis m. durmitorensis: Breuning, 1935: 1291 (Korab Mt.); Breuning, 1978: 257. Carabus croaticus: Horion, 1941: 41; Tabaković-Tošić, 1988: 135; Březina, 1999: 74. Carabus croaticus durmitorensis: Březina, 1999: 74 (North Albanian Alps); Bousquet et al., 2003: 163. Carabus croaticus durmitorensis var. babinjensis: Březina, 1999: 74 (North Albanian Alps). Carabus croaticus durmitorensis var. ljubetensis: Březina, 1999: 74. Carabus croaticus ljubetensis: Turin et al., 2003: 254. Notes. Apfelbeck published three consecutive descriptions of Carabus croaticus ljubetensis (Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930), the second one has long time been accredited to be correct. However it is wrong and the original description must be assigned to the oldest one. The same author gave also two descriptions of Carabus croaticus babinjensis (Apfelbeck, 1919a; Apfelbeck, 1919b), as for the present it is not clear which of both has priority. 10.6. Carabus violaceus Linnaeus, 1758 10.6.1. Carabus violaceus azurescens Dejean, 1826 [= rilvensis Kolbe, 1887; zabljakensis Eidam, 1927; shljebensis Mandl, 1985] References. Carabus violaceus azurescens n. zabljakensis: Breuning, 1935: 1261 (Maranajt Mt.; Merdita); Breuning, 1978: 254. Carabus violaceus azurescens m. zabljakensis: Štěrba, 1945: 152 (Maranajt Mt.; Merdita). Carabus violaceus: Hieke & Wrase, 1988: 15. Carabus violaceus azurescens var. shljebensis: Březina, 1999: 77 (North Albanian Alps). Carabus violaceus azurescens: Turin et al., 2003: 260. 32


10.6.2. Carabus violaceus dryas Gistl, 1857 [= merditanus Apfelbeck, 1917; shardagensis Apfelbeck, 1917; korabensis Csiki, 1944] References. Carabus violaceus merditanus: Apfelbeck, 1917: 220 (Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1918a: 75-76 (Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1930: 162 (Merdita: Zepë Mt., loc. typ.); Cleu, 1968: 28 (Merdita); Cleu, 1969: 167. Carabus violaceus azurescens m. merditana: Breuning, 1935: 1262 (Vermosh; Zepë Mt. near Oroshi; Peshtrik Mt.; Gjalicë e Lumes); Breuning, 1978: 254 (Zepë Mt. near Oroshi). Carabus violaceus zabljakensis v. merditanus: Arnold, 1939: 186. Carabus violaceus var. shardagensis: Csiki, 1940: 210 (Gjalicë Mt.; Korab Mt.). Carabus violaceus var. korabensis: Csiki, 1944: 50-51 (Korab Mt., loc. typ.). Carabus violaceus azurescens m. merditanus: Štěrba, 1945: 152 (Vermosh; Zebë Mt.). Carabus violaceus shardagensis: Cleu, 1968: 28 (Korab Mt.); Cleu, 1969: 167 (Korab Mt.). Carabus picenus peristericus: Tarrier, 1974: 242 (Korab Mt.). Carabus violaceus dryas: Březina, 1999: 77; Bousquet et al., 2003: 165; Turin et al., 2003: 260. Carabus violaceus dryas var. merditanus: Březina, 1999: 77 (Merdita: Zepë Mt.; Gjalicë e Lumes). Carabus violaceus dryas var. korabensis: Březina, 1999: 77 (Korab Mt.). Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 1 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000 m, 14.VII.1993, rests of 1 s., A. Gobetti leg. (cAVT). Notes. Apfelbeck published three descriptions of both C. violaceus merditanus and C. violaceus shardagensis, respectively in Serbo-Croatian, in Latin and in German (Apfelbeck, 1917; Apfelbeck, 1918a; Apfelbeck, 1930), the second ones have long time been accredited to be correct. However it is incorrect and the original descriptions must be assigned to the oldest ones. A revision of the type of the var. shardagensis Apfelbeck, 1917 is needed to elucidate to which subspecies, azurescens or dryas, it belongs. Csiki (1944: 50, Fig. 1a) represented a figure of this variation which let me assume that it belongs rather to the former subspecies. # Carabus violaceus exasperatus Duftschmid, 1812 [= vlasuljensis Apfelbeck, 1894] References. Carabus violaceus vlasuljensis: Apfelbeck, 1907a: 502 (Merdita); Apfelbeck, 1918c: 160, 163 (North Albanian Alps, Vermosh). Notes. The easternmost known records of ssp. exasperatus come from Bosnia Herzegovina, so that the literature data for it should be referred to some of both subspecies that really live in Albania. Subgenus Oreocarabus Géhin, 1876 Notes. In my opinion, C. hortensis, C. neumeyeri and C. preslii represent separate species by reason of its clear and constant differences in both the overall aspect of the pronotum and the penis (Casale et al., 1982: 185, 190 – fig. 75, 195). Externally, the first species can be easily distinguished from the other two in the more quadrate aspect of the pronotum (vs. subovate or trapezium-shaped), the lateral border of pronotum posteriorly sinuate (vs. straight to slightly sinuate) and in the lateral lobes of pronotum clearly horizontal or slightly oblique (vs. oblique). Additionally C. neumeyeri has lateral borders of pronotum posteriorly more convergent than those in C. hortensis. The following nomenclatorial acts are proposed: Carabus (Oreocarabus) neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri Schaum, 1856; Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii Dejean, 1830 (see Casale et al., 1982: 194).

COLEOPTERA: CARABIDAE: CARABINAE

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10.7. Carabus hortensis Linnaeus, 1758 [= rhodopensis Apfelbeck, 1904] References. Carabus hortensis: Apfelbeck, 1918c: 163 (North Albanian Alps: Vermosh). Carabus hortensis rhodopensis: Apfelbeck, 1918c: 164 (North Albanian Alps: Grebeni near Vermosh, 11001750 m). Carabus hortensis hortensis: Breuning, 1933: 709 (Peshtrik Mt.; Vermosh); Bousquet et al., 2003: 179. Material included (studied A. Vigna Taglianti). Tropoia, Valle di Valbona, 900 m, 816.VII.1993, 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 1900 m, 10.VII.1993, 1 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 1000 m, 14.VII.1993, 1 f., L. Latella leg. (cAVT). 10.8. Carabus neumeyeri Schaum, 1856 [= herzegowinensis Apfelbeck, 1904; pindicus Apfelbeck, 1904; albanicola Breuning, 1965; hilleri Mandl, 1964] References. Carabus hortensis neumeyeri: Apfelbeck, 1904: 42 (Velipoja); Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m); Apfelbeck, 1909: 297 (Berat); Schatzmayr, 1943b: 94 (Shendelliut); Březina, 1999: 46; Bousquet et al., 2003: 179. Carabus hortensis neumeyeri n. neumeyeri: Breuning, 1933: 710 (Sarisaltik near Kruja; Vlorë;); Breuning, 1978: 198. Carabus hortensis neumeyeri n. herzegowinensis: Breuning, 1933: 712 (Velipoja near Lezhë). Carabus hortensis: Hieke & Wrase, 1988: 19. Carabus hortensis neumeyeri var. pindicus: Březina, 1999: 46. Carabus hortensis neumeyeri var. albanicola: Březina, 1999: 46 (Llogora). Carabus hortensis neumeyeri var. hilleri: Březina, 1999: 46 (Reza e Kanalit near Llogora). Carabus preslii neumeyeri: Turin et al., 2003: 228. Material studied. Kurbneshi, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., 2 f., P. Beron & B. Petrov leg. (NMNHS). 10.9. Carabus preslii Dejean, 1830 [= jonicus Apfelbeck, 1904] References. Carabus hortensis preslii m. jonica: Breuning, 1933: 713 (Gjirokastër); Breuning, 1978: 198. Material studied. Pogyara, 1027 m, 4.VI.1994, 2 f., G. Blagoev leg. (NMNHS); Leskovac, 650 m, 2.VI.1995, 1 f., V. Sakalian leg. (NMNHS). Notes. Bousquet et al. (2003: 179) has omitted the taxon for Albania, so that it is confirmed here for the country. Subgenus Pachystus Motschulsky, 1866 10.10. Carabus cavernosus I. Frivaldszky von Frivald, 1838 10.10.1. Carabus cavernosus cavernosus I. Frivaldszky von Frivald, 1838 [= sterbai Breuning, 1936] References. Carabus cavernosus cavernosus: Breuning, 1933: 798 (Korab Mt.; Gjalicë e Lumes); Guéorguiev et al., 1997: 8; Březina, 1999: 35; Bousquet et al., 2003: 181; Turin et al., 2003: 234. Carabus cavernosus var. sterbai: Csiki, 1940: 210 (Korab Mt.). Carabus cavernosus n. cavernosus: Breuning, 1978: 207. Carabus cavernosus: Hieke & Wrase, 1988: 15.

34


10.11. Carabus graecus Dejean, 1826 10.11.1. Carabus graecus morio Mannerheim, 1830 [= winkleri Breuning, 1933; neowinkleri Deuve, 1991] References. Carabus graecus winkleri: Breuning, 1933: 802 (Maja e Tartarit Mt., 800 m, loc. typ.). Carabus graecus n. winkleri: Breuning, 1978: 207 (Maja e Tartarit Mt.). Carabus graecus: Hieke & Wrase, 1988: 14; Březina, 1999: 36. Carabus graecus morio: Březina, 1999: 36; Bousquet et al., 2003: 182. Carabus graecus morio var. winkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus morio var. neowinkleri: Březina, 1999: 36 (Maja e Tartarit Mt.). Carabus graecus graecus: Turin et al., 2003: 236. ? subgenus Platycarabus A. Morawitz, 1886 ? Carabus irregularis Fabricius, 1792 ? Carabus irregularis ramanus Sokolář, 1909 [= bucephalus Kraatz, 1879]] References. Carabus irregularis bucephalus: Turin et al., 2003: 249. Notes. Turin et al. (2003) showed the species range reaching North Albania. According to Březina (1999: 53) ssp. ramanus is known from the Bjelasièa Mt. in Northeastern Montenegro not very far from the studied territory. The species may live in Albania. Subgenus Procerus Dejean, 1821 10.12. Carabus gigas Creutzer, 1799 10.12.1. Carabus gigas gigas Creutzer, 1799 [= parnassicus Kraatz, 1884] References. Carabus gigas parnassicus: Březina, 1999: 72. Carabus gigas gigas: Bousquet et al., 2003: 185. Material studied. Malesia e Polisit Mt., 1850 m, near Sopoti, VI.1966, 1 s. (cXM). Notes. First detailed record for Albania. Subgenus Procrustes Bonelli, 1810 10.13. Carabus coriaceus Linnaeus, 1758 10.13.1. Carabus coriaceus cerisyi Dejean, 1826 [= hopffgarteni Kraatz, 1877; subrugosus Kraatz, 1877; weiratherianus Breuning, 1936] References. Carabus coriaceus cerisyi m. weiratheriana: Breuning, 1937: 1406 (Korab Mt.); Breuning, 1978: 270. Carabus coriaceus subrugosus v. weiratherianus: Arnold, 1939: 192. Carabus coriaceus var. hopffgarteni: Csiki, 1940: 209 (Kula Ljums; Gjalicë Mt.; Korab Mt.). Carabus coriaceus subrugosus: Hieke et al., 1986: 183 (Borshi near Vlorë). Carabus coriaceus: Hieke & Wrase, 1988: 13. 10.13.2. Carabus coriaceus excavatus Charpentier, 1825 [= rugosus Dejean, 1826; dalmaticus Géhin, 1885; joniensis Breuning, 1932; albanicus Breuning, 1943] References. Carabus coriaceus rugosus: Apfelbeck, 1904: 20 (Velipoja; Berat; Vlorë); Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 297 (Ardenica; Vlorë; Pojani). Carabus coriaceus rugosus n. joniensis m. albanicus: Breuning, 1932: 20 (Maja e Tartarit Mt., loc. typ.); Mařan, 1949: 3 (Maja e Tartarit Mt.). Carabus coriaceus excavatus m. dalmatica: Breuning, 1937: 1403 (Velipoja; Kruja; Durrës; Merdita; Tiranë; Berat; Vlorë; south of Vlorë); Breuning, 1978: 269. Carabus coriaceus excavatus n. joniensis: Breuning, 1937: 1404 (Vlorë; Gjirokastër); Magistretti, 1965: 55; Breuning, 1978: 269. Carabus coriaceus excavatus: Arnold, 1939: 191; Hieke COLEOPTERA: CARABIDAE: CARABINAE

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et al., 1986: 183 (Mal i Dajtit Mt.: Shkall Priskë); Březina, 1999: 62; Bousquet et al., 2003: 187; Turin et al., 2003: 278. Carabus coriaceus excavatus dalmaticus: Schatzmayr, 1943b: 94 (Lushnje; Karbunara e Madhe; Dukati, south of Vlorë). Carabus coriaceus joniensis: Březina, 1999: 62; Bousquet et al., 2003: 187. Carabus coriaceus joniensis: var. albanicus: Březina, 1999: 62 (Maja e Tartarit Mt.). Carabus coriaceus mediterraneus: Turin et al., 2003: 278. Material studied. Sauk, 18.II.1960, 1 m. (MNST); Ibë, 3.VII.1962, 1 f. (MNST); Bizë, 27.VII.1966, 1 f. (MNST); Tiranë, 27.III.1968, 1 f. (MNST); Tiranë Park, 22-31.X.1997, 2 s., R. Ward leg. (CMNH); Tiranë Park, 1-4.I.1998, 2 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m, 11.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 2000 m, 14.VII.1993, 1 f., A. Gobetti leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 2 m., P. Crucitti & C. Cavalieri leg. (cAVT). Note: According to Turin et al. (2003) C. coriaceus mediterraneus Born, 1906 has transionian distribution and lives in Albania. Subgenus Tachypus Weber, 1801 10.14. Carabus cancellatus Illiger, 1798 10.14.1. Carabus cancellatus alessiensis Apfelbeck, 1901 [= hypsobius Apfelbeck, 1918] References. Carabus cancellatus var. alessiensis: Apfelbeck, 1901b: 121 (Lezhë, loc. typ.). Carabus cancellatus alessiensis: Apfelbeck, 1904: 30; 33 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë River Outfall); Breuning, 1933: 594 (Velipoja near Lezhë; Shkodër; Pulaj); Arnold, 1939: 161 (North Albania); Breuning, 1978: 187 (Velipoja); Březina, 1999: 41 (Velipoja near Lezhë); Bousquet et al., 2003: 195; Turin et al., 2003: 194. Carabus cancellatus subgraniger var. alessiensis: Bernau, 1913: 198 (Lezhë). Carabus cancellatus emarginatus: Bernau, 1913: 200. Carabus cancellatus: Hieke & Wrase, 1988: 17. Material studied. Shkodër, 14.V.1963, 2 m. (MNST). Notes. The type locality of the form hypsobius Apfelbeck, 1918c: 163 is Vunšaj (in Albanian) or Vusanje (in Serbo-Croatian) in the North Albanian Alps. It is situated in the Montenegrin territory very closely to Albania. C. cancellatus emarginatus Duftschmid, 1812 is a valid subspecies reaching to Bosnia Herzegovina in the east. Subgenus Tomocarabus Reitter, 1896 10.15. Carabus convexus Fabricius, 1775 10.15.1. Carabus convexus dilatatus Dejean, 1826 [= chionophilus Apfelbeck, 1904; miliusi Breuning, 1927] References. Carabus convexus var.: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m). Carabus convexus dilatatus: Breuning, 1933: 874 (Vlorë; Gjirokastër); Bousquet et al., 2003: 198; Turin et al., 2003: 230. Carabus convexus dilatatus n. miliusi: Breuning, 1933: 876 (North Albanian Alps; Merdita); Breuning, 1978: 214. Carabus convexus miliusi: Arnold, 1939: 178. Carabus convexus var. chionophilus: Csiki, 1940: 210 (Korab Mt.). Carabus convexus: Hieke & Wrase, 1988: 19. Carabus convexus dilatatus var. miliusi: Březina, 1999: 33. Material studied. Mal i Dajtit Mt., 11.V.1965, 1 m. (MNST); North Albanian Alps, Maj’e Radohimes, 1900-2200, 28.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); 36


North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 3 f., P. Beron & B. Petrov leg. (NMNHS). ? subgenus Trachycarabus Géhin, 1876 ? Carabus scabriusculus Olivier, 1795 ? Carabus scabriusculus bulgarus Lapouge, 1908 References. Carabus scabriusculus bulgarus: Turin et al., 1993: 42. Notes. This species is questionable for Albania due to lack of exact records. On the other side, it was quoted for the neighboring countries Montenegro and Serbia.

Tribe CYCHRINI Laporte, 1834 11. Genus Cychrus Fabricius, 1794 11.1. Cychrus semigranosus Palliardi, 1825 # Cychrus semigranosus semigranosus Palliardi, 1825 References. Cychrus semigranosus semigranosus: Häckel, 2003: 205. Notes. This record is referred to the ssp. montenegrinus because the nominotypical race does not populate Albania. # Cychrus semigranosus balcanicus Hopffgarten, 1881 References. Cychrus semigranosus n. balcanicus: Mařan, 1940c: 133-134 (North Albanian Alps). Notes. This East Balkan race does not inhabit Albania, therefore it is removed from the species list. # Cychrus semigranosus montanus Winkler, 1939 References. Cychrus semigranosus montanus: Häckel, 2003: 205. Notes. Winkler (1939: 199) described this subspecies on material from two different regions - the Eastern Carpathians and Western Balkans. Mandl (1989: 69) designated lectotype and paralectotypes for the specimens from the former region, and syntypes of montanus from the latter region were assigned to C. semigranosus dinaricus Mandl, 1989 (“Cvrstnica”, loc. typ.). Presumably, the subspecies mentioning for Albania (Häckel, 2003: 205) is based on incorrect interpretation of the locality “Zljeb planina nördlich der Nord-albanischen Alpen” (Winkler, 1939: 199; Mařan, 1940c: 133; Mandl, 1989: 68). In fact the Zljeb Mt. is situated in the Kosovo Region of Serbia. 11.1.1. Cychrus semigranosus montenegrinus Apfelbeck, 1904 [= albanicus Hopp, 1929; dinaricus Mandl, 1989] References. Cychrus semigranosus semigranosus: Röschke, 1907: 255; Häckel, 2003: 205. Cychrus semigranosus: Apfelbeck, 1907a: 498, 503 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 210 (Shkëlzen Mt.; Korab Mt.). Cychrus semigranosus albanicus: Hopp, 1929: 2 (Tomor Mt, 2000 m, loc. typ.); Schatzmayr, 1943b: 94 (Tomor Mt.); Mandl, 1989: 69 (Peshtrik Mt.; Gjalicë Mt.; Çukall Mt.; Merdita); Häckel, 2003: 205. Cychrus semigranosus ssp.: Schatzmayr, 1943b: 93 (Shendelliut). Cychrus semigranosus montenegrinus: Winkler, 1939: 200 (Llogora; Tomor Mt.; Kruja). Cychrus semigranosus n. montenegrinus: Mařan, 1940c: 128, 130-132, 135-136 (Llogora). Cychrus semigranosus montanus: Häckel, 2003: 205. Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exp.’18. Pashtrik 4-14.7”, 3 s. (NMW); “Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Giallica” / “Albania leg. Bischoff 1936”, 1 s. (NMW). COLEOPTERA: CARABIDAE: ELAPHIRINAE

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Notes. Winkler (1939) synonymized the ssp. albanicus with ssp. montenegrinus. Mařan (1940c) accepted this point of view, but Mandl (1989) treated both as separate subspecies. Samples of the ssp. montenegrinus, ssp. albanicus, as well as the holotype and paratypes of the ssp. dinaricus have been studied in NMW. Differences between specimens from different localities exist mostly in the sizes of the body and shape of the pronotum, but they also have occurred in the limits of one and the same population. In reality characters that can support well-defined geographical races in the Western Balkans, excluding the well-differentiated ssp. travnikanus Apfelbeck, 1904 (type series in NMW) and the not studied ssp. jakupicensis Mařan, 1943, are absent. Therefore the following synonymy is proposed: Cychrus semigranosus albanicus Hopp, 1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck, 1904; Cychrus semigranosus dinaricus Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck, 1904. In MNHUB, I have furthermore seen 69 s. from different localities, which remained unregistered.

Subfamily ELAPHRINAE Erichson, 1837 Tribe ELAPHRINI Erichson, 1837 12. Genus Elaphrus Fabricius, 1775 Subgenus Neoelaphrus Hatch, 1951 12.1. Elaphrus uliginosus Fabricius, 1792 References. Elaphrus uliginosus: Murraj, 1968: 33 (Bizë). Material studied. “N 88”, 1 f. (MNST). Notes. The record of Murraj (1968) is included here with minimal reservation because of his correct determination of the female above. Goulet (2003: 207) omitted the species for Albania, but herewith it is confirmed for the country.

Subfamily OMOPHRONINAE Bonelli, 1810 Tribe OMOPHRONINI Bonelli, 1810 13. Genus Omophron Latreille, 1802 Subgenus Omophron s.str. 13.1. Omophron limbatum (Fabricius, 1777) References. Omophron limbatum: Hieke et al., 1986: 183 (Bizë near Shëngjergji); Hieke & Wrase, 1988: 23; Hůrka, 2003: 208. Material studied. “Skutari Bojana”, 3 s. (NMW); “Oboti Bojana” / “Apfelb. Alban. spt. IV-V.05”, 1 s. (NMW); “Berat” / “Albania leg. Bischoff 1931”, 17 s. (MNHUB); “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tepelene” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Vojusa 38


Dragot” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS).

Subfamily BRACHININAE Bonelli, 1810 Tribe BRACHININI Bonelli, 1810 14. Genus Aptinus Bonelli, 1810 Subgenus Aptinus s.str. 14.1. Aptinus acutangulus Chaudoir, 1876 Material studied. North Albanian Alps, Bogë, 1400-1800 m, 1.6.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.6.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Theth, VII.1993, 15 m., 14 f., P. Dioli leg. (cAVT). Notes. New species for the fauna of Albania. 14.2. Aptinus bombarda (Illiger, 1800) References. Aptinus bombarda: Apfelbeck, 1918c: 168 (North Albanian Alps, Vermosh, 1200 m); Hůrka, 1988: 290-291 (Shkëlzen Mt.); Hrdlička, 2003: 212. Material studied. “Vermosa’14 Penther”, 1 m. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 f. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 m. (NMW); “Maja e Jezerce Albania sept.”, 1 m. (NMW); “Shkëlzën” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); North Albanian Alps, Bogë, 14001800 m, 1.6.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2000 m, 11.VII.1993, 2 m., L. Latella leg. (cAVT); Tropoia, Valbona, Kollatës Mt., 1000 m, 14.VII.1993, 1 m., L. Latella leg. (cAVT). Notes. A. bombarda occurs sympatrically with A. acutangulus in the North Albanian Alps (see also Casale & Vigna Taglianti, 1983: 36; Hůrka, 1988: 289); the former lives at the same altitude or higher than the latter. The study of the penis reveals that the male from Bogë has somewhat differing apex compared with those from Vermosa and Maja e Jezercoë. The penis apex in the latter is identical with the illustration of Casale & Vigna Taglianti (1983: 32, Fig. 8). Very likely the population from the massif of Bogë forms a separate race. 14.3. Aptinus merditanus Apfelbeck, 1917 14.3.1. Aptinus merditanus merditanus Apfelbeck, 1917 [= granulipennis Apfelbeck, 1917] References. Aptinus lugubris rufipes: Apfelbeck, 1907a: 499, 501 (Mal i Shenjt Mt.: top region, 1500 m). Aptinus merditanus: Apfelbeck, 1917: 227 (Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1918b: 89 (Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1930: 171-172 COLEOPTERA: CARABIDAE: BRACHININAE

39

(Merdita: Zepë Mt. near Fandi, loc. typ.); Casale & Vigna Taglianti, 1983: 29, 39, 41 (Merdita: Zepë Mt.; Shen Thanas Mt. near Vlorë); Hůrka, 1988: 293-294 (Mal i Shenjt Mt.; Zepë Mt.; Mal i Dajtit Mt.; Mal i Krujës Mt.; Shen Ili; Maja e Tartarit Mt.; Cafa Kjorës; Tomor Mt.; Shen Thanas Mt.; Llogora); Wrase, 1991: 18. Aptinus merditanus var. granulipennis: Apfelbeck, 1917: 227 (South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.); Apfelbeck, 1918b: 89 (South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.); Apfelbeck, 1930: 172 (South Albania: Shen Thanas Mt. near Vlorë, loc. typ.; Northwestern Greece: Peristeri Mt. near Joannina, loc. typ.). Aptinus merditanus merditanus: Hrdlička, 2003: 212. Material studied. “Merdita M. Schèit” / “Coll. Apfelbeck” / “merditanus Typ. Apf.”, 1 m. (HNHM); 1 s. from “Çafa Kjores”, 53 s. from “Llogora” and “Logara, 8 s. from “Maja e Tartarit”, 40 s. from “Mal i Dajtit”, 1 s. from “Mal i Krujes”, 10 s. from “Shen Ili”, 1 s. from “Tomor” (MNHUB). Notes. Apfelbeck published three consecutive descriptions for both A. merditanus and A. merditanus var. granulipennis (Apfelbeck, 1917; Apfelbeck, 1918b; Apfelbeck, 1930) and the second ones have long time been accredited to be valid. Actually this is incorrect and the original descriptions must be assigned to the oldest ones. Aptinus lugubris rufipes (Apfelbeck, 1907a: 499) is nomen nudum (see also Hůrka, 1988: 292). In that case, Apfelbeck had most probably the intention to introduce a name for A. merditanus earlier than the year of 1917.

15. Genus Brachinus Weber, 1801 Subgenus Brachinus s.str. # Brachinus alexandri F. Battoni, 1984 References. Brachinus alexandri: Guéorguiev & Guéorguiev, 1995: 239. Notes. Dealing with B. plagiatus data in the paper of Hùrka (1988), I noted that B. alexandri was recorded by mistake for Albania (Guéorguiev & Guéorguiev, 1995). Most likely Hrdlička(2003: 213) followed this mistake marking the species for the country. Here B. alexandri is removed from the list of Albanian ground beetles. 15.1. Brachinus crepitans (Linnaeus, 1758) [= fallax Apfelbeck, 1904] References. Brachinus crepitans: Apfelbeck, 1904: 348 (Vlorë; Berat); Apfelbeck, 1909: 298 (Vlorë; Berat); Csiki, 1940: 222 (Kula Ljums; Gjalicë Mt.); Hieke & Wrase, 1988: 163; Hrdlička, 2003: 213. Brachinus crepitans ab. fallax: Apfelbeck, 1904: 348 (Vlorë); Csiki, 1940: 222 (Kula Ljums). Material studied. “Avlona”, 1 s. (HNHM); “Han Holfit 5.-10. Mai” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban-Exp.’18. Pashtrik, 4-14.7.”, 1 s. (NMW); “Dures” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lume i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Iballe” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); “Albania Fush-Dukati” / “7.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 f., G. Blagoev leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2 m., 1 f., D. Antonelli leg. (cAVT). 40


15.2. Brachinus ejaculans Fischer von Waldheim, 1828 References. Brachinus ejaculans: Apfelbeck, 1904: 351 (Velipoja; Vlorë); Hieke & Wrase, 1988: 165; Hrdlička, 2003: 213. Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Miloti, Alb. Matzenauer”, 3 s. (NMW); “Shullaz” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 11 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB). 15.3. Brachinus elegans Chaudoir, 1842 [= ganglbaueri Apfelbeck, 1904] References. Brachynus psophiae or ganglbaueri: Klaptocz, 1910: 399 (Shkodër); Schatzmayr, 1943b: 113 (Lushnje). Brachinus ganglbaueri: Hieke & Wrase, 1988: 164. Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Hrdlička (2003: 213) has omitted the species for Albania. Here it is confirmed for the country. 15.4. Brachinus plagiatus Reiche, 1868 References. Brachinus plagiatus: Apfelbeck, 1904: 350 (Velipoja; Vlorë); Horion, 1941: 353; Schatzmayr, 1943b: 112 (Skrofotina); Hieke & Wrase, 1988: 165; Hùrka, 1988: 205 (Tiranë). Material studied. “Avlona”, 6 s. (HNHM); “Durazzo”, 2 s. (HNHM); “Alban. Wgth. Valona”, 9 s. (NMW); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB). Notes. Hrdlička (2003: 213) has omitted the species for Albania. Here it is confirmed for the country. 15.5. Brachinus psophia Audinet-Serville, 1821 References. Brachinus psophia: Apfelbeck, 1904: 349 (Vlorë); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988: 164; Hrdlička, 2003: 213. “Brachynus psophiae aut Ganglbaueri”: Schatzmayr, 1943b: 113 (Lushnje). Material studied. “Avlona”, 1 s. (HNHM); “Alban. Wgth. Valona”, 2 s. (NMW); Radona, Shkiela, 21.VI.1994, 1 m., B. Guéorguiev leg. (NMNHS). Subgenus Brachynidius Reitter, 1919 15.6. Brachinus bodemeyeri Apfelbeck, 1904 References. Brachinus bodemeyeri: Apfelbeck, 1904: 352 (Vlorë); Hieke & Wrase, 1988: 166; Hrdlička, 2003: 213. Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB); “Alban. Wgth. Valona”, 4 s. (NMW); “Dukati” / “ex Coll. Scheibel”, 1 s. (MNHUB).

COLEOPTERA: CARABIDAE: BRACHININAE

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15.7. Brachinus brevicollis Motschulsky, 1844 [= peregrinus Apfelbeck, 1904] References. Brachinus peregrinus: Apfelbeck, 1904: 351 (Vlorë, loc. typ.); Reitter, 1919: 140; Schatzmayr, 1935: 245; Schatzmayr, 1943b: 112 (Tërpan; Dukati; Lushnje; Vlorë); Hieke & Wrase, 1988: 165. Brachinus brevicollis: Hrdlička, 2003: 213. Material included (studied A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT). 15.8. Brachinus explodens Duftschmid, 1812 [= glabratus Dejean, 1824; strepens Fischer von Waldheim, 1828] References. Brachinus explodens: Apfelbeck, 1904: 353 (Vlorë; Berat); Apfelbeck, 1909: 298 (Vlorë; Berat); Csiki, 1940: 222 (Ura i Lopez); Schatzmayr, 1943b: 113 (Dukati); Hieke et al., 1986: 190 (North Albanian Alps near Theth); Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214. Brachinus explodens ab. strepens: Csiki, 1940: 222 (Ura i Lopez; “Stičen” near Gjalicë Mt.). Brachinus explodens ab. glabratus: Csiki, 1940: 222 (Kula Ljums; Ura i Lopez). Material studied. “Avlona”, 2 s. (HNHM); “Skutari-Holcekai” / “Alban. montenegr. Grenze, Penther’14”, 12 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura i Lopez” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 8 s. (NMW); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Cerrik” / “16.IV.1958 leg. Xh. Murraj”, 7 s. (HNHM); Tiranë, 7.IV.1966, 1 s. (MNST); Vlorë, 1.V.1994, 2 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg. (NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 14.I.1998 / 20-25.I.1998 / 1.III.1998 / 8.III.1998, 13 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M. Komnenov leg. (cSH). Notes. I have seen furthermore 160 s. from different localities in MNHUB, which remained unregistered. 15.9. Brachinus sichemita Reiche & Saulcy, 1855 References. Brachinus explodens sichemita: Apfelbeck, 1904: 353 (Berat; Vlorë). Notes. Hrdlička (2003: 214) has omitted the species for Albania, therefore its occurrence in the country needs further confirmation. Subgenus Brachynolomus Reitter, 1919 15.10. Brachinus immaculicornis Dejean, 1826 15.10.1. Brachinus immaculicornis immaculicornis Dejean, 1826 References. Brachinus immaculicornis: Apfelbeck, 1909: 298 (Vlorë). 42


Notes. Hrdlička (2003: 214) has omitted the species for Albania; therefore its occurrence in the country needs further confirmation. Subgenus Cnecostolus Reitter, 1919 15.11. Brachinus bayardi Dejean, 1831 References. Brachinus bayardi: Schatzmayr, 1943b: 112 (Mal i Krujes). Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Notes. Hrdlička (2003: 214) has omitted the species for Albania. Here it is confirmed for the country. 15.12. Brachinus bipustulatus Quensel, 1806 References. Brachinus bipustulatus: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909: 298 (Vlorë); Reitter, 1919: 135; Hieke & Wrase, 1988: 166; Hrdlička, 2003: 214. Material studied. “Scutari” / “collectio Paganetti”, 1 s. (NMW); “Mustajbeg Velipoja Albania”, 1 s. (MNHUB); “Durazzo Apfelbeck”, 1 s. (HNHM); “Oboti Bojana”, 4 s. (HNHM); “Alban. Wgth. Valona”, 1 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 15.13. Brachinus exhalans (P. Rossi, 1792) References. Brachinus exhalans: Apfelbeck, 1904: 354 (Velipoja; Vlorë); Apfelbeck, 1909: 298 (Vlorë); Schatzmayr, 1943b: 112 (Lushnje; Mal i Krujes); Hrdlička, 2003: 214. Material studied. “Avlona”, 3 s. (HNHM); “Avlona Albania mer.”, 1 s. (HNHM); “Albanien Avlona v. Oertzen.”, 5 s. (MNHUB); “Durazzo Apfelbeck”, 2 s. (HNHM); “Alban. Wgth. Valona”, 6 s. (NMW); “Murshi” / “Albania leg. Bischoff 1936”, 111 s. (MNHUB); Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS). Species insertae sedis 15.14. Brachinus nigricornis Gebler, 1829 [= incertus Brullé, 1834] References. Brachinus incertus: Apfelbeck, 1904: 350 (Velipoja near Lezhë); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988: 165. Brachinus nigricornis: Hrdlička, 2003: 216. Material studied. “Durazzo”, 1 s. (HNHM); “Albania Avlona”, 5 s. (HNHM); “Bojana Mundung”, 3 s. (MNHUB); “Alban. Wgth. Valona”, 12 s. (NMW).

COLEOPTERA: CARABIDAE: SIAGONINAE

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Subfamily SIAGONINAE Bonelli, 1813 Tribe SIAGONINI Bonelli, 1813 16. Genus Siagona Latreille, 1804 16.1. Siagona europaea Dejean, 1826 16.1.1. Siagona europaea europaea Dejean, 1826 [= depressa Bedel, 1887 (nec Fabricius, 1798)] References. Siagona europaea: Apfelbeck, 1904: 358 (Vlorë); Schatzmayr, 1943b: 95 (Mal i Krujes; Lushnje; Skrofotina); Hieke, 1981: 76; Hieke & Wrase, 1988: 27. Siagona depressa: Apfelbeck, 1909: 297 (Vlorë). Siagona europaea europaea: Baehr, 2003a: 218. Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Liqen i Butrinten Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1937”, 2 s. (MNHUB). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., M. Komnenov leg. (cSH). ? Siagona europaea oberleitneri Dejean, 1830 References. Siagona europaea var. oberleitneri: Ganglbauer, 1904: 648. Notes. This race is endemic for the Ionic Islands. Taking into account the proximity of these islands to South Albania further collectings are needed to determine the subspecies status of the populations in the studied territory.

Subfamily SCARITINAE Bonelli, 1810 Tribe CLIVININI Rafinesque, 1815 Subtribe CLIVININA Rafinesque, 1815 17. Genus Clivina Latreille, 1802 17.1. Clivina collaris (Herbst, 1784) References. Clivina collaris: Balkenohl, 2003: 219. Material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 816.VII.1993, 1 m., L. Latella leg. (cAVT). Notes. First detailed record for Albania.

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17.2. Clivina fossor (Linnaeus, 1758) 17.2.1. Clivina fossor fossor (Linnaeus, 1758) References. Clivina fossor: Apfelbeck, 1904: 68 (Velipoja near Lezhë; Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke et al., 1986: 184 (Shkodër); Hieke & Wrase, 1988: 27. Clivina fossor fossor: Balkenohl, 2003: 219. Material studied. “Scutary Bojana”, 3 s. (NMW); “Scutary i. See Ufer”, 2 s. (NMW); “Scutary Vraka”, 3 s. (NMW); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). 17.3. Clivina laevifrons Chaudoir, 1842 References. Clivina laevifrons: Apfelbeck, 1904: 69 (Velipoja); Rambousek, 1912: 69; Hieke & Wrase, 1988:28; Balkenohl, 2003: 220. Material studied. “Scutary Bojana”, 1 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.”, 1 s. (NMW); “Scutary i. See Ufer”, 2 s. (NMW); “Scutary Vraka”, 1 s. (NMW).

Subtribe REICHEINA Jeannel, 1957 ! genus Dalmatoreicheia Magrini & Bulirsch, 2005 ! Dalmatoreicheia sp. Material studied. “Kruja, Alban., Mader”, 1 m. (NMW). Notes. The specimen belongs to a new species.

18. Genus Reicheadella Reitter, 1913 Notes. The Albanian species of this subtribe are object of taxonomic work (Bulirsch & Guéorguiev, in preparation). Subgenus Reicheadella s.str. 18.1. Reicheadella bischoffi (Meschnigg, 1933) References. Reicheia bischoffi: Meschnigg, 1933:172 (Tomor Mt. near Berat, loc. typ.); Turin, 1981: 72. Reicheadella bischoffi: Jeannel, 1957: 179; Casale et al., 1998: 95; Balkenohl, 2003: 222. Material studied. “Tomor Juli 1932” / “Albania leg. Bischoff 1932”, 2 syntypes (MNHUB); “Tomor: Tomor Alban. mer.” / “lg. Winkler Mai 1931” / “bischoffi Meschn. det. A. Winkler”, 2 m. (NMW). ! Reicheadella sp. Material studied. “Albania mer. Liqueni i Butrintit, 1.V.1958; A. Smetana leg.”, 1 f. (NMPC). Notes. The specimen belongs to a new species.

COLEOPTERA: CARABIDAE: SCARITINAE

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Subtribe DYSCHIRIINI W. Kolbe, 1880 19. Genus Dyschirius Bonelli, 1810 Subgenus Chiridysus Fedorenko, 1996 19.1. Dyschirius strumosus Erichson, 1837 References. Dyschirius strumosus: Balkenohl, 2003: 224. Subgenus Dyschiriodes Jeannel, 1941 Species group “integer” 19.2. Dyschirius intermedius Putzeys, 1846 References. Dyschirus intermedius: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 78 (Oboti); Hieke & Wrase, 1988: 31; Fedorenko, 1993: 819; Balkenohl, 2003: 225. Species group “nitidus” 19.3. Dyschirius agnatus Motschulsky, 1844 [= lucidus Putzeys, 1846] References. Dyschirius lucidus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës). Dyschirius agnatus: Balkenohl, 2003: 224. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 9 s. (HNHM); “Tepelene” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 5 s. (MNHUB). 19.4. Dyschirius nitidus (Dejean, 1825) 19.4.1. Dyschirius nitidus nitidus (Dejean, 1825) References. Dyschirius nitidus: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 63-64 (Berat); Csiki, 1940: 212 (Kula Ljums); Horion, 1941: 98; Ieniștea, 1979: 224; Gryuntal, 1984b: 514; Hieke & Wrase, 1988: 29. Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB). Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here it is confirmed for the country. Species group “minutus” 19.5. Dyschirius aeneus (Dejean, 1825) 19.5.1. Dyschirius aeneus aeneus (Dejean, 1825) References. Dyschirius aeneus: Apfelbeck, 1904: 72 (Velipoja near Lezhë); Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 76; Hieke & Wrase, 1988: 31. Dyschirius aeneus aeneus: Balkenohl, 2003: 224. 46


Material studied. “Durrazo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). 19.6. Dyschirius apicalis Putzeys, 1846 References. Dyschirius apicalis: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 77 (Durrës); Hieke & Wrase, 1988: 31; Balkenohl, 2003: 224. Material studied. “Alban. Wgth. Valona”, 1 s. (HNHM); “Skutari Bojana” / “Mündg. Apf. 08.”, 3 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Liqen Butrintës” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS). [Dyschirius auriculatus (Wollaston, 1867)] Notes. An expected species for Albania due to its occurring in Montenegro (P. Bulirsch, personal communication) 19.7. Dyschirius bacillus Schaum, 1857 19.7.1. Dyschirius bacillus arbensis J. Müller, 1911 References. Dyschiriodes bacillus arbensis: Fedorenko, 1996: 191 (Vlorë); Balkenohl, 2003: 224. Dyschirius bacillus bacillus: Balkenohl, 2003: 224. Material studied. Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS). Notes. According to Fedorenko (1996: 191) the specimen from Vlorë shows transitional characters between the subspecies arbensis and lucanus (Focarile, 1959). P. Bulirsch (personal communication) identified the specimen from Shkiela and referred it to spp. arbensis. 19.8. Dyschirius cylindricus (Dejean, 1825) 19.8.1. Dyschirius cylindricus hauseri A. Fleischer, 1898 References. Dyschirius cylindricus: Apfelbeck, 1907a: 496 (Buenë River Outfall). Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 21 s. (MNHUB). Notes. Balkenohl (2003: 225) has omitted the species for Albania. Here it is confirmed for the country. 19.9. Dyschirius laeviusculus Putzeys, 1846 References. Dyschirius laeviusculus: Hieke et al., 1986: 184 (Kula e Lumës near Kukës). Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM). Notes. Balkenohl (2003: 226) has omitted the species for Albania. Here D. laeviusculus is confirmed for the country. The locality Kula e Lumës outlines the southern limit of the species distribution in the Balkan Peninsula. 19.10. Dyschirius luticola Chaudoir, 1850 19.10.1. Dyschirius luticola luticola Chaudoir, 1850 Material studied. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. COLEOPTERA: CARABIDAE: SCARITINAE

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19.11. Dyschirius macroderus Chaudoir, 1850 19.11.1. Dyschirius macroderus macroderus Chaudoir, 1850 References. Dyschirius macroderus: Apfelbeck, 1907a: 496 (Buenë River Outfall). Notes. Balkenohl (2003: 226) has omitted the species for Albania. Actually D. macroderus needs further confirmation for the country. 19.12. Dyschirius minutus (Dejean, 1825) 19.12.1. Dyschirius minutus albanicus J. Müller, 1922 References. Dyschirius minutus albanicus: Müller, 1922b: 48, 82-83 (Berat, loc. typ.); Müller, 1937: 122; Focarile, 1959: 51; Guéorguiev & Guéorguiev, 1995: 71; Guéorguiev et al., 1997: 9. Dyschirius punctatus albanicus: Müller, 1923: 107 (Osum River near Berat, loc. typ.); Drovenik & Peks, 1994: 25 (Berat); Drovenik & Peks, 1999: 29 (Berat). Dyschirius minutus: Balkenohl, 2003: 226. Material studied. “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura-Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Notes. Müller published two descriptions of the subspecies albanicus. First he described D. minutus albanicus (Müller, 1922), and next D. punctatus albanicus (Müller, 1923). Drovenik and Peks (Drovenik & Peks, 1994; Drovenik & Peks, 1999) erroneously indicated the locality Berat for Montenegro. Fedorenko (2001) confirmed the subspecific status of albanicus and his point of view is followed here. 19.13. Dyschirius morio Putzeys, 1866 [= simulator J. Müller, 1922] References. Dyschirius simulator: Müller, 1922b: 48, 90 (Berat, loc. typ.; Shkodër, loc. typ.); Müller, 1923: 105-107 (Osum River near Berat, loc. typ.); Ieniștea, 1979: 220 (Berat); Hieke & Wrase, 1988: 32. Dyschirius morio: Guéorguiev & Guéorguiev, 1995: 66; Balkenohl, 2003: 226. Material studied. “Berat” / “Albania leg. Bischoff 1931”, 15 s. (MNHUB). Notes. Müller published two description of D. simulator (Müller, 1922; Müller, 1923), the older one is correctly treated as valid. # Dyschirius punctatus (Dejean, 1825) References. Dyschirius punctatus: Magistretti, 1965: 107; Balkenohl, 2003: 227. Notes. A primarily West-Mediterranean species (Fedorenko, 1996: 161-162, Fig. 382) which presence in Albania is highly doubtful. Likely the confusion about its presence there is due to the view of Magistretti (1965: 106-107) who treated D. minutus as synonym of D. punctatus. Species removed from the list of the Albanian Carabidae. [Dyschirius pusillus (Dejean, 1825)] Notes. An expected species for Albania as it was recently collected in North Greece (P. Bulirsch, personal communication)

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Species group “chalybeus” 19.14. Dyschirius chalybeus Putzeys, 1846 19.14.1. Dyschirius chalybeus gibbifrons Apfelbeck, 1899 References. Dyschirius gibbifrons: Apfelbeck, 1899: 290; Focarile, 1959: 48 (Shkodër; Vlorë). Dyschirius chalybeus gibbifrons: Balkenohl, 2003: 225. Material studied. “Alban. Wgth. Valona”, 7 s. (HNHM; MNHUB); “Skutari l. See-Ufer”, 9 s. (HNHM; MNHUB); “Skutari 29.3.16”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen Butrintës” / “Albania leg. Bischoff 1936”, 10 s. (MNHUB); Tiranë Park, 22.II.1998, 1 s., from grasses along small sluggish stream, R. Ward leg. (CMNH). 19.15. Dyschirius salinus Schaum, 1843 19.15.1. Dyschirius salinus striatopunctatus Putzeys, 1846 References. Dyschirius salinus: Apfelbeck, 1907a: 496 (Buenë River Outfall); Müller, 1922b: 72 (Durrës); Hieke & Wrase, 1988: 30. Dyschirius salinus striatopunctatus: Balkenohl, 2003: 227. Material studied. “Berat” / “Albania leg. Bischoff 1931”, 14 s. (MNHUB); “Durrazo” / “Albania leg. Bischoff 1931”, 21 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB): “Durrës” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB). Subgenus Dyschirius s.str. Notes. New subgenus for the fauna of Albania. 19.16. Dyschirius latipennis Seidlitz, 1867 Material included (studied P. Bulirsch). Labinoti-Fushe (Elbasan District), Shkumbinit River, 2.VII.2001, 4 s., Fencl & Moravec leg. (cPB). Notes. New species for the fauna of Albania. Subgenus Eudyschirius Fedorenko, 1996 Species group “globosus” 19.17. Dyschirius globosus (Herbst, 1784) References. Dyschirius globosus: Balkenohl, 2003: 229. Species group “lafertei” 19.18. Dyschirius gracilis (Heer, 1837) 19.18.1. Dyschirius gracilis gracilis (Heer, 1837) [= lafertei Putzeys, 1846] References. Dyschirius lafertei: Apfelbeck, 1907a: 495 (Buenë River). COLEOPTERA: CARABIDAE: SCARITINAE

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Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB). Notes. Balkenohl (2003: 229) omitted this species for the Balkans in spite of at least three localities from the region that were showed by Fedorenko (1996: 129, Fig. 355). P. Bulirsch (personal communication) informed me that there is a reliable record of D. gracilis from Northwestern Greece. The species is confirmed here for the fauna of Albania. 19.19. Dyschirius importunus Schaum, 1857 19.19.1. Dyschirius importunus importunus Schaum, 1857 References. Dyschirius importunus: Apfelbeck, 1904: 74 (Velipoja near Lezhë); Gryuntal, 1984a: 292; Hieke & Wrase, 1988: 33. Dyschirius importunus importunus: Balkenohl, 2003: 229. Material studied. “Alban. Wgth. Valona”, 3 s. (HNHM). Subgenus Paradyschirius Fedorenko, 1996 Species group “substriatus” 19.20. Dyschirius parallelus Motschulsky, 1844 19.20.1. Dyschirius parallelus ruficornis Putzeys, 1846 References. Dyschirius ruficornis: Apfelbeck, 1907a: 495 (Buenë River); Müller, 1922b: 79 (Shkodër; Berat); Messner, 1964: 277; Hieke & Wrase, 1988: 31. Dyschiriodes parallelus ruficornis: Fedorenko, 1996: 202 (Vijosë River, Tepelenë District). Dyschirius parallelus ruficornis: Balkenohl, 2003: 230. Material studied. “Berat” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB). 19.21. Dyschirius substriatus (Duftschmid, 1812) 19.21.1. Dyschirius substriatus priscus J. Müller, 1922 References. Dyschirius substriatus priscus: Müller, 1922b: 80 (Berat); Horion, 1941: 104; Ieniștea, 1979: 221; Hieke et al., 1986: 184 (Kula e Lumës near Kukës); Balkenohl, 2003: 230. Dyschirius substriatus: Hieke & Wrase, 1988: 32. Dyschiriodes substriatus: Fedorenko, 1996: 204.

Tribe SCARITINI Bonelli, 1810 Subtribe SCARITINA Bonelli, 1810 20. Genus Scarites Fabricius, 1775 Subgenus Parallelomorphus Motschulsky, 1849 20.1. Scarites laevigatus Fabricius, 1792 References. Scarites laevigatus: Apfelbeck, 1904: 67 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke & Wrase, 1988: 27; Balkenohl, 2003: 232. 50


Material studied. “Durazzo” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Himara Juni” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); 2 km N Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Shkodër, 12.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS). 20.2. Scarites terricola Bonelli, 1813 20.2.1. Scarites terricola terricola Bonelli, 1813 Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albania Sauk” / “16.VI.1958 leg. Xh. Murraj”, 1 s. (HNHM); Saranda District, Butrinti, 16.IV.1994, 1 s., S. Beshkov leg. (NMNHS); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS). Notes. New species for the fauna of Albania. Subgenus Scallophorites Motschulsky, 1858 20.3. Scarites buparius (Forster, 1771) References. Scarites buparius: Hieke et al., 1986: 184 (Vlorë). Material examined. “Vom Alb. Amerik Institut…” / “Zweifellos Alb. Konvenienc”, 1 s. (MNHUB); “S Albanien Vlora, Strand p.7.58 leg. Grebenscikov”, 1 s. (DEI). Additional material included (studied A. Vigna Taglianti). Tiranë, botanical garden, 6.VI.1998, 1 s., A. Paparistos leg. (cAVT). Notes. Balkenohl (2003: 233) has omitted the species for Albania. Here it is confirmed for the country.

Subfamily APOTOMINAE LeConte, 1853 Tribe APOTOMINI LeConte, 1853 21. Genus Apotomus Illiger, 1807 21.1. Apotomus rufithorax Pecchioli, 1837 References. Apotomus rufithorax: Schatzmayr, 1943b: 99 (Mal i Krujes; Karbunara e Madhe); Wrase, 1992: 93 (Durrës); Wrase, 2003a: 237. Material studied. “Durres” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 1 m., 2 f., D. Antonelli leg. (cAVT). 21.2. Apotomus rufus (P. Rossi, 1790) References. Apotomus rufus: Wrase, 1992: 77 (Lum i Tiranë). Notes. Wrase (2003a: 237) has omitted the species for Albania, therefore it occurrence in the country needs further confirmation. COLEOPTERA: CARABIDAE: TRECHINAE

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Subfamily TRECHINAE Bonelli, 1810 Tribe BEMBIDIINI Stephens, 1827 Subtribe BEMBIDIINA Stephens, 1827 22. Genus Asaphidion Gozis, 1886 Notes. Hartmann has revised the material studied under this genus from HNHM. ? Asaphidion cyanicorne (Pandellé, 1867) ? Asaphidion cyanicorne cyanicorne (Pandellé, 1867) Material studied. “Korab” / “A. cyanicornis cyanicornis det. H. Schweiger” / “Fundort wahrsch. falsch!”, 1 m. with extracted genitalia (NMW); “Korab, Alb. Bischoff 1935” / “A. cyanicornis cyanicornis det. H. Schweiger”, 1 m. with extracted genitalia (NMW). Notes. Schweiger noted on the label that this locality might be untrustworthy. For the time being his remark is taken into consideration despite A. cyanicorne has been recorded for Croatia and Bulgaria (Mařan, 1933: 89; Marggi et al., 2003: 242). 22.1. Asaphidion flavipes (Linnaeus, 1761) References. Asaphidion flavipes: Apfelbeck, 1904: 79 (Velipoja near Lezhë); Schatzmayr, 1943b: 95 (Tërpan); Hieke & Wrase, 1988: 42. Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Mamuras Alban. Kruja Dr. J. Müller”, 5 s. (NMW); “Argirocastro Albania Weirather”, 2 s. (NMW); “Lumi Bençes Tepelene, Alb. m.”, 1 s. (NMW); “Berat Alb. Dr. Müller’18”, 1 s. (NMW); “Elbasan Alb., Mader”, 16 s. (HNHM; NMW); “Tomor-Buranj Juli 1932” / “Albania leg. Bischoff 1932”, 12 s. (MNHUB); “Bojana Shkoder 1934”, 8 s. (NMW); “Gjinokastrë Umg. Albania 1934 Bischoff ”, 1 ex. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Berat” / “Albania leg. Bischoff 1936”, 5 s. (NMW; MNHUB); Saranda District, Lyukove, 5.V.1994, 1 m., P. Stoev leg. (NMNHS); Përmeti District, Pelombasi, 2.VI.1994, 1 f., G. Blagoev leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 f., L. Penev leg. (NMNHS); Tiranë Park, 12-19.XI.1997, 1 f., R. Ward leg. (CMNH). Notes. Marggi et al. (2003: 242) have omitted the species for Albania. Here A. flavipes is confirmed for the country. 22.2. Asaphidion nebulosum (P. Rossi, 1792) 22.1.1. Asaphidion nebulosum balcanicum Netolitzky, 1918 References. Asaphidion caraboides: Apfelbeck, 1904: 78 (Durrës). Asaphidion caraboides balcanicus: Netolitzky, 1918: 215 (Buenë River near Shkodër; Berat); Schatzmayr, 1943b: 95 (Ura Zezes); Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 41; Sakalian & Guéorguiev, 1997: 46; Marggi et al., 2003: 242. Asaphidion caraboides var. balcanicus: Csiki, 1940: 212 (Kula Ljums; Gjalicë Mt.). Bembidion elegans: Murraj, 1968: 33 (Tomor Mt.). 52


Material studied. “Shkëlzën Alban.”, 3 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 14 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 62 s. (NMW); “Skutari, Alb. Klenka”, 3 s. (NMW); “Vojusa – Shkoder 1934”, 1 s. (NMW); “Dajtit Alb. Bischoff 1935”, 8 s. (NMW); “Cucale” / “Albania leg. Bischoff 1936”, 1 s. (NMW); Tomor Mt., 21.IX.1961, 2 m., 3 f., Xh. Murraj leg. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS). Notes. Bonavita & Vigna Taglianti (2005) concluded that A. caraboides and A. nebulosum are two distinct species, the first monotypic and living in Central Europe, the second polytypic (with four subspecies) and living in South Europe and Anatolia. The record of Murraj (1968) about Bembidion elegans is based on misidentification. Except the material mentioned above the present author has seen further more than 40 s. in MNHUB, which remained unregistered. 22.3. Asaphidion rossii (Schaum, 1857) References. Asaphidion rossii: Hieke et al., 1986: 186 (Uji Ftohte, south of Tepelenë); Marggi et al., 2003: 243. Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tiranë” / “Albania leg. Bischoff 1931”, 2 s. (NMW; MNHUB); “Bushek Alb. Elbasan Bischoff 1932”, more than 45 s. (NMW; MNHUB); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 2 s. (NMW; MNHUB); “Gjirakastrë Ung. Albania 1937 Bischoff 1934”, 3 s. (NMW); “Dajtit, Albania Bischoff 1935”, 8 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 11 s. (MNHUB); “Berat” / “Albania leg. Bischoff 1936”, 2 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Vojusa – Tepelene 1936”, 4 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Lem i Kuri Ura Mes” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1939”, 2 s. (MNHUB). 22.4. Asaphidion stierlini (Heyden, 1880) References. Asaphidion flavipes: Csiki, 1940: 212 (Kula Ljums). Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM). Notes. New species for the fauna of Albania.

23. Genus Bembidion Latreille, 1802 Subgenus Bembidion s.str. ? Bembidion humerale Sturm, 1825 References. Bembidion humerale: Murraj, 1968: 33 (Tomor Mt.). Notes. The record of Murraj (1968) might be doubtful because of some misidentifications found in his collection. Unfortunately, the cited material cannot be found. ? Bembidion quadrimaculatum (Linnaeus, 1761) ? Bembidion quadrimaculatum quadrimaculatum (Linnaeus, 1761) [= quadriguttatum (Fabricius, 1775) nec Pontoppidan, 1763] References. Bembidion quadrimaculatum: Csiki, 1940: 213 (Kula Ljums). Bembidion quadriguttatum (Fabricius): Csiki, 1940: 213 (Kula Ljums). COLEOPTERA: CARABIDAE: TRECHINAE

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Notes. In HNHM, I found single specimen identical with that cited by Csiki (1940) to belong to B. quadripustulatum. The occurrence of B. quadrimaculatum in Albania is very probable due to the distribution of this common species on the Balkans. 23.1. Bembidion quadripustulatum Audinet-Serville, 1821 23.1.1. Bembidion quadripustulatum quadripustulatum Audinet-Serville, 1821 [= quadriguttatum (Olivier, 1795) nec Fabricius, 1775] Material studied. “Skutari Bojana” / “Apfelbeck”, 1 s. (NMW); “Alban., Wgth. Valona” / “Apfelbeck”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Skutari, Alb. Klenka”, 3 s. (NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Bojana - Ufer b. Shkoder, Alb.” / “Alb. Winkler 1934”, 1 s. (NMW); “Durazzo, Alb. Bischoff ”, 1 s. (NMW); Tiranë Park, 1.III.1998, 1 s., R. Ward leg. (CMNH). Notes. New species for the fauna of Albania. Subgenus Bembidionetolitzkya E. Strand, 1929 Notes. This is one of the most complex subgenera from the genus in the Balkans. Apparently, the center of diversity of this subgenus in the Peninsula is situated in the northern half of Albania and the adjacent territories of Montenegro and Kosovo. The taxonomical problems with several yet not specified forms inhabiting the studied territory deserve special endeavour. Species group “tibiale” 23.2. Bembidion rhodopense Apfelbeck, 1902 References. Bembidion rhodopense: Hieke & Wrase, 1988: 47 (Korab Mt.); Guéorguiev & Guéorguiev, 1995: 106; Guéorguiev et al., 1997: 14; Sakalian & Guéorguiev, 1997: 46. Material studied. “…Albanien”, 5 s. (NMW); “Shkëlzën Alb. Bischoff 1935”, 1 s. (NMW); “Korab” / “Albania leg. Bischoff 1935”, 2 s. (NMW; MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 5 s. (NMW; MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 27 s. (NMW; MNHUB). Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for the country. 23.3. Bembidion tibiale (Duftschmid, 1812) References. Bembidion tibiale: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke & Wrase, 1988: 47. Material studied. “Shkëlzën Alban.”, 1 s. (NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9 s. (MNHUB). Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for the country.

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Species group “complanatum-geniculatum” 23.4. Bembidion geniculatum Heer, 1837 23.4.1. Bembidion geniculatum geniculatum Heer, 1837 References. Bembidion geniculatum: Hieke & Wrase, 1988: 48. Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Korab” / “Albania Bischoff 1935”, 1 s. (MNHUB); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (NMW). Notes. Marggi et al. (2003: 245) have omitted the species for Albania. Here B. geniculatum is confirmed for the country with first detailed records. 23.5. Bembidion justinae Meschnigg, 1947 References. Bembidion justinae: Meschnigg, 1947: 94 (North Albanian Alps: Dedaj, 780 m, south of Maja e Radohines Massif, near a little brook, loc. typ.); Turin, 1981: 81; Marggi et al., 2003: 245. Notes. Finding and revising the type of B. justinae is needed to clarify its systematic position and relationships with the related species. ? Bembidion relictum Apfelbeck, 1904 References. Bembidion relictum: Netolitzky, 1943a: 116; Hieke & Wrase, 1988: 47; Nitzu, 1995: 158. Material studied. “Dr. Penther Prokletija” / “Sammlung Apfelbeck”, 1 f. (HNHM). Notes. Hieke & Wrase (1988) and Nitzu (1995) noted this species for Albania. Obviously, these authors have followed the record “Albanian (Drin, Orsk leg. Winkler)” (Netolitzky, 1943a: 116). Actually this locality is situated in Western Kosovo close to the border with Montenegro. Due to lack of exact records B. relictum is still regarded as questionable for Albania; the material studied above is without definite state attribution. ! Bembidion sp. (aff. complanatum Heer, 1937) Material studied. “Jezerce K. Wgth.18”, 1 f. (NMW). Notes. Netolitzky has identified the female above as B. complanatum Heer, 1837. I examined it and think also that this specimen is externally most similar to B. complanatum. The female at hand has pointed apex of elytra and a body-size of 6.6-6.7 mm. The specimen from Maja e Jezerzë differs from B. justinae and B. relictum in the black antennomeres, excluding the reddish scapus. The study of more male specimens, including such from the same locality, and their comparison with specimens of B. complanatum, B. relictum and the type of B. justinae, could establish the correct status of this form. Species group “varicolor” 23.6. Bembidion conforme (Dejean, 1831) References. Bembidion conforme: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Horion, 1941: 132. Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Marggi et al. (2003: 245) have omitted for Albania. Here it is confirmed for the country. COLEOPTERA: CARABIDAE: TRECHINAE

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23.7. Bembidion meschniggi Netolitzky, 1943 References. Bembidion meschniggi: Netolitzky, 1943a: 113 (Dedaj, loc. typ.); Turin, 1981: 82; Marggi et al., 2003: 246. Material studied. “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” / “meschniggi n. sp. det. Netolitzky” / “Type” / “Genital - Nr 458 präpariert G. Müller 1978”, 1 syntype (NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Paul Meyer” / “Cotypus det. Netolitzky” / “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus anno 1996 revidit Miloš Fassati”, 1 syntype (NMW); “Albania Markdedaj Ing. Meschnigg” / “coll. Netolitzky” / “Cotypus det. Netolitzky” / “Type” / “meschniggi n. sp. det. Netolitzky” / “meschniggi Cotypus anno 1996 revidit Miloš Fassati”, 1 syntype (NMW). Notes. I consider B. meschniggi being a distinct species and most probably a sister taxon of B. atrocaeruleum (Stephens, 1828), but further study should differentiate the former from the latter and to prove its species status. There are in Albania two places with the name Dedaj (? = Markdedaj), situated not far from each other. The first place is north of the Drin River (Shkodër District), at the southern slopes of the North Albanian Alps, at 1012 meters altitude, whereas the second one is south of the Drin River (Pukë District), at 500 meters altitude. For the present one cannot specify what of both the type locality of B. meschniggi is. 23.8. Bembidion varicolor (Fabricius, 1803) 23.8.1. Bembidion varicolor varicolor (Fabricius, 1803) [= tricolor (Fabricius, 1801)] References. Bembidion tricolor: Apfelbeck, 1904: 87 (Velipoja); Hieke & Wrase, 1988: 46. Material studied. “Shkëlzën Alban.”, 2 s. (NMW); “Shkëlzën Alb. Alfons Bischoff ”, 1 s. (NMW); “Shkëlzën” / “Albania Bischoff 1935”, 9 s. (MNHUB). Notes. Marggi et al. (2003: 246) have omitted the species for Albania. Here it is confirmed for the country. Species group “fasciolatum” # Bembidion ascendens K. Daniel, 1902 References. Bembidion ascendens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi et al., 2003: 245. Notes. Sure records for this species from the Balkan Peninsula are known from Slovenia, Croatia, and Western Bosnia Herzegovina. Since no reliable records are known eastwards of the city of Sarajevo, B. ascendens is removed from the list of the Albanian ground beetles. Most likely, the literature data for it refer to B. brunoi. 23.9. Bembidion brunoi (Bonavita, 2001) References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 468 (Kiri River near Mesi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Marggi et al., 2003: 245. Bembidion ascendens: Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al., 2003: 245. Material studied. “Skutari Mesi”, 1 f. (NMW); “Merdita Oroshi” / “Alban. spt. IV.V.’05”, 2 m., 4 f. (NMW); “Mesi Kiri Ufer” / “Albania leg. Bischoff 1931”, 1 m., 2 f. (MNHUB); 5 km from Tepelenë, Drina River, 3.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS). 56


Notes. In both the shape and the internal structure of the median lobe this species seems most related to B. fasciolatum. On the other hand, in the wider margin of the pronotum and bigger size of the body, it resembles B. ascendens. 23.10. Bembidion concoeruleum Netolitzky, 1943 References. Bembidion coeruleum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Hieke & Wrase, 1988: 47. Bembidion concoeruleum: Netolitzky, 1943a: 114 (Oroshi; Shkodër; Dukati); Marggi et al., 2003: 245. Bembidion coeruleum concoeruleum: Hieke et al., 1986: 185 (Borshi near Vlorë; Uji Ftohte, south of Tepelenë). Ocydromus concoeruleus: Bonavita & Vigna Taglianti, 1993: 84. Material studied. “Orosi (Ptr.) Alban. spt.”, 5 s. (HNHM; NMW); “Skutari Mesi” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Prokletija”, 1 s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Dukati”, 1 s. (NMW); “Skutari, Alb. Klenka”, 4 s. (NMW); “Skutari, Alb. Mader”, 1 s. (NMW); “Lumi, Benges Tepelene, Alb. m.”, 2 s. (NMW); “Argirokastron Albania mai 1929 Weirather”, 6 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff V.1931”, 1 s. (NMW); “Korab, Alb. Guri Kuq Bischoff 1936”, 2 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 2 m., G. Blagoev leg. (NMNHS). Notes. All previously published data for B. coeruleum Audinet-Serville, 1821 from the Balkans refer to this species (see also Bonavita & Vigna Taglianti, 1993). # Bembidion fasciolatum (Duftschmid, 1812) References. Bembidion fasciolatum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte near Tepelenë); Marggi et al., 2003: 245. Notes. I failed to find material of this species from the Balkan Peninsula in the collections, excluding a single specimen from the locality “Travnik”. There are two places with this name - one in Slovenia and one in Bosnia Herzegovina. B. fasciolatum is removed from the list of the Albanian Carabidae. The records cited for it in the past most probably concern either an unnamed species (the record near Oroshi) or B. brunoi (the records by Mesi River and in South Albania). ! Bembidion sp. (ex. gruppo “fasciolatum”) References. Bembidion fasciolatum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Marggi et al., 2003: 245. Material studied. “Orosi (Ptr.) Alban. spt.”, 6 s. (DEI; HNHM; MNHUB). Notes. The specimens from DEI and HNHM have been identified as Bembidion merditanum Apfelbeck as two of them from the first depository were incorrectly designated as syntypes of this form. In fact, Bembidion merditanum Apfelbeck is nomen nudum. The series from Oroshi may belong to separate species. [subgenus Bracteon Bedel, 1879] [Bembidion litorale (Olivier, 1790) [= elegans Germar, 1824]] References. Bembidion elegans: Murraj, 1968: 33 (Tomor Mt.). Notes. Murraj (1968) cited this species from the Tomor Mt. Among the material loaned from MNST, I found five specimens from this locality bearing label data as the published ones and COLEOPTERA: CARABIDAE: TRECHINAE

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identified by the author as B. elegans. In fact, all the specimens concern Asaphidion nebulosum balcanicum. For the present B. litorale is treated as an expected species for Albania because it inhabits most of the Balkan countries. Subgenus Chlorodium Motschulsky, 1864 23.11. Bembidion splendidum Sturm, 1825 23.11.1. Bembidion splendidum pincum De Monte, 1957 References. Bembidion splendidum: Apfelbeck, 1907a: 495 (Buenë River). Material studied. “Scutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 3 s. (NMW). Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for the country. According to Bonavita and Vigna Taglianti (2005) the Balkan populations of the species belong to ssp. pincum. Subgenus Diplocampa Bedel, 1896 23.12. Bembidion assimile Gyllenhal, 1810 References. Bembidion assimile: Apfelbeck, 1904: 111 (Velipoja); Hieke & Wrase, 1988: 57. Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM). Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here it is confirmed for the country. 23.13. Bembidion fumigatum (Duftschmid, 1812) References. Bembidion fumigatum: Apfelbeck, 1904: 111 (Velipoja near Lezhë); Hieke & Wrase, 1988: 57. Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Actually, further confirmation for the country is needed. Subgenus Emphanes Motschulsky, 1850 23.14. Bembidion axillare (Motschulsky, 1844) 23.14. Bembidion axillare occiduum Margi & Huber, 2001 [= rivulare (Dejean, 1831)] References. Bembidion rivulare: Hieke & Wrase, 1988: 58. Material studied. “Durrës Alban.”, 1 s. (NMW); “Albanien Ruskuli 24.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Ruskuli Alb., Mader”, 1 s. (NMW); “Durazzo, Alb. Ravasini, Lona”, 2 s. (HNHM); “Durrazo, Alb. Bischoff ”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 9 s. (MNHUB); “Valona, Alb. Bischoff 1936”, 2 s. (NMW); Radona, Shkiela, 21.VI.1994, 3 s., B. Guéorguiev leg. (NMNHS). Notes. Marggi et al. (2003: 248) have omitted the species for Albania. Here B. axillare is confirmed for the country with first detailed records. 58


23.15. Bembidion azurescens Dalla Torre, 1877 23.15.1. Bembidion azurescens azurescens Dalla Torre, 1877 References. Bembidion azurescens: Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Kula e Lumës near Kukës); Hieke & Wrase, 1988: 58. Bembidion azurescens azurescens: Marggi et al., 2003: 248. Material studied. “Oboti Schirel” / “Apfelbeck”, 1 s. (NMW); “Berat, Alb. Ravasini, Lona”, 3 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 8 s. (NMW). 23.16. Bembidion latiplaga Chaudoir, 1850 23.16.1. Bembidion latiplaga latiplaga Chaudoir, 1850 References. Bembidion latiplaga: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988: 59. Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Oboti Schirei” / “Apfelbeck”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW; MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 3 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb. Winkler 1937”, 1 s. (NMW); “BojanaUfer b. Shkoder, Alb.” / “dch. Winkler XI 1942”, 1 s. (NMW); “Bojana Oboti, Alb.” / “ex coll. Winkler 1948”, 1 s. (NMW); “Tirana Ungb. Albania” / “ex coll. Winkler 1948”, 2 s. (NMW). Notes. Marggi et al. (2003: 249) have omitted the species for Albania. Here it is confirmed for the country. 23.17. Bembidion normannum Dejean, 1831 23.17.1. Bembidion normannum mediterraneum Csiki, 1928 [= meridionale Ganglbauer, 1891] References. Bembidion normannum meridionale: Apfelbeck, 1904: 103 (Velipoja near Lezhë). Bembidion normannum apfelbecki: Müller-Motzfeld, 1986a: 262 (Raskuli); Hieke & Wrase, 1988: 59. Bembidion normannum mediterraneum: Hieke & Wrase, 1988: 59; Marggi et al., 2003: 249. Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM). Notes. Hieke & Wrase (1988) have noted that both subspecies of B. normannum inhabit Albania. However, I consider the material studied and reference data as belonging to ssp. mediterraneum (see also Marggi et al., 2003: 249). 23.18. Bembidion tenellum Erichson, 1837 23.18.1. Bembidion tenellum tenellum Erichson, 1837 References. Bembidion tenellum: Csiki, 1940: 213 (Kula Ljums). Material studied. “Skutari Vraka” / “Apfelbeck”, 1 s. (NMW); “Oboti Schiren” / “Apfelbeck”, 1 s. (NMW); “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 7 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. Dr. J. Müller”, 3 s. (NMW); “Berat, Alb. Ravasini, Lona”, 1 s. (NMW); “Liqen Butrintës Alb. 1936. Bischoff ”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb. Winkler 1937”, 1 s. (NMW); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 22.II.1998, 2 s., from grasses along small sluggish stream, R. Ward leg. (CMNH). COLEOPTERA: CARABIDAE: TRECHINAE

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Notes. Marggi et al. (2003: 249) have omitted the species for Albania. Here it is confirmed for the country. Subgenus Euperyphus Jeannel, 1941 23.19. Bembidion combustum Ménétries, 1832 23.19.1. Bembidion combustum combustum Ménétries, 1832 References. Bembidion combustum: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 48. Bembidion combustum combustum: Marggi et al., 2003: 249. Material studied. “Albania, Berat”, 1 s. (NMW); “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 9 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 11 s. (HNHM; MNHUB); “Berat, Alb. Ravasini, Lona”, 3 s. (NMW); “Skutari, Alb. Klenka”, 1 s. (NMW); “Tiranë, Alb. Alf. Bischoff ”, 2 s. (NMW); “Cucale Geb.” / “Albania leg. Bischoff 1931”, 1 s. (NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranës Alb. V.1934 Mjr. Strupi”, 2 s. (NMW); “Lum Erzen Alb. V.1934 Mjr. Strupi”, 2 s. (NMW); “Tiranë” / “lgt. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “lgt. Bischoff Albania 1935”, 2 s. (MNHUB); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST). 23.20. Bembidion scapulare Dejean, 1831 23.20.1. Bembidion scapulare oblongum Dejean, 1831 References. Bembidion oblongum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi). Bembidion scapulare oblongum: Netolitzky, 1943b: 17; Hieke & Wrase, 1988: 50. Material studied. “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW); “Berat Alb. Dr. J. Müller’18”, 1 s. (NMW); “Elbasan, Alb. 4.IX.18. Dr. H. Priesner”, 1 s. (NMW); “Elbasan Alb., Mader”, 3 s. (NMW); “Lumi Dukati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Elbasan Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomorica” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 2 s. (NMW; MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS). Notes. Marggi et al. (2003: 250) have omitted the species for Albania. Here it is confirmed for the country. Presently the subspecific status of the Balkan populations remains unsettled. 23.21. Bembidion testaceum (Duftschmid, 1812) 23.21.1. Bembidion testaceum testaceum (Duftschmid, 1812) Material studied. “Berat, Alb. Ravasini, Lona”, 1 s. (NMW); “Bojana-Schkodër Alb., Bischoff 1934”, 1 s. (NMW). Notes. New species for the fauna of Albania. Subgenus Metallina Motschulsky, 1850 23.22. Bembidion lampros (Herbst, 1784) References. Bembidion lampros: Apfelbeck, 1904: 83 (Velipoja; Vlorë); Apfelbeck, 1909: 297 (Vlorë; Sop); Csiki, 1940: 212 (Gjalicë Mt.); Hieke & Wrase, 1988: 44; Marggi et al., 2003: 251. 60


Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); North Albanian Alps, Bogë, 1000-1100 m, 30.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1600 m, near cave B 52, 20.VI.1994, 1 s., T. Ivanova leg. (NMNHS); Tiranë Park, 22-31.X.1997 / 1-9.IX.1997 / 8.II.1998 / 11.II.1998 / 15.II.1998 / 22.II.1998, 9 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. Except the specimens mentioned above, I have seen additionally nearly 100 s. in NMW and MNHUB, which remained unregistered. 23.23. Bembidion properans (Stephens, 1828) References. Bembidion lampros var. properans: Csiki, 1940: 212 (Kula Ljums). Bembidion properans: Hieke & Wrase, 1988: 44. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan Alb., Mader”, 1 s. (HNHM); “Lumi i Tiranës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB); Tiranë Park, 8.II.1998, 2 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here it is confirmed for the country. Except the specimens mentioned above, in NMW the present author has seen other specimens, which remained not registered. Subgenus Microserrullula Netolitzky, 1921 23.24. Bembidion quadricolle (Motschulsky, 1844) [= inserticeps Chaudoir, 1850; cyaneomicans Mařan, 1933] References. Bembidion inserticeps: Apfelbeck, 1907a: 495 (Buenë River). Bembidion quadricolle: Hieke & Wrase, 1988: 42; Nitzu, 2003: 180. Material studied. “Albania Scutari”, 1 s. (MNHUB); “Skutari Bojana”, 2 s. (HNHM); “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 5 s. (NMW; MNHUB); “Berat, Alb. Ravasini, Lona”, 1 s. (NMW); “Berat, Alb. Ravasini, Lona” / “cyaneomicans det. Paul Meyer”, 1 s. (NMW); “Berat” / “Albania leg. Bischoff 1931”, 1 s. (NMW); “Lumi Berati Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tepelene” / “Albania leg. Bischoff 1932”, 5 s. (NMW; MNHUB); “Ura Erzen” / “Albania leg. Bischoff. 1932”, 2 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1934”, 2 s. (NMW). Notes. Marggi et al. (2003: 251) have omitted the species for Albania. Here B. quadricolle is confirmed for the country. Subgenus Nepha Motschulsky, 1864 23.25. Bembidion callosum Küster, 1847 23.25.1. Bembidion callosum callosum Küster, 1847 [= laterale Dejean, 1831; pseudolaterale Netolitzky, 1943a: 103] References. Bembidion laterale: Apfelbeck, 1904: 108 (Velipoja). Bembidion laterale pseudolaterale: Netolitzky, 1943a: 103. COLEOPTERA: CARABIDAE: TRECHINAE

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Material included (studied P. Bonavita). “Maja e Tartarit” / “Albania leg. Bischoff 1936”, 1 f. (NMW). Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for the country. 23.26. Bembidion caucasicum (Motschulsky, 1844) [= substriatum Chaudoir, 1846] References. Bembidion substriatum: Apfelbeck, 1907a: 501 (Merdita); Csiki, 1940: 213 (Shkëlzen Mt.; Gjalicë Mt.). Bembidion caucasicum: Müller, 1937: 124; Hieke & Wrase, 1988: 56. Material studied. “Merdita Munela”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania, Mts Gyalicalums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Gjallica, Alb. Bischoff ”, 1 s. (NMW); “Maja e Jezercë Albania Bischoff leg.”, 2 s. (NMW); “Korab, Guri Kua Alban. Bischoff 35”, 5 s. (NMW); “Shkëlzen, Alb. Bischoff 1935”, 1 s. (NMW); “Maja e Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (NMW); “Korab” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Shkëlzen, Alb. Bischoff 5.8.1936”, 3 s. (HNHM; NMW); North Albanian Alps, Maj’e Radohimes, 2200-2400 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 2 f., L. Latella leg. (cAVT). Notes. Marggi et al. (2003: 252) have omitted the species for Albania. Here it is confirmed for the country. Besides the material studied above, in NMW and MNHUB I have seen other numerous specimens from the massifs of the North Albanian Alps, Korab Mt., Koritnik Mt., Gjalicë Mt. and Maja e Policanit Mt. 23.27. Bembidion vseteckai Mařan, 1936 23.27.1. Bembidion vseteckai dissimile G. Müller, 1943 References. Bembidion genei illigeri: Müller, 1943: 13; Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Kula e Lumës near Kukës); Hieke & Wrase, 1988: 55. Bembidion illigeri: Marggi et al., 2003: 253. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Elbasan, Alb. Leop. Mader leg.”, 2 s. (NMW); “Kruja Alb. Leop. Mader”, 2 s. (NMW); “Terbaci Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); Mal i Dajtit Mt., 6.V.1962, 6 s. (MNST). Notes. Except the specimens studied above I have seen several additional ones in NMW, which remained unregistered. Bonavita (2005), as well P. Bonavita and A. Vigna Taglianti (personal communication), noticed that all records or literature data about B. genei illigeri Netolitzky, 1914 from the southern parts of Balkan Peninsula concern B. vseteckai dissimile. Subgenus Notaphemphanes Netolitzky, 1920 23.28. Bembidion ephippium (Marsham, 1802) References. Bembidion ephippium: Hieke & Wrase, 1988: 46. Material studied. “Valona, Albania”, 4 s. (NMW); “Durazzo Alb. Dr. J. Müller”, 5 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB). 62


Notes. Marggi et al. (2003: 253) have omitted the species for Albania. Here it is confirmed for the country with first detailed records. Subgenus Notaphus Dejean, 1821 23.29. Bembidion varium (Olivier, 1795) [= rumelicum Apfelbeck, 1902; dyrrhachii De Monte, 1947] References. Bembidion varium: Apfelbeck, 1894: 547; Apfelbeck, 1904: 85 (Velipoja near Lezhë); Hieke & Wrase, 1988: 46; Marggi et al., 2003: 254. Bembidion varium dyrrhachii: De Monte, 1947: 52 (Durrës, loc. typ.). Bembidion rumelicum: Guéorguiev & Guéorguiev, 1995: 100. Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Oboti Schirei” / “Apfelbeck”, 1 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 12 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 7 s. (MNHUB); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW); Radona, Shkiela, 21.VI.1994, 4 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 22.II.1998, 30 s., from grasses along small sluggish stream, R. Ward leg. (CMNH). Notes. Except the material studied above the present author has seen more specimens in NMW, which remained unregistered. Subgenus Ocydromus Clairville, 1806 23.30. Bembidion aetolicum Apfelbeck, 1901 23.30.1. Bembidion aetolicum aetolicum Apfelbeck, 1901 References. Bembidion aetolicum: Netolitzky, 1943b: 12 (Tomor Mt.). Bembidion aetolicum aetolicum: Müller-Motzfeld, 1996b: 164 (Tomor Mt.: Buranj; Tomor Mt.: Kulmak); Marggi et al., 2003: 254. Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor Buranj Alb. 10.4.1932 Alf. Bischoff ”, 4 s. (NMW); “Barshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 2 s. (NMW). 23.31. Bembidion decorum (Zenker in Panzer, 1799) 23.31.1. Bembidion decorum decorum (Zenker in Panzer, 1799) References. Bembidion decorum: Apfelbeck, 1907a: 496, 498 (Kiri River near Mesi; Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 49. Bembidion decorum decorum: Müller-Motzfeld, 1996b: 153 (Shkodër; Mal i Dajtit Mt.; Shkëlzen Mt.; Bushek near Elbasan; Tiranë; Zeza-Brüche near Kruja; Lumi i Bençes near Tepelenë; Tomor Mt.: Kulmak; Miloti; Gjirokastër; Prespa Lake; Cafa Stams; Kiri River near Mesi); Marggi et al., 2003: 255. Material studied. Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Except the specimens studied above, I have seen several more ones in NMW, which remained unregistered.

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23.32. Bembidion siculum Dejean, 1831 23.32.1. Bembidion siculum smyrnense Apfelbeck, 1904 References. Bembidion siculum: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576 m; Fani i Madh River near Oroshi); Hieke et al., 1986: 185 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 49. Bembidion siculum smyrnense: Müller-Motzfeld, 1996b: 170 (Shkodër; Maje Shbogut; Himara; Llogora; Radavac; Sarapta; Lura Mt.: Lurë e Vjetër); Marggi et al., 2003: 256. Material studied. “Merdita Oroschi”, 3 s. (NMW); “Oroši (Ptr.) Alban. spt.”, 5 s. (HNHM); “Winkler, Lona, Bischoff, V.1931” / “Terbaci Alban. mer.”, 1 s. (NMW); “Llogora” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Himara Juni” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); Përmeti District, Pelombashi, 2.VI.1994, 3 s., G. Blagoev leg. (NMNHS). Notes. Besides the material studied above, I have seen other more in NMW, which remained unregistered. Subgenus Ocyturanes Müller-Motzfeld, 1986 23.33. Bembidion balcanicum Apfelbeck, 1899 23.33.1. Bembidion balcanicum balcanicum Apfelbeck, 1899 References. Bembidion balcanicum: Müller, 1918b: 85 (Munella Mt.); Müller, 1926: 61 (Munella Mt.); Csiki, 1940: 213 (Gjalicë Mt.; Korab Mt.); Horion, 1941: 136; Netolitzky, 1943a: 42 (Korab Mt.); Hieke & Wrase, 1988: 52. Bembidion balcanicum balcanicum: Marggi et al., 2003: 256. Material studied. “Latif Koritnik”, 11 s. (HNHM; NMW); “Merdita Munela”, 1 s. (HNHM); “Merdita Munela” / “Wgth 1906 Merdita”, 10 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther,’14”, 2 s. (NMW); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania, Mts. Gyalicalums” / “Akad. Balk. Exp. Csiki 1918”, 21 s. (HNHM); “Alban Exped. Gialica Ljums 17.-26.VI’18”, 2 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Jezerce Albania Bischoff leg.”, 4 s. (NMW); “Maja e Jezerce Alban. sept.” / “Alb. Winkler 1934”, 1 s. (NMW); “Albania Korab 1935 Bischoff leg.”, 1 s. (NMW); “Giallica, Alb. Bischoff 1935”, 3 s. (NMW); “Tomor - Tojk Bischoff Alban.”, 3 s. (NMW); “Tomor - Tojk Bischoff 1935”, 3 s. (NMW); “Maja Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 3 s. (NMW); “Shkelzën, Alb. Bischoff, 5.8.1936” / “Albania leg. Bischoff 1936”, 8 s. (HNHM; NMW). Notes. Besides the material studied above, in NMW and MNHUB I have seen numerous other specimens from the massifs of Shkëlzën Mt., Gjalicë Mt., Korab Mt. and Tomor Mt., which remained unregistered. 23.34. Bembidion gudenzii (Neri, 1981) References. Bembidion gudenzii: Müller-Motzfeld, 1985: 225 (“Hochgebirge bei Valona (Epyrus)”). Notes. Marggi et al. (2003: 256) have omitted B. gudenzii for Albania. Actually, the species needs further confirmation for the country. 23.35. Bembidion pindicum Apfelbeck, 1901 References. Bembidion pindicum: Netolitzky, 1943a: 49 (Korab Mt.); Guéorguiev, 1999: 71. 64


Material studied. “Korab, Guri-Kua Alban. Bischoff 35”, 1 s. (NMW); “Tomor-Tojk Alban 1935 Bischoff leg.” / “pindicum var.? dtt. Netolitzky”, 1 s. (NMW); “Tomor - Tojk Alban. 1935 Bischoff leg.”, 2 s. (NMW). Notes. Marggi et al. (2003: 257) have omitted the species for Albania. The specimens studied have been found to be conspecific with three syntypes of B. pindicum labelled “Apfelbeck Peristeri” / “(Peryphus) pindicum typ. Apf.” / “Typus” (NMW), therefore the species is here confirmed for Albania. 23.36. Bembidion praeustum Dejean, 1831 23.36.1. Bembidion praeustum praeustum Dejean, 1831 References. Bembidion praeustum: Hieke et al. 1986: 186 (Uji Ftohte, soth of Tepelenë); Hieke & Wrase, 1988: 52. Bembidion praeustum praeustum: Marggi et al., 2003: 257. Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Tomor: Kulmak Albania mer.” / “Ravasini & Lona”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.” / “Alb. Winkler”, 1 s. (NMW); “Bushek b. Elbasan, Alb. Alf. Bischoff ”, 1 s. (NMW); “Maj’e Shbogut Alnania mer.” / “Winkler, Lona, Bischoff, V.1931”, 2 s. (NMW); “Humara juni” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB); “Maranaj, Alb. Bischoff, 1934”, 1 s. (NMW); “Lumi i Tiranës Alb. V.1934 Mjr. Strupi”, 1 s. (NMW); “Köleyrë” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Boge ober d. Quelle” / “Albania leg. Mueller 1939”, 1 s. (NMW); “Borshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 8 s. (HNHM; NMW); “Borshë, Alb. Bischoff 1940”, 1 s. (NMW); “Tirane, Alb. dch. Winkler 42”, 1 s. (NMW); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Përmeti District, Pelombasi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS). Notes. A part of the material in NMW has been determined as B. praeustum viridifluum J. Müller, 1929. However, I refer provisionally all the specimens studied to the nominotypical race. 23.37. Bembidion signatipenne Jacquelin du Val, 1852 References. Bembidion signatipenne: Hieke & Wrase, 1988: 53. Notes. Marggi et al. (2003: 257) have omitted B. signatipenne for Albania. Actually, the species needs further confirmation for the country. 23.38. Bembidion stolfai J. Müller, 1943 References. Bembidion stolfai: Müller, 1943: 12 (North Albanian Alps: Okol near Bogë, loc. typ.; Maja e Jezercë); Netolitzky, 1943a: 41 (North Albanian Alps: Okol near Bogë; Maja e Jezercë); Marggi et al., 2003: 257. Material studied. “Maja e Jezerce Albania sept.” / “Alb. Winkler 1934” / “coll. Netolitzky” / “stolfai m. n. sp. det. J. Müller” / “Co-typus” / “Coll. Netolitzky”, 1 f. syntype (NMW). Notes. The syntype of B. stolfai was studied and compared with specimens of B. reiseri reiseri Apfelbeck, 1902 from Bosnia Herzegovina (including a syntype labelled “reis. 1900 Cvrstica” / “Testediolum bosnicum typ. Apf.” / “Reiseri” / “Typus”), topotypical material of B. reiseri vranense Apfelbeck, 1902 and extensive material of B. balcanicum balcanicum. B. stolfai is distinct from the other two species in the pronotum wider than long being narrower in the anterior part and in COLEOPTERA: CARABIDAE: TRECHINAE

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the elytra having distinct transverse microsculpture. Besides, it differs from B. reiseri s.l. in the wrinkles on outer basal fovea of pronotum clear and long (vs. reduced), the shoulders of elytra distinct (vs. round) and the sides of elytra after middle parallel (vs. slightly round). B. stolfai is also living sympatrically with B. balcanicum balcanicum in the massif of Maja e Jezercë. Subgenus Odontium LeConte, 1848 23.39. Bembidion foraminosum Sturm, 1825 References. Bembidion foraminosum: Hieke & Wrase, 1988: 43. Material studied. “Tiranë” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB). Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for the country with first detailed record. 23.40. Bembidion striatum (Fabricius, 1792) References. Bembidion striatum: Netolitzky, 1918: Beilage zu Heft 4-6 (Shkodër; Oboti); Geisthardt, 1975: 8. Bembidion striatul sic!: Panin, 1942: 567 (Shkodër). Material studied. “Oboti Schirei”, 1 s. (NMW); “Skutari Bojana”, 5 s. (NMW); “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 4 s. (NMW). Notes. Marggi et al. (2003: 257) have omitted the species for Albania. Here it is confirmed for the country. Subgenus Omoperyphus Netolitzky, 1931 23.41. Bembidion hypocrita Dejean, 1831 23.41.1. Bembidion hypocrita illyricum Netolitzky, 1918 References. Bembidion hypocrita illyricum: Müller, 1918b: 89; Müller, 1926: 66; Netolitzky, 1943a: 122. Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Mader”, 1 s. (NMW); “Argyrókastron, Alb. leg. Weirather”, 1 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 3 s. (NMW; MNHUB); “Shkelzën, Alb. Bischoff, 5.8.1936”, 1 s. (NMW); “Bischoff 21.VII.1940 Börshë-i-Vjetër Wildbach-Quellen”, 7 s. (NMW); “Börshë-i-Vjetër Alb. 21.7.1940 Bischoff ”, 1 s. (MNHUB). Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for the country with first detailed records. Subgenus Peryphanes Jeannel, 1941 23.42. Bembidion brunnicorne Dejean, 1831 23.42.1. Bembidion brunnicorne brunnicorne Dejean, 1831 References. Bembidion brunnicorne: Netolitzky, 1943a: 50; Csiki, 1940: 213 (Kula Ljums). Material studied. “Albania Kula Ljums”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 7 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Kruja, Alban., Mader”, 1 s. (NMW); “Tomor: Kulmak Albania mer.”, 3 s. (NMW); “Mal i Dajtit” / 66


“Albania leg. Bischoff 1935”, 2 s. (HNHM; MNHUB); “Maranaj 1937” / “Albania bei Skutari”, 1 s. (NMW); “Börshë i Vjetër Wildbach-Quellen Bischoff, 21.VII.1940”, 4 s. (NMW). Notes. Marggi et al. (2003: 258) have omitted the species for Albania. Here it is confirmed for the country. Besides the material listed above, I have seen additionally more than 70 s. from different localities in MNHUB, which remained unregistered. All the material in the last museum was identified as B. milleri. 23.43. Bembidion dalmatinum Dejean, 1831 23.43.1. Bembidion dalmatinum dalmatinum Dejean, 1831 References. Bembidion dalmaticum sic!: Anonymous, 1907: 286. Bembidion dalmatinum: Apfelbeck, 1907a: 498 (Fani i Madh River near Oroshi); Csiki, 1940: 213 (Gjalicë Mt.); Schatzmayr, 1943b: 95 (Dukati; Lushnje); Hieke & Wrase, 1988: 53. Chlaenius nigricornis: Murraj, 1968: 33 (Tiranë; Tomor Mt.). Bembidion dalmatinum dalmatinum: Marggi et al., 2003: 258. Material studied. Tomor Mt., 21.IX.1961, 9 s., Xh. Murraj leg. (MNST); Theth, 800-1200 m, 28.V.1993, 7 s., P. Beron & B. Petrov leg. (cPN; NMNHS); North Albanian Alps, Bogë, 14001800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District, Merkurth, 11.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); Përmeti District, Pelombashi, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Tepelenë District, Vijosë River, 2.VI.1994, 27 s., G. Blagoev leg. (NMNHS); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 1 s., L. Penev leg. (NMNHS); Leskoec - Barmashi, 880 m, 2.VI.1995, 2 s., G. Blagoev leg. (NMNHS); 5 km Tepelenë, Drina River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Ljukove near Vlorë, 6.VI.1995, 4 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH). Notes. About the record of Murraj (1968) above see the notes under Chlaenius nigricornis. Besides the material studied, the present author has seen numerous other specimens in NMW and MNHUB, which remained unregistered. 23.44. Bembidion deletum Audinet-Serville, 1821 23.44.1. Bembidion deletum deletum Audinet-Serville, 1821 Material studied. “Borshë, Alb. Bischoff 1940”, 1 s. (NMW). Notes. New species for the fauna of Albania. 23.45. Bembidion grandipenne Schaum, 1862 23.45.1. Bembidion grandipenne grandipenne Schaum, 1862 References. Bembidion grandipenne: Apfelbeck, 1907a: 498, 501 (Fani i Madh River near Oroshi); Hieke & Wrase, 1988: 54. Material studied. “Oroschi (Ptr.) Alban. spt.”, 4 s. (HNHM; MNHUB); “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Albania Malčija” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Skutari Alb., Mader”, 1 s. (NMW); “Lumi i Bençes Tepelene, Alb. m.”, 2 s. (NMW); “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 2 s. (NMW; MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shkëlzën, Alb. Bischoff 1935”, 1 s. (NMW); “Borshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 4 s. (NMW; MNHUB); “Borshë, Alb. Bischoff 1940”, 1 s. (NMW). COLEOPTERA: CARABIDAE: TRECHINAE

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Notes. Marggi et al. (2003: 259) have omitted the species for Albania. Here it is confirmed for the country. # Bembidion milleri Jacquelin du Val, 1852 # Bembidion milleri milleri Jacquelin du Val, 1852 References. Bembidion milleri: Hieke & Wrase, 1988: 54. Notes. According to Müller-Motzfeld (personal communication in 1994) B. milleri does not live in the Balkans. P. Bonavita and A. Vigna Taglianti (personal communication) stated that it occurs only in Slovenia and probably in Serbia. Here the upper opinions are followed, though Marggi et al. (2003: 258-259) noted the species from the Balkan Peninsula. Subgenus Peryphus Dejean, 1821 23.46. Bembidion bualei Jacquelin du Val, 1852 23.46.1. Bembidion bualei bualei Jacquelin du Val, 1852 [= albanicum J. Müller, 1935] References. Bembidion andreae: Apfelbeck, 1904: 92 (Durrës). Bembidion andreae albanicum: Müller, 1935: 152 (Berat, loc. typ.; Tiranë; Shkodër); Netolitzky, 1943a: 27 (Berat); Hieke et al. 1986: 186 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 51; Guéorguiev & Guéorguiev, 1995: 111; Guéorguiev et al., 1997: 15. Bembidion andreae bualei: Panin, 1942: 582 (Berat); Schatzmayr, 1943b: 95 (Ura Zezes). Bembidion cruciatum albanicum: Marggi et al., 2003: 260. Material studied. “Alban. Exped. Gjalica Ljuma 17.-26.VI.’18”, 6 s. (NMW); “Bojana Shkodër Alb., Bischoff 1937” / “Albania leg. Bischoff 1937”, 1 s. (HNHM). Notes. According to a recent study (Coulon, 2006) the Balkan populations of the species related to B. cruciatum Dejean, 1831 belong to B. bualei bualei, and the form albanicum is a synonym of the latter. Except the material studied above, I have seen more specimens from different localities in NMW and MNHUB, which remained unregistered. 23.47. Bembidion femoratum Sturm, 1825 23.47.1. Bembidion femoratum femoratum Sturm, 1825 Material studied. “Scutari, Alb. Klenka”, 1 s. (NMW); “Bojana, Alb. Shkodër Bischoff 1934”, 31 s. (NMW); “Albania leg. Bischoff 1937”, 1 s. (NMW); “Bojana Shkodër Albania leg. Bischoff 1937”, 4 s. (NMW); 5 km from Tepelenë, Drina River, 3.VI.1995, 2 s., G. Blagoev leg. (NMNHS). Notes. New species for the fauna of Albania. 23.48. Bembidion subcostatum (Motschuslky, 1850) 23.48.1. Bembidion subcostatum vau Netolitzky, 1913 [= javurkovae Fassati, 1944] References. Bembidion subcostatum: Hieke & Wrase, 1988: 52. Bembidion subcostatum javurkovae: Hieke et al. 1986: 186 (Borshi near Vlorë; Lurë e Vjetër near Lura Mt.); Marggi et al., 2003: 263. Material studied. “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 4 s. (NMW); “Scutari Alb. Dr. J. Müller”, 1 s. (NMW); “Berat, Alb. sch. Winkler”, 1 s. (NMW); “Tirana Alb. dch. Winkler 42”, 2 s. (NMW); “Bushek Juni” / “Berat Alban. mer.” / “Albania leg. Bischoff 1932”, 4 s. (NMW); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS). 68


Notes. Netolitzky identified the specimens in NMW as ssp. vau, ssp. javurkovae Fassati, 1944 or ssp. spartanum Fassati, 1944. However, in a forthcoming paper these three taxa are synonymized (Guéorguiev & Lobo, in press). In HNHM I have seen numerous additional specimens mostly from the localities “Korab” and “Kula Lums”, and in MNHUB - other more than 200 s., as all those data remained unregistered. 23.49. Bembidion tetracolum Say, 1823 23.49.1. Bembidion tetracolum tetracolum Say, 1823 References. Bembidion ustulatum (Linnaeus) (sic!): Csiki, 1940: 213 (Kula Ljums; Korab Mt.); Murraj, 1968: 33 (Tomor Mt.). Material studied. “Dukati” / “Apfelbeck”, 1 s. (NMW); “Merdita M. Scheit” / “Mali Scheit Apfelbeck”, 1 s. (NMW); “Alban. Vermosa Penther”, 1 s. (NMW); “Kruja, Alban., Mader”, 4 s. (NMW); “Berat Alban. mer.”, 2 s. (NMW); “Albania leg. Bischoff 1936”, 2 s. (NMW); “Kèlcyrë” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Schkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW); Tomor Mt., 21.IX.1961, 1 m., Xh. Murraj leg. (MNST); Ljukova, 100 m, 5.VI.1995, 1 m., 1 f., G. Blagoev leg. (NMNHS). Notes. Marggi et al. (2003: 263) have omitted the species for Albania. Here it is confirmed for the country. The record of Murraj (1968) is included here since the revision of one male from the Tomor Mt., having the same label data as the published material, has been found to be correct. Subgenus Philochthus Stephens, 1828 23.50. Bembidion decolor Apfelbeck, 1911 References. Bembidion decolor: Apfelbeck, 1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916: 340-341 (Buenë River; Vlorë); Netolitzky, 1943a: 63 (Elbasan; Kruja); Hieke & Wrase, 1988: 62; Marggi et al., 2003: 263. Material studied. “Alban. Wgth Valona” / “Coll. Apfelbeck”, 1 m. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM). Notes. Apfelbeck published two descriptions of this species, the first in Latin and SerboCroatian and second one in Latin and German (Apfelbeck, 1911; Apfelbeck, 1916). The older one is correctly treated as valid. 23.51. Bembidion guttula (Fabricius, 1792) 23.51.1. Bembidion guttula guttula (Fabricius, 1792) References. Bembidion guttula: Hieke & Wrase, 1988: 61. Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Alb. Winkler leg.”, 1 s. (NMW); “Maje Tartarit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for the country with first detailed records.


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23.52. Bembidion inoptatum Schaum, 1857 References. Bembidion inoptatum: Apfelbeck, 1904: 112 (Velipoja near Lezhë; Vlorë); Apfelbeck, 1911: 210 (Buenë River; Vlorë); Apfelbeck, 1916: 341 (Buenë River; Vlorë); Csiki, 1940: 213 (Kula Ljums); Hieke & Wrase, 1988: 62. Material studied. “Bojana Oboti, Alb.” / “ex coll. Winkler 1948”, 1 s. (NMW). Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for the country. 23.53. Bembidion iricolor Bedel, 1879 References. Bembidion iricolor: Apfelbeck, 1909: 297 (Vlorë). Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Actually, it needs further confirmation for the country. 23.54. Bembidion lunulatum (Geoffroy, 1785) References. Bembidion lunulatum: Schatzmayr, 1943b: 96 (Stan-Karbunarë). Material studied. “Elbasan Alb., Mader”, 4 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 2 s. (NMW); “Tirana, Alb., dch. Winkler 42”, 1 s. (NMW); Tiranë Park, 22.II.1998, 1 s., from grasses along small sluggish stream, R. Ward leg. (CMNH). Notes. Marggi et al. (2003: 264) have omitted the species for Albania. Here it is confirmed for the country. 23.55. Bembidion mannerheimii C.R. Sahlberg, 1827 Material studied. “Tomor: Kulmak Albania mer.”, 1 s. (NMW); “Tomor: Kulmak Albania mer.” / “lg. Winkler mai 1931”, 5 s. (NMW). Notes. New species for the fauna of Albania. Netolitzky has identified the material from the Balkans either as “Bembidion neresheimeri” or as “cfr. neresheimeri”. I determined the material from Albania as B. mannerheimii. According to D.W. Wrase (personal communication) B. neresheimeri J. Müller, 1929 is a distinct species that also may inhabit the Balkan Peninsula; Marggi et al. (2003: 264) have cited the latter only for Slovenia. Subgenus Phyla Motschulsky, 1844 23.56. Bembidion tethys Netolitzky, 1926 References. Bembidion tethys: Focarile, 1959: 58; Hieke & Wrase, 1988: 61. Material studied. “Elbasan Alb. 3.IX.18 Dr. H. Priesner”, 4 s. (NMW; MNHUB); “Elbasan Alb., Mader”, 2 s. (NMW); “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW); “Lum Erzen Alb. V.1934 Mjr. Strupi”, 1 s. (NMW); Ljukove, 100 m, 5.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 23-28.II.1998, 3 s., R. Ward leg. (CMNH). Notes. Marggi et al. (2003: 265) have omitted the species for Albania. Here it is confirmed for the country with first detailed records for the country.

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Subgenus Princidium Motschulsky, 1864 23.57. Bembidion punctulatum Drapiez, 1820 23.57.1. Bembidion punctulatum punctulatum Drapiez, 1820 References. Bembidion punctulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke et al., 1986: 185 (Uji Ftohte, south of Tepelenë; Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 44. Material studied. “Tirane” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Tiranës” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); Tepelenë District, Drina River, 3.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Marggi et al. (2003: 266) have omitted the species for Albania. Here it is confirmed for the country. Besides the material studied above, the present author has seen some more specimens in NMW, which remained unregistered. Subgenus Talanes Motschulsky, 1864 Notes. New subgenus for the fauna of Albania. 23.58. Bembidion subfasciatum Chaudoir, 1850 Material studied. “Bojana - Umg. Pulaj” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Ruskuli Alb., Mader”, 1 s. (NMW); “Liqen Butrintes” / “leg. Bischoff. 1936”, 3 s. (NMW). Notes. New species for the fauna of Albania. Subgenus Terminophanes Müller-Motzfeld, 1998 Notes. New subgenus for the faunas of the Balkan Peninsula and Albania. 23.59. Bembidion terminale Heer, 1841 ! Bembidion terminale ssp. Material studied. “Albania, Alban Alps, Okol, 800 m, 15-20.VII.1996, leg. G. Holzinger”, 1 f. (HNHM). Notes. New species for the faunas of the Balkan Peninsula and Albania. For the time being the subspecific status of the local population remains unspecified. The specimen in question has a somewhat different shape of the pronotum, compared with specimens from the Alps and North Iran. Subgenus Testedium Motschulsky, 1864 23.60. Bembidion bipunctatum (Linnaeus, 1761) 23.60.1. Bembidion bipunctatum nivale Heer, 1837 References. Bembidion bipunctatum var. pyritosum: Csiki, 1940: 212 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.). Bembidion bipunctatum: Hieke & Wrase, 1988: 45. Material studied. “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, more than 50 s. (HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania Mts. COLEOPTERA: CARABIDAE: TRECHINAE

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Korab” / “Akad. Balk. Exp. Csiki 1918”, 15 s. (HNHM); “Korab Preshk, Alb. Bischoff 1936”, 1 s. (HNHM). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 4 m., 1 f., L. Latella leg. (cAVT). Notes. Marggi et al. (2003: 269) have omitted the species for Albania. Here it is confirmed for the country. Besides the material studied above, I have seen other numerous specimens from different localities in NMW and MNHUB, which remained unregistered. Subgenus Trepanes Motschulsky, 1864 23.61. Bembidion articulatum (Panzer, 1796) References. Bembidion articulatum: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Marggi et al., 2003: 270. Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari Alb. Dr. J. Müller”, 1 s. (NMW); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 3 s. (NMW). 23.62. Bembidion maculatum Dejean, 1831 23.62.1. Bembidion maculatum maculatum Dejean, 1831 Material studied. “Tirana, Alb. Ravasini, Lona”, 1 s. (NMW). Notes. New species for the fauna of Albania. 23.63. Bembidion octomaculatum (Goeze, 1777) References. Bembidion octomaculatum: Apfelbeck, 1904: 110 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Kiri River near Mesi); Hieke & Wrase, 1988: 60. Material studied. “Alban. Wgth. Valona” / “Apfelbeck”, 1 s. (NMW); “Fieri, Albania a Semeni, Müller”, 4 s. (NMW); “Skutari, Alb. Prof. Dr. Müller”, 2 s. (NMW); “Bojana-Ufer b. Shkoder, Alb.” / “Alb. Winker 1934”, 1 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW). Notes. Marggi et al. (2003: 270) have omitted the species for Albania. Here it is confirmed for the country.

24. Genus Ocys Stephens, 1829 24.1. Ocys harpaloides (Audinet-Serville, 1821) References. Ocys harpaloides: Apfelbeck, 1904: 117 (Vlorë); Apfelbeck, 1909: 297 (Pasha Liman). Material studied. “Elbasan Alb., Mader”, 1 s. (NMW); “Scutari Alb., Mader”, 2 s. (NMW); “Tiranë” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Borshë-i-Vjetër Wildbach-Quellen Bischoff 21.VII.1940”, 1 s. (NMW). Notes. Marggi et al. (2003: 272) have omitted the species for Albania. Here it is confirmed for the country.

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25. Genus Sinechostictus Motschulsky, 1864 Subgenus Pseudolimnaeum Kraatz, 1888 Notes. New subgenus for the fauna of Albania. 25.1. Sinechostictus doderoi (Ganglbauer, 1891) Material studied. “Tomor Buranj Alb. 10.7.1932 Alf. Bischoff ”, 1 s. (NMW); “Sckëlzën” / “Albania Bischoff 1935”, 2 s. (NMW). Notes. New species for the fauna of Albania. Subgenus Sinechostictus s.str. 25.2. Sinechostictus tarsicus (Peyron, 1858) References. Bembidion elongatum var. effluviorum: Csiki, 1940: 213 (Kula Ljums). Bembidion elongatum Dejean: Hieke & Wrase, 1988: 56. Material studied. “Albania Buneševce” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 6 s. (HNHM); “Miloti Alban. Matzenauer”, 1 s. (NMW); “Elbasan Alb., Mader”, 1 s. (NMW); “Lumi Berati Alban. mer.”, 1 s. (NMW); “Lumi Beratit Alban. mer. Lona. Winkler.”, 1 s. (NMW); “Llogora” / “Albania leg. Bischoff 1932”, 8 s. (MNHUB); “Llogora, Alb. Alf. Bischoff 2.VI.1932”, 3 s. (NMW); “Llogora” / “Albania leg. Bischoff 1935”, 1 s. (NMW). Notes. Marggi et al. (2003: 267 sub Bembidion elongatum tarsicum) have omitted the species for Albania. Here it is confirmed for the country. Bonavita & Vigna Taglianti (2005) concluded that in the Balkan Peninsula is present only S. tarsicus, a distinct species, while S. elongatus (Ganglbauer, 1891) is a West European species. S. effluviorus (Peyron, 1858) is a separate species confined only to the territory of Asian Turkey. 25.3. Sinechostictus millerianus (Heyden, 1883) Material studied. “Shkëlzen” / “Albania leg. Bischoff 1936”, 1 s. (NMW). Notes. New species for the fauna of Albania.

Subtribe TACHYINA Motschulsky, 1862 26. Genus Lymnastis Motschulsky, 1862 Notes. New genus for the fauna of Albania. 26.1 Lymnastis galilaeus Piochard de la Brûlerie, 1876 Material studied. “Alban. Wgth. Valona”, 2 s. (MNHUB); “Mysege” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. COLEOPTERA: CARABIDAE: TRECHINAE

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27. Genus Paratachys Casey, 1918 27.1. Paratachys bistriatus (Duftschmid, 1812) References. Tachys bistriatus: Apfelbeck, 1904: 117 (Velipoja); Murraj, 1968: 33 (Vau i Dejës); Hieke et al., 1986: 184 (Borshi near Vlorë); Hieke & Wrase, 1988: 62. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Albania Miloti”, 3 s. (HNHM); “Miloti, Alb. Matzenauer”, 1 s. (HNHM); “Elbasan Alb., Mader”, 7 s. (HNHM); “Lim i Tiranes” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Burelle” / “Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Mursi-Kara Alibey” / “Albania leg. Bischoff 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1934”, 1 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Librazhd, Prenjas, 750 m, 7.V.1995, 1 s., L. Penev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 8.II.1998 / 28.II.1998, 54 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. Kopecký (2003: 275) has omitted the species for Albania, so that it is here confirmed for the country. The record of Murraj (1968) is included here with great reservation in view of possible misidentification (it was found a male of Bembidion praeustum from the Tomor Mt. determinated by Murraj as Tachys bistriatus). 27.2. Paratachys fulvicollis (Dejean, 1831) References. Tachys fulvicollis: Apfelbeck, 1904: 118 (Velipoja near Lezhë); Hieke & Wrase, 1988: 63. Material studied. “Mustajbeg Velipoja Albania”, 2 s. (HNHM); “Tirane” / “Albania leg. Bischoff 1932”, 2 s. (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (NMW); Tiranë Park, 1.III.1998, 9 s., R. Ward leg. (CMNH). Notes. Kopecký (2003: 275) has omitted the species for Albania. Here it is confirmed for the country. 27.3. Paratachys micros (Fischer von Waldheim, 1828) References. Tachys micros: Hieke & Wrase, 1988: 63. Material studied. “Ibe Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. Kopecký (2003: 276) has omitted the species for Albania. Here P. micros is confirmed for the country with first detailed record.

28. Genus Polyderis Motschulsky, 1862 Notes. New genus for the fauna of Albania. 28.1. Polyderis cardioderus (Chaudoir, 1850) Material studied. Tiranë Park, 8.II.1998, 1 s., grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. New species for the fauna of Albania.

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29. Genus Sphaerotachys J. Müller, 1926 29.1. Sphaerotachys haemorrhoidalis (Ponza, 1805) References. Tachys haemorrhoidalis: Apfelbeck, 1904: 120 (Velipoja); Hieke et al., 1986: 184 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 65. Material studied. “Bojana Shkoder”, 10 s. (NMW); “Medua, Alb. 10.5.914”, 2 s. (NMW); “Elbasan. Alb. 29.VIII.18 Dr. H. Priesner”, 3 s. (MNHUB); “Berat Alb. Dr. Müller 18”, 2 s. (NMW); “Skutari, Alb. Klenka”, 2 s. (NMW); “Elbasan Alb., Mader”, 7 s. (NMW); “Vorra, Alban., Mader”, 5 s. (NMW); “Bushek Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lumi i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 3 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 8.II.1998, 89 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. Kopecký (2003: 278) has omitted the species for Albania. Here it is confirmed for the country.

30. Genus Tachys Dejean, 1821 30.1. Tachys scutellaris Stephens, 1828 References. Tachys scutellaris: Hieke & Wrase, 1988: 63. Material studied. “Durazzo, Alb. Dr. J. Müller”, 9 s. (NMW); “Rusculi Alb., Mader”, 1 s. (NMW); “Durres” / “Albania leg. Bischoff 1932”, 18 s. (MNHUB); “Humara Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania leg. Bischoff 1934”, 5 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 6 s. (MNHUB); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, more than 30 s. (NMW). Notes. Kopecký (2003: 276) has omitted the species for Albania. Here it is confirmed for the country with first detailed records.

31. Genus Tachyta Kirby, 1837 Subgenus Tachyta s.str. 31.1. Tachyta nana (Gyllenhal, 1810) References. Tachyta nana: Apfelbeck, 1904: 121 (Velipoja); Hieke et al., 1986: 184 (Bizë near Shëngjergji); Hieke & Wrase, 1988: 66. Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Ricavac 1300 m 25.VI.2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Mamuras Alban. Kruja Dr. J. Müller”, 1 s. (NMW); “Kap Rodeni Alb., leg. Mader”, 1 s. (NMW); “Kruja, Alban., Mader”, 5 s. (NMW); “Llogora” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB). Notes. Kopecký (2003: 277) has omitted the species for Albania. Here it is confirmed for the country.


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32. Genus Tachyura Motschulsky, 1862 Subgenus Tachyura s.str. 32.1. Tachyura diabrachys (Kolenati, 1845) References. Tachys sexstriatus var. bisbimaculatus: Csiki, 1940: 214 (Kula Ljums). Tachys inaequalis: Hieke et al., 1986: 184 (Borshi near Vlorë). Tachys diabrachys: Hieke & Wrase, 1988: 65. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Elbasan, Alb. 4.IX.18 Dr. H. Priesner”, 1 s. (MNHUB); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 10 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (NMW); “Drin Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (NMW); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Librazhd, Prenjas, 750 m, 7.V.1995, 7 s., L. Penev leg. (NMNHS); Leskovac - Barmashi, 880 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Notes. Kopecký (2003: 279) has omitted the species for Albania. Here it is confirmed for the country. The most part of the literature data and older identifications of Tachys inaequalis (Reitter, 1884, nec Kolenati, 1845), T. sexstriatus (Duftschmid, 1812) and T. bisbimaculatus (Chevrolat, 1860) from the Balkans concern this species. 32.2. Tachyura parvula (Dejean, 1831) References. Tachys parvulus: Hieke & Wrase, 1988: 64. Material studied. “Elbasan Alb. 29.VII.18 Dr. H. Priesner”, 1 s. (NMW); “Elbasan Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Skutari, Alb. Klenka”, 3 s. (NMW); “Skutari Alb., Mader”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB). Notes. Kopecký (2003: 279) has omitted the species for Albania. Here T. parvula is confirmed for the country with first detailed records. # Tachyura sexstriata (Duftschmid, 1812) Notes. I have not seen any specimen from this species from the Balkan Peninsula, excluding Crete. The older records for it from the region refer to T. diabrachys.

Tribe PATROBINI Kirby, 1837 33. Genus Deltomerus Motschulsky, 1850 Subgenus Deltomerus s.str. ? Deltomerus malissorum Apfelbeck, 1918 References. Deltomerus malissorum: Zamotailov, 2003: 283. Material studied. Montenegro. “Vunsaj, Alban. leg. Plec, 1914” / “Deltomerus mallisorum Typus! Apfelb.” / “Typus”, 1 m. syntype (NMW); “Wgth. 1918 Kom VIII.”, 1 m., 2 f. (NMW); “Zljeb Pl. Kulina” / “Montenegro leg. Bischoff 1931”, 1 m., 3 f. (MNHUB).

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Notes. After the original description, only Zamotailov (2003: 283) noted this species for Albania without any detailed records. However, the type locality, Vunšaj (in Albanian) or Vusanje (in Serbo-Croatian) is situated in the Montenegrin section of the North Albanian Alps very close to Albanian territory (Apfelbeck, 1918c: 160-161, 164-165). The massifs of Kom Mt. and Zljeb Mt. also lie outside the investigated territory. Due to lack of any reliable record from Albania D. malissorum is treated here as questionable. Very probably it inhabits the country. 33.1. Deltomerus parumpunctatus Zamotailov, 1990 References. Deltomerus parumpunctatus: Zamotailov, 1990: 133, 135 (North Albanian Alps: “Čafa Borit”, loc. typ.); Zamotailov, 1992: 56; Zamotailov, 2003: 283. Material studied. “Wgth. 1918 Menina pl. N. alb. alp.” / “Deltomerus parumpunctatus m. paratype V. N. Kurnakov det. 1970”, 1 m. designated by mistake as paratype (NMW); “Wgth. 1918 Menina pl.” / “Deltomerus parumpunctatus m. alotype V. N. Kurnakov det. 1970”, 1 f. designated by mistake as paratype (NMW). Notes. Zamotailov (1990: 135) have noted no paratypes in the description of D. parumpunctatus. For this reason, the two specimens in NMW designated as paratypes should not be considered as part of the type series. Subgenus Paradeltomerus Apfelbeck, 1908 33.2. Deltomerus paradoxus Apfelbeck, 1908 Notes. Apfelbeck published three consecutive descriptions for both the subgenus and the species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest ones are correctly treated as valid. ? Deltomerus paradoxus paradoxus Apfelbeck, 1908 References. Deltomerus paradoxus: Zamotailov, 1992: 56. Deltomerus paradoxus paradoxus: Zamotailov, 2003: 283. Notes. Hieke (1981: 80) stated by mistake that D. paradoxus is described from Albania. In reality, the nominotypical form was described from the alpine zone of the Shar Planina Mts. (Apfelbeck, 1907c; Apfelbeck, 1908). Dinchev & Atanasov (1998) stated that only 1 % of the territory of the last mountain massif is situated in Albania. Actually all specimens of the nominotypical race in the collections originate from Macedonia or the Kosovo Region of Serbia. Due to lack of reliable data the subspecies is treated as doubtful for Albania. 33.2.1. Deltomerus paradoxus korabensis J. Müller, 1936 References. Deltomerus paradoxus korabensis: Müller, 1936: 108-110 (Korab Mt., loc. typ.; Bjeshka Matroxh); Zamotailov, 2003: 283. Deltomerus paradoxus: Zamotailov, 1992: 56. Material studies. Nearly 15 s. from Korab Mt. (MNHUB).


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Tribe POGONINI Laporte, 1834 34. Genus Pogonistes Chaudoir, 1871 Notes. New genus for the fauna of Albania. 34.1. Pogonistes convexicollis Chaudoir, 1871 Material examined. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. For the time being, the locality near Durrës is the westernmost locality in the species range. 34.2. Pogonistes gracilis (Dejean, 1828) Material examined. “Durres” / “Albania leg. Bischoff 1932”, 29 s. (MNHUB). Notes. New species for the fauna of Albania. For the present, the locality near Durrës is one of the easternmost finds in the species range.

35. Genus Pogonus Dejean, 1821 Subgenus Pogonus s.str. 35.1. Pogonus littoralis (Duftschmid, 1812) References. Pogonus litoralis sic!: Apfelbeck, 1904: 149 (Velipoja); Hieke & Wrase, 1988: 67. Material studied. “Alban. Wgth. Valona”, 2 s. (HNHM); “Latif Pulaj”, 1 s. (HNHM); “Latif Pulaj” / “Latif Alban. sept.”, 5 s. (NMW). Notes. Bousquet (2003a: 287) has omitted the species for Albania. Here it is confirmed for the country. Besides the material studied above, in MNHUB I have seen more than 60 s. (most of them come from the vicinity of Durrës), which remained unregistered. ? Pogonus luridipennis (Germar, 1823) References. Pogonus luridipennis: Murraj, 1968: 33 (Kamëz). Notes. The data of Murraj (1968) are doubtful because of possible misidentification (among the materials loaned from MNST, I found a male of Bembidion combustum from the Tomor Mt. which had been determinated by Murraj as P. luridipennis). 35.2. Pogonus riparius Dejean, 1828 References. Pogonus riparius: Apfelbeck, 1904: 149 (Velipoja near Lezhë; Shkodër); Hieke & Wrase, 1988: 67; Bousquet, 2003a: 287. Material studied. “Latif Pulaj”, 6 s. (HNHM); “Latif Pulaj” / “Latif Alban. sept.”, 7 s. (NMW); “Merdita, Alb. Matzenauer”, 2 s. (NMW); “Durazzo” / “Albania leg. Bischoff 1931”, 6 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Valona” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). 78


Tribe TRECHINI Bonelli, 1810 Subtribe PERILEPTINA Sloane, 1903 36. Genus Perileptus Schaum, 1860 Subgenus Perileptus s.str. 36.1. Perileptus areolatus (Creutzer, 1799) 36.1.1. Perileptus areolatus areolatus (Creutzer, 1799) References. Perileptus areolatus: Apfelbeck, 1907a: 496 (Kiri River near Mesi); Csiki, 1940: 214 (Kula Ljums); Hieke & Wrase, 1988: 34. Perileptus areolatus areolatus: Moravec et al., 2003: 288. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Skutari Alb., Mader”, 1 s. (HNHM); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Leskovac - Barmashi, 880 m, 2.VI.1995, 4 s., G. Blagoev leg. (NMNHS). Subtribe TRECHINA Bonelli, 1810 ! genus Adriaphaenops Noesske, 1928 ! Adriaphaenops sp. References. Aphaenopsis sp.: Latella, 1990: 148 (North Albanian Alps: Mali i Thatë Mt., cave “Shpella e Haxhisë”, 1720 m). Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave “Shpella e Haxise”, 8.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. According to Vigna Taglianti (personal communication) the female mentioned above belongs to a hitherto undescribed species.

37. Genus Albanotrechus Casale & V. B. Guéorguiev, 1994 37.1. Albanotrechus beroni Casale & V. B. Guéorguiev, 1994 References. Albanotrechus beroni: Casale & Guéorguiev, 1994: 418-420 (cave “Spela Merkurth” near to village of Merkurth, Rreshen District); Moravec et al., 2003: 290.

38. Genus Duvalius Delarouzée, 1859 [subgenus Bicharotrechus Bokor, 1922] [species group “pilifer”] [Duvalius droveniki Magrini, 1998] Notes. This species was described from the locality “Hajla Prokletije” in the Montenegrin part of the North Albanian Alps (Magrini, 1998), very close to the Albanian border. D. droveniki might live in Albania too.


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# Duvalius maglicensis Winkler, 1933 References. Duvalius maglicensis: Turin, 1981: 112. Notes. This species lives only in Bosnia Herzegovina and Montenegro (see also Moravec et al., 2003: 299). Species deleted from the list of the Albanian ground beetles. Subgenus Duvalius s.str. Notes. The arrangement of the separate groups is in accordance with the works of Casale (1983), Casale & Laneyrie (1983), Casale & Vigna Taglianti (1990) and Casale et al. (1996). The specificity of the group of “fodori” is discussed in a coming paper (Guéorguiev, in preparation). I. South Aegean groups Species group “krueperi” 38.1. Duvalius didonnai Magrini, Vanni & Ingusio, 1996 References. Duvalius didonnai: Magrini et al., 1995a: 27-29 (cave “Shpella Lekpetes”, 1150 m, near to Kurvelesh, loc. typ.; cave “Shpella e Kasarit”, 1100 m, near to Kurvelesh); Moravec et al., 2003: 302. Species group “georgii” 38.2. Duvalius bischoffi Meschnigg, 1936 References. Duvalius bischoffi: Meschnigg, 1936: 48-49 (cave “Homberg-Höhle” at Maja e Tanasit Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b: 99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 110; Casale, 1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of Vlorë); Moravec et al., 2003: 301. 38.3. Duvalius georgii (J. Müller, 1922) References. Duvalius georgi: J. Müller, 1922a: 87-88 (hot pole in Tomor Mt., loc. typ.); Jeannel, 1928: 567-570 (Tomor Mt.: Toik, Kulmak); Scheibel, 1937a: 18, 20 (Tomor Mt.); Schatzmayr, 1943b: 98 (Tomor Mt.: Toik, Kulmak); Casale, 1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (hot pole and cave at Tomor Mt., east of Berat); Moravec et al., 2003: 303. Material studied. “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona” / “ex Coll. Bokor”, 1 f. (HNHM); “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona”, 3 s. (NMW); “Tomor: Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 3 s. (NMW; MNHUB). 38.4. Duvalius muelleri Winkler, 1933 References. Duvalius muelleri: Winkler: 1933a: 75-76 (Shen Elias Mt., cave “Stere gropa e Vasos”, 800 m, south of Vlorë, loc. typ.); Scheibel, 1937a: 18, 20 (Shen Elias Mt., south of Vlorë); Schatzmayr, 1943b: 98 (Shen Elias Mt., cave “Stere gropa e Vasos”); Turin, 1981: 112; Casale, 80


1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave at Shen Elias Mt., south of Vlorë); Magrini et al., 1995b: 25 (Llogora Mt., cave “Stera Kullëxhive”, south of Vlorë); Moravec et al., 2003: 304. Material studied. “Stere gropa de Vasos” / “Reza e Kanalit, Alban. m.” / “Winkler, Lona, Bischoff, V.1931”, 8 syntypes (NMW); “Shen-E’Lias Juni 1932” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Shen-Elias” / “leg. Bischoff Albania 1934”, 1 f. (HNHM); “Shen-E’Lias Juni 1932” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB). Species group “fuchsi” 38.5. Duvalius fuchsi Scheibel, 1937 References. Duvalius fuchsi: Scheibel, 1937a: 15-18, 20 (nameless cave near Llogora, south of Vlorë, loc. typ.); Turin, 1981: 111; Casale, 1983: 249; Casale & Laneyrie, 1982: 199; Genest & Juberthie, 1994: 592 (cave at Llogora, south of Vlorë); Moravec et al., 2003: 303. Material studied. “Höhle in der Llogora” / “Albania leg. Bischoff 1935”, 1 syntype (MNHUB); “Höhle i.-d. Llgora” / “Bischoff mai 1935”, 4 syntypes (MNHUB). 37.6. Duvalius pominii Schatzmayr, 1943 References. Duvalius pominii: Schatzmayr, 1943b: 96-97, 99 (cave “Stere e Vogel” and cave “Stere e Madhe” near Dukati, south of Vlorë, loc. typ.); Turin, 1981: 112; Genest & Juberthie, 1994: 592 (cave “Stere e Vogel” and cave “Stere e Madhe” near Dukati); Moravec et al., 2003: 305. 38.7. Duvalius regiszogui Meschnigg, 1936 References. Duvalius regiszogui: Meschnigg, 1936: 46-47 (cave “Homberg-Höhle” Maja e Tanasit Mt., loc. typ.); Scheibel, 1937a: 18, 20 (Maja e Tanasit Mt., south of Vlorë); Schatzmayr, 1943b: 99 (Maja e Tanasit Mt.: cave “Homberg”); Turin, 1981: 113; Casale, 1983: 249; Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (cave “Homberg”, Maja e Tanasit Mt., south of Vlorë); Moravec et al., 2003: 305. II. North Aegean groups Species group “leonhardi” 38.8. Duvalius sturanyi (Apfelbeck, 1904) 38.8.1. Duvalius sturanyi stenocephalus (Apfelbeck, 1918) References. Trechus stenocephalus: Apfelbeck, 1918c: 164 (North Albanian Alps, Vermosh, 1100 m, loc. typ.). Duvalius leonhardi sturanyi: Jeannel, 1928: 548 (North Albanian Alps: Vermosh, 1100 m). Duvalius sturanyi stenocephalus: Scheibel, 1937a: 18-20 (North Albanian Alps: Vermosh, Menina Planina Mt.); Pretner, 1978: 130, 135, 148, 167-168 (Vermosh; Menina Planina Mt.); Moravec et al., 2003: 305. Duvalius leonhardi stenocephalus: Mařan, 1939c: 60 (Vermosh). Duvalius sturanyi: Casale & Laneyrie, 1982:199; Genest & Juberthie, 1994: 592 (North Albanian Alps: Menina Planina Mt. near Vermosh). COLEOPTERA: CARABIDAE: TRECHINAE

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Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14” / “Typus” / “Trechus stenocephalus Typus! Apfelb.”, 1 f. syntype of Trechus stenocephalus (NMW). Notes. Jeannel (1928: 582-584) synonymized D. stenocephalus with D. leonhardi sturanyi, but later Scheibel (1937a: 19-20) proved that the former is a subspecies of D. sturanyi. Further studies are needed to test the status of D. sturanyi sturanyi and D. sturanyi stenocephalus, having in mind the different forms of their aedeagus and copulatory pieces (Scheibel, 1937a). The same concerns the remote populations of the former near Foča and from the Komovi Mt. as well as the status of D. sturanyi matejkai Mařan, 1939 described from the Montenegrin section of the North Albanian Alps. Species group “fodori” Notes. The treatment of this species group is in accordance with a separate author’s paper in preparation. It treats Duvalius fodori Scheibel, 1937, D. strupii Scheibel, 1937 and D. vignai Casalei, 1983. 38.9. Duvalius strupii Scheibel, 1937 References. Duvalius strupii: Scheibel, 1937b: 282-283 (Korab Mt., 2000 m, loc. typ.); Schatzmayr, 1943b: 98 (Korab Mt.); Turin, 1981: 113; Casale & Laneyrie, 1982:191; Casale, 1983: 250 (Korab Mt.); Genest & Juberthie, 1994: 592 (Korab Mt.); Moravec et al., 2003: 305. Material studied. “Korab” / “Albania leg. Bischoff 1936”, 17 specimens, incl. 12 syntypes (MNHUB). Subgenus Euduvalius Jeannel, 1929 Species group “gogalai” 38.10. Duvalius zhalovi B. V. Guéorguiev, 2005 References. Duvalius zhalovi: Guéorguiev, 2005: 33-40 (cave “Peshtnik”, 1100 m near Ostreni i Madh, loc. typ.).

39. Genus Neotrechus J. Müller, 1913 Species group “dalmatinus” 39.1. Neotrechus lonae (J. Müller, 1914) 39.1.1. Neotrechus lonae lonae (J. Müller, 1914) References. Trechus lonae: J. Müller, 1914: 1003-1005 (Çukall Mt.: cave “Scpedha n’Giuka Kunors”, loc. typ.; Çukall Mt.: cave “Scpedha n’Ciuk tugit Vukai”, loc. typ.). Neotrechus lonae: Winkler, 1926: 86, 96 (caves in Çukall Mt.); Scheibel, 1936: 229 (Çukall Mt.); Scheibel, 1937a: 20 (Çukall Mt.). Neotrechus lonai sic!: Jeannel, 1928: 296-297 (Çukall Mt.: cave “Scpedha n’ giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Schatzmayr, 1943b: 97 (Çukall Mt.: cave “Scpedha n’ giuka kunors” and cave “Scpedha n’ Ciuk tugit Zukali”); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (caves in Çukall Mt., Rapshë). Neotrechus lonai sic! lonai sic!: Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall Mt.); Pretner, 1978: 147, 166 (Çukall Mt.). Neotrechus lonae lonae: Moravec et al., 2003: 315. 82


Material studied. “Cukali Gebirge N-Albanien Lona leg. 1914”, 4 s. (NMW); “Scpedha n’Giuka Zukali Albanien Lona Giuno 1914”, 1 s. (DEI); “Cukale Albania sept.” / “Bischoff 1930 coll. A. Winkler”, 5 s. (NMW); “Cucale Geb. Höhle von Vukaj” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Shpella Kodra Çorit” / “Cukale Geb.” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shpella Rasa-Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Shpella Çuka Vjit” / “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. N. lonae lonae is sympatric and syntopic with N. malissorum (J. Müller, 1914) in most of the caves where they have been found. The report of this subspecies from the area of Rapshë (Genest & Juberthie, 1994: 592) is erroneous since this region is populated by N. lonae atavus. The nominotypical subspecies differs from both ssp. atavus and ssp. shkodrensis in the wider and more rounded, almost not narrowed to apex, elytra. 39.1.2. Neotrechus lonae atavus J. Müller, 1934 [= bischoffi Meschnigg, 1936] References. Neotrechus bischoffi: Meschnigg, 1936: 44-46 (North Albanian Alps: cave “Shpella e lugstit, Hot-Brigje” near Rapshë, loc. typ.); Schatzmayr, 1943b: 98 (cave “Shpella e lugstit, Hot-Brigje” near Rapshë); Casale & Laneyrie, 1983: 180. Neotrechus lonae atavus: Scheibel, 1936: 227-228 (cave “Shpella e lugstit, Hot Brgje” near Rapshë); Scheibel, 1937a: 20 (cave “Hot Brigje”); Moravec et al., 2003: 315. Neotrechus lonai sic! atavus: Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Rapshë); Pretner, 1978: 147, 166 (cave “Shpella e lugstit, Hot Brgje” near Rapshë); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (cave “Shpella e Lugstit”, cave “Hot Brigje” near Rapshë). Material studied. “Shpella e Lugstit” / “Brigie Hot” / “leg. Bischoff Albania 1934”, 7 syntypes of Neotrechus bischoffi (MNHUB); “Shpella e Lugstit” / “Brigie Hot” / “Albania leg. Bischoff 1935”, 11 syntypes of Neotrechus bischoffi (MNHUB). Notes. Scheibel (1936) established the synonymy of N. bischoffi with N. lonae atavus. Regardless of that, Schatzmayr (1943b) and Casale & Laneyrie (1983) listed the former as valid species. The type localities of both N. lonae atavus and N. suturalis pretneri Scheibel, 1936 coincide and are located in Montenegro very close to the Albanian border. 39.1.3. Neotrechus lonae shkodrensis Scheibel, 1936 References. Neotrechus lonae shkodrensis: Scheibel, 1936: 228-229 (cave “Shpella kufinit, KoplikutHotit” near Ivanaj, north of Liqen i Shkodrës, loc. typ.); Scheibel, 1937a: 20 (Ivanaj); Moravec et al., 2003: 315. Neotrechus lonai sic! shkodrensis: Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Ivanaj); Pretner, 1978: 147, 166 (cave “Shpella kufinit, Koplikut-Hotit” near Ivanaj); Casale & Laneyrie, 1983: 180; Drovenik & Peks, 1994: 45; Genest & Juberthie, 1994: 592 (cave “Kufinit et Kopikut, Hotit peés” near Ivanaj, north of Liqen i Shkodrës); Drovenik & Peks, 1999: 51. Material studied. “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1935”, 4 syntypes (MNHUB); “Shpella Kufin Hot Koplicut Ivanaj” / “Albania leg. Bischoff 1937”, 4 s. (MNHUB). Notes. Drovenik & Peks (1994: 45) and Drovenik & Peks (1999: 51) indicated that the type locality of this subspecies is situated in Montenegro. Actually it is located in Albania, very close to the state border.


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39.2. Neotrechus malissorum (J. Müller, 1914) References. Trechus malissorum: Müller, 1914: 1005 (Çukall Mt.: cave “Scpedha n’ kunor Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ Ciuk tujit Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha e Bacit Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha n’ rasst’ kunors”, loc. typ.; Çukall Mt.: cave “Scpedha n’ giuka kunors”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs n’ gur t’ barzodhs”, loc. typ.). Neotrechus dalmatinus malissorum: Winkler, 1926: 86, 96 (caves in Çukall Mt.); Scheibel, 1937a: 20 (Çukall Mt.). Neotrechus malissorum: Jeannel, 1928: 290-291 (Çukall Mt.: cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave “Scpedha e bacit Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”; cave “Scpedha Nre Gegs n’ gur t’ barzodhs Zukali”); Schatzmayr, 1943b: 98 (Çukall Mt.: cave “Scpedha n’ kunor Zukali”, cave “Scpedha n’ Ciuk tujit Zukali”, cave “Scpedha e bacit Zukali”; cave “Scpedha n’ rasst’ kunors”; cave “Scpedha n’ giuka kunors”; cave “Scpedha Nre Gegs n’ gur t’ barzodhs Zukali”); Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (Çukall Mt.); Casale & Laneyrie, 1983: 180; Genest & Juberthie, 1994: 592 (Çukall Mt., cave “n’Kunor Vukai”, cave “n’Ciuk tujit Vukai”, cave “Lacit Vukai”, cave “n’Rast’Kunors”, cave “n’Giuka Kunors”); Moravec et al., 2003: 315. Material studied. “Scpedha n’Giuka Zukali Albanien Lona Giuno 1914”, 6 s. (DEI); “ScpedhaNre Geg N’gur Bazzodha Zukali Albanien”, 1 s. (HNHM); “Scpedha-Nre Geg N’gur Barzodha Zukali Albanien Lona Giugno 1914”, 4 s. (MNHUB); “Cukali Gebirge N-Albanien Lona leg. 1914”, 4 s. (NMW); “Albania m. Cucali C. Lona”, 2 s. (HNHM); “Cukali Albania sept.” / “leg. Bischoff 1931”, 2 s. (NMW); “Shpellë Quku i Konors Vukaj Cucale” / “Albania leg. Bischoff 18.VI.31”, 2 s. (MNHUB); “Shpella Kodra Carit” / “Cukale Geb.” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Shpella Kodra Carit Thep” / “Cukale” / “Albania leg. Bischoff 1935”, 24 s. (NMW; MNHUB); “Shpella Luga Ìen Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Shpella Rasa Kunors Vukaj” / “Cukale” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Shpella Gollë” / “Cukale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-iThatë Mt., cave “Shpella e Haxise”, 8.VII.1993, rests of 3 s., A. Gobetti leg. (cAVT). 39.3. Neotrechus suturalis (Schaufuss, 1864) 39.3.1. Neotrechus suturalis pentheri Winkler, 1926 References. Trechus suturalis: Apfelbeck, 1918c: 164 (North Albanian Alps: small cave near Rapshë). Neotrechus dalmatinus suturalis var. pentheri: Winkler, 1926: 85-86, 96 (near Rapshë, loc. typ.). Neotrechus dalmatinus pentheri: Scheibel, 1937a: 20 (Rapshë). Neotrechus suturalis pentheri: Jeannel, 1928: 295 (North Albanian Alps: caves near Rapshë); Absolon & Mařan, 1947: 2; Mařan, 1947: 11 (North Albanian Alps: Rapshë); Pretner, 1978: 146-147, 167 (Rapshë); Casale & Laneyrie, 1983: 180; Drovenik & Peks, 1994: 46; Genest & Juberthie, 1994: 592 (North Albanian Alps: caves at Rapshë); Drovenik & Peks, 1999: 53; Moravec et al., 2003: 316. Material studied. “Alban. montenegr. Grenze, Penther’14” / “Höhle bei Rapsa, 727 m”, 15 syntypes of Trechus suturalis (DEI; MNHUB); “Albania Rapsa Dr. Grabowski”, 86 s. (HNHM); “Höhlen bei Rapsa leg. Bischoff 1931”, 1 s. (DEI); “Höhlen bei Rapsa 20.VII.31” / “Albania leg. Bischoff 1931”, 4 s. (MNHUB); “Rapsh” / “Albania leg. Bischoff 1935”, 40 s. (MNHUB). Notes. Drovenik & Peks (1994: 46) and Drovenik & Peks (1999: 53) noted by mistake the type locality in Montenegro. In fact, the village of Rapshë is situated in Albania. 84


40. Genus Trechus Clairville, 1806 Subgenus Trechus s.str. Species group “quadristiatus” 40.1. Trechus obtusus Erichson, 1837 40.1.1. Trechus obtusus obtusus Erichson, 1837 References. Trechus obtusus: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9. Trechus obtusus obtusus: Jeannel, 1927: 309 (“Petrov planina”, 2000 m); Moravec et al., 2003: 337. Material studies. “Latif Koritnik”, 1 s. (NMW); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). 40.2. Trechus quadristriatus (Schrank, 1781) References. Trechus quadristriatus: Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1943b: 96 (Lushnje; cave “Stere e Madhe” near Dukati); Hieke et al., 1986: 184 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 35; Nonveiller et al., 1994: 9; Moravec et al., 2003: 339. Material studied. “Elbasan Alb., Mader”, 2 s. (HNHM); “Kruja, Alban., Mader”, 4 s. (HNHM); “Sarisaltik, 1180 m Alb., leg. Mader”, 3 s. (HNHM); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997 / 23-28.XI.1997, 47 s., R. Ward leg. (CMNH). Notes. Except the specimens studied above, in MNHUB the present author has seen additionally about 210 s., which remained unregistered. Species group “babinjensis” 40.3. Trechus babinjensis Jeannel, 1927 References. Trechus babinjensis: Jeannel, 1927: 267-268 (North Albanian Alps, loc. typ.); Casale & Laneyrie, 1983: 141; Moravec et al., 2003: 327. 40.4. Trechus valbonensis Jeannel, 1927 References. Trechus valbonensis: Jeannel, 1927: 268 (North Albanian Alps, loc. typ.); Casale & Laneyrie, 1983: 141; Moravec et al., 2003: 343. Material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT). Species group “subnotatus” 40.5. Trechus subnotatus Dejean, 1831 40.5.1. Trechus subnotatus subnotatus Dejean, 1831 References. Trechus subnotatus subnotatus: Apfelbeck, 1904: 127 (Vlorë); Jeannel, 1927: 446 (Kruja; Tomor Mt.: Kulmak); Pawlowski, 1973: 232; Moravec et al., 2003: 341. Trechus subnotatus: Jeannel, COLEOPTERA: CARABIDAE: TRECHINAE

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1925: 23 (Tomor Mt.: Kulmak); Horion, 1941: 181; Schatzmayr, 1943b: 96 (Dukati; cave “Stere e Madhe” near Dukati); Casale & Laneyrie, 1983: 147; Hieke & Wrase, 1988: 36; Nonveiller et al., 1994: 9, 15; Guéorguiev & Guéorguiev, 1995: 75; Guéorguiev et al., 1997: 11; Sakalian & Guéorguiev, 1997: 46. Material studied. “Kruja, Alban., Mader”, 2 s. (HNHM). Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 1 m., D. Antonelli leg. (cAVT). Notes. In MNHUB I have seen also additional 39 s., which remained not registered. 40.5.2. Trechus subnotatus ljubetensis Apfelbeck, 1908 References. Trechus subnotatus ljubetensis: Moravec et al., 2003: 341. Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Albania” / “ljubetensis Apf. R. Jeannel”, 1 m. (NMW). Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. The identification of the two specimens from Vlorë is doubtful since the ssp. ljubetnesis is normally treated as local endemic of the Shar Planina Mts. ? species group “cardioderus” ? Trechus irenis Csiki, 1912 [= cardioderus balcanicus Jeannel, 1927] References. Trechus cardioderus balcanicus: Tabaković-Tošić, 1988: 137. Notes. Tabaković-Tošić (1988) noted the range of this taxon reaching Albania. Due to lack of reliable records from the country the species is treated as questionable for Albania. Species group “pulchellus” 40.6. Trechus merditanus Apfelbeck, 1906 References. Trechus merditanus: Apfelbeck, 1906: 1163 (Merdita: Mal i Shenjt Mt. and Zepë Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Jeannel, 1927: 539-540 (Merdita: Mal i Shenjt Mt. and Zepë Mt.); Pawlowski, 1972: 312-314; Casale & Laneyrie, 1983: 151; Moravec et al., 2003: 336. Material studied. “Merdita M. Scheit” / “merditanus n. sp. Apfelb.”, 2 s. probably syntypes (NMW). Notes. According to Jeannel (1927), T. merditanus is a vicarious species of T. priapus. 40.7. Trechus priapus K. Daniel, 1902 40.7.1. Trechus priapus priapus K. Daniel, 1902 References. Trechus priapus: Meixner, 1939: 309; Pawlowski, 1973: 249; Hieke & Wrase, 1988: 38; Nonveiller et al., 1994: 9; Guéorguiev & Guéorguiev, 1995: 77; Guéorguiev et al., 1997: 10. Trechus priapus priapus: Jeannel, 1927: 537 (North Albanian Alps; Gjalicë Mt.); Moravec et al., 2003: 338. Material studied. “Alban. Exped. Gjalica Ljums 17.-26.VI’18”, 2 s. (NMW). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT). Notes. Except the two specimens above, in MNHUB I have seen additionally nearly 800 s., which remained unregistered. 86


Species group “egreginus” 40.8. Trechus egregius Jeannel, 1927 References. Trechus egregius: Jeannel, 1927: 551-552 (North Albanian Alps, loc. typ.); Casale & Laneyrie, 1983: 152; Moravec et al., 2003: 330. Species group “tristis” 40.9. Trechus nigrinus Putzeys, 1847 [= tristis (Duftschmid, 1812)] References. Trechus nigrinus: Apfelbeck, 1904: 129 (Vlorë); Apfelbeck, 1907a: 498-499 (Mal i Shenjt Mt.: top region, 1500 m); Jeannel, 1927: 555 (Vlorë); Moravec et al., 2003: 336. Trechus tristis: Csiki, 1940: 214 (Gjalicë Mt.); Pawlowski, 1973: 250; Hieke & Wrase, 1988: 38; Nonveiller et al., 1994: 9. Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp.’18. Pashtrik 4-14.7”, 1 s. (NMW); “Kruja, Alban. Mader”, 3 s. (HNHM); “Tomor” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”, 67 s. (MNHUB); “Mal i Gjer” / “Albania leg. Bischoff 1937”, 15 s. (MNHUB). Notes. Except the material studied above, in MNHUB the present author has seen more 48 s., which remained unregistered. Species group “obtusiusculus” 40.10. Trechus albanicus Apfelbeck, 1905 References. Trechus albanicus: Apfelbeck, 1905: 241 (Northeastern Albania / Kosovo border region: alpine belt of Koritnik Mt., loc. typ.); Apfelbeck, 1907b: 636-637 (alpine belt of Koritnik Mt., loc. typ.); Jeannel, 1927: 563-564 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 214 (Gjalicë Mt.; Korab Mt.); Casale & Laneyrie, 1982: 153; Drovenik & Peks, 1994: 39 (Koritnik Mt.); Drovenik & Peks, 1999: 44 (Koritnik Mt.); Moravec et al., 2003: 326. Material studied. “Latif Koritnik” / “Trechus albanicus Apfb. Typus! Coll. Apfelbeck”, 1 f. syntype (HNHM); “Latif Koritnik”, 2 syntypes (DEI; NMW); “Latif Alban. spt.”, 1 s. (DEI); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Alban. Exped. Gjalica Ljums 17.-26.VI.’18”, 1 s. (NMW); “Gjalica” / “Albania leg. Bischoff 1936”, 2 s. (NMW); “Gjallica” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB). Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b) and the latter has long time been accredited to be correct (see also Casale & Laneyrie, 1982; Drovenik & Peks, 1994: 39; Drovenik & Peks, 1999: 44; Moravec et al., 2003: 326). However, it is incorrect and the original description must be assigned to the older one. 40.11. Trechus kobingeri Apfelbeck, 1902 40.11.1. Trechus kobingeri kobingeri Apfelbeck, 1902 [= distinctissimus Roubal, 1920] References. Trechus kobingeri: Jeannel, 1925: 23 (Çukall Mt.); Jeannel, 1927: 566 (Çukall Mt.); Casale & Laneyrie, 1982: 153; Nonveiller et al., 1994: 8, 18 (near to Shkodër). Trechus distinctissimus: COLEOPTERA: CARABIDAE: TRECHINAE

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Casale & Laneyrie, 1982: 153. Trechus kobingeri bulgaricus: Guéorguiev et al., 1997: 10. Trechus kobingeri kobingeri: Moravec et al., 2003: 333. Material studied. “Maja e Jezerce Albania sept.”, 1 m. (NMW); “Cucale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Tråchus kobingeri bulgaricus Pawlowski, 1972 is a junior synonym of T. kobingeri pawlowskianus P. Moravec & Lompe, 2003, and its record for Albania is erroneous. This race is known only from a limited region in Western Bulgaria. # Trechus obtusiusculus Ganglbauer, 1889 References. Trechus obtusiusculus: Nonveiller et al., 1994: 9. Notes. The species is recorded with detailed records only from Bosnia Herzegovina and Montenegro. The data for Albania is too doubtful, so that it is removed from the species list. 40.12. Trechus ravasinianus Lorenz, 1998 [= ravasinii Jeannel, 1925] References. Trechus ravasinii: Jeannel, 1925: 24-25 (Tomor Mt.: Kulmak, Toik, Abbas Ali, loc. typ.); Jeannel, 1927: 570-571 (Tomor Mt.: Abbas Ali, Kulmak, Toik); Casale & Laneyrie, 1982: 153. Trechus ravasinianus: Moravec et al., 2003: 339. Material studied. “Tomor: Tomor Albania mer.” / “lg. Winkler Mai 1931”, 1 f. (HNHM); “Tomor: Kulmak Albania mer.” / “lg. Winkler Mai 1931”, 12 s. (NMW). Notes. Except the localities mentioned above, in MNHUB I have seen more than 100 s., which remained unregistered. ! Trechini gen. indet. ! Trechini gen. & sp. indet. Material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Mali-i-Thatë Mt., cave “Shpella e Haxise”, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT). Notes. According to Vigna Taglianti (personal communication) the parts found in the cave “Shpella e Haxise” may belong to an unknown genus close to Pheggomisetes Knirsch, 1923.

Subtribe TRECHODINA Jeannel, 1926 41. Genus Thalassophilus Wollaston, 1854 41.1. Thalassophilus longicornis (Sturm, 1825) References. Thalassophilus longicornis: Apfelbeck, 1907a: 496 (Kiri River near Mesi). Notes. Moravec et al. (2003: 346) have omitted the species for Albania. Actually, it needs further confirmation for the country.

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Subfamily HARPALINAE Bonelli, 1810 Tribe CHLAENIINI Brullé, 1834 Subtribe CALLISTINA Laporte, 1834 42. Genus Callistus Bonelli, 1810 42.1. Callistus lunatus (Fabricius, 1775) 42.1.1. Callistus lunatus lunatus (Fabricius, 1775) References. Callistus lunatus: Schatzmayr, 1943b: 99 (Lushnje); Murraj, 1968: 33 (Hibë; Sauk); Hieke et al., 1986: 190 (Ibë near Krraba). Callistus lunatus lunatus: Kirschenhofer, 2003: 347. Material studied. “Lomi Beratit Albanien”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Hani Grabom Mai” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); Sauk, 9.VII.1961, 1 f. (MNST). Notes. The records of Murraj (1968) are included here with minimal reservation since the revision of a female from Sauk having the same label data as the published material has been found to be correctly identified by him.

Subtribe CHLAENIINA Brullé, 1834 43. Genus Chlaenius Bonelli, 1810 Subgenus Chlaeniellus Reitter, 1908 43.1. Chlaenius flavipes Ménétries, 1832 References. Chlaenius flavipes: Apfelbeck, 1907a: 496, 498, 501 (Kiri River near Mesi; Fani i Madh River near Oroshi); Schatzmayr, 1943b: 100 (Zezes River near Ura Zezes); Hieke et al., 1986: 190 (Borshi near Vlorë); Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 349. Material studied. “Oboti Bojana”, 1 s. (NMW); “Skutari Mesi”, 1 s. (NMW); “Merdita Oroschi” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.’18”, 1 s. (NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 8 s. (NMW); “Elbasan Alb., Mader”, 8 s. (NMW; MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); Tepelenë District, Vijosë River, 2.VI.1994, 2 s., G. Blagoev leg. (NMNHS). 43.2. Chlaenius nigricornis (Fabricius, 1787) References. Chlaenius nigricornis: Apfelbeck, 1904: 158 (Durrës); Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 350. Material studied. “Scutari l. See-Ufer”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Albania Rittm. Matoha”, 1 s. (NMW); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 2 s. (MNHUB). COLEOPTERA: CARABIDAE: HARPALINAE

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Notes. Murraj (1968: 33) cited this species from two localities (Tiranë; Tomor Mt.). Among the material loaned from MNST, I found out 9 s. from the second place having the same label data as the material published by Murraj as Chlaenius nigricornis. Actually, all the nine specimens refer to Bembidion dalmatinum. 43.3. Chlaenius nitidulus (Schrank, 1781) References. Chlaenius nitidulus: Hieke & Wrase, 1988: 149; Kirschenhofer, 2003: 350. Material studied. “Maja ë Qori 2050 m, Albania”, 1 s. (NMW); Sauk, 5.III.1961, 1 m. (MNST). Notes. First detailed records for Albania. 43.4. Chlaenius terminatus Dejean, 1826 References. Chlaenius terminatus: Schatzmayr, 1943b: 100 (Stan-Karbunarë; Skrofotina; Vlorë). Material studied. “Oboti Bojana”, 3 s. (HNHM). Notes. Kirschenhofer (2003: 350) has omitted the species for Albania. Here it is confirmed for the country. 43.5. Chlaenius tristis (Schaller, 1783) 43.5.1. Chlaenius tristis tristis (Schaller, 1783) References. Chlaenius tristis: Hieke & Wrase, 1988: 149. Chlaenius tristis tristis: Kirschenhofer, 2003: 350. Material studied. “Oboti Bojana”, 5 s. (HNHM); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. First detailed records for Albania. 43.6. Chlaenius vestitus (Paykull, 1790) References. Chlaenius vestitus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (StanKarbunarë); Kirschenhofer, 2003: 351. Material studied. “Bojana Oboti, Alb.”, 1 s. (NMW); “Lumi Dukatit Alban. mer.”, 4 s. (NMW); “Maj’e Tartarit Albania mer.”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Kula Ljums 14.20.VII.’18”, 1 s. (NMW); “Elbasan, Alb. 15.IX.18 Dr. H. Priesner”, 4 s. (NMW); “Alessio, Alb. Matzenauer”, 2 s. (NMW); “Miloti, Alb. Matzenauer”, 1 s. (NMW); 40 s. from different localities (MNHUB); Librazhd, Prenjas, 750 m, 7.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Drina River, 3.VI.1995, 1 s., V. Sakalain leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT). Subgenus Chlaenites Motschulsky, 1860 43.7. Chlaenius spoliatus (P. Rossi, 1792) 43.7.1. Chlaenius spoliatus spoliatus (P. Rossi, 1792) References. Chlaenius spoliatus: Schatzmayr, 1943b: 99 (Durrës); Hieke & Wrase, 1988: 147. Chlaenius spoliatus spoliatus: Kirschenhofer, 2003: 351. 90


Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Oboti Bojana”, 2 s. (HNHM); “Durazzo” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Himara juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen i Butrintes” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB); “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i That” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT). Subgenus Chlaenius s.str. 43.8. Chlaenius festivus (Panzer, 1796) 43.8.1. Chlaenius festivus festivus (Panzer, 1796) References. Chlaenius festivus: Apfelbeck, 1904: 157 (Vlorë; Velipoja near Lezhë); Apfelbeck, 1909: 298 (Vlorë); Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 100 (Lushnje); Murraj, 1968: 33 (Radenjë); Hieke & Wrase, 1988: 148. Chlaenius festivus festivus: Gridelli, 1925: 120 (Shkodër; Berat); Kirschenhofer, 2003: 351. Material studied. “Alban., Wgth. Valona”, 1 s. (NMW); “Oboti Bojana”, 1 s. (HNHM); “Skutari l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); nearly 40 s. from different localities (MNHUB); Sauk, 18.II.1961, 1 f. (MNST); Rodenj, 26-28.VII.1962, 1 s. (MNST); Tepelenë District, Vijosë River, 2.VI.1994, 1 m., G. Blagoev leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 4 m., 2 f., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. The specimen collected at Liqen i Shkodrës was determined as C. velutinus (Duftschmid, 1812) by Kirschenhofer, but I refer this record to the nominotypical subspecies. The material published by Murraj (1968) was found to be correctly determinated. Subgenus Dinodes Bonelli, 1810 43.9. Chlaenius cruralis Fischer von Waldheim, 1829 References. Chlaenius cruralis: Kirschenhofer, 2003: 352. Material studied. “Skutari” / “Strasser”, 3 s. (NMW); Ibë, 4.VII.1963, 1 m. (MNST). Notes. First detailed records for Albania. 43.10. Chlaenius decipiens (L. Dufour, 1820) [= azureus Duftschmid, 1812, nec Fabricius, 1775; laticollis Chaudoir, 1843; ambiguus Csiki, 1931] References. Chlaenius decipiens: Apfelbeck, 1904: 154 (Vlorë); Hieke et al., 1986: 190 (Ibë near Krraba); Hieke & Wrase, 1988: 148; Kirschenhofer, 2003: 352. Chlaenius azureus: Apfelbeck, 1909: 298 (Vlorë). Chlaenius decipiens var. ambiguus: Csiki, 1940: 219 (Gjalicë Mt.). Chlaenius decipiens laticollis: Schatzmayr, 1943b: 100 (Lushnje). Material studied. “Elbasan Albanien”, 1 s. (NMW); “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tepelene Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tirane” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. COLEOPTERA: CARABIDAE: HARPALINAE

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Bischoff 1936”, 1 s. (MNHUB); “Ivan Geb Korce” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); Tiranë, 27.IX.1958, 1 m. (MNST); Tiranë, 10.V.1962, 1 m. (MNST). Subgenus Epomis Bonelli, 1810 43.11. Chlaenius circumscriptus (Duftschmid, 1812) References. Chlaenius circumscriptus: Kirschenhofer, 2003: 353. Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. First detailed records for Albania. 43.12. Chlaenius dejeani (Dejean, 1831) References. Chlaenius dejeani: Apfelbeck, 1904: 154 (Vlorë); Schatzmayr, 1943b: 99 (Lushnje); Hieke & Wrase, 1988: 147. Epomis dejeani: Apfelbeck, 1909: 298 (Vlorë). Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB). Notes. Kirschenhofer (2003: 353) has omitted the species for Albania. Here it is confirmed for the country. Subgenus Trichochlaenius Seidlitz, 1887 43.13. Chlaenius aeneocephalus Dejean, 1826 43.13.1. Chlaenius aeneocephalus aeneocephalus Dejean, 1826 References. Chlaenius aeneocephalus: Apfelbeck, 1904: 156 (Vlorë); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988: 148. Notes. Kirschenhofer (2003: 355) has omitted the species for Albania, so that it needs of further confirmation for the country.

Tribe CYCLOSOMINI Laporte, 1834 Subtribe MASOREINA Chaudoir, 1870 44. Genus Masoreus Dejean, 1821 Subgenus Masoreus s.str. 44.1. Masoreus wetterhallii (Gyllenhal, 1813) 44.1.1. Masoreus wetterhallii wetterhallii (Gyllenhal, 1813) References. Masoreus wetterhallii: Hieke & Wrase, 1988: 152. Material studied. “Tomor Tojk” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. Bousquet (2003b: 357) has omitted the species for Albania. Here it is confirmed for the country with a first detailed record. 92


Tribe DRYPTINI Bonelli, 1810 45. Genus Drypta Latreille, 1796 Subgenus Drypta s.str. 45.1. Drypta dentata (P. Rossi, 1790) References. Drypta dentata: Apfelbeck, 1904: 346 (Velipoja); Hieke & Wrase, 1988: 153; Baehr, 2003b: 358. Material studied. “Albanien Avlona v. Oertzen.”, 1 s. (MNHUB).

Tribe HARPALINI Bonelli, 1810 Subtribe AMBLYSTOMINA Fauvel, 1889 46. Genus Amblystomus Erichson, 1837 46.1. Amblystomus levantinus Reitter, 1883 46.1.1. Amblystomus levantinus levantinus Reitter, 1883 References. Amblystomus levantinus: Apfelbeck, 1904 (Velipoja); Schatzmayr, 1943b: 101 (StanKarbunarë; Lushnje); Hieke & Wrase, 1988: 165 (Velipoja). Amblystomus levantinus levantinus: Jaeger & Wrase, 2003: 360. Notes. According to D. Wrase (personal communication) the “metallescens-levantinus” group needs revision. This notable specialist on the Palaearctic Harpalini wrote me: “I don’t know really what levantinus is, as I haven’t checked yet the type. I have several species, clearly different in genitalia belonging to that group. For the time being the best solution of that problem it to continue the classification we have now.”. 46.2. Amblystomus metallescens (Dejean, 1829) References. Amblystomus metallescens: Csiki, 1940: 219 (Kula Ljums; Gjalicë Mt.); Schatzmayr, 1943b: 101 (Lushnje; Mal i Krujes); Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Oboti Bojana”, 1 s. (HNHM); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). 46.3. Amblystomus niger (Heer, 1841) References. Amblystomus niger: Hieke et al., 1986: 188 (Ibë near Krraba; Bizë near Shëngjergji); Hieke & Wrase, 1988: 146; Jaeger & Wrase, 2003: 360. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 8.II.1998 / 15.II.1998, 4 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). COLEOPTERA: CARABIDAE: HARPALINAE

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Subtribe ANISODACTYLINA Lacordaire, 1854 47. Genus Anisodactylus Dejean, 1829 Subgenus Anisodactylus s.str. 47.1. Anisodactylus binotatus (Fabricius, 1787) [= spurcaticornis Dejean, 1829] References. Anisodactylus binotatus: Apfelbeck, 1909: 298 (Pashaliman); Hieke et al., 1986: 186 (North Albanian Alps near Theth); Hieke & Wrase, 1988: 112; Ito, 2003a: 360. Anisodactylus binotatus ab. spurcaticornis: Apfelbeck, 1909: 298 (Pasha Liman). Material studied. “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Theth, 1-8.VIII. 1961, 1 m. (MNST); Theth, 800-1200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS). 47.2. Anisodactylus nemorivagus (Duftschmid, 1812) References. Anisodactylus nemorivagus: Apfelbeck, 1904: 212 (Velipoja); Horion, 1941: 247; Hieke & Wrase, 1988: 112. Notes. Ito (2003a: 361) has omitted the species for Albania. Actually, it needs further confirmation for the country. 47.3. Anisodactylus pueli Schauberger, 1933 47.3.1. Anisodactylus pueli pueli Schauberger, 1933 References. Anisodactylus pueli: Schauberger, 1933: 124 (Vlorë, loc. typ.); Sciaky, 1979: 15 (Vlorë); Turin, 1981: 150; Wrase, 1991: 12; Noonan, 1996: 34, 109 (Vlorë). Anisodactylus pueli pueli: Ito, 2003a: 361. Material studied. “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT). Subgenus Pseudodichirius Lutshnik, 1921 47.4. Anisodactylus intermedius Dejean, 1829 References. Anisodactylus intermedius: Hieke & Wrase, 1988: 113; Ito, 2003a: 361. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Rodeni, 2-22.VII.1962, 1 s. (MNST); Gjirokastra, 6.V.1994, 1 m., P. Stoev leg. (NMNHS). Notes. First detailed records for Albania.

48. Genus Diachromus Erichson, 1837 48.1. Diachromus germanus (Linnaeus, 1758) References. Diachromus germanus: Ito, 2003a: 362. Material studied. “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 2 s. (HNHM). Notes. First detailed record for Albania. 94


49. Genus Gynandromorphus Dejean, 1829 49.1. Gynandromorphus etruscus (Quensel, 1806) 49.1.1. Gynandromorphus etruscus etruscus (Quensel, 1806) References. Gynandromorphus etruscus: Schatzmayr, 1943b: 105 (Dukati); Hieke & Wrase, 1988: 113. Gynandromorphus etruscus etruscus: Ito, 2003a: 362. Material studied. “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS).

Subtribe DITOMINA Bonelli, 1810 50. Genus Carterus Dejean, 1830 Subgenus Carterus s.str. 50.1. Carterus dama (P. Rossi, 1792) References. Ditomus dama: Apfelbeck, 1909: 298 (Vlorë). Carterus dama: Hieke & Wrase, 1988: 145; Wrase, 1989: 39 (Mursi - Kara Ali Bey); Wrase, 2003b: 363. Material studied. “Murrsi - Kara Ali Bey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS). Additional material included (studied D.W. Wrase). Oricum, south of Vlorë, X.1994, 2 f., K. Resl leg. (cWR). 50.2. Carterus gilvipes (Piochard de la Brûlerie, 1873) References. Carterus gilvipes: Schatzmayr, 1943b: 101 (Dukati); Wrase, 2003b: 364. 50.3. Carterus rotundicollis Rambur, 1837 Material studied. Borsh, 15-28.V.1961, 1 f. (MNST). Notes. New species for the fauna of Albania.

51. Genus Ditomus Bonelli, 1810 51.1. Ditomus calydonius (P. Rossi, 1790) 51.1.1. Ditomus calydonius calydonius (P. Rossi, 1790) References. Ditomus calydonius calydonius: Wrase, 2003b: 364. 51.2. Ditomus tricuspidatus (Fabricius, 1792) References. Ditomus tricuspidatus: Ganglbauer, 1889: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë); Wrase, 2003b: 365. Carterus tricuspidatus: Apfelbeck, 1904: 173 (Vlorë).

COLEOPTERA: CARABIDAE: HARPALINAE

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Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB); “Brataj Ramica Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “M. e Tartarit juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Borshne” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); Borsh, 15-28.V.1961, 1 f. (MNST); Ljukove, 100 m, 5.VI.1995, 1 m., G. Blagoev leg. (NMNHS). Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 2 s., M. Komnenov leg. (cSH).

52. Genus Dixus Billberg, 1820 52.1. Dixus clypeatus (P. Rossi, 1790) References. Aristus clypeatus: Apfelbeck, 1909: 298 (Berat). Ditomus clypeatus: Hieke et al., 1986: 186 (Borshi near Vlorë; Mali me Gropë near Livadhet e Selitës; Mal i Dajtit Mt.: Shkall Priskë); Hieke & Wrase, 1988: 146. Dixus clypeatus: Wrase, 2003b: 365. Material studied. “Kruja, Alban., Mader”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 3 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Albanien Dajti…6.6.55 Burmeister”, 1 s. (MNHUB); “N 111”, 1 s. (MNST), Sauk, 10-20.VI.1961, 1 s. (MNST). Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M. Komnenov leg. (cSH). 52.2. Dixus eremita (Dejean, 1825) References. Ditomus eremita: Schatzmayr, 1943b: 102 (Skrofotina; Lushnje). Dixus eremita: Wrase, 2003b: 365; Wrase, 2005: 820. 52.3. Dixus obscurus (Dejean, 1825) References. Aristus obscurus: Apfelbeck, 1909: 298 (Vlorë; Berat). Ditomus obscurus: Schatzmayr, 1943b: 101 (Lushnje); Hieke et al., 1986: 186 (Borshi near Vlorë; Poliçan near Tomor Mt.). Dixus obscurus: Wrase, 2003b: 365. Material studied. “Serandë” Piqeros” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB).

53. Genus Graniger Motschulsky, 1864 53.1. Graniger femoralis (Coquerel, 1859) References. Carterophonus femoralis: Sciaky & Franzini, 1976: 26 (Berat).

96


54. Genus Odontocarus Solier, 1835 Notes. New genus for the fauna of Albania. 54.1. Odontocarus robustus (Dejean, 1831) Material studied. “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Butrinti, 4.VI.1995, 1 s., V. Sakalian leg. (NMNHS). Notes. New species for the fauna of Albania. [genus Pachycarus Solier, 1835] [subgenus Mystropterus Chaudoir, 1842] [Pachycarus macedonicus V.B. Guéorguiev & B.V. Guéorguiev, 1997] Notes. This species has been described from the Macedonian part of the Galičica Mt. very close to the Albanian border, so that it may inhabit Albania too.

Subtribe HARPALINA Bonelli, 1810 55. Genus Acinopus Dejean, 1821 Subgenus Acinopus s.str. 55.1. Acinopus baudii A. Fiori, 1913 55.1.1. Acinopus baudii hellenicus Heinz, 1982 References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995: 205; Kataev & Wrase, 2003: 368. Acinopus baudii hellenicus: Heinz, 1982: 16 (Tepelenë; Maja e Konispol, Seraqin); Kataev & Wrase, 2003: 368. Material studied. “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Murthe bis Kara Alibey” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB). 55.2. Acinopus picipes (Olivier, 1795) [= emarginatus Chaudoir, 1842] References. Acinopus emarginatus: Apfelbeck, 1904: 180 (Vlorë). Acinopus picipes: Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1943b: 102 (Karbunara e Madhe; Tërpan; Lushnje); Hieke & Wrase, 1988: 143; Kataev & Wrase, 2003: 368. Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Mal i That” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Mal i Dajtit Mt.: Shkall Priskë, 26-30. VI.1961, 1 f. (MNST); Ibë, 4.VII.1962, 1 m. (MNST); Ibë, 8.VII.1966, 1 f. (MNST); “AL (Berat). Paß ca. 5 km S. Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 s. (MNHUB); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg. (NMNHS). Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., Heinz & Peks leg. (cWR); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH); Dhermi, 200 m, 12.VII.2006, 2 s., M. Komnenov leg. (cSH). COLEOPTERA: CARABIDAE: HARPALINAE

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# Acinopus subquadratus Brullé, 1832 References. Acinopus subquadratus: Magistretti, 1965: 272; Guéorguiev & Guéorguiev, 1995: 205; Kataev & Wrase, 2003: 368. Notes. Guéorguiev & Guéorguiev (1995) mentioned this taxon for Albania, following an older indication for South Albania (Magistretti, 1965: 272). Kataev & Wrase (2003: 368) have most probably followed also the same sources. However, according to Heinz (1982) A. subquadratus inhabits only the southern and central regions of Greece and the data on its presence in Albania concerns A. baudii hellenicus; both species were often mixed in the past. The species is removed from the list of the Albanian Carabidae. Subgenus Oedematicus Bedel, 1897 55.3. Acinopus megacephalus (P. Rossi, 1794) References. Acinopus megacephalus: Apfelbeck, 1904: 180 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Puel, 1934: 79; Schatzmayr, 1943b: 102 (Dukati; Skrofolina); Jeanne, 1984: 24; Hieke & Wrase, 1988: 143; Kataev & Wrase, 2003: 369. Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 3 s. (MNHUB); “Durrës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Myzeqe” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 3 s. (MNHUB). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Lushnja-Divjaka, 21.VII.1993, 1 m., L. Latella leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 3 s., leg. M. Komnenov (cSH). Subgenus Osimus Fischer von Waldheim, 1829 55.4. Acinopus ammophilus Dejean, 1829 References. Acinopus ammophilus: Apfelbeck, 1907a: 498, 501 (Oroshi). Notes. Kataev & Wrase (2003: 369) have omitted the species for Albania. Actually, it needs further confirmation for the country.

56. Genus Harpalus Latreille, 1802 Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982 56.1. Harpalus tenebrosus Dejean, 1829 [= centralis Schauberger, 1929] References. Harpalus tenebrosus var. centralis: Csiki, 1940: 220 (Kula Ljums). Harpalus tenebrosus: Hieke & Wrase, 1988: 138; Kataev & Wrase, 2003: 370. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB).

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Subgenus Harpalus s.str. Species group “honestus” 56.2. Harpalus attenuatus Stephens, 1828 References. Harpalus attenuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 220 (Shkëlzen Mt.; Kula Ljums); Schatzmayr, 1943b: 104 (“Poggio Boschetto”); Hieke et al., 1986: 187 (Borshi near Vlorë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 139; Kataev & Wrase, 2003: 373. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 4 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja ae Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Vlesh” / “30.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albanien-Exp. DEI Borshi südl. Vlora 14.-27.V.1961”, 1 s. (MNHUB); Mal i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 2 f. (MNST); 2 km N from Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS). 56.3. Harpalus honestus (Duftschmid, 1812) 56.3.1. Harpalus honestus honestus (Duftschmid, 1812) References. Harpalus honestus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 220 (Korab Mt.); Hieke & Wrase, 1988: 138. Harpalus honestus honestus: Kataev & Wrase, 2003: 377. Material studied. “Vermosa Penther”, 20 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. Kula Ljums 18-28.V.’18”, 1 s. (NMW); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Ibë, 18.V.1960, 1 m. (MNST). 56.4. Harpalus rubripes (Duftschmid, 1812) References. Harpalus rubripes: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 220 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke & Wrase, 1988: 133; Kataev & Wrase, 2003: 381. Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Alban. Exp. 1918 Korab, 2331.VII.”, 1 s. (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal COLEOPTERA: CARABIDAE: HARPALINAE

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i Shenjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “AL (Tirana) 8 km N Psene. E Tirana 1450 m. 30.IV.1993 leg. H. + S. Peks”, 2 m. (MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); Cafasan, 1230 m, 27.VII.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS). 56.5. Harpalus rufipalpis Sturm, 1818 56.5.1. Harpalus rufipalpis rufipalpis Sturm, 1818 [= rufitarsis (Duftschmid, 1812)] References. Harpalus rufitarsis: Apfelbeck, 1904: 196 (Berat); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Berat); Hieke et al., 1986: 188 (Bizë near Shëngjergji); Hieke & Wrase, 1988: 138. Harpalus rufipalpis rufipalpis: Kataev & Wrase, 2003: 381. Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB); “Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 3 s. (HNHM). Notes. In addition to the material studied above, in NMW I have seen more specimens, which remained unregistered. 56.6. Harpalus sulphuripes Germar, 1824 56.6.1. Harpalus sulphuripes sulphuripes Germar, 1824 References. Harpalus sulphuripes: Apfelbeck, 1904: 196 (Vlorë); Anonymous, 1907: 284; Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Schauberger, 1926: 32 (Vlorë; Pasha Liman); Hieke et al., 1986: 188 (Tomor Mt.: Abbas Ali, 2401 m); Hieke & Wrase, 1988: 139. Harpalus sulphuripes sulphuripes: Kataev & Wrase, 2003: 383. Material studied. More than 50 s. from different localities (MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS); Saranda District, Ljukove, 5.V.1995, 1 s., P. Stoev leg. (NMNHS); Gjirokastra, 6.V.1994, 1 s., P. Stoev leg. (NMNHS); 12 km NE Erseka, 900 m, 12.V.1995, 1 s., L. Penev leg. (NMNHS). Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M. Komnenov leg. (cSH). Species group “atratus” 56.7. Harpalus atratus Latreille, 1804 [= subsinuatus (Duftschmid, 1812)] References. Harpalus atratus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Hieke et al., 1986: 187 (Mal i Dajtit Mt.: Shkall Priskë); Hieke & Wrase, 1988: 137; Kataev & Wrase, 2003: 373. Harpalus atratus var. subsinuatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m). Material studied. “Albania Zalle-Dardha” / “23.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); more than 60 s. from different localities (MNHUB).

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Additional material included (studied A. Vigna Taglianti). 5 km S of Tepelenë (crossroads Permet), 350 m, 21-22.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. In addition to the material studied above, in NMW I have seen more specimens, which remained unregistered. Species group “quadripunctatus” 56.8. Harpalus laevipes Zetterstedt, 1828 [= quadripunctatus Dejean, 1829] References. Harpalus quadripunctatus: Hieke & Wrase, 1988: 133. Harpalus laevipes: Kataev & Wrase, 2003: 378. Material studied. “Gjallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. First detailed record for Albania. Species group “serripes” 56.9. Harpalus flavicornis Dejean, 1829 56.9.1. Harpalus flavicornis flavicornis Dejean, 1829 References. Harpalus flavicornis: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988: 136. Harpalus flavicornis flavicornis: Kataev & Wrase, 2003: 376. Material studied. “Mal i Dajtit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Lumi i Erzen” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Mal i That” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albanien - Tirana Dajti-Vorberge 3.V.39…Müller J.”, 2 s. (MNHUB); Mal i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 m. (MNST); Zooteknika, 14.V.1963, 1 s. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 1 s. (MNHUB); “AL (Berat). Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB); Cafasan, 1230 m, 27.VII.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 4 s., G. Blagoev leg. (NMNHS); 5 km Leskovac, 650 m, 2.VI.1995, 1 s., G. Blagoev leg. (NMNH); Tiranë Park, 11.I.1998 / 8.II.1998, 4 s., R. Ward leg. (CMNH). 56.10. Harpalus serripes (Quensel, 1806) 56.10.1. Harpalus serripes serripes (Quensel, 1806) [= subchalybaeus Reitter, 1900] References. Harpalus serripes: Apfelbeck, 1904: 197 (Vlorë); Anonymous, 1907: 283; Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Csiki, 1940: 220 (Korab Mt.); Hieke et al., 1986: 187 (Ibë near Krraba); Hieke & Wrase, 1988: 135. Harpalus serripes ab. subchalybaeus: Schauberger, 1926: 25 (Dukati i Ri). Harpalus serripes serripes: Kataev & Wrase, 2003: 382. Material studied. L. Selitës, 3-9.VII.1961, 1 m. (MNST); Ibë, 4.VII.1963, 1 f. (MNST); “AL (Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. + S. Peks”, 10 s. (MNHUB); 2 km N from Durrës, 24.V.1993, 2 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Cafasan, 1230 m, 27.VII.1993, 3 s., P. Stoev, D. Zaprjanova leg. (NMNHS); Lushnja, Divjaka National Park, 10.V.1995, 1 s., S. COLEOPTERA: CARABIDAE: HARPALINAE

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Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m, 6.VI.1995, 3 s., G. Blagoev leg. (NMNHS). Additional material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). Notes. In addition to the material studied above, in NMW and ZNHU I have seen more than 130 s. from different localities, which remained unregistered. Species group “pumilus” [Harpalus picipennis (Duftschmid, 1812)] References. Harpalus picipennis: Apfelbeck, 1904: 200; Hieke & Wrase, 1988: 141; Kataev & Wrase, 2003: 380. Notes. Hieke & Wrase (1988) noted the species for Albania following Apfelbeck’s record “Prisren” (Apfelbeck, 1904: 200). In fact this place is situated in the Kosovo Region of Serbia. Most likely, Kataev & Wrase (2003) have used the same sources. For the time being the species is treated here as probable but not yet proven for Albania (having in mind recent data from the north parts of Republic of Macedonia, unpublished information). 56.11. Harpalus pumilus Sturm, 1818 [= vernalis Fabricius, 1801] References. Harpalus vernalis: Müller, 1931: 69; Hieke & Wrase, 1988: 140; Kataev & Wrase, 2003: 380. Material studied. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB). Notes. First detailed record for Albania. Species group “anxius” 56.12. Harpalus anxius (Duftschmid, 1812) References. Harpalus anxius: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Kataev & Wrase, 2003: 373. Material studied. “Latif Cucali”, 1 s. (NMW); “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). Notes. Hieke & Wrase (1988: 139) noted this species for Albania apparently following the record “Jannina” (Apfelbeck, 1904: 199). In fact this locality is situated in Greece. 56.13. Harpalus subcylindricus Dejean, 1829 References. Harpalus subcylindricus: Hieke & Wrase, 1988: 140; Kataev & Wrase, 2003: 382. Material studied. “Skutari” / “Strasser”, 1 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rapsh” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Mal i Dajtit Mt., 10.V.1965, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë ~ 600 m 27.IV.1993 leg. Heinz u. Sonja Peks”, 3 s. (MNHUB); “AL (Berat). Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 m., 1 f. (MNHUB); North Albanian Alps, Maj’e Radohimes, 2400-2550 102


m, 29.V.1993, 3 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Radohimes, 1000-1100 m, 5.IX.1993, 2 s., P. Beron, B. Petrov leg. (NMNHS). Notes. First detailed records for Albania. Species group “flavescens” 56.14. Harpalus flavescens (Piller et Mitterpacher, 1783) Material studied. “Elbasan, Alb. 11.IX.18 Dr. H. Priesner”, 1 s. (NMW). Notes. New species for the fauna of Albania. Species group “tardus” 56.15. Harpalus albanicus Reitter, 1900 References. Harpalus albanicus: Reitter, 1900: 114 (? Shkodër, loc. typ.); Apfelbeck, 1904: 200 (Shkodër); Rambousek, 1912: 86; Horion, 1941: 230; Roudier, 1949: 176 (Shkodër); Magistretti, 1965: 308; Hieke & Wrase, 1988: 134; Kataev & Wrase, 2003: 372. Material studied. “Skutari” / “coll. Reitter”, 1 f. syntype (HNHM). Notes. Tschitschérine (1901: 154) expressed doubt about the correct interpretation of the species type locality “Scutari”. It may as well represent the present city of Üsküdar located on the Asian shore of the Bosporus right opposite the peninsula of Istanbul, or the present Albanian City of Shkodër. 56.16. Harpalus tardus (Panzer, 1796) References. Harpalus tardus: Apfelbeck, 1904: 198 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Hieke & Wrase, 1988:134; Kataev & Wrase, 2003: 383. Material studied. “Latif Oroshi”, 1 s. (NMW); “Maranaj” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Alban. Exped. Kula Ljums 18-28.V.1918”, 1 s. (NMW). Species group “latus” 56.17. Harpalus latus (Linnaeus, 1758) References. Harpalus latus: Kataev & Wrase, 2003: 378. Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & P. Stoev leg. (NMNHS). Notes. First detailed records for Albania. 56.18. Harpalus solitaris Dejean, 1829 References. Harpalus solitaris: Kataev & Wrase, 2003: 382. Material studied. North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 f., P. Beron, B. Petrov leg. (NMNHS). Notes. First detailed record for Albania. COLEOPTERA: CARABIDAE: HARPALINAE

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Species group “smaragdinus” 56.19. Harpalus smaragdinus (Duftschmid, 1812) References. Harpalus smaragdinus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 382. Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); Theth, 1-8.VIII.1961, 1 m. (MNST); “AL (Shkodar). Paß SE Puke 850 m 2.V.1993 leg. H. & S. Peks”, 1 m. (MNHUB); North Albanian Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Rapsh, 15.V.2002, 3 m., 1 f., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). Species group “autumnalis” # Harpalus anatolicus Tschitschérine, 1898 # Harpalus anatolicus anatolicus Tschitschérine, 1898 [= skutariensis Reitter, 1900] References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”). Notes. Reitter (1900: 107) indicated as type locality of H. skutariensis the site “Scutari” (see also Kataev & Wrase, 1997). “Scutari” was the Greek name for the district of Istanbul now known as Üsküdar, located on the Asian shore of the Bosporus right opposite the peninsula of Istanbul. The same name was also used in the past for the present city of Shkodër. Tschitschérine (1901: 146) has stated that H. skutariensis Reitter is identical with H. tristis. Apfelbeck (1904: 199) accepted this opinion despite Daniel & Daniel (1902: 250-251) had already synonymized H. tristis to H. danieli. Later on, Mlynář (1979: 83) considered H. skutariensis as synonym of H. anatolicus. Kataev & Wrase (1997) confirmed this synonymy and correctly gave data only for the Anatolian Peninsula. The species is excluded from the list of the Albanian Carabidae. 56.20. Harpalus autumnalis (Duftschmid, 1812) References. Harpalus autumnalis: Hieke & Wrase, 1988: 136; Kataev & Wrase, 2003: 373. Material studied. “Skutari Bojana”, 3 s. (NMW); “Lum i Tiranes” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB). Notes. First detailed records for Albania. # Harpalus danieli Reitter, 1900 [= tristis Tschitschérine, 1898] References. Harpalus tristis: Apfelbeck, 1904: 199 (“Scutari”); Müller, 1931: 65 (“Skutari”). Notes. Tschitschérine (1901: 146) stated H. skutariensis Reitter is synonym of H. tristis, but a bit later Daniel & Daniel (1902: 250-251) synonymized H. tristis to H. danieli. Apfelbeck (1904: 199) accepted the opinion of Tschitschérine. Later on, Mlynář (1979) solved the problem (see notes under H. anatolicus). Recently, Kataev & Wrase (2003: 375) have indicated H. danieli for European Turkey and former Yugoslavia (Serbia and Montenegro). I consider, however, due to lack of reliable records, the species should be excluded from the faunas of Albania and adjacent territories in the north. 104


[Harpalus triseriatus A. Fleischer, 1897] Notes. This Balkan endemic species lives in Bosnia Herzegovina, Macedonia, Bulgaria and Greece (Kataev & Wrase, 2003: 383) and most probably inhabits Albania as well. Species group “cupreus” 56.21. Harpalus cupreus Dejean, 1829 56.21.1. Harpalus cupreus fastuosus Faldermann, 1836 References. Harpalus cupreus fastuosus: Schatzmayr, 1943b: 104 (Lushnje). Material studied. “Skutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “MurrsiKara Alibey” / “leg. Bischoff Albania 1934” / “cotype” / “Harpalus bischoffi Schaub. det. Dr. E. Schaub. 35” / “(H. bischoffi Schb. i. l.) = H. cupreus Z. Mlynar det. 1976”, 1 m. with extracted genitalia (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Notes. Kataev & Wrase (2003: 374) have omitted the species for Albania. Here it is confirmed for the country. Harpalus bischoffi Schauberger is nomen nudum. 56.22. Harpalus euchlorus Ménétriés, 1839 Material studied. “Skutari” / “Strasser” / “collectio Paganetti”, 2 s. (NMW). Notes. New species for the fauna of Albania. B. Kataev (personal communication) expressed the opinion that H. euchlorus may be a synonym of the preceding species. Species group “dimidiatus” 56.23. Harpalus dimidiatus (P. Rossi, 1790) References. Harpalus dimidiatus: Apfelbeck, 1904: 193 (Vlorë; Berat); Apfelbeck, 1907a: 495 (northeast of Shkodër); Apfelbeck, 1909: 298 (Vlorë; Berat); Csiki, 1940: 220 (Kula Ljums); Schatzmayr, 1943b: 104 (“Poggio Boschetto”; Lushnje; Dukati); Hieke et al., 1986: 187 (North Albanian Alps near Theth); Hieke & Wrase, 1988: 132; Kataev & Wrase, 2003: 375. Material studied. “Merdita M. Scheit”, 2 s. (NMW); “Skutari Mesi”, 2 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Alban. Exp. Kula Ljums 1828.V.’18”, 5 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 2 s. (NMW); more than 60 s. from different localities (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 2 s. (HNHM); “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 12 s. (HNHM); “Albania Shkoder” / “21.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); Ibë, 24.IV.1962, 2 f. (MNST); Ibë, 4.VII.1963, 2 f. (MNST); Mal i Dajtit Mt., 10.V.1965, 3 m., 1 f. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m. (MNST); Tiranë, 7.IV.1966, 1 m. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 10 s. (MNHUB); “AL (Vlore) Llogora-Paß 1025 m 29.IV.1993 leg. F. + S. Peks”, 5 s. (MNHUB); “AL (Shkodar) Paß SE Puke 850 m 2.V.1993”, 1 s. (MNHUB); Hotolisht - Librazhd, 300 m, 7.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Librazhd - Prenjas, 750 m, 7.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 3 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Dukati Mt., 30 km S Vlorë, 450 m, 11.V.1995, 2 s., L. Penev leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 2 s., G. Blagoev leg. (NMNHS); Tiranë Park, 22.II.1998, 1 s., R. Ward leg. (CMNH). COLEOPTERA: CARABIDAE: HARPALINAE

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Additional material included (studied S. Hristovski, A. Vigna Taglianti and D.W. Wrase). Tiranë, XI.58, 1 s. (collector illegible) (cWR); Tiranë District, Kffaba-Paß, Südseite, 500 m, 29.IV.1993, 5 s., H. & S. Peks leg. (cWR); Korcë District, Mai i That Mt., Zwezde-Sattel, 10001100 m, 4.V.1993, 6 s., Heinz & Peks leg. (cWR); Fier, 4.VII.1993, 1 f., L. Latella leg. (cAVT); Tiranë, botanical garden, 4.VI.2002, 1 f., A. Paparistos leg. (cAVT); Tiranë, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). Species group “dispar” 56.24. Harpalus punctatostriatus Dejean, 1829 References. Harpalus punctatostriatus: Apfelbeck, 1909: 298 (Vlorë; Sop); Schatzmayr, 1943b: 103 (Lushnje); Hieke & Wrase, 1988: 128; Kataev & Wrase, 2003: 380. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Albania 1918”, 1 s. (HNHM); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 56.25. Harpalus pygmaeus Dejean, 1829 References. Harpalus pygmaeus: Apfelbeck, 1904: 195 (Vlorë; Berat); Apfelbeck, 1909: 298 (Berat); Hieke et al., 1986: 188 (Ibë near Krraba); Hieke & Wrase, 1988: 139; Kataev & Wrase, 2003: 380. Material studied. “Albania 1918”, 1 s. (HNHM); “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 10 s. (NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Ura Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Dajti” / “6.V.1959 leg. Xh. Murraj”, 1 s. (HNHM); Sauk, 9.VII.1961, 3 s. (MNST); Tiranë, 10.V.1962, 1 s. (MNST); Tiranë Park, 22-31.X.1997, 2 s., R. Ward leg. (CMNH); Tiranë Park, 16-21.II.1998, 2 s., R. Ward leg. (CMNH). Notes. The specimens from the locality “Tiranë Park” are somewhat bigger (7.1-7.4 mm), compared with specimens that inhabit Central Europe (cfr. Hůrka, 1996). Species group “hospes” 56.26. Harpalus hospes Sturm, 1818 56.26.1. Harpalus hospes hospes Sturm, 1818 Material studied. “AL (Korcë) Qaf e Qarrit – 1100 m 4.V.1993, leg. Heinz u. S. Peks”, 9 s. (MNHUB). Notes. New species for the fauna of Albania.

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Species group “affinis” 56.27. Harpalus affinis (Schrank, 1781) [= aeneus (Fabricius, 1775); viridulus (Geoffroy, 1785); limbopunctatus Fuss, 1858] References. Harpalus aeneus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1943b: 103 (Lushnje). Harpalus affinis: Csiki, 1940: 219 (Gjalicë Mt.); Kataev & Wrase, 2003: 371. Harpalus affinis ab. viridulus: Csiki, 1940: 219-220 (Gjalicë Mt.; Korab Mt.); Harpalus affinis ab. limbopunctatus: Csiki, 1940: 220 (Gjalicë Mt.; Korab Mt.). Material studied. Korcë, 10-20. VIII.1962, 1 m., 1 f. (MNST); North Albanian Alps, Theth, 800-1200 m, 28.V.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Radohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, 5-9.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT); Tropoia, valley of Valbona River, 900 m, 816.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT). Notes. In addition to the material studied above, in NMW and MNHUB I have seen more specimens, which remained unregistered. Species group “distinguendus” 56.28. Harpalus distinguendus (Duftschmid, 1812) 56.28.1. Harpalus distinguendus distinguendus (Duftschmid, 1812) References. Harpalus distinguendus: Apfelbeck, 1904: 192 (Vlorë; Durrës); Apfelbeck, 1909: 298 (Vlorë; Berat); Schatzmayr, 1943b: 103 (Dukati; Lushnje); Hieke & Wrase, 1988: 131. Harpalus distinguendus distinguendus: Kataev & Wrase, 2003: 376. Material studied. Ibë, 18.V.1960, 1 f. (MNST); Sauk, 5.III.1961, 1 m. / 23.IV.1963, 1 f. (MNST); Rojan, 6.III.1961, 1 f. (MNST); Rodeni, 26-28.VII.1962, 1 f. (MNST); Tiranë, V.1963, 1 m. / 7.IV.1966, 5 m., 6 f. (MNST); Mal i Dajtit Mt., 10.V.1965, 1 f. (MNST); Kurbneshi, VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 m., 2 f., P. Stoev leg. (NMNHS); Saranda District, Qeparo, 4-5.V.1994, 2 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 1 m., G. Blagoev leg. (NMNHS); Bureli, 22.VI.1994, 1 f., B. Guéorguiev leg. (NMNHS); Hotolisht - Librazhd, 300 m, 7.V.1995, 1 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Librazhd - Prenjas, 750 m, 7.V.1995, 1 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 1 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 22-31.X.1997 / 8.II.1998 / 23-28.II.1998, 41 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 m., 1 f., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. In addition to the material studied above, in NMW and MNHUB I have seen some more specimens, which remained unregistered.


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56.29. Harpalus saxicola Dejean, 1829 [= fugax Faldermann, 1836] References. Harpalus fugax: Apfelbeck, 1907a: 498-499, 501 (Oroshi; Mal i Shenjt Mt.: top region, 1500 m). Harpalus saxicola: Kataev & Wrase, 2003: 381. Material studied. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 7 s. (NMW); “Rapsa 1018.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i That” / “Albania Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Albania Tirana” / “28.IX.1958 leg. Xh. Murraj”, 1 s. (HNHM). Species group “oblitus” 56.30. Harpalus oblitus Dejean, 1829 56.30.1. Harpalus oblitus oblitus Dejean, 1829 References. Harpalus oblitus: Hieke & Wrase, 1988: 130. Harpalus oblitus oblitus: Kataev, 1993: 89, fig. 106; Kataev & Wrase, 2003: 379. Notes. In NMW and MNHUB there are several specimens with labels indicating they come from Albania. Additional labels with detailed localities revealed however that the material originates from sites located in the valley of Sava River (Croatia and Serbia). Most probably, Hieke & Wrase (1988: 130) have based their data for the species occurrence in Albania on the wrongly labeled specimens in question. In spite of that Kataev (1993) displayed on map one sure locality for the country. Subgenus Pseudophonus Motschulsky, 1844 56.31. Harpalus griseus (Panzer, 1796) References. Harpalus griseus: Csiki, 1940: 219 (Kula Ljums); Schatzmayr, 1943b: 103 (Dukati). Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 3 s. (NMW); Rodeni, 222.VII.1962, 1 m. (MNST); Tiranë, Selit e Vogel, 17.IX.1993, 2 s., S. Beshkov leg. (NMNHS); “Albania, Alban Alps, Okol, 900 m, swept singled, 16-19.VII.1996, leg. A. Orosz”, 1 m. (HNHM). Notes. Kataev & Wrase (2003: 385) have omitted the species for Albania. Here it is confirmed for the country. 56.32. Harpalus rufipes (DeGeer, 1774) [= pubescens (O.F. Müller, 1776)] References. Harpalus pubescens: Apfelbeck, 1904: 187 (Vlorë; Berat); Schatzmayr, 1943b: 103 (Dukati). Ophonus pubescens: Apfelbeck, 1909: 298 (Berat). Harpalus rufipes: Csiki, 1940: 219 (Kula Ljums); Kataev & Wrase, 2003: 385. Pseudophonus rufipes: Hieke et al., 1986: 187 (Vlorë; North Albanian Alps near Theth; Shkodër); Hieke & Wrase, 1988: 126. Material studied. “Elbasan, Alb., 28.VIII.18 Dr. H. Priesner”, 1 s. (NMW); “Alban., Ricavac Penther”, 1 s. (NMW); “Alban. Exped. Kula Ljums 14.20.VII’18”, 1 s. (NMW); Tiranë, 28.IX.1958, 1 f. (MNST); Ibë, 18.V.1960, 1 m., 1 f. (MNST); Theth, 1-8.VIII.1961, 3 f. (MNST); Tiranë, 10.V.1961, 1 m., 2 f. (MNST); Rushkull, 6.IV.1962, 1 f. (MNST); Rodeni, 2-28.VII.1962, 2 m., 2 f. (MNST); Shkodër, 14.V.1963, 2 m. (MNST); Tiranë, 11.IX.1963, 2 f. (MNST); Shkodër, 12. VII.1965, 1 m. (MNST); Mal i Dajtit Mt., 24.V.1966, 1 m. (MNST); North Albanian Alps, 108


Bogë, 1000-1100 m, 5-9.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District, Merkurth, 11.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995, 1 m., 2 f., D. Antonelli leg. (cAVT); Tiranë, Mal i Dajtit Mt., 7.VI.2002, 1 f., A. Paparistos leg. (cAVT); M. Polisit, 8.VII.2004, 1 f., A. Paparistos leg. (cAVT); Malj Cojepit 1300-1400, 17.VII.2004, 1 m., A. Paparistos leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 815.VII.2006, 3 s., leg. M. Komnenov (cSH). Notes. Besides the material studied above, in NMW and MNHUB I have seen more specimens, which remained unregistered.

57. Genus Ophonus Dejean, 1821 Subgenus Brachyophonus Sciaky, 1987 57.1 Ophonus krueperi Apfelbeck, 1902 References. Ophonus krueperi: Kataev & Wrase, 2003: 388. Material included (studied D.W. Wrase). “AL (Kolonjë) Paß ca. 26 km n. Leskovik -1000 m 5.V.1993 Heinz leg.”, 1 m. (cWR; probably more specimens in cWH). Notes. Apfelbeck published two descriptions for this taxon. First he described Ophonus krueperi (Apfelbeck, 1902b: 543), and after than – Harpalus krueperi (Apfelbeck, 1904: 185). Incorrectly the second description has long time been accredited to be the valid. Actually, this is incorrect and the older description must be assigned as valid. Müller (1931: 48 sub Harpalus krueperi) and Sciaky (1987: 69 sub Ophonus krueperi) followed the record “Chani Driskos” (Apfelbeck, 1904) when they cited the species for Albania. However, this locality lies in Northwestern Greece. According to D.W. Wrase (personal communication), Kataev & Wrase (2003: 388) used the record near Leskovac, marking the species for Albania. First detailed record for the country. Subgenus Hesperophonus Antoine, 1959 57.2. Ophonus azureus (Fabricius, 1775) [= oberthuri (Pater, 1938)] References. Harpalus azureus: Apfelbeck, 1904: 184 (Vlorë); Csiki, 1940: 219 (Kula Ljums; Ura i Lopez). Ophonus azureus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë); Hieke et al., 1986: 187 (North Albanian Alps near Theth; Bizë near Shëngjergji); Hieke & Wrase, 1988: 125; Kataev & Wrase, 2003: 388. Harpalus azureus oberthuri: Schatzmayr, 1943b: 102 (Lushnje). Material studied. “Latif Orosi”, 1 s. (NMW); “Merdita Orosi” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Merdita M. Sheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Ura e Loper” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Kula Ljums 18.-28.V.’18”, 12 s. (NMW); “Alban. Exped. 1918 Bicaj 14.15.VI.”, 8 s. (NMW); “Alban. Exped.’18 Pashtrik, 4-14.7.”, 2 s. (NMW); 53 s. from different localities (MNHUB); Ibë, 17-25.VI.1961, 1 f. (MNST); Rodens, 2-22. VII.1962, 1 s. (MNST); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. + S. Peks”, 2 m., 1 f. (MNHUB); “AL (Korce) Qaf e Qarrit – 1100 m, 4.v.1993 leg. Heinz u. COLEOPTERA: CARABIDAE: HARPALINAE

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Sonja Peks”, 1 m. (MNHUB); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Rhesen District, Merkurth, 11.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Saranda District, Lyukove, 5.V.1994, 2 s., P. Stoev leg. (NMNHS); Saranda District, Qeparo, 4-5.V.1995, 1 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 5.V.1995, 1 s., P. Stoev leg. (NMNHS); Dukati Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS); Kiari Pass, 1160 m, 1.VI.1995, 5 s., G. Blagoev leg. (NMNHS); Tiranë Park, 15.II.1998, 1 s., from grass and soil around tree trunks, R. Ward leg. (CMNH); Tiranë District, Petrela, 28.XI.2000, 1 s., A. Zhalov leg. (NMNHS). Notes. In the region of Gllavë O. azureus was found to lives sympatrically with O. similis. 57.3. Ophonus cribricollis (Dejean, 1829) References. Harpalus cribricollis: Apfelbeck, 1904: 184-185 (Vlorë; Berat); Csiki, 1940: 219 (Shkëlzen Mt.; Kula Ljums; Gostilë). Ophonus cribricollis: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Vlorë; Berat); Hieke & Wrase, 1988: 126; Kataev & Wrase, 2003: 389. Material studied. “Merdita Oroshi”, 3 s. (NMW); “Skutari Mesi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Pasha-Liman Albanien V.1908 H. Hopp”, 1 s. (MNHUB); “Alban. Exped. Kula Ljums 18-28.V.1918”, 3 s. (NMW); “Devoll-Tal juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tiranë” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Dukati Mt., 30 km S Vlorë, 450 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District), 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. In addition to the material studied above, in HNHM the present author has seen more specimens from the localities “Kula Lums”, “Delvine”, “Skölsen” and “Mont Peklen”. Sciaky (1987: 66) cited the locality “Štimlje” for Albania, but actually this place is situated in the Kosovo Region of Serbia. 57.4. Ophonus similis (Dejean, 1829) Material studied. “Hani Grabom Mai” / “Alban. montenegr. Grenze “Alban. Exped. 1918 Bicaj 14.15.VI.”, 1 s. (NMW); “AL (Berat). Paß ca. 5 km S Gllava 600 m, 27.IV.1993 leg. H. + S. Peks”, 2 f. (MNHUB). Notes. New species for the fauna of Albania. 57.5. Ophonus subquadratus (Dejean, 1829) [= meridionalis (Dejean, 1829)] References. Ophonus meridionalis: Apfelbeck, 1909: 298 (Berat; Gramata). Material examined. “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Additional material included (studied S. Hristovski). Dhermi, 200 m, 12.VII.2006, 1 s., M. Komnenov leg. (cSH). Notes. Kataev & Wrase (2003: 389) have omitted the species for Albania. Here it is confirmed for the country. 110


Subgenus Incisophonus Sciaky, 1987 57.6. Ophonus incisus (Dejean, 1829) References. Ophonus incisus: Hieke & Wrase, 1988: 121; Kataev & Wrase, 2003: 389. Material studied. “Murthi” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Albania Cerric” / “16.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS). Notes. First detailed records for Albania. Subgenus Metophonus Bedel, 1897 57.7. Ophonus brevicollis (Audinet-Serville, 1821) [= albanensis (Schauberger, 1933)] References. Harpalus parallelus albanensis: Schauberger, 1933: 65 (Vlorë, loc. typ.). Harpalus albanensis: Turin, 1981: 156. Ophonus brevicollis: Sciaky, 1987: 92-93 (Vlorë); Kataev & Wrase, 2003: 389. 57.8. Ophonus cordatus (Duftschmid, 1812) References. Ophonus cordatus: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Kataev & Wrase, 2003: 389. Material examined. “Mal i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB). 57.9. Ophonus laticollis Mannerheim, 1825 [= punctatulus (Duftschmid, 1812)] References. Harpalus punctatulus: Csiki, 1940: 219 (Gjalicë Mt.). Ophonus punctulatus sic!: Hieke et al., 1986: 187 (Mali me Gropë near Livadhet e Selitës). Ophonus laticollis: Kataev & Wrase, 2003: 390. Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. Hieke & Wrase (1988: 122 sub O. punctatulus) noted this species for Albania apparently following the record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus punctatulus). Actually this locality is situated in the Kosovo Region of Serbia. 57.10. Ophonus melletii (Heer, 1837) Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. 57.11 Ophonus parallelus (Dejean, 1829) [= albanicus Schauberger, 1933] References. Ophonus parallelus albanicus: Schauberger, 1933a: 65 (Vlorë, loc. typ.). Ophonus parallelus: Hieke & Wrase, 1988: 122; Kataev & Wrase, 2003: 390. Material studied. “Alban. Exp.’18 Kruma 5.VI.”, 1 s. (NMW); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “AL (Berat) Paß ca. 5 km s. Gllavë 600 m, 27.IV.1993 leg. Heinz u. S. Peks”, 1 m. (MNHUB). COLEOPTERA: CARABIDAE: HARPALINAE

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57.12. Ophonus puncticeps Stephens, 1828 References. Harpalus puncticeps: Horion, 1941: 212; Magistretti, 1965: 280; Geisthardt, 1975: 8. Ophonus puncticeps: Hieke et al., 1986: 187 (North Albanian Alps near Theth); Kataev & Wrase, 2003: 390. Material studied. “Mali i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “AL (Berat). Paß ca. 5 km S Gllava 600 m. 27.IV.1993 leg. H. + S. Peks”, 1 f. (MNHUB). 57.13. Ophonus puncticollis (Paykull, 1798) References. Ophonus puncticollis: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 298 (Berat); Kataev & Wrase, 2003: 390. Harpalus puncticollis: Müller, 1931: 50 (Tomor Mt.); Csiki, 1940: 219 (Kula Ljums); Hieke & Wrase, 1988: 122. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM). 57.14. Ophonus rufibarbis (Fabricius, 1792) References. Ophonus rufibarbis: Kataev & Wrase, 2003: 391. Material studied. “Berat…Alban.”, 1 s. (MNHUB); “AL (Vlorë) Llogora-Paß (N-Seite) 800 m, 29.IV.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB). Notes. Hieke & Wrase (1988: 124) noted the species for Albania apparently following the record “Prisren” (Apfelbeck, 1904: 183 sub Harpalus rufibarbis). However, this place is situated in the Kosovo Region of Serbia. First detailed records for Albania. 57.15. Ophonus rupicola (Sturm, 1818) References. Harpalus rupicola: Apfelbeck, 1904: 183 (Velipoja); Hieke & Wrase, 1988: 124. Ophonus rupicola: Kataev & Wrase, 2003: 391. Material studied. Kjari Pass, 1160 m, 1.VI.1995, 1 s., G. Blagoev leg. (NMNHS). 57.16. Ophonus schaubergerianus (Puel, 1937) References. Ophonus schaubergerianus: Kataev & Wrase, 2003: 391. Material studied. “Mal i Shenit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB). Notes. First detailed record for Albania. Subgenus Ophonus s.str. 57.17. Ophonus ardosiacus (Lutshnik, 1922) References. Ophonus ardosiacus: Kataev & Wrase, 2003: 391. Material studied. Ibë, 4.VII.1962, 1 f. (MNST). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2 m., D. Antonelli leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 815.VII.2006, 3 s., leg. M. Komnenov (cSH). Notes. First detailed records for Albania. 112


57.18. Ophonus diffinis (Dejean, 1829) References. Harpalus diffinis: Schatzmayr, 1943b: 102 (Mal i Krujes). Ophonus diffinis: Kataev & Wrase, 2003: 391. [Ophonus franziniorum Sciaky, 1987] Notes. According to D.W. Wrase (personal communication), the occurrence of this winged species in Albania is very probable. Except its proven presence in Bulgaria (Sciaky, 1987: 49; Wrase, 1991: 14), there are reliable records for its occurrence in the Greek Ionic Islands and European Turkey (D.W. Wrase, B. Guéorguiev, unpublished data). 57.19. Ophonus sabulicola (Panzer, 1796) [= columbinus (Germar, 1817)] References. Ophonus sabulicola: Apfelbeck, 1909: 298 (Vlorë; Berat); Kataev & Wrase, 2003: 392. Harpalus sabulicola: Csiki, 1940: 219 (Kula Ljums). Harpalus sabulicola columbinus: Schatzmayr, 1943b: 102 (Tërpan; Dukati). Ophonus sabulicola columbinus: Hieke et al., 1986: 187 (Poliçan near Tomor Mt.). Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); 11 s. from different localities (MNHUB); Ibë, 4.VII.1963, 1 f. (MNST); “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 2 s. (MNHUB); “AL (Korcë) Qaf e Qarrit – 1100 m 4.V.1993, leg. Heinz u. S. Peks”, 18 s. (MNHUB). Additional material included (studied S. Hristovski and D.W. Wrase). Korcë District, Mai i That Mt., Zwezde-Sattel, 1000-1100 m, 4.V.1993, 1 f., W. Heinz & S. Peks leg. (cWR); “AL (Kolonjë) Paß ca. 2 km ne. Leskovik -1000 m 5.V.1993 H. & S. Peks leg.”, 16 s. (cWR); 2 km N from Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS); Hotel “Edva”, Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). 57.20. Ophonus stictus Stephens, 1828 Material included (studied S. Hristovski). Hotel “Edva” at Radhime (Vlorë District), 815.VII.2006, 1 f., leg. M. Komnenov (cSH). Notes. New species for the fauna of Albania.

58. Genus Parophonus Ganglbauer, 1891 Subgenus Ophonomimus Schauberger, 1923 58.1. Parophonus hirsutulus (Dejean, 1829) References. Ophonus hirsutulus: Apfelbeck, 1902a: 99 (Velipoja near Lezhë). Harpalus hirsutulus: Apfelbeck, 1904: 186 (Velipoja). Parophonus hirsutulus: Apfelbeck, 1907a: 495 (Buenë River); Sciaky, 1992: 50. Ophonomimus hirsutulus: Hieke & Wrase, 1988: 141. Material studied. “Oboti Bojana”, 1 s. (HNHM); “Oboti Schirei”, 2 s. (HNHM). Notes. Ito (2003b: 392) has omitted the species for Albania. Here it is confirmed for the country.


113

Subgenus Parophonus s.str. 58.2. Parophonus dejeani (Csiki, 1932) References. Parophonus dejeani: Ito, 2003b: 393. 58.3. Parophonus laeviceps (Ménétriés, 1832) [= fallax (Peyron, 1858); planicollis Apfelbeck, 1904 non (Dejean, 1829)] References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë). Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Tachyophonus fallax: Schatzmayr, 1943b: 104 (Stan-Karbunarë). Parophonus laeviceps: Hieke & Wrase, 1988: 142; Ito, 2003b: 393. 58.4. Parophonus maculicornis (Duftschmid, 1812) References. Harpalus maculicornis: Apfelbeck, 1904: 186 (Velipoja; Vlorë; Berat). Ophonus maculicornis: Apfelbeck, 1909: 298 (Vlorë; Berat). Trichotichnus maculicornis: Csiki, 1940: 220 (Kula Ljums). Parophonus maculicornis: Hieke et al., 1986: 186 (Borshi near Vlorë); Hieke & Wrase, 1988: 142; Ito, 2003b: 393. Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Pasha-Liman Albanien V.1908 H. Hopp”, 2 s. (MNHUB); “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Elbasan, Alb. 2.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Logara juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Lum i Erzen Burelle” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “MurrsiKara Alibey” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 1 s. (HNHM); Lushnja, Divjaka National Park, 10.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (cWR; NMNHS); Tiranë Park, 1.II.1998 / 5.II.1998 / 8.II.1998, 12 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). 58.5. Parophonus mendax (P. Rossi, 1790) References. Harpalus mendax: Apfelbeck, 1904: 187 (Vlorë). Ophonus mendax: Apfelbeck, 1909: 298 (Vlorë). Tachyophonus mendax: Schatzmayr, 1943b: 104 (Stan-Karbunarë). Parophonus mendax: Sciaky, 1992: 55; Hieke & Wrase, 1988: 143; Ito, 2003b: 393. Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alessio, Alb. Matzenauer”, 1 s. (NMW); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Tiranë Park, 1-4.I.1998 / 11.I.1998 / 1.II.1998, 10 s., R. Ward leg. (CMNH). 58.6. Parophonus planicollis (Dejean, 1829) References. Ophonus planicollis: Apfelbeck, 1903: 54 (Vlorë); Apfelbeck, 1909: 298 (Vlorë). Harpalus planicollis: Apfelbeck, 1904: 186 (Vlorë). Parophonus planicollis: Ito, 2003b: 393. Material studied. “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj” / “suturifer Rttr. det. Jedlicka”, 1 f. (HNHM). 114


Notes. Hieke & Wrase (1988: 142) noted that the Apfelbeck’s figures for P. planicollis (Apfelbeck, 1904: 187) refer to P. laeviceps.

59. Genus Trichotichnus A. Morawitz, 1863 Subgenus Trichotichnus s.str. 59.1. Trichotichnus laevicollis (Duftschmid, 1812) 59.1.1. Trichotichnus laevicollis laevicollis (Duftschmid, 1812) References. Trichotichnus laevicollis: Schatzmayr, 1926: 35 (Çukall Mt.); Horion, 1941: 233; Hieke & Wrase, 1988: 141. Trichotichnus laevicollis laevicollis: Ito, 2003b: 396. Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 3 f., P. Beron & B. Petrov leg. (NMNHS).

Subtribe STENOLOPHINA Kirby, 1837 60. Genus Acupalpus Latreille, 1829 Subgenus Acupalpus s.str. 60.1. Acupalpus brunnipes (Sturm, 1825) Material studied. “Valona”, 1 s. (HNHM); “Oboti Bojana”, 1 s. (HNHM); “Albania Vlesh” / “30.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM). Notes. New species for the fauna of Albania. 60.2. Acupalpus elegans (Dejean, 1829) References. Acupalpus elegans: Hieke & Wrase, 1988: 116; Jaeger, 1992: 224; Jaeger, 1999: 957; Jaeger & Kataev, 2003: 398. Material studied. “Skutari Bojana”, 1 s. (HNHM); “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Liqen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS). Notes. First detailed records for Albania. 60.3. Acupalpus luteatus (Duftschmid, 1812) References. Acupalpus luteatus: Hieke & Wrase, 1988: 118; Jaeger, 1992: 225; Jaeger, 1999: 959; Jaeger & Kataev, 2003: 398. Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Oboti Bojana”, 1 s. (HNHM); “Skutari Bojana”, 1 s. (HNHM); “Skutari Drin”, 1 s. (HNHM). Additional material included (studied B. Jaeger). “Velipoja” / “Coll. Paganetti”, 1 s. (NMW). Notes. First detailed records for Albania. COLEOPTERA: CARABIDAE: HARPALINAE

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60.4. Acupalpus maculatus (Schaum, 1860) [= immundus Reitter, 1900] References. Acupalpus immundus: Apfelbeck, 1904: 206 (? Shkodër). Acupalpus maculatus: Csiki, 1940: 221 (Kula Ljums); Horion, 1941: 238; Focarile, 1959: 81; Hieke & Wrase, 1988: 118; Jaeger, 1992: 225; Jaeger, 1999: 960; Jaeger & Kataev, 2003: 398. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Busher Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Presba See” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS). Additional material included (studied B. Jaeger). “Elbasan, VIII.1918, Priesner”, 2 s. (MTKD); “Elbasan, Mader”, 1 s. (MTKD); Shkodër, VIII. 1941, 12 s., Capra leg. (MHNG; cBJ); Rrogozhinë, 1 s. (cBJ). Notes. The record “Scutari” (Apfelbeck, 1904) is questionable because the material of Strasser could be collected from the city of Üsküdar - in the Asian seacoast of Bosporus. 60.5. Acupalpus meridianus (Linnaeus, 1761) References. Acupalpus meridianus: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226; Jaeger, 1999: 960; Jaeger & Kataev, 2003: 398. Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB). Notes. First detailed record for Albania. 60.6. Acupalpus notatus Mulsant & Rey, 1861 References. Acupalpus notatus: Horion, 1941: 238; Focarile, 1959: 82; Hieke et al., 1986: 188 (Uji Ftohte, south of Tepelenë; Ibë near Krraba); Hieke & Wrase, 1988: 118; Jaeger, 1988: 20 (Ljum, Shkumbin; Durrës: Rogozina; Elbasan; Mati: Burrelë; Tepelenë: Uji Ftohte; Ibë near Krraba; Tiranë; Vlorë); Jaeger, 1992: 226; Jaeger, 1999: 961; Jaeger & Kataev, 2003: 398. Material studied. “Elbasan Alb., Mader”, 3 s. (NMW; MNHUB); “Vorra, Alb. leg. Mader”, 2 s. (NMW); “Ljum i Shkumbin, Juni 1932” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Burelle” / “Albania leg. Bischoff 1934”, 16 s. (MNHUB); “Shullaz” / “Albania leg. Bischoff 1934”, 5 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 3 s. (MNHUB); Tiranë Park, 22.II.1998, 30 s., grasses along small sluggish stream, R. Ward leg. (CMNH). Additional material included (studied B. Jaeger and A. Vigna Taglianti). “Elbasan, Mader”, 4 s. (MTKD, cBJ); “Elbasan, Priesner”, 4 s. (MTKD); Shkodër, VIII. 1941, 2 s., Capra leg. (MHNG); Durrës, Plepa, 15.VIII.1995, 1 m., D. Antonelli leg. (cAVT). 60.7. Acupalpus paludicola Reitter, 1884 [= puncticollis Auct. nec (Coquerel, 1859)] References. Acupalpus paludicola: Apfelbeck, 1904: 206 (Vlorë; Velipoja); Apfelbeck, 1909: 298 (Vlorë-Malket); Jaeger, 1989: 148 (Vlorë; Lum i Tiranes; Shkodër); Jaeger, 1992: 226; Jaeger, 1999: 961; Jaeger & Kataev, 2003: 399. Acupalpus puncticollis: Hieke et al., 1986: 188 (Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 117. Material studied. “Albania Avlona v. Oertzen”, 3 s. (MNHUB); “Skutari l. See-Ufer”, 1 s. (NMW); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB; cBJ). 116


Notes. All older references and collection records in the museums about A. puncticollis (Coquerel, 1859) from Albania refer to A. paludicola. 60.8. Acupalpus parvulus (Sturm, 1825) [= dorsalis (Fabricius, 1787)] References. Acupalpus dorsalis: Apfelbeck, 1904: 206 (Velipoja; Vlorë); Apfelbeck, 1909: 298 (Vlorë); Csiki, 1940: 221 (Kula Ljums); Hieke & Wrase, 1988: 117. Material studied. “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); Tiranë Park, 8.II.1998 / 1.III.1998, 3 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Notes. Jaeger & Kataev (2003: 399) have omitted the species for Albania. Here it is confirmed for the country. According to B. Jaeger and D.W. Wrase (personal communication) the records from Albania may concern aberrant specimens of A. maculatus without darkened humeral macula, but the only certainty for a correct identification is the study of male genitalia. 60.9. Acupalpus planicollis (Schaum, 1857) References. Acupalpus planicollis: Apfelbeck, 1904: 205 (Velipoja); Apfelbeck, 1909: 298 (VlorëMalket); Hieke & Wrase, 1988: 116; Guéorguiev & Guéorguiev, 1995: 177; Guéorguiev et al., 1997: 20; Jaeger & Kataev, 2003: 399. Material studied. “Liqen i Butrintes vivari” / “leg. Bischoff Albani 1934”, 1 s. (MNHUB). Additional material included (studied B. Jaeger). “Velipoja, Mustjabeg”, 1 s. (ZSM). 60.10. Acupalpus suturalis Dejean, 1829 60.10.1. Acupalpus suturalis suturalis Dejean, 1829 References. Acupalpus suturalis: Hieke & Wrase, 1988: 117; Jaeger, 1992: 226. Acupalpus suturalis suturalis: Jaeger, 1999: 962; Jaeger & Kataev, 2003: 399. Material studied. “Oboti Bojana”, 1 s. (HNHM); Tiranë Park, 22-31.X.1997, 1 s., R. Ward leg. (CMNH); Tiranë Park, 22.II.1998, 3 s., R. Ward leg. (CMNH). Additional material included (studied B. Jaeger). “Scutari, VIII. 1941, Capra”, 1 s. (MHNG). Notes. First detailed records for Albania.

61. Genus Anthracus Motschulsky, 1850 61.1. Anthracus longicornis (Schaum, 1857) References. Anthracus longicornis: Apfelbeck, 1904: 207 (Velipoja); Hieke & Wrase, 1988: 118; Jaeger & Kataev, 2003: 400. Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Oboti Bojana”, 4 s. (HNHM); “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 4 s. (MNHUB). 61.2. Anthracus quarnerensis (Reitter, 1884) References. Anthracus quarnerensis: Apfelbeck, 1904: 208 (Velipoja); Hieke & Wrase, 1988: 119; Jaeger & Kataev, 2003: 400. COLEOPTERA: CARABIDAE: HARPALINAE

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62. Genus Bradycellus Erichson, 1837 Subgenus Bradycellus s.str. 62.1. Bradycellus caucasicus (Chaudoir, 1846) [= collaris (Paykull, 1798)] References. Bradycellus collaris: Csiki, 1940: 221 (Shkëlzen Mt.). Material studied. “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM). Notes. Jaeger & Kataev (2003: 400) have omitted the species for Albania. Here it is confirmed for the country. 62.2. Bradycellus distinctus (Dejean, 1829) References. Bradycellus distinctus: Jaeger, 1990: 10; Jaeger, 1999: 966; Jaeger & Kataev, 2003: 401. Material studied. “Skutari Bojana”, 5 s. (NMW). Additional material included (studied B. Jaeger). “Skutari Bojana”, 12 s. (MNHUB; DEI; NMPC; cBJ). Notes. First detailed record for Albania. 62.3. Bradycellus ganglbaueri Apfelbeck, 1904 References. Bradycellus ganglbaueri: Jaeger, 1997: 352 (Mal i Krujes); Jaeger & Kataev, 2003: 401. 62.4. Bradycellus verbasci (Duftschmid, 1812) References. Bradycellus verbasci: Hieke & Wrase, 1988: 120; Jaeger & Kataev, 2003: 401. Material studied. Velipoja, 19.IX.1993, 8 s., S. Beshkov leg. (NMNHS; ZISP); Tiranë Park, 19.XI.1997, 1 s., R. Ward leg. (CMNH). Additional material included (studied B. Jaeger). “Velipoja Avola v. Oertzen”, 1 s. (MNHUB). Notes. First detailed records for Albania.

63. genus Dicheirotrichus Jaquelin du Val, 1857 Subgenus Dicheirotrichus s.str. 63.1. Dicheirotrichus obsoletus (Dejean, 1829) References. Dicheirotrichus obsoletus: Apfelbeck, 1904: 211 (Velipoja); Hieke & Wrase, 1988: 120; Jaeger & Kataev, 2003: 402. Material studied. “Durres” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB). Notes. According to B. Jaeger (personal communication) E. Schauberger identified the material from MNHUB as D. obsoletus, but a revision of that is needed.

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64. Genus Loxoncus Schmidt-Göbel, 1846 64.1. Loxoncus procerus (Schaum, 1858) References. Anoplogenius procerus: Apfelbeck, 1904: 202 (Velipoja near Lezhë); Apfelbeck, 1907a: 496 (Buenë River Outfall); Schatzmayr, 1943b: 105 (Lushnje). Loxoncus procerus: Wrase, 1991: 12; Kataev, 2002: 364 (Buenë River; Buenë River at Oboti); Jaeger & Kataev, 2003: 404. Material studied. “Apfelb. Albanien”, 2 s. (MNHUB); “Apfelb. Albanien” / “Bojana Mündung”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM); “Skutari Bojana”, 3 s. (HNHM); “Liqen i Butrintes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 65. Genus Stenolophus Dejean, 1821 Subgenus Egadroma Motschulsky, 1855 65.1. Stenolophus marginatus Dejean, 1829 References. Egadroma marginata: Apfelbeck, 1904: 204 (Vlorë; Berat); Apfelbeck, 1907a: 496 (Buenë River); Hieke et al., 1986: 188 (Durrës; Uji Ftohte, south of Tepelenë); Hieke & Wrase, 1988: 116. Stenolophus marginatus: Apfelbeck, 1909: 298 (Vlorë); Jaeger & Kataev, 2003: 405. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Oboti Bojana”, 3 s. (HNHM); “Skumbi Elbasan Alb. 10.8.1918 Dr. H. Priesner”, 3 s. (NMW); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Myzeqe Ebene” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); “Albanien-Exp. DEI Uji Ftohte, 200 m, südl. Tepelene lux 29.-31.V.1961”, 1 s. (MNHUB). Subgenus Stenolophus s.str. 65.2. Stenolophus abdominalis Gené, 1836 65.2.1. Stenolophus abdominalis persicus Mannerheim, 1844 Material studied. Tiranë Park, 22.II.1998, 1 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH); Tiranë Park, 1.III.1998, 2 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. New species for the fauna of Albania. 65.3. Stenolophus discophorus (Fischer von Waldheim, 1823) References. Stenolophus discophorus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988: 114; Jaeger & Kataev, 2003: 405. Acupalpus discophorus: Csiki, 1940: 221 (Kula Ljums). Material studied. “Oboti Bojana”, 5 s. (NMW); “Albania Kula Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 4 s. (MNHUB); Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS). COLEOPTERA: CARABIDAE: HARPALINAE

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65.4. Stenolophus mixtus (Herbst, 1784) References. Stenolophus mixtus: Apfelbeck, 1904: 203 (Velipoja); Hieke & Wrase, 1988: 114; Jaeger & Kataev, 2003: 406. 65.5. Stenolophus proximus Dejean, 1829 References. Stenolophus proximus: Apfelbeck, 1904: 202 (Velipoja); Hieke & Wrase, 1988: 114; Jaeger & Kataev, 2003: 406. 65.6. Stenolophus skrimshiranus Stephens, 1828 References. Stenolophus skrimshiranus: Apfelbeck, 1904: 203 (Buenë River at Velipoja); Apfelbeck, 1909: 298 (Pasha Liman); Hieke & Wrase, 1988: 115; Jaeger & Kataev, 2003: 406. Material studied. “Ljum i Schkumbin Juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). 65.7. Stenolophus teutonus (Schrank, 1781) References. Stenolophus teutonus: Apfelbeck, 1904: 203 (Velipoja); Apfelbeck, 1909: 298 (Vlorë); Murraj, 1968: 33 (Ibë); Hieke et al., 1986: 188 (Vlorë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 115; Jaeger & Kataev, 2003: 406. Acupalpus teutonus: Csiki, 1940: 221 (Kula Ljums); Schatzmayr, 1943b: 105 (Fieri). Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB); “Alban. Exp.’18 Kula Ljums 26.VI.-3.VII.”, 2 s. (NMW); “Elbasan, Alb. 16.IX.18 Dr. H. Priesner”, 6 s. (NMW); “Berat, Juli”, “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Busher, Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Shkumbin, Juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Maia e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tepelene” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Burelle” / “Albania leg. Bischoff 1934”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1934”, 7 s. (MNHUB); “Shullaz” / “Albania leg. Bischoff 1934”, 4 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Tiranë, 24.VI.1961, 1 f. (MNST); Rodeni, 2-22.VII.1962, 1 s. (MNST); Tiranë Park, 22.II.1998, 1 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. The record of Murraj (1968) is included here with reservation because of the misidentification found in his collection.

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Tribe LEBIINI Bonelli, 1810 Subtribe CYMINDIDINA Laporte, 1834 66. Genus Cymindis Latreille, 1806 Subgenus Cymindis s.str. 66.1. Cymindis angularis Gyllenhal, 1810 Material studied. “Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 15 s. (MNHUB); “Taja” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 20 s. (MNHUB); “Taja” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Notes. New species for the faunas of the Balkan Peninsula and Albania. Most likely, the populations from the high mountains in North Albania are glacial relics. 66.2. Cymindis axillaris (Fabricius, 1794) 66.2.1. Cymindis axillaris axillaris (Fabricius, 1794) References. Cymindis axillaris: Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Klaptocz, 1910: 399 (Oroshi); Hieke et al., 1986: 190 (North Albanian Alps near Theth). Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mali i Shenjit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Boge” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania Mueller leg. 1939”, 9 s. (MNHUB); “Iballe” / “Albania Mueller leg. 1939”, 1 s. (MNHUB). Notes. Kabak (2003: 411) has omitted the species for Albania. Here it is confirmed for the country. 66.2.2. Cymindis axillaris adriatica J. Müller, 1921 References. Cymindis axillaris adriatica: Müller, 1921a: 139 (Shkodër). 66.3. Cymindis coadunata Dejean, 1825 66.3.1. Cymindis coadunata coadunata Dejean, 1825 Material studied. “Çafa Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 6 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB). Notes. New species for the fauna of Albania. 66.4. Cymindis humeralis (Geoffroy, 1785) References. Cymindis humeralis: Csiki, 1940: 222 (Shkëlzen Mt.); Hieke & Wrase, 1988: 161. COLEOPTERA: CARABIDAE: HARPALINAE

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Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban. Exp.’18 Pashtrik, 4-14.7.”, 1 s. (NMW); 68 s. from different localities (MNHUB). Notes. Kabak (2003: 413) has omitted the species for Albania. Here it is confirmed for the country. 66.5. Cymindis lineata (Quensel, 1806) References. Cymindis lineata: Apfelbeck, 1904: 343 (Vlorë); Apfelbeck, 1907a: 499 (Mal i Shenjt Mt.: top region, 1500 m); Hieke & Wrase, 1988: 162; Kabak, 2003: 413. References. “Merdita Oroshi” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Alban. Exped. Kula Ljums 18.28.V.1918”, 2 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 3 s. (NMW); “Mal i Dajtit Tirane” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Scheinjit” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Borsne” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “AL (Korcë) Vlacisht n. Korcë, 800 m 4.V.1993, leg. Heinz u. S. Peks (ex. coll H. Peks)”, 1 s. (MNHUB). 66.6. Cymindis lineola L. Dufour, 1820 References. Cymindis axillaris lineola: Apfelbeck, 1907a: 495 (Buenë River). Material studied. “Latif Skutari”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 2 s. (HNHM). Notes. Kabak (2003: 414) has omitted the species for Albania. Here it is confirmed for the country. ? Cymindis pindicola Apfelbeck, 1901 Material studied. “Neres” / “Apfelb. 1906 Albania or.”, 1 s. (HNHM). Notes. Apfelbeck described this species from the mountains of Peristeri, Veluchi and Oxya situated in Northwestern and Central Greece. The specimen from “Neres” has been found to be conspecific with a single syntype of C. pindicola labeled “Veluchi Gb. Stralcia” / “Coll. Apfelbeck” (HNHM). However, I failed to find a place named “Neres” in Albania or in the adjacent regions. Because of uncertain localization, this record remains questionable for Albania. 66.7. Cymindis scapularis Schaum, 1857 66.7.1. Cymindis scapularis scapularis Schaum, 1857 References. Cymindis scapularis var.: Apfelbeck, 1909: 298 (Vlorë-Fieri). Notes. Kabak (2003: 411) has omitted the species for Albania. Actually, it needs further confirmation for the country. Subgenus Menas Motschulsky, 1864 Notes. New subgenus for the fauna of Albania.

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66.8. Cymindis miliaris (Fabricius, 1801) Material studied. “Albania Rapsa Dr. Grabowski”, 3 s. (HNHM). Notes. New species for the fauna of Albania. Subgenus Tarulus Bedel, 1906 66.9. Cymindis vaporariorum (Linnaeus, 1758) References. Cymindis vaporariorum: Hieke & Wrase, 1988: 162. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Korab” / “Albania leg. Bischoff 1935”, 20 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Notes. Kabak (2003: 418) has omitted the species for Albania. Here it is confirmed for the country with first detailed records.

Subtribe DROMIINA Bonelli, 1810 67. Genus Dromius Bonelli, 1810 Notes. New genus for the fauna of Albania. Subgenus Dromius s.str. Notes. New subgenus for the fauna of Albania. 67.1. Dromius quadrimaculatus (Linnaeus, 1758) Material studied. “Elbasan, Alb. 19.IX.18. Dr. H. Priesner”, 1 s. (NMW). Notes. New species for the fauna of Albania.

68. Genus Microlestes Schmidt-Göbel, 1846 68.1. Microlestes corticalis (L. Dufour, 1820) [= escorialensis (Brisout de Barneville, 1885)] References. Microlestes plagiatus escorialensis: Holdhaus, 1904: 335 (Velipoja; Vlorë). Microlestes corticalis escorialensis: Schatzmayr, 1943b: 112 (Lushnje). Microlestes corticalis: Hieke & Wrase, 1988: 160; Kabak, 2003: 422. Material studied. “Albanien Avlona v. Oertzen”, 1 s. (MNHUB). 68.2. Microlestes fissuralis (Reitter, 1901) References. Microlestes fissuralis: Holdhaus, 1904: 332 (Velipoja; “Ardenica” near Vlorë); Csiki, 1940: 221 (Gjalicë Mt.); Horion, 1941: 343; Hieke et al., 1986: 190 (Kula e Lumës near Kukës); Hieke & Wrase, 1988: 159; Kabak, 2003: 422. COLEOPTERA: CARABIDAE: HARPALINAE

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Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Kara Alibey” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); Librazhd Prenjas, 750 m, 7.V.1995, 1 s., L. Penev leg. (NMNHS). 68.3. Microlestes luctuosus Holdhaus, 1904 68.3.1. Microlestes luctuosus luctuosus Holdhaus, 1904 References. Microlestes exilis luctuosus: Holdhaus, 1904: 331 (Berat). Microlestes luctuosus: Hieke & Wrase, 1988: 159. Microlestes apterus: Wrase, 1991: 18. Material studied. “Albanien Avlona v. Oertzen”, 2 s. (MNHUB); “Kjori” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Murrsi - Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. According to Wrase (1991) the material cited as M. exilis Schmidt-Göbel, 1846 and M. exilis luctuosus (Apfelbeck, 1904: 330-331) may concern either M. apterus or M. luctuosus. I refer it provisionally to the latter species, which was proved to inhabit the country. Kabak (2003: 423) has omitted or M. luctuosus for Albania. Here, the species is confirmed for the country. 68.4. Microlestes maurus (Sturm, 1827) 68.4.1. Microlestes maurus maurus (Sturm, 1827) References. Microlestes maurus: Holdhaus, 1904: 332 (Velipoja; Vlorë); Hieke & Wrase, 1988: 159; Kabak, 2003: 423. 68.5. Microlestes minutulus (Goeze, 1777) References. Blechrus minutulus: Apfelbeck, 1909: 298 (Vlorë; Berat). Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i Krujes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the country. 68.6. Microlestes negrita (Wollaston, 1854) 68.6.1. Microlestes negrita apfelbecki Holdhaus, 1904 References. Microlestes minutulus apfelbecki: Holdhaus, 1904: 330 (Velipoja). Microlestes negrita: Hieke & Wrase, 1988: 158. Material studied. “Cafa Liqenit Iballe” / “Albania leg. Mueller 1939”, 1 s. (MNHUB). Notes. Kabak (2003: 423) has omitted the species for Albania. Here it is confirmed for the country. 68.7. Microlestes plagiatus (Duftschmid, 1812) References. Microlestes plagiatus: Apfelbeck, 1907a: 495 (Buenë River); Csiki, 1940: 221 (Korab Mt.). Notes. Kabak (2003: 423) has omitted the species for Albania. Actually it needs further confirmation for the country. 124


69. Genus Paradromius Fowler, 1887 Subgenus Manodromius Reitter, 1905 69.1. Paradromius linearis (Olivier, 1795) 69.1.1. Paradromius linearis linearis (Olivier, 1795) References. Dromius linearis: Hieke & Wrase, 1988: 155. Material studied. “Vorra, Alb. 25.IX.18 Dr. H. Priesner”, 1 s. (NMW); “Bushek juni” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Durres juni” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranës” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Lum i Tiranës” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); Tiranë Park, 8.II.1998, 2 s., R. Ward leg. (CMNH). Notes. Kabak (2003: 424) has omitted the species for Albania. Here P. linearis is confirmed for the country with first detailed records.

70. Genus Philorhizus Hope, 1838 subgenus Philorhizus s.str. 70.1. Philorhizus notatus (Stephens, 1827) References. Dromius notatus: Hieke & Wrase, 1988: 156. Material studied. “Oboti Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM); “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. Kabak (2003: 426) has omitted the species for Albania. Here it is confirmed for the country with first detailed records. 70.2. Philorhizus sigma (P. Rossi, 1790) References. Dromius sigma: Apfelbeck, 1904: 337 (Velipoja); Hieke & Wrase, 1988: 157; Kabak, 2003: 426.

Subtribe LEBIINA Bonelli, 1810 71. Genus Lebia Latreille, 1802 Subgenus Lamprias Bonelli, 1810 Notes. New subgenus for the fauna of Albania. 71.1. Lebia cyanocephala (Linnaeus, 1758) 71.1.1. Lebia cyanocephala cyanocephala (Linnaeus, 1758) Material studied. “Skutari Drin”, 1 s. (HNHM); “Tirane” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB). COLEOPTERA: CARABIDAE: HARPALINAE

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Notes. The mention of Hieke & Wrase (1988: 153) for Albania is apparently result of a following the record “Prisren” (Apfelbeck, 1904: 323). However, this locality is situated in the Kosovo Region of Serbia. Kabak (2003: 427) has most likely followed one of these sources, too. Since there is no previous reliable record from Albania, L. cyanocephala is treated here as new species for the fauna of Albania. Subgenus Lebia s.str. 71.2. Lebia cruxminor (Linnaeus, 1758) 71.2.1. Lebia cruxminor cruxminor (Linnaeus, 1758) References. Lebia cruxminor: Hieke et al., 1986: 190 (North Albanian Alps near Theth; Kula e Lumës near Kukës); Hieke & Wrase, 1988: 153. Material studied. “Skutari Drin”, 1 s. (HNHM); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Cucale Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Kjori” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albanien-Exp. DEI Nordalban. Thethi, 1.-4.VIII.61”, 1 s. (MNHUB). Notes. Kabak (2003: 428) has omitted the species for Albania. Here L. cruxminor is confirmed for the country. 71.3. Lebia humeralis Dejean, 1825 References. Lebia humeralis: Hieke & Wrase, 1988: 154. Material studied. “Albania Djakova” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Alban. Exped. Gjalica Ljums 17-26.VI.’18”, 1 s. (NMW); “Lum i Tiranës” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Kabak (2003: 429) has omitted the species for Albania. Here it is confirmed for the country with first detailed records. 71.4. Lebia scapularis (Geoffroy, 1785) 71.4.1. Lebia scapularis scapularis (Geoffroy, 1785) References. Lebia scapularis: Hieke & Wrase, 1988: 154. Material studied. “Serandë Piqeros” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. Kabak (2003: 429) has omitted the species for Albania. Here L. scapularis is confirmed for the country with a first detailed record. 71.5. Lebia trimaculata (Villers, 1789) Material studied. 12 km NE Erseka, 900 m, 12.V.1995, 2 s., L. Penev leg. (NMNHS). Notes. New species for the fauna of Albania.

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Subtribe LIONYCHINA Jeannel, 1948 72. Genus Apristus Chaudoir, 1846 72.1. Apristus reticulatus Schaum, 1857 Material studied. “Mesi Kiri-Ufer” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. New species for the fauna of Albania. 72.2. Apristus subaeneus Chaudoir, 1846 References. Apristus subaeneus: Apfelbeck, 1907a: 495 (Liqen i Shkodrës); Müller, 1921a: 139; Müller, 1924: 139; Hieke & Wrase, 1988: 161. Material studied. “Skutari Drin”, 1 s. (NMW); “Skutari Vraka”, 6 s. (HNHM); “Skutari Vraka” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW). Notes. Kabak (2003: 431) has omitted the species for Albania. Here it is confirmed for the country. 73. Lionychus Wissmann, 1846 Notes. New genus for the fauna of Albania. Subgenus Lionychus s.str. Notes. New subgenus for the fauna of Albania. 73.1. Lionychus maritimus Fairmaire, 1862 Material studied. “Himara juni” / “Albania leg. Bischoff 1932”, 17 s. (MNHUB). Notes. New species for the faunas of the Balkan Peninsula and Albania. 73.2. Lionychus quadrillum (Duftschmid, 1812) Material studied. “Medua, Alb. 10.5.914”, 1 s. (HNHM). Notes. New species for the fauna of Albania.

74. Genus Syntomus Hope, 1838 74.1. Syntomus impressus (Dejean, 1825) 74.1.1. Syntomus impressus impressus (Dejean, 1825) References. Metabletus impressus: Apfelbeck, 1907a: 495 (Buenë River). Syntomus impressum sic!: Hieke & Wrase, 1988: 158. Material studied. “Albanien Avlona v. Oertzen.”, 2 s. (MNHUB); Saranda District, Ljukove, 3-4.VI.1994, 5 s., G. Blagoev leg. (NMNHS). Notes. Kabak (2003: 432) has omitted the species for Albania. Here it is confirmed for the country. COLEOPTERA: CARABIDAE: HARPALINAE

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74.2. Syntomus obscuroguttatus (Duftschmid, 1812) References. Metabletus obscuroguttatus: Apfelbeck, 1904: 327 (Vlorë; Velipoja); Apfelbeck, 1909: 298 (Vlorë). Syntomus obscuroguttatus: Hieke & Wrase, 1988: 157; Kabak, 2003: 433. Material studied. “Oboti Bojana”, 4 s. (HNHM); “Skutari Bojana”, 1 s. (HNHM); “Logara” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); Velipoja, 19.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 1 s., G. Blagoev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Tiranë Park, 15.II.1998, 320 s., from grass and soil around tree trunks, R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Durrës, Plepa, 15.VIII.1995, 2 m., D. Antonelli leg. (cAVT). 74.3. Syntomus pallipes (Dejean, 1825) References. Metabletus pallipes: Apfelbeck, 1909: 298 (Vlorë). Syntomus pallipes: Hieke & Wrase, 1988: 157. Material studied. “Okol Shkalla” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Notes. Kabak (2003: 433) has omitted the species for Albania. Here it is confirmed for the country. 74.4. Syntomus truncatellus (Linnaeus, 1761) References. Metabletus truncatellus: Csiki, 1940: 221 (Korab Mt.). Syntomus truncatellus: Hieke & Wrase, 1988: 158. Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 3 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Korab” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB). Notes. Kabak (2003: 433) has omitted the species for Albania. Here it is confirmed for the country.

Tribe LICININI Bonelli, 1810 Subtribe LICININA Bonelli, 1810 75. Genus Badister Clairville, 1806 Subgenus Badister s.str. 75.1. Badister bullatus (Schrank, 1798) Material studied. “Scutari Bojana”, 1 s. (HNHM); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW). Notes. New species for the fauna of Albania. 75.2. Badister unipustulatus Bonelli, 1813 References. Badister unipustulatus: Apfelbeck, 1904: 163 (Velipoja); Murraj, 1968: 33 (Tiranë); Hieke & Wrase, 1988: 151; Baehr, 2003c: 440. 128


Material studied. “Oboti Bojana”, 2 s. (HNHM). Notes. The record of Murraj (1968) is included here with reservation due to misidentifications found in his collection. Subgenus Baudia Ragusa, 1884 75.3. Badister dilatatus Chaudoir, 1837 References. Badister dilatatus: Apfelbeck, 1904: 164 (Velipoja); Hieke & Wrase, 1988: 151; Baehr, 2003c: 441. 75.4. Badister peltatus (Panzer, 1796) 75.4.1. Badister peltatus peltatus (Panzer, 1796) References. Badister peltatus peltatus: Baehr, 2003c: 441.

76. Genus Licinus Latreille, 1802 Subgenus Licinus s.str. 76.1. Licinus cassideus (Fabricius, 1792) 76.1.1. Licinus cassideus cassideus (Fabricius, 1792) [= latus Dejean, 1831] References. Licinus cassideus latus: Magistretti, 1965: 263; Hieke & Wrase, 1988: 150. Licinus cassideus cassideus: Baehr, 2003c: 442. Material studied. “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomor-Tojk juli 1932” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Boge Aufstieg z. Rab” / “Albania leg. Mueller 1939”, 1 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Vasjar (Tepelenë District), 18.IV.2003, 2 f., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. Csiki (1931: 911-912) treated dohrni, graecus and submarginatus as synonyms of cassideus. Here, I follow the arrangement of Baehr (2003c: 442). In spite of that, a revision of the Balkan taxa close to L. cassideus is needed. First detailed records for Albania. 76.1.2. Licinus cassideus dohrni Fairmaire, 1863 References. Licinus cassideus dohrhi: Fairmaire, 1863: 131 (Albania, loc. typ.); Reitter, 1900: 150; Apfelbeck, 1904: 162; Baehr, 2003c: 442. Material studied. “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 1 s. (MNHUB). 76.2. Licinus graecus Apfelbeck, 1901 [= submarginatus J.R. Sahlberg, 1903] References. Licinus graecus: Apfelbeck, 1901a: 431 (Berat); Apfelbeck, 1909: 298 (Berat). Licinus cassideus graecus: Apfelbeck, 1904: 162 (Berat). Licinus submarginatus: Schatzmayr, 1943b: 100 (Tërpan); Baehr, 2003c: 442. COLEOPTERA: CARABIDAE: HARPALINAE

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Notes. Apfelbeck published two descriptions of L. graecus. First he described L. graecus (Apfelbeck, 1901a: 431), and after that L. cassideus graecus (Apfelbeck, 1904: 162). Incorrectly the second taxon has been accredited to be the valid, which create a future complication. In 1904, Apfelbeck (1904: 399, 402 note 1) synonymized L. submarginatus J.R. Sahlberg, 1903 with L. cassideus graecus Apfelbeck, 1904, not with L. graecus Apfelbeck, 1901. The only explanation of this fact is that this outstanding specialist seemed to have forgotten his earlier description. Later, the subsequent authors (Schatzmayr, 1943b; Baehr, 2003c) followed this view, but treated L. submarginatus as valid species, and L. cassideus graecus Apfelbeck, 1904 – for its synonym. I believe, however, that both Licinus - species described by Apfelbeck are identical based on identical type localities. According to the rules of ISZN (1999) an original description of a taxon is the oldest one. Therefore, the following nomenclatorial act is proposed: Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn. nov. of Licinus (Licinus) graecus Apfelbeck, 1901. 76.3. Licinus silphoides (P. Rossi, 1790) References. Licinus silphoides: Apfelbeck, 1909: 298 (Vlorë-Fieri); Schatzmayr, 1943b: 100 (Karbunara e Madhe; Stan-Karbunarë); Focarile, 1959: 76; Hieke & Wrase, 1988: 150; Baehr, 2003c: 442. Subgenus Neorescius Bedel, 1906 76.4. Licinus oertzeni Reitter, 1889 76.4.1. Licinus oertzeni oertzeni Reitter, 1889 References. Licinus oertzeni: Apfelbeck, 1907a: 499, 501,503 (Mal i Shenjt Mt.: top region, 1500 m). 76.4.2. Licinus oertzeni albanicus J. Müller, 1932 References. Licinus oertzeni albanicus: Müller, 1932: 209 (Tomor Mt. near Berat, loc. typ.; Northwestern Greece, “Buč-Gebirge”, loc. typ.); Baehr, 2003c: 443. Material studied. “Merdita Orochi”, 1 s. (HNHM); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Çafa Kjores” / “Albania leg. Bischoff 1935” / “Licin. Oertzeni albanicus m det. J. Müller”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. The distinction of the two subspecies of L. oertzeni s.l. is very easy due to their clear external differences. Probably both taxa deserve species status.

Tribe ODACANTHINI Laporte, 1834 Notes. New tribe for the fauna of Albania.

77. Genus Odacantha Paykull, 1798 Notes. New genus for the fauna of Albania. 130


Subgenus Odacanta s.str. Notes. New subgenus for the fauna of Albania. 77.1. Odacantha melanura (Linnaeus, 1767) Material studied. “Skutari Bojana” / “Coll. Apfelbeck”, 1 s. (HNHM). Notes. New species for the fauna of Albania.

Tribe OODINI LaFerté-Sénectère, 1851 78. Genus Oodes Bonelli, 1810 78.1. Oodes gracilis A. Villa & G.B. Villa, 1833 References. Oodes gracilis: Apfelbeck, 1904: 159 (Velipoja); Focarile, 1959: 75; Hieke & Wrase, 1988: 150; Bousquet, 2003c: 445. Material studied. Shkodër District, Mali Tarabosh Mt., 100 m, 24.XI.2000, 3 m., limestone terrain, A. Zhalov leg. (NMNHS). 78.2. Oodes helopioides (Fabricius, 1792) 78.2.1. Oodes helopioides helopioides (Fabricius, 1792) References. Oodes helopioides: Apfelbeck, 1904: 159 (Velipoja near Lezhë); Hieke & Wrase, 1988: 150; Bousquet, 2003c: 445.

Tribe OMPHREINI Ganglbauer, 1891 79. Genus Omphreus Dejean, 1828 Notes. This genus needs revision. The author’s examinations revealed only a small number of specimens of the most species and these in bad condition (excluding O. morio s.l.) in the largest Central European collections (DEI; HNHM; NMW). Contrary, in MNHUB the collection of Omphreus is better represented. Subgenus Omphreus s.str. 79.1. Omphreus lonai Winkler, 1933 References. Omphreus lonai: Winkler, 1933b: 118 (Llogora, 1200 m, southeast of Vlorë, loc. typ.); Bousquet, 2003d: 446. Material studied. “Llogora” / “Albania leg. Bischoff 1935”, 1 m. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 3 f. (MNHUB).


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79.2. Omphreus morio Dejean, 1828 79.2.1. Omphreus morio albanicus Apfelbeck, 1906 References. Omphreus albanicus: Apfelbeck, 1906: 1666-1667 (Merdita: Mal i Shenjt Mt., loc. typ.; Zepë Mt., loc. typ.); Apfelbeck, 1907a: 498 (Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1918c: 160, 165 (Mal i Shenjt Mt.; Zepë Mt.; North Albanian Alps, 1300-2000 m); Hieke, 1981: 82. Omphreus morio: Anonymous, 1907: 286. Omphreus morio albanicus: Winkler, 1933b: 117 (Mal i Dajtit Mt.; Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Peshtrik Mt.); Drovenik & Peks, 1994: 75 (Peshtrik Mt.); Drovenik & Peks, 1999: 87 (Peshtrik Mt.). Material studied. “Merdita M. Scheit” / “Coll. Apfelbeck”, 4 s. (HNHM); “Merdita M. Scheit”, 3 s. (NMW); “Merdita Zebia”, 6 s. (HNHM; NMW; MNHUB); “Merdita Munela”, 1 s. (HNHM); “Oroši (Lat.) Alban. spt.” / “Zebja”, 2 s. (DEI; MNHUB); “Merdita Munela” / “Wgth., 1906 Merdita”, 1 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Alban. Exped. Gjalica Ljums 17.-26.VI’18”, 2 s. (NMW); “Mal i Dajtit juni” / “Albania leg. Bischoff 1932”, 2 m., 1 f. (MNHUB); “Mal i Krujes” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 1 m. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 m. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 f. (MNHUB); Reshen District, near Merkurth, 920-980 m, 11.VI.1993, 1 s., grassland, P. Beron & B. Petrov leg. (NMNHS); Mal i Dajtit Mt., 20 km east of Tiranë, 1000 m, 9.V.1995, 3 s., L. Penev leg. (NMNHS). Notes. Bousquet (2003d: 446) has omitted O. morio for Albania. Here the species is confirmed for the country. 79.2.2. Omphreus morio malissorum Winkler, 1933 References. Omphreus morio malissorum: Winkler, 1933b: 117 (North Albanian Alps: Maja e Jezercë, loc. typ.; Starca; Çukall Mt.). Material studied. “Maja e Jezerce Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1 syntype (NMW); “Maja e Jezercoë” / “Albania leg. Bischoff 1931”, 1 syntype (MNHUB); “Cukale Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 1 syntype (NMW); “Cucale Geb.” / “Albania leg. Bischoff 1931”, 3 syntypes (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 9 s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg. 1939”, 3 s. (MNHUB); “M. Raba Massiv” / “Albania Mueller leg. 1939”, 6 s. (MNHUB); North Albanian Alps, 1400-1800 m, 1.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS). 79.3. Omphreus ravasinii J. Müller, 1923 References. Omphreus ravasinii: Müller, 1923: 111-112 (Tomor Mt., alpine zone, loc. typ.); Winkler, 1933b: 118 (Tomor Mt.); Bousquet, 2003d: 446. Material studied. “Tomor-Abas Ali juli 1932” / “Albania leg. Bischoff 1932”, 1 f. (MNHUB). ! Omphreus ravasinii ssp. Material studied. “Korab” / “Albania leg. Bischoff 1936”, 1 f. (MNHUB). Notes. It seems this specimen belong to a vicarious form close to O. ravasinii. Unfortunately its abdomen is destroyed and further material is needed to specify its right status. 132


Tribe PANAGAEINI Bonelli, 1810 80. Genus Panagaeus Latreille, 1802 Subgenus Panagaeus s.str. 80.1. Panagaeus cruxmajor (Linnaeus, 1758) [= schaumi Ganglbauer, 1892] References. Panagaeus cruxmajor ab. schaumi: Apfelbeck, 1904: 153 (Velipoja near Lezhë). Panagaeus cruxmajor: Hieke & Wrase, 1988: 152; Baehr, 2003d: 447. Material studied. “Scutari Drin”, 3 s. (HNHM); “Oboti Bojana”, 2 s. (HNHM).

Tribe PLATYNINI Bonelli, 1810 81. Genus Agonum Bonelli, 1810 Subgenus Agonum s.str. 81.1. Agonum afrum (Duftschmid, 1812) Material studied. Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 7 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. New species for the fauna of Albania. 81.2. Agonum carbonarium Dejean, 1828 81.2.1. Agonum carbonarium carbonarium Dejean, 1828 [= alpestre (Heer, 1838)] References. Agonum alpestre: Schmidt, 1995: 168 (Korab Mt.). Notes. Bousquet (2003e: 448 sub A. alpestre) has omitted A. carbonarium for Albania. The species occurrence in the central area of the Balkans is interesting from the biogeographic point of view since the Albanian locality is distant from the main Western European range of the species, as well as from its Asian shelters. 81.3. Agonum duftschmidi J. Schmidt, 1994 [= moestum (Duftschmid, 1812)] References. Platynus viduus var. moestus: Apfelbeck, 1904: 292 (Velipoja). Platynus viduus moestus: Apfelbeck, 1907a: 495 (Liqen i Shkodrës). Platynus moestus: Apfelbeck, 1909: 297 (Vlorë). Agonum moestum: Hieke & Wrase, 1988: 86. Agonum duftschmidi: Bousquet, 2003e: 450. 81.4. Agonum extensum Ménétriés, 1849 [= binaghii Burlini, 1939] References. Agonum monachum var. binaghii: Burlini, 1939: 60 (Vlorë). Agonum extensum: Schatzmayr, 1943b: 111 (Skrofotina). Material studied. “Apfelb. Avlona”, 1 s. (HNHM); “Myzeqe Ebene” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for the country. COLEOPTERA: CARABIDAE: HARPALINAE

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81.5. Agonum gisellae Csiki, 1931 [= angustatum Dejean, 1828] References. Agonum angustatum: Hieke & Wrase, 1988: 86; Schmidt, 1994: 27 (Buenë River near Shkodër). Agonum gisellae: Bousquet, 2003e: 450. 81.6. Agonum hypocrita (Apfelbeck, 1904) References. Agonum hypocrita: Csiki, 1940: 218 (Korab Mt.). Material studied. “Scutari l. See-Ufer”, 3 s. (HNHM); “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Bushnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB). Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for the country. 81.7. Agonum longicorne Chaudoir, 1846 [= holdhausi (Apfelbeck, 1904)] References. Platynus holdhausi: Apfelbeck, 1904: 294 (Velipoja); Apfelbeck, 1907a: 495 (Buenë River). Agonum holdhausi: Hieke & Wrase, 1988: 87. Agonum longicorne: Schmidt, 1994: 32 (Buenë River near Oboti); Bousquet, 2003e: 450. Material studied. “Latif Pulaj”, 2 s. (HNHM); “Oboti Wald” / “Apfelb. Alban. spt. IV.V.’05”, 1 paralectotype of Platynus holdhausi (NMW); “Bojana Shkoder” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB). Notes. In NMW I found a paralectotype not listed by Schmidt (1994: 31) among the type material of A. longicorne, though the author included the record of this material (Oboti) in his list of species distribution. 81.8. Agonum marginatum (Linnaeus, 1758) References. Agonum marginatum: Hieke & Wrase, 1988: 85. Material studied. “Skutari Vraka” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Valona” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); Tiranë Park, 8.II.1998, shaded lake edge under leaves and wet, rotten wood and 22.II.1998, from grasses along small sluggish stream, 22 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Humelicë (between Tepelenë and Gjirokastër), 350 m, 23.IV.2003, 1 m., P. Crucitti & C. Cavalieri leg. (cAVT). Notes. Bousquet (2003e: 450) has omitted the species for Albania. Here it is confirmed for the country with first detailed records. 81.9. Agonum muelleri (Herbst, 1784) Material studied. “Skutari l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Alban. Vermosa Penther”, 3 s. (NMW). Notes. The mention of Hieke & Wrase (1988: 85) for Albania is apparently a result of following the record “Prisren” (Apfelbeck, 1904: 290 sub Platynus muelleri). However, currently this locality 134


is situated in the Kosovo Region of Serbia. Bousquet (2003e: 451) has most likely followed one of these sources, too. Since there is no previous reliable record from Albania, A. muelleri is treated here as new species for the fauna of Albania. 81.10. Agonum permoestum Puel, 1938 References. Agonum permoestum: Puel, 1938: 194 (Velipoja); Schmidt, 1994: 25 (Buenë River near Shkodër; Bushnesh; Mamurash; Murrsi-Kara Alibey; Buenë River near Oboti; Rushkull; Liqen i Shkodrës; Sukthi near Gurse); Bousquet, 2003e: 451. Material studied. “Scutari l. See-Ufer”, 1 s. (HNHM); “Skutari l. See-Ufer” / “Apfelb. Alban. spt. IV.V.’05”, 8 s. (NMW); “Oboti Bojana”, 1 s. (HNHM). 81.11. Agonum sexpunctatum (Linnaeus, 1758) References. Agonum sexpunctatum: Csiki, 1940: 218 (Korab Mt.); Hieke et al., 1986: 189 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 84. Material studied. “Alban. Vermosa Penther”, 1 s. (NMW); “Ricavac 1300 m, 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Albania Tirana” / “3.VI.1958 leg. Xh. Murraj”, 3 s. (HNHM). Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for the country. 81.12. Agonum sordidum Dejean, 1828 81.12.1. Agonum sordidum sordidum Dejean, 1828 References. Platynus sordidus: Apfelbeck, 1904: 290 (Velipoja; Vlorë); Apfelbeck, 1909: 297 (Vlorë). Agonum sordidum: Schatzmayr, 1943b: 111 (Stan-Karbunarë); Hieke et al., 1986: 189 (Poliçan near Tomor Mt.); Hieke & Wrase, 1988: 85. Material studied. “Elbasan, Alb. IX… Dr. H. Priesner”, 1 s. (NMW); “Liquen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB). Notes. Bousquet (2003e: 451) has omitted the species for Albania. Here it is confirmed for the country. 81.13. Agonum viridicupreum (Goeze, 1777) 80.13.1. Agonum viridicupreum viridicupreum (Goeze, 1777) References. Platynus viridicupreus: Apfelbeck, 1904: 290 (Velipoja). Agonum viridicupreum: Hieke et al., 1986: 189 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 85. Material studied. “Murrsi-Kara Alibey” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Liquen i Butrintes Vivari” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB); “Han-Hot” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); “Cafa Liqenit Iballe” / “Albania Mueller leg. 1939”, 5 s. (MNHUB); Tiranë Park, 11.I.1998 / 1.III.1998 / 8.III.1998, 4 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). COLEOPTERA: CARABIDAE: HARPALINAE

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Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for the country. Subgenus Europhilus Chaudoir, 1859 81.14. Agonum antennarium (Duftschmid, 1812) References. Platynus antennarius: Apfelbeck, 1904: 295 (Velipoja). Agonum antennarium: Hieke & Wrase, 1988: 87. Material studied. “Mustajbeg Velipoja Albania”, 2 s. (MNHUB); “Maja Rogamit” / “Albania leg. Bischoff 1935”, 11 s. (MNHUB); Tiranë Park, 22.II.1998 / 1.III.1998 / 8.III.1998, 8 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. Bousquet (2003e: 452) has omitted the species for Albania. Here it is confirmed for the country. 81.15. Agonum thoreyi Dejean, 1828 81.15.1. Agonum thoreyi thoreyi Dejean, 1828 References. Platynus thoreyi: Apfelbeck, 1904: 296 (Velipoja). Agonum thoreyi: Hieke & Wrase, 1988: 87; Bousquet, 2003e: 453.

82. Genus Anchomenus Bonelli, 1810 Subgenus Anchomenus s.str. 82.1. Anchomenus dorsalis (Pontoppidan, 1763) References. Platynus dorsalis: Apfelbeck, 1904 (Vlorë; Berat; Velipoja); Apfelbeck, 1909: 297 (Vlorë; Berat); Hieke & Wrase, 1988: 83. Platynus dorsale: Schatzmayr, 1943b: 111 (Lushnje); Hieke et al., 1986: 189 (North Albanian Alps near Theth). Material studied. “Albanien Avlona v. Oertzen”, 3 s. (MNHUB); “Alban. Vermos Penther”, 3 s. (NMW); “Skutari Kolcekai” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Bushek bei Elbasan” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bushek juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Lum i Tiranes” / “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Liquen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 12 s. (MNHUB); “Murrsi-Kara Ali Bey” / “leg. Bischoff Albania 1934”, 5 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Am Wege von Murthi nach Konispol” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB); “Cafa e Ceromis Iballe” / “Albania Mueller leg. 1939”, 2 s. (MNHUB); “Albanien Tirana Dajti-Vorberge 3.V.39…Mueller”, 1 s. (MNHUB); “Albania Cerrik” / “18.IV.1958 leg. Xh. Murraj”, 5 s. (HNHM); Sauk, 14.III.1961, 1 f. (MNST); Rhesen District, Merkurth, 11.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Ljukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS). Additional material included (studied S. Hristovski and A. Vigna Taglianti). Qesarat, near Memaliaj (Tepelenë District), valley of Vijosë River, 300-360 m, 18.IV.2003, 1 m., 1 f., P. 136


Crucitti & C. Cavalieri leg. (cAVT); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). Notes. Bousquet (2003e: 455) has omitted the species for Albania. Here it is confirmed for the country.

83. Genus Olisthopus Dejean, 1828 83.1. Olisthopus fuscatus Dejean, 1828 References. Olisthopus fuscatus: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88. Material studied. “Alban. Vermosa Penther”, 2 s. (NMW); “Ibe Erzen” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Piqeros” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda District, Lyukove, 5.V.1994, 1 s., P. Stoev leg. (NMNHS); Saranda District, Lyukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). Notes. Bousquet (2003e: 461) has omitted the species for Albania. Here it is confirmed for the country. 83.2. Olisthopus glabricollis (Germar, 1817) References. Olisthopus glabricollis: Schatzmayr, 1943b: 111 (Lushnje); Hieke & Wrase, 1988: 88. Material studied. “Alban. Exp. Kula Ljums 18-28.V.’18”, 4 s. (NMW); “Alban. Exp.’18. Kruma 5.VI.”, 4 s. (NMW); “Burelle” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB). Notes. Bousquet (2003e: 461) has omitted the species for Albania. Here it is confirmed for the country. ? Olisthopus rotundatus (Paykull, 1790) ? Olisthopus rotundatus rotundatus (Paykull, 1790) References. Olisthopus rotundatus: Horion, 1941: 310; Hieke & Wrase, 1988: 88. Olisthopus rotundatus rotundatus: Bousquet, 2003e: 461. Notes. Horion (1941: 310) and Hieke & Wrase (1988) noted this species for Albania apparently following the record “Prisren” (Apfelbeck, 1904: 287). Actually this locality is situated in the Kosovo Region of Serbia. Most probably, Bousquet (2003e: 461) has followed the above sources. For the time being P. rotundatus is regarded as questionable (possible but yet not proven) for Albania.

84. Genus Paranchus Lindroth, 1974 Notes. New genus for the fauna of Albania. 84.1. Paranchus albipes (Fabricius, 1796) Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW). Notes. New species for the fauna of Albania.


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85. Genus Platynus Bonelli, 1810 Subgenus Batenus Motschulsky, 1865 85.1. Platynus scrobiculatus (Fabricius, 1801) 85.1.1. Platynus scrobiculatus turcicus Apfelbeck, 1904 References. Platynus scrobiculatus turcicus: Schmidt, 1996: 347. Material studied. “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze Penther’14”, 5 s. (NMW); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 26 s. (MNHUB). Notes. Apfelbeck (1904: 288) described P. scrobiculatus turcicus from Bosnia Herzegovina. In the same year, Csiki (1904) described P. serbicus Csiki, 1904 from Southeast Serbia. Recently, Bousquet (2003e: 463) treated both taxa as different species. However, judging by the description of these taxa and based on research on different populations from the Eastern Balkans (Guéorguiev & Muilwijk, 2001), I consider the diagnostic characters indicated by Apfelbeck and Csiki reflect rather geographical variations than being of specific importance. On account of that, P. turcicus is treated here as conspecific with P. scrobiculatus (Fabricius, 1801): Platynus (Batenus) scrobiculatus turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus Apfelbeck, 1904. Recently, we also treated Csiki’s taxon as subspecies of P. scrobiculatus, omitting formal change in rank (Guéorguiev & Muilwijk, 2001). Here this act is introduced: Platynus (Batenus) scrobiculatus serbicus Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904. The Albanian populations are provisionally assigned to the ssp. turcicus. Bousquet (2003e: 463 sub Platynus turcicus) has omitted the species for Albania. Here it is confirmed for the country with first detailed records. Subgenus Platynus s.str. 85.2. Platynus assimilis (Paykull, 1790) References. Platynus assimilis: Apfelbeck, 1904: 296 (Vlorë; Berat; Velipoja); Hieke & Wrase, 1988: 84. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 19 s. (NMW); “Bushnesh” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1935”, 5 s. (MNHUB); “Pylla Thumana” / “Albania leg. Bischoff 1935”, 10 s. (MNHUB). Notes. Bousquet (2003e: 464) has omitted the species for Albania. Here it is confirmed for the country.

Tribe PTEROSTICHINI Bonelli, 1810 86. Genus Abax Bonelli, 1810 Subgenus Abacopercus Ganglbauer, 1891 86.1. Abax carinatus (Duftschmid, 1812) 86.1.1. Abax carinatus carinatus (Duftschmid, 1812) [= latus (Dejean, 1828)] References. Abax carinatus var. latus: Apfelbeck, 1904: 216 (Velipoja). Abax carinatus: Hieke & Wrase, 1988: 79. Abax carinatus carinatus: Bousquet, 2003f: 469. Material studied. “Velipoja Albania”, 1 s. (HNHM); 18 s. from different localities (MNHUB). 138


Subgenus Abax s.str. 86.2. Abax ovalis (Duftschmid, 1812) References. Abax ovalis: Hieke & Wrase, 1988: 79; Bousquet, 2003f: 470. Material studied. “Koritnik” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB). Notes. First detailed records for Albania.

87. Genus Molops Bonelli, 1810 Notes. The species recognition in this group stay difficult due to the great external resemblance of the adults that often resulted in impossibility to distinguish the females of closely related species. Besides, a great similarity and variations in the shape of the aedeagus in close species occur. The current systematics of the genus must not be treated as definitive (Mlynář, 1977). Except the “spartanus” group, all the other groups of species of Molops s.l. display North Aegean origin. The type material used by Müller (1917; 1918a) to describe five Albanian taxa of Molops s.str. is still preserved in NMW; the same concerns the type material published by Apfelbeck (1918c). Subgenus Molops s.str. Species group “robustus” 87.1. Molops malshentianus Apfelbeck, 1917 [= maderi J. Müller, 1933] References. Molops malshetianus: Apfelbeck, 1917: 224 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Apfelbeck, 1918b: 88-89 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Apfelbeck, 1930: 167-168 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Mlynář, 1977: 144 (Mal i Shenjt Mt. near Oroshi). Molops maderi: Müller, 1933: 217 (Kruja Mt., 600 m, loc. typ.); Mařan, 1940c: 192 (Kruja Mt.); Mařan, 1945: 60 (Kruja Mt.); Mlynář, 1977: 39-40 (Kruja Mt.); Turin, 1981: 130; Bousquet, 2003f: 473. Material studied. “Kruja, Alban., Mader”, 1 s. (NMW); “Ibe-Erzen” / “Albania leg. Bischoff 1932”, 1 f. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 f. (MNHUB); “Mal i Krujes” / “leg. Bischoff Albania 1934”, 3 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 f., L. Penev leg. (NMNHS). Notes. Apfelbeck published three consecutive descriptions for M. malshentianus (Apfelbeck, 1917; Apfelbeck, 1918b; Apfelbeck, 1930), the second one has been accredited to be the valid. Actually this is incorrect and the original description must be assigned to the oldest one. According to Bousquet (2003f: 473) M. malshentianus is synonym of M. maderi, but since the former has been described earlier, M. malshentianus must be treated as senior synonym: Molops (Molops) maderi syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917. Notwithstanding, revision of the types is needed to elaborate this association and to designate lectotypes. The record from “Maja e Tartarit” should be treated with caution since it is based on a single female specimen. COLEOPTERA: CARABIDAE: HARPALINAE

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# Molops weiratheri J. Müller, 1930 [= alinae J. Müller, 1932] References. Molops parnassicola alinae: Müller, 1932: 220 ((“Südalbanien (Epiros)”); Mařan, 1940c: 190, 192. Notes. Mařan (1940c) specified that the Müller’s record for Albania (Müller, 1932) refers to Greece. M. weiratheri inhabits the mountains in South Macedonia and North Greece reaching in the south the massif of Olympus Mt. (Mlynář, 1977: 41-42). Species group “spartanus” 87.2. Molops spartanus (Schaum, 1862) ? Molops spartanus parnassicola Kraatz, 1875 References. Molops cf. parnassicola: Müller, 1936: 105 (Gjirokastër). Notes. As Müller (1936) has determinated his material only to species level, the subspecific treatment of the population near Gjirokastër needs further specification. 87.2.1. Molops spartanus valonensis J. Müller, 1936 References. Molops spec.? parnassicola: Ganglbauer, 1889: 54 (Vlorë). Molops parnassicola: Apfelbeck, 1904: 217, 220 (Vlorë); Apfelbeck, 1909: 297 (Kanina near Vlorë; Hagion Elias = Keraun); Müller, 1918a: 7 (Vlorë). Molops parnassicola valonensis: Müller, 1936: 105 (Reza e Kanalit Mt. near Llogora, loc. typ.; Maja e Tartarit Mt.; Shëndelli Mt.); Mařan, 1940c: 190, 192 (Llogora); Mařan, 1945: 59 (Vlorë). Molops spartanus valonensis: Mlynář, 1977: 47-48 (Reza e Kanalit Mt. near Llogora; Llogora; Maja ë Qori; Cafa ë Qori); Bousquet, 2003f: 475. Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Llogora, Albania mer.”, 4 s. (MNHUB); “Maj’e Tartarit Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Reza e Kanalit Logara, Albania m.” / “Winkler, Lona, Bischoff, V.1931”, 5 s. (NMW); “Logara” / “Albania leg. Bischoff 1932”, 158 s. (MNHUB); “Maja e Qori juni 1932” / “Albania leg. Bischoff 1932”, 10 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 36 s. (MNHUB); “Mato-Hasanaj juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen-Ili juni” / “Albania leg. Bischoff 1932”, 35 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 48 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 30 s. (MNHUB); Ibë, 24.IV.1962, 1 m. (MNST). Notes. Before the description of this subspecies, all records from South Albania had been attributed to M. spartanus parnassicola Kraatz, 1875. Bousquet (2003f: 475) indicated an incorrect page of the original description of ssp. valonensis. Species group “simplex” 87.3. Molops simplex Chaudoir, 1868 # Molops simplex simplex Chaudoir, 1868 References. Molops simplex simplex: Bousquet, 2003f: 475. Notes. Mlynář (1977: 54) noted that the older data for this subspecies from Albania concerns M. simplex meridionalis, because the nominotypical subspecies inhabits regions removed from the borders of the country.

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87.3.1. Molops simplex meridionalis Mlynář, 1977 References. Molops simplex: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Müller, 1918a: 7 (Maranajt Mt.; Rapshë). Molops simplex meridionalis: Mlynář, 1977: 55; Bousquet, 2003f: 474. Notes: The subspecies occurs only in the marginal northwestern part of the country. Species group “alpestris” 87.4. Molops alpestris (Dejean, 1828) 87.4.1. Molops alpestris imitator J. Müller, 1917 References. Molops alpestris: Apfelbeck, 1907a: 497-498 (Maranajt Mt.: top region, 1500-1576 m); Hieke & Wrase, 1988: 81. Molops alpestris imitator: Müller, 1917: 347-348 (Rapshë, loc. typ.); Müller, 1918a: 11 (North Albanian Alps: Rapshë); Mlynář, 1977: 80-81 (Rapshë); Hieke, 1981: 81. Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 3 syntypes (NMW); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 8 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS). Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here, it is confirmed for the country. Species group “elatus” 87.5. Molops parreyssi Kraatz, 1875 87.5.1. Molops parreyssi lonae J. Müller, 1921 References. Molops parreyssi: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Klaptocz, 1910: 399 (Theth); Müller, 1918a: 7 (Maranajt Mt.). Molops parreyssi lonae: Müller, 1921b: 28-30 (Çukall Mt., loc. typ.); Mlynář, 1977: 96 (Çukall Mt.: cave “Kunor Vukai”); Bousquet, 2003f: 473. Material studied. “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Cucale Albania sept.” / “leg. Bischoff 1931”, 4 s. (NMW); nearly 100 s. from locality “Cucale” (MNHUB); 4 s. from locality “Maranay” (MNHUB). Notes. The species occurs only in the marginal northwestern parts of Albania. Species insertae sedis 87.6. Molops merditanus Apfelbeck, 1906 87.6.1. Molops merditanus merditanus Apfelbeck, 1906 References. Molops merditanus: Apfelbeck, 1906: 1664 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.; Merdita: Zepë Mt. near Fandi, loc. typ.); Apfelbeck, 1907a: 499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 10 (Merdita: Mal i Shenjt Mt. near Oroshi; Merdita: Zepë Mt. near Fandi); Müller, 1930 (Merdita). Molops merditanus merditanus: Mlynář, 1977: 106-107 (Mal i Shenjt Mt. near Oroshi; Zepë Mt.); Bousquet, 2003f: 473. COLEOPTERA: CARABIDAE: HARPALINAE

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Material studied. “Merdita M. Scheit”, 1 paralectotype (DEI); “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05” / “Molops merditanus typ. † Apfelb.”, 1 f. (NMW); “Merdita Zebia”, 2 paralectotypes (DEI); “Merdita Zebia” / “Molops merditanus typ. ‡ Apfelb.”, 1 f. (NMW). Notes. Mlynář (1977: 106) noted that the lectotype of M. merditanus is in the collection of HNHM, but I could not find it there. 87.6.2. Molops merditanus intermedius J. Müller, 1917 References. Molops obtusangulus intermedius: Müller, 1917: 348-349 (Montenegro: Podgorica, loc. typ.; Albania / Montenegro border: Ricavac, loc. typ; Albania: Vermosh, loc. typ). Molops merditanus intermedius: Mlynář, 1977: 107-108 (Ricavac; Vermosh); Drovenik & Peks, 1994: 71 (Ricavac); Drovenik & Peks, 1999: 83 (Ricavac); Bousquet, 2003f: 473. Material studied. “Ricavac juni” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW); “Vermosa 10-21.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype (NMW). Notes. Each of the three localities mentioned in the original description (Müller, 1917) could be a potential “locus typicus”, however a lectotype designation still has to be done. Species group “curtulus” Notes. This taxonomically difficult species complex is treated here differently than previously (Mlynář, 1977): Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri reiseri Apfelbeck, 1904 Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri corallipes J. Müller, 1936 Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri pseudoalpestris Mlynář, 1977 Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri victoris J. Müller, 1918 Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Müller, 1917 87.7. Molops albanicus Apfelbeck, 1904 References. Molops albanicus: Apfelbeck, 1906: 1665 (Mal i Shenjt Mt.; Zepë Mt.); Apfelbeck, 1907a: 499, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 8 (Merdita: Mal i Shenjt Mt.; Merdita: Zepë Mt.); Müller, 1930: 124 (Merdita: Zepë Mt.); Mlynář, 1977: 118, 120 (Mal i Shenjt Mt.; Zepë Mt.; Oroshi); Bousquet, 2003f: 472. Material studied. “Merdita Zebia”, 9 s. (DEI; HNHM; NMW); “Merdita M. Scheit” / “Coll. Apfelbeck”, 1 m. (HNHM); “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 21 s. (DEI; NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 3 s. (NMW); “Alban. Exped. Pashtrik’18 4-17.7”, 1 s. (NMW); 9 s. from locality “Merdita” (MNHUB); 33 s. from locality “Llogora” (MNHUB); 11 s, from locality “Mal i Shenjit” (MNHUB); 3 s. from locality “Iballe” (MNHUB); 1 s. from locality “Mal i Gjer” (MNHUB); 3 s. from locality “Shen-Ili” (MNHUB). Additional material included (studied A. Vigna Taglianti). Valbona, Hellit Kuq Mt., south slope, 930 m, 6.VII.1993, 1 m., L. Latella leg. (cAVT). 142


Notes. Apfelbeck (1904: 222) described this species basing on a single male from the type locality “Prizren”. This place is situated in the Kosovo Region of Serbia. The binomen Molops transfuga Apfelbeck (Müller, 1930: 124; Mlynář, 1977: 120) is nomen nudum. 87.8. Molops apfelbecki Ganglbauer, 1891 Notes. In the past, M. apfelbecki and M. reiseri have been treated as separate species. Probable cause for this state is that Apfelbeck (1904: 223), while describing M. reiseri, missed to compare it with M. apfelbecki. Later, Mlynář (1977) similarly did not compare the aedeagus of both species, but noticed the resemblance in their female genital structures. The present author has studied the aedeagus of M. reiseri holdhausi (syntype, NMW), M. reiseri (lectotype, HNHM), M. reiseri victoris (syntype, NMW; specimens from Koprivnik Mt. and Shkëlzen Mt., HNHM) and M. apfelbecki (lectotype, HNHM) and found that M. reiseri holdhausi is different in the shape of the median lobe from the other three taxa. In fact, the last taxa have identical aedeagus. Therefore, M. reiseri reiseri and M. apfelbecki apfelbecki are considered as geographical races of single species, and M. holdhausi – as valid species which occurs together with M. apfelbecki s.l. 87.8.1. Molops apfelbecki victoris J. Müller, 1918 References. Molops victoris: Müller, 1918a: 1-3, 6 (Albania / Montenegro border: North Albanian Alps, 1300-2000 m, loc. typ.). Molops reiseri victoris: Mlynář, 1977: 114-115 (North Albanian Alps; Shkëlzen Mt.). Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” / “victoris n. det. J. Müller”, 2 m. syntypes of Molops victoris (NMW); “Albania Skölsen” / “Akad. Balk. Exp. Csiki 1917”, 1 m. with extracted penis (HNHM); “Maja Peis” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 5 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 22 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Bischoff, Mueller 1939” / “Type” / “Apfelbecki sbsp. Bischoffi m. det. Müller”, 1 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Bischoff, Mueller 1939” / “Type”, 5 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Bischoff, Mueller 1939” / “M. Apfelbecki sbsp. Bischoffi m. det. G. Müller 1940”, 1 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Bischoff, Mueller 1939”, 5 s. (MNHUB). Additional material studied. Bosnia Herzegovina / Montenegro. “Apfelbeck Volujak” / “Coll. Apfelbeck”, 1 m. lectotype and 3 paralectotypes of Molops apfelbecki (HNHM). Montenegro. “Vunšaj, Cafa glava, Alban. Penther” / “reiseri Apf.”, 1 m. of Molops apfelbecki victoris (NMW). Serbia (Kosovo Region). “Montes Koprivnik” / “Akad. Balk. Exp. Csiki 1917”, 1 m. of Molops apfelbecki victoris (HNHM). Serbia. “Reiser 1898 Kopaonik” / “Coll. Apfelbeck”, 1 m. lectotypus and 2 paralectotypes of Molops reiseri (HNHM); “Reiser 1899 Kopaonik” / “reiseri Apf. B& typus” / “type”, 1 m. of Molops apfelbecki reiseri (NMW); “Reiser 1899 Kopaonik” / “reiseri Apf. Serb. turk. Grence” / “type”, 1 m., 1 f. of Molops apfelbecki reiseri (NMW); “Reisser 1899 Kopaonik”, 1 m., 1 f. of Molops apfelbecki reiseri (MNHUB). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. Müller (1930: 126) revised the material of Apfelbeck (1918c: 166 sub M. reiseri from Vunšaj, Cafa glava) and determinated it as M. victoris. However, Bousquet (2003f: 474) has omitted M. reiseri victoris (currently M. apfelbecki victoris, see above) for Albania in spite of the fact that Mlynář (1977: 115) gave a detailed record. Here the subspecies is confirmed for the country. The trinomen M. apfelbecki bischoffi G. Müller 1940, found on labels of two specimens, is nomen nudum. COLEOPTERA: CARABIDAE: HARPALINAE

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87.9. Molops curtulus Ganglbauer, 1891 87.9.1. Molops curtulus cakorensis Mlynář, 1977 References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps). Molops curtulus: Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Müller, 1918a: 10 (Maranajt Mt.; North Albanian Alps: Rapshë, Grebeni, Vermosh, Ricavac); Csiki, 1940: 217 (Shkëlzen Mt.). Molops curtulus cakorensis: Mlynář, 1977: 111. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa Penther”, 1 s. (NMW); “13.15.VI.14 Vermosa ca 1000 m”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 8 s. (NMW); “Grebeni 16.VI.” / “Alban. montenegr. Grenze, Penther’14”, 5 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Rogamit” / “Albania leg. Bischoff 1935”, 3 f. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 9 s. (MNHUB); “Boge Aufstieg z. Rab” / “Albania leg. Bischoff, Mueller 1939”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS). Notes. Bousquet (2003f: 472) has omitted the species for Albania. Here it is confirmed for the country. 87.9.2. Molops curtulus winklerianus J. Müller, 1936 References. Molops winklerianus: Müller, 1936: 103 (Tomor Mt.: Kulmak, loc. typ.; Shëndelli Mt. near Tepelenë; Llogora); Molops curtulus winklerianus: Mlynář, 1977: 111 (Tomor Mt.: Kulmak; Llogora); Bousquet, 2003f: 472. Material studied. “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW). Notes. Mlynář (1977: 111) stated in a footnote that the populations of M. curtulus from Central Albania may belong to a separate race, so further investigations of material from this region of the country will reveal more details for elucidation that statement. 87.10. Molops holdhausi J. Müller, 1917 References. Molops holdhausi: Müller, 1917: 352-353 (Albania / Montenegro border: North Albanian Alps, 1300-2000 m, loc. typ.); Müller, 1918a: 6 (North Albanian Alps); Müller, 1930: 126 (North Albanian Alps, Grebeni near Vermosh). Molops reiseri: Apfelbeck, 1918c: 166 (North Albanian Alps, Grebeni near Vermosh). Molops reiseri holdhausi: Mlynář, 1977: 112, 114 (North Albanian Alps). Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14” / “holdhausi n. det. J. Müller”, 1 m. syntype of Molops holdhausi (NMW); “Grebeni, Vermoša Alban., Penther 16 Juni, am Schnee” / “reiseri Apf.”, 2 m. (NMW). Additional material studied. Montenegro. “Durmitor”, 4 s. (HNHM); “Montenegro Durmitor, Penther’04”, 1 m., 1 f. (NMW); “Jugoslavia mer. Prokletija Gusinje Matìjka VI.1937”, 1 m., 1 f. (MNHUB); “Jugoslavia mer. Prokletja Gusinje Kodym 20.VI.1937”, 2 s. (HNHM). Notes. Müller described M. holdhausi and M. victoris as separate species (Müller 1917; Müller 1918a). Mlynář (1977: 112-113) neglected the fact that M. reiseri holdhausi and M. reiseri victoris occur sympatrically and expressed doubt about the status of both taxa due to the lack of enough specimens at hand (Mlynář (1977: 115). The present author compared the penis structures of several males, previously determined as ssp. holdhausi or ssp. victoris, and ascertained 144


the first is good species - Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Müller, 1917. Bousquet (2003f: 474) has omitted M. holdhausi for Albania though Müller (1930) gave a detailed record. Here the species is confirmed for the country. 87.11. Molops plurisetosus J. Müller, 1918 [= malissorum Apfelbeck, 1918] References. Molops apfelbecki plurisetosus: Müller, 1918a: 3-4, 11 (Albania / Montenegro border: North Albanian Alps, 1200-2000 m, loc. typ.; Albania / Montenegro border: Ricavac, loc. typ.; Albania: Vermosh, loc. typ.). Molops malissorum: Apfelbeck, 1918c: 160, 167-168 (Montenegro: North Albanian Alps, Vusanje, loc. typ.; Albania: pastures over Theth). Molops plurisetosus: Mlynář, 1977: 120-121 (North Albanian Alps; Ricavac; Vermosh). Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4 syntypes of Molops plurisetosus (NMW); “Ricavac Juni” / “Alban. montenegr. Grenze, Penther’14”, 1 syntype of Molops plurisetosus (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 syntypes of Molops plurisetosus (NMW); North Albanian Alps, Theth, 800-1200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 3 m., 2 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, cave “No 25”, 23.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Shkodër District, Kozuja, 1800 m, 27.VIII.1996, 1 m., 1 f., T. Ivanova leg. (NMNHS). Additional material studied. Montenegro. “Vunšaj, Prokletija-Geb., Albania Dr. Penther” / “Molops malissorum n. sp. Apf.”, 2 syntypes of Molops malissorum (NMW). Notes. Bousquet (2003f: 474) has omitted the species for Albania. Here it is confirmed for the country. Currently, the exact publishing dates of the descriptions of M. plurisetosus and M. malissorum were not found, so the problem with the priority of one of the two names is pending. Each of the different localities mentioned in the original descriptions of M. plurisetosus (Müller, 1918a) could be a potential “locus typicus”, however lectotype designation still has to be done. Species group “rufipes” 87.12. Molops osmanilis Apfelbeck, 1904 [Molops osmanilis osmanilis Apfelbeck, 1904] Notes. The record from “Prisren” (Apfelbeck, 1904: 229) refers to the Kosovo Region of Serbia, so perhaps Bousquet (2003f: 473) has followed this source when listing M. osmanilis osmanilis for Albania. The nominotypical subspecies is treated here as likely to be found in the country. 87.12.1. Molops osmanilis tomoricensis J. Müller, 1923 References. Molops tomoricensis: Müller, 1923: 109-110 (Tomor Mt., alpine zone, loc. typ.). Molops osmanilis tomoricensis: Mlynář, 1977: 123 (Tomor Mt.: Tomor, Abbas Ali); Bousquet, 2003f: 473. Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 5 s. (NMW; MNHUB); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Kulmak Albania mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 3 (NMW); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 7 s. (NMW); more than 90 s. from Tomor Mt. (MNHUB); “Mal i Dajtit Tirane” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara COLEOPTERA: CARABIDAE: HARPALINAE

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juni” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Mal i Dajtit Tirane” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 13 s. (MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 2 s. (MNHUB); “AL (Tirana) 8 km N psene E Tirana 1450 m. 30.IV.1993 leg. H. & S. Peks”, 2 s. (MNHUB). Notes. All the specimens examined from Albania are put to this race more or less provisionally. However, for instance specimens from Southwest Albania (localities “Logara” and “Maja e Dhembelit”) have a shape of the pronotum like spp. osmanilis, whereas these from Central Albania (localities “Mal i Dajtit” and “Mal i Krujes”) – like ssp. tomoricensis. 87.13. Molops rufipes Chaudoir, 1843 87.13.1. Molops rufipes rufipes Chaudoir, 1843 References. Molops rufipes: Rambousek, 1912: 93; Mlynář, 1977: 124; Hieke, 1981: 81; Hieke & Wrase, 1988: 82. Molops rufipes rufipes: Mlynář, 1977: 124 (Mal i Dajtit Mt.). Material studied. “Mal i Daitit b. Tirana, Alb. C.” / “leg. Bischoff 1931 coll. A. Winkler”, 2 s. (NMW); “Tomor: Tomor Alban. mer.”, 1 m. (MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 m. (MNHUB); more than 50 s. from locality “Iballe” (MNHUB); more than 50 s. from locality “Mal i Dajtit” (MNHUB); Tiranë, 26.VII.1965, 1 f. (MNST); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 4 m., 3 f., L. Penev leg. (NMNHS). Notes. Bousquet (2003f: 474) has omitted the subspecies for Albania. Here it is confirmed for the country. The material of Rambousek (1912) was not revised and therefore it is placed in the nominotypical race provisionally. # Molops rufipes klisuranus Apfelbeck, 1902 References. Molops klisuranus: Apfelbeck, 1918c: 166 (North Albanian Alps). Notes. Two wrong records (Müller, 1930: 125-126, revised as M. curtulus s.l.) for this taxon have been recorded from the Western Balkan (South Serbia, Sandzak: Raška near Novi Pazar, Apfelbeck, 1904: 222; North Albanian Alps, Apfelbeck, 1918c: 166). Therefore the taxon is removed from the list of the Albanian Carabidae. 87.13.2. Molops rufipes munelensis Apfelbeck, 1906 References. Molops sturanyi munelensis: Apfelbeck, 1906: 1665-1666 (Munella Mt., loc. typ.); Müller, 1918a: 9 (Merdita: Munella Mt.). Molops rufipes munelensis: Mlynář, 1977: 128, 130 (Munella Mt.; Munella Mt.: Oroshi; Berishe); Bousquet, 2003f: 474. Material studied. “Orosi”, 5 s. (HNHM); “Oroši (Ptr.) Alban. spt.” / “Munela” / “Coll. Apfelbeck”, 1 lectotype (HNHM); “Merdita Munela”, 3 paralectotypes (HNHM); “Merdita Munela”, 9 s. (DEI; MNHUB); “Merdita Munela Apfb.”, 4 s. (MNHUB); “Merdita” / “Apfb.”, 2 s. (DEI); “Merdita Munela” / “Wgth. 1906 Merdita”, 21 s. (NMW); “Merdita Zebia”, 9 s. (NMW); “Alban. Exp.’18 Pashtrik, 29.V.-4.VI.”, 9 s. (NMW); “Alban. Exp.’18 Pashtrik, 4-17.7”, 5 s. (NMW); “M. Mulecifit” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Berishe” / “Albania leg. Mueller 1939”, 6 s. (MNHUB); “AL (Shkodar). Paß SE Puke 850 m 2.V.1993” / “leg. H. & S. Peks”, 1 m. (MNHUB). [Molops rufipes pseudoosmanilis Mlynář, 1977] Notes. Since this subspecies was hitherto recorded only from the Macedonian section of the Korabi Mt., it is expected to live in Albania, too. 146


87.13.3. Molops rufipes steindachneri Apfelbeck, 1908 References. Molops rufipes steindachneri: Mlynář, 1977: 128 (Korab Mt.). Notes. Bousquet (2003f: 474) has omitted this subspecies for Albania. Actually its occurrence in the country needs confirmation. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), the oldest one is correctly treated as valid. 87.13.4. Molops rufipes sturanyi Apfelbeck, 1906 References. Molops sturanyi: Apfelbeck, 1906: 1665 (Merdita: Mal i Shenjt Mt., loc. typ.; Merdita: Munella, loc. typ.; Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 497, 499, 503 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Müller, 1918a: 9 (Merdita: Zepë Mt.; Merdita: Mal i Shenjt Mt.; Maranajt Mt.); Müller, 1930: 123 (Gjalicë Mt.). Molops steindachneri: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Molops rufipes sturanyi: Mlynář, 1977: 128 (Mal i Shenjt Mt.; Zepë Mt.; Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 474. Material studied. “Albania” / “Merdita”, 2 s. (DEI); “Merdita M. Schèit” / “Apfelb. Alban. spt. IV.V.’05”, 1 lectotype (HNHM); “Merdita Zebia”, 4 paralectotypes (HNHM); “Merdita Zebia”, 6 s. (DEI; MNHUB); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 11 s. (HNHM); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 49 s. (NMW); 15 s. from locality “Bjeshka Matroxh” (MNHUB); 20 s. from localaity “Derzë” (MNHUB); more than 300 s. from locality “Gjallica” (MNHUB); more than 500 s. from locality “Korab” (MNHUB); 2 s. from locality “Taja” (MNHUB). Notes. The record from the Maranajt Mt. (Müller, 1918a: 9) has to be treated cautiously because it is the only record of this species north of the Drin River. Species group “piceus” 87.14. Molops piceus (Panzer, 1793) 87.14.1. Molops piceus balcanicus Mlynář, 1977 References. Molops austriacus: Müller, 1918a: 9 (Rapshë; North Albanian Alps); Horion, 1941: 302. Molops piceus balcanicus: Mlynář, 1977: 136; Hieke, 1981: 81. Molops piceus: Hieke & Wrase, 1988: 82. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB). Notes. Bousquet (2003f: 473) has omitted the species for Albania. Here it is confirmed for the country. Subgenus Stenochoromus L. Miller, 1866 87.15. Molops montenegrinus (L. Miller, 1866) 87.15.1. Molops montenegrinus cavernicola J. Müller, 1916 References. Molops montenegrinus cavernicola: Müller, 1916: 81 (Çukall Mt.: cave “Sckamit”, loc. typ.); Müller, 1918a: 11-12 (Çukall Mt.: cave “Sckamit”; North Albanian Alps: Ricavac, Vermosh); Bousquet, 2003f: 475. COLEOPTERA: CARABIDAE: HARPALINAE

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Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, more than 20 s. (NMW); “Kunor Vukaj Zukali Albanien Lona Giugno 1914”, 1 s. (DEI); “Starka Alb. sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 6 s. (NMW); “Maja e Jezerce Albania sept.” / “leg. Bischoff 1931 coll. A. Winkler”, 2 s. (NMNHS); “Maja e Jezercoë” / “Albania leg. Bischoff 1933”, 9 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 27 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 95 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 3 s. (MNHUB); “Boge Aufstieg Z. Rab” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 7 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 2 m., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes, 1800-2200 m, 28.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Mali-i-Thatë Mt., cave “Shpella e Haxise, 8.VII.1993, rests of 1 s., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 1 m., 2 f., L. Latella leg. (cAVT). Notes. All the data from Albania are placed provisionally to the ssp. cavernicola, since no intraspecific revision of M. montenegrinus s.l. is made. In addition to the material studied above, in NMW I have seen more specimens, which remained unregistered.

88. Genus Myas Sturm, 1826 Subgenus Myas s.str. 88.1. Myas chalybaeus (Palliardi, 1825) [= rugosicollis Brullé, 1832] References. Myas chalybaeus: Tschitschérine, 1899: 70; Anonymous, 1907: 284; Hieke & Wrase, 1988: 69; Bousquet, 2003f: 475. Myas rugosicollis: Tschitschérine, 1899: 70. Material studied. “Alban. Exp.’18. Kruma 5.VI.”, 1 s. (NMW); “Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit Tiranë” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 7 s. (MNHUB); “Valbona Çerem” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Berishe” / “Albania Mueller leg. 1939”, 3 s. (MNHUB); “Boge Ober D. Quelle” / “Albania Mueller leg. 1939”, 3 s. (MNHUB); “Çafa e Ceromis Iballe” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); 5 km Leskoec, 2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Stebleva, cave “Shpella e Hadzhi i Malit”, 5-8.V.2006, 1 s., A. Zhalov, D. Raychev & J. Tsvetanov leg. (NMNHS). Notes. First detailed records for Albania.

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89. Genus Pedius Motschulsky, 1850 89.1. Pedius inquinatus (Sturm, 1824) References. Pterostichus inquinatus: Apfelbeck, 1904: 257 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1930: 198; Horion, 1941: 279; Schatzmayr, 1942: 65; Schatzmayr, 1943b: 110 (Mal i Krujes); Hieke & Wrase, 1988: 71. Pedius inquinatus: Bousquet, 2003f: 478. Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Lum i Tiranes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 2231.X.1997, 3 s., R. Ward leg. (CMNH). 89.2. Pedius longicollis (Duftschmid, 1812) References. Pedius longicollis: Bousquet, 2003f: 478.

90. Genus Poecilus Bonelli, 1810 Subgenus Ancholeus Dejean, 1828 90.1. Poecilus puncticollis (Dejean, 1828) References. Pterostichus puncticollis: Schatzmayr, 1943b: 110 (Kruja; Skrofotina; Stan-Karbunarë). Poecilus puncticollis: Hieke & Wrase, 1988: 71; Bousquet, 2003f: 480. Material studied. “Liquen i Butrintes” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Subgenus Poecilus s.str. 90.2. Poecilus cupreus (Linnaeus, 1758) 90.2.1. Poecilus cupreus cupreus (Linnaeus, 1758) References. Pterostichus cupreus: Apfelbeck, 1904: 253 (Velipoja); Apfelbeck, 1907a: 495 (Buenë River); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1930: 184 (Tomor Mt.); Schatzmayr, 1943b: 109 (Kruja; Stan-Karbunarë; Lushnje). Poecilus cupreus: Hieke & Wrase, 1988: 70. Poecilus cupreus cupreus: Bousquet, 2003f: 482. Material studied. “Oboti Bojana”, 1 s. (NMW); “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Alban. Exp.’18 Pashtrik 4-14.7.”, 7 s. (NMW); “Albanien Ura Gjolés Umg. Tirana Marz 1942 Muller Ausbeute”, 1 s. (MNHUB); “Albania Ostrovice” / “7.VII.1959 leg. Xh. Murraj”, 2 s. (HNHM); Sauk, 14.V.1961, 1 m. (MNST); Sauk, 3.IV.1963, 1 m. (MNST); Saranda District, Butrinti, 16.IV.1994, 3 s., S. Beshkov leg. (NMNHS); Tiranë Park, 8.II.1998, 2 m., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A. Paparistos leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT). Notes. In addition to the material studied, in MNHUB I have seen other more 24 s. from different localities, which remained unregistered. COLEOPTERA: CARABIDAE: HARPALINAE

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90.3. Poecilus cursorius (Dejean, 1828) 90.3.1. Poecilus cursorius cursorius (Dejean, 1828) References. Pterostichus cursorius: Schatzmayr, 1930: 187; Schatzmayr, 1943b: 110 (Lushnje). Poecilus cursorius: Hieke & Wrase, 1988: 71. Poecilus cursorius cursorius: Bousquet, 2003f: 482. Material studied. “Tirane” / “Albania leg. Bischoff 1933”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Sukthi bei Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 90.4. Poecilus koyi Germar, 1824 90.4.1. Poecilus koyi koyi Germar, 1824 References. Pterostichus koyi dinaricus: Schatzmayr, 1930: 176 (Tomor Mt.); Schatzmayr, 1942: 55 (Tomor Mt.). Pterostichus koyi: Schatzmayr, 1943b: 109 (Tërpan). Poecilus koyi: Csiki, 1940: 216 (Korab Mt.); Hieke et al., 1986: 188 (Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 69. Poecilus koyi koyi: Bousquet, 2003f: 482. Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 18 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exped. Korab 1918 24.31.VII.”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 1 s. (HNHM); “Albania Zalle-Dardha” / “23.V.1958 leg. Xh. Murraj”, 3 s. (HNHM); Bizë, 9-16.VII.1961, 2 f. (MNST); Mal i Dajtit Mt., 10-11.V.1965, 5 m., 1 f.; Mal i Dajtit Mt., 24.V.1966, 2 f. (MNST); Tiranë, 30.V. 1965, 2 m. (MNST); 2 km N Durrës, 24.V.1993, 1 s., P. Stoev & D. Zaprjanova leg. (NMNHS). 90.5. Poecilus lepidus (Leske, 1785) 90.5.1. Poecilus lepidus lepidus (Leske, 1785) References. Pterostichus lepidus: Csiki, 1940: 216 (Korab Mt.); Schatzmayr, 1942: 56. Poecilus lepidus: Hieke & Wrase, 1988: 70. Poecilus lepidus lepidus: Bousquet, 2003f: 483. Material studied. “Alban. Vermosa Penther”, 4 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). 90.6. Poecilus rebeli (Apfelbeck, 1904) References. Pterostichus rebeli: Apfelbeck, 1904: 254 (Velipoja; Vlorë); Apfelbeck, 1907a: 495 (Buenë River); Scatzmayr, 1930: 184; Schatzmayr, 1942: 59; Bousquet, 2003f: 483. Material studied. “Velipoja Albania” / “Coll. Apfelbeck”, 1 syntype (HNHM); “Mustajbeg Velipoja Albania”, 2 syntypes (NMW); “Oboti Albania”, 1 s. (NMW); “Scutari Bojana” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW). 90.7. Poecilus striatopunctatus (Duftschmid, 1812) References. Poecilus striatopunctatus: Apfelbeck, 1907a: 495 (Buenë River); Hieke & Wrase, 1988: 71; Bousquet, 2003f: 484. Material studied. “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). 150


90.8. Poecilus versicolor (Sturm, 1824) [= coerulescens (Linnaeus, 1758)] Material studied. “Alban. Ricavac Penther”, 2 s. (NMW); “Alban. Vermosa Penther”, 4 s. (NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Albania Mali-Thate” / “17.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Sauk” / “27.VII.1959 leg. Xh. Murraj”, 1 s. (HNHM); Golem, 10.VII. 1965, 1 s. (MNST); Tiranë, 26.VIII.1965, 1 m. (MNST). Notes. The mention of Hieke & Wrase (1988: 70) for Albania bases apparently on the record “Prisren” (Apfelbeck, 1904: 255 sub Pterostichus coerulescens). However, currently this locality is situated in the Kosovo Region of Serbia. Bousquet (2003f: 484) has most likely followed one of these sources too. Since there is no previous reliable record from Albania, P. versicolor is treated here as new species for the fauna of Albania.

91. Genus Pterostichus Bonelli, 1810 Subgenus Argutor Dejean, 1821 91.1. Pterostichus cursor (Dejean, 1828) References. Pterostichus vernalis cursor: Apfelbeck, 1904: 257 (Velipoja). Pterostichus cursor: Hieke & Wrase, 1988: 73; Bousquet, 2003f: 486. Material studied. “Skutari Bojana”, 1 s. (NMW); “Skutari l. See-Ufer”, 1 s. (NMW); “Skutari Mesi”, 1 s. (NMW); “Skutari Vraka”, 1 s. (NMW); “Vermosa’14 Penther”, 1 s. (NMW); “Liquen i Butrintes vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Rushkull” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Subgenus Bothriopterus Chaudoir, 1835 91.2. Pterostichus oblongopunctatus (Fabricius, 1787) 91.2.1. Pterostichus oblongopunctatus oblongopunctatus (Fabricius, 1787) References. Pterostichus oblongopunctatus: Schatzmayr, 1943a: 85; Hieke & Wrase, 1988: 74. Pterostichus oblongopunctatus oblongopunctatus: Bousquet, 2003f: 487. Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Maja Rogamit” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Cucale” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB). Notes. First detailed records for Albania. Subgenus Cheporus Latreille, 1829 91.3. Pterostichus burmeisteri Heer, 1838 91.3.1. Pterostichus burmeisteri burmeisteri Heer, 1838 [= metallicus (Fabricius, 1792)] References. Pterostichus metallicus: Klaptocz, 1910: 399 (Theth). Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Maja e Jezercë Çafa Ostars” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Shkëlzën” / “Albania COLEOPTERA: CARABIDAE: HARPALINAE

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leg. Bischoff 1935”, 18 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja Rogamit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Boge Aufstieg z. Rab” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); “Boge Çafa Shtegot” / “Albania Mueller leg. 1939”, 1 s. (MNHUB); “Maja e Dragomirit” / “Albania Mueller leg. 1939”, 1 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 15.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. Bousquet (2003f: 488) has omitted the species for Albania. Here it is confirmed for the country. Subgenus Feronidius Jeannel, 1942 91.4. Pterostichus melas (Creutzer, 1799) 91.4.1. Pterostichus melas depressus (Dejean, 1828) References. Pterostichus melas: Apfelbeck, 1904: 265 (Vlorë; Velipoja); Apfelbeck, 1909: 297 (Vlorë; Berat); Schatzmayr, 1943b: 109 (Kruja; Dukati; Tërpan); Hieke et al., 1986: 189 (Tiranë; Borshi near Vlorë; Poliçan near Tomor Mt.). Pterostichus melas depressus: Hieke & Wrase, 1988: 76; Bousquet, 2003f: 495. Material studied. “Mustajbeg Velipoja Albania”, 4 s. (HNHM); “Sudalbanien Kopstein”, 1 s. (NMW); “Skutari Penther”, 4 s. (NMW); “Hani Grabom mai” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Elbasan, Alb. VIII.-IX.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Rapsa Dr. Grabowski”, 1 s. (HNHM); “Albania Kamze” / “23.III.1958 leg. Xh. Murraj”, 3 s. (HNHM); “Albania Sauk” / “16.VI.1958 leg. Xh. Murraj”, 1 s. (HNHM); “Albania Terbac” / “6.VI.1960 leg. Xh. Murraj”, 1 s. (HNHM); Sauk, 9.III.1961, 1 m. (MNST); Tepelenë, 1-3.VI.1961, 1 m. (MNST); Rodenj, 2628.VII.1962, 2 m. (MNST); Kurbneshi, VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); Tiranë, Selit e Vogel, 17.IX.1993, 1 s., S. Beshkov leg. (NMNHS); Saranda District, Ljukove, 5.V.1994, 6 s., P. Stoev leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 22-31.X.1997 / 1-9.XI.1997, 5 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 f., A. Paparistos leg. (cAVT); Rapsh, Reç, 15.V.2002, 1 f., A. Paparistos leg. (cAVT). Notes. Besides the material studied above, in MNHUB I have seen other more 63 s. from Albania, which remained unregistered. Subgenus Melanius Bonelli, 1810 91.5. Pterostichus atterimus (Herbst, 1784) 91.5.1. Pterostichus atterimus atterimus (Herbst, 1784) References. Pterostichus atterimus: Apfelbeck, 1904: 258 (Velipoja); Schatzmayr, 1943a: 87. Pterostichus atterimus atterimus: Bousquet, 2003f: 499. Material studied. “Gjormi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 91.6. Pterostichus elongatus (Duftschmid, 1812) References. Pterostichus elongatus: Schatzmayr, 1930: 210 (Shkodër); Hieke & Wrase, 1988: 76; Bousquet, 2003f: 499. 152


Material studied. “Mustajbeg Velipoja Albania”, 1 s. (HNHM); “Mamurash” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Subgenus Morphnosoma Lutshnik, 1915 91.7. Pterostichus melanarius (Illiger, 1798) 91.7.1. Pterostichus melanarius melanarius (Illiger, 1798) References. Pterostichus melanarius: Hieke & Wrase, 1988: 75. Pterostichus melanarius melanarius: Bousquet, 2003f: 499. Material studied. “Alban. Exp. 1918 Gjallica Ljums 17-26.VI.”, 1 s. (NMW); “Lesh” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 8-16.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT). Notes. First detailed records for Albania. # subgenus Oreoplatysma Jakobson, 1907 # Pterostichus lodosi Heinz, 1977 References. Pterostichus lodosi: Bousquet, 2003f: 505. Notes. Most probably, Bousquet (2003f: 505) has listed this Caucasian endemic species for the country by mistake. Therefore, it is removed from the list of the Albanian ground beetles. Subgenus Parahaptoderus Jeanne, 1969 Notes. New subgenus for the fauna of Albania. 91.8. Pterostichus brevis (Duftschmid, 1812) 91.8.1. Pterostichus brevis brevis (Duftschmid, 1812) Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze Penther’14”, 7 s. (NMW). Notes. New species for the fauna of Albania. Subgenus Phonias Gosis, 1886 91.9. Pterostichus apfelbecki Csiki, 1908 [= convexiusculus Apfelbeck, 1904] References. Pterostichus convexiusculus: Apfelbeck, 1904: 261-262 (Velipoja, loc. typ.); Magistretti, 1965: 380; Jeanne, 1988: 74. Pterostichus apfelbecki: Schatzmayr, 1943a: 82 (Velipoja); Bousquet, 2003f: 507. Argutor convexiusculus: Jeanne, 1988: 74. 91.10. Pterostichus diligens (Sturm, 1824) References. Pterostichus diligens: Hieke & Wrase, 1988: 72; Bousquet, 2003f: 507. Material studied. “Vermosa’14 Penther”, 2 s. (NMW); “Çafa Markofçes” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB). Notes. First detailed records for Albania. COLEOPTERA: CARABIDAE: HARPALINAE

153

91.11. Pterostichus strenuus (Panzer, 1796) References. Pterostichus strenuus: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 234; Hieke & Wrase, 1988: 72; Bousquet, 2003f: 508. Argutor strenuus: Jeanne, 1988: 74. Material studied. “Skutari Albania”, 3 s. (NMW); “Mustajbeg Velipoja Albania”, 5 s. (HNHM; NMW); “Buschnesh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Pyla Thumana” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Subgenus Platypterinus Reitter, 1908 91.12. Pterostichus ottomanus Apfelbeck, 1908 91.12.1. Pterostichus ottomanus ottomanus Apfelbeck, 1908 References. Pterostichus ottomanus: Schatzmayr, 1943a: 106. Pterostichus ottomanus ottomanus: Bousquet, 2003f: 508. Material studies. “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalle Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 4 s. (MNHUB). Notes. Apfelbeck published three consecutive descriptions of this taxon (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. First detailed records for Albania. Subgenus Platysma Bonelli, 1810 91.13. Pterostichus niger (Schaller, 1783) 91.13.1. Pterostichus niger niger (Schaller, 1783) References. Pterostichus niger: Schatzmayr, 1930: 221 (Tomor Mt.); Csiki, 1940: 216 (Kula Ljums). Material studied. “Gjormi” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); Bizë, 27.VII.1966, 1 m. (MNST). Additional material included (studied A. Vigna Taglianti). Fier, 4.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. Bousquet (2003f: 509) has omitted this species for Albania. Here it is confirmed for the country. Subgenus Pseudomaseus Chaudoir, 1838 91.14. Pterostichus anthracinus (Illiger, 1798) 91.14.1. Pterostichus anthracinus anthracinus (Illiger, 1798) References. Pterostichus anthracinus: Apfelbeck, 1909: 297 (Pasha Liman); Hieke & Wrase, 1988: 74. Pterostichus anthracinus anthracinus: Bousquet, 2003f: 510. Material studied. 36 s. from different localities (MNHUB); “Albania Shkoder” / “25.V.1958 leg. Xh. Murraj”, 1 s. (HNHM); Tiranë Park, 22.II.1998 / 8.III.1998, 6 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). 154


91.14.2. Pterostichus anthracinus biimpressus (Küster, 1853) References. Pterostichus anthracinus biimpressus: Apfelbeck, 1904: 260 (Velipoja; Vlorë); Bucciareli & Sopracordevole, 1958: 198 (Mamurash); Bousquet, 2003f: 510. Notes. Magistretti (1965: 378) noted by mistake that the type locality of ssp. biimpressus is “Almissa (Albania)”. In fact this Italian name concerns the town of Omiš at the Dalmatian coast of Croatia. 91.15. Pterostichus gracilis (Dejean, 1828) 91.15.1. Pterostichus gracilis gracilis (Dejean, 1828) References. Pterostichus gracilis: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 219; Schatzmayr, 1943a: 84; Hieke & Wrase, 1988: 74. Pterostichus gracilis gracilis: Bousquet, 2003f: 511. 91.16. Pterostichus minor (Gyllenhal, 1827) 91.16.1. Pterostichus minor minor (Gyllenhal, 1827) References. Pterostichus minor: Apfelbeck, 1904: 261 (Velipoja); Schatzmayr, 1930: 220; Hieke & Wrase, 1988: 74. Pterostichus minor minor: Bousquet, 2003f: 511. 91.17. Pterostichus nigrita (Paykull, 1790) Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. New species for the fauna of Albania. Subgenus Pterostichus s.str. [= Aphaon Heyden, 1906, syn. nov.] Notes. The principal character used to separate Pterostichus s.str. and Aphaon (replacement name for Phaon Heyden, 1883) is the number of lateral setiferous pores on pronotum (Heyden, 1883). However, this character is inapplicable to divide the species of the two subgenera. For instance, Pterostichus fasciatopunctatus (Creutzer, 1799), type species of the genus and nominotypical subgenus has sometimes 1-3 marginal seta on the lateral border of pronotum. On the other hand, P. rhilensis Rottenberg, 1874, the type species of Aphaon also has 1-3 (sometimes 4) such setae. In fact, no other species from Aphaon shares this character. The remaining four diagnostic features of Aphaon, e.g. transverse keel on the last visible sternum in the males, number of the marginal pores on the same sternum in both sexes, reduction of the scutellar stria and sharp basal border of elytra, cannot be of subgeneric importance since these present also in other species of Pterostichus s.str. For this reason Aphaon is treated as junior synonym of the latter subgenus and the following nomenclatorial acts are introduced: Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment from Pterostichus (Aphaon) bischoffianus Jedlička, 1936 Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment from Pterostichus (Aphaon) brucki Schaum, 1859


155

Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment from Pterostichus (Aphaon) latifianus latifianus Apfelbeck, 1906 Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment from Pterostichus (Aphaon) latifianus strupii J. Müller, 1935 Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment from Pterostichus (Aphaon) lumensis lumensis Apfelbeck, 1905 Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment from Pterostichus (Aphaon) lumensis litae Csiki, 1940 Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment from Pterostichus (Aphaon) lumensis ljubetensis Apfelbeck, 1908 Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment from Pterostichus (Aphaon) malissorum Apfelbeck, 1905 Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment from Pterostichus (Aphaon) meisteri Reitter, 1885 Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment from Pterostichus (Aphaon) merklii (J. Frivaldszky, 1879) Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment from Pterostichus (Aphaon) pentheri Apfelbeck, 1918 Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment from Pterostichus (Aphaon) reiseri Ganglbauer, 1889 Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment from Pterostichus (Aphaon) rhilensis rhilensis Rottenberg, 1874 Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment from Pterostichus (Aphaon) rhilensis kourili Mařan, 1933 Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment from Pterostichus (Aphaon) rhilensis vitosensis Mařan, 1933 Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment from Pterostichus (Aphaon) walteri Reitter, 1883 91.18. Pterostichus bischoffianus Jedlička, 1936 References. Pterostichus bischoffianus: Jedlička, 1936: 167 (Gjalicë Mt., loc. typ.); Schatzmayr, 1943a: 109 (Gjalicë Mt.); Bousquet, 2003f: 515. Material studied. “Gjallica” / “Albania leg. Bischoff 1936” / “Typus” / “bischoffianus sp.n. type det. Ing. Jedlička”, 1 m. syntype (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 2 m. (MNHUB). 91.19. Pterostichus brucki Schaum, 1859 References. Pterostichus brucki: Apfelbeck, 1918c: 166 (North Albanian Alps, Vermosh, 1100 m); Jedlička, 1936: 167 (Koritnik Mt.; Gjalicë Mt.); Csiki, 1940: 216 (Gjalicë Mt.); Schatzmayr, 1943a: 107 (Koritnik Mt.; Gjalicë Mt.); Hieke, 1981: 80; Hieke & Wrase, 1988: 78; Bousquet, 2003f: 511. Material studied. “Vermoša 1100 m” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Gjalica” / “Albania leg. Bischoff 1935”, 1 s. (NMW); “Koritnik” / “Albania leg. Bischoff 1935”, 8 s. (NMW; MNHUB); “Gjallica” / “Albania leg. 156


Bischoff 1935”, 1 s. (MNHUB); “Çafa Markofçes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 32 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). 91.20. Pterostichus latifianus Apfelbeck, 1906 91.20.1. Pterostichus latifianus latifianus Apfelbeck, 1906 [= munella Jedlička, 1936] References. Pterostichus latifianus: Apfelbeck, 1906: 1663 (North Albania, Merdita: Mal Shenjt Mt. and Munella Mt., loc. typ.); Apfelbeck, 1907a: 498-499, 502 (Mal i Shenjt Mt.: top region, 1500 m); Schatzmayr, 1930: 256; Jedlička, 1936: 168 (Merdita; Dërze); Jedlička, 1939: 52; Schatzmayr, 1943a: 109 (Merdita; Dërze). Pterostichus malissorum ab. munella: Jedlička, 1936: 169 (Munella Mt., loc. typ.); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Munella Mt.). Pterostichus latifianus latifianus: Bousquet, 2003f: 485. Material studied. “Apfelb. Albanien”, 2 syntypes (MNHUB); “Latif Alban. sept.”, 4 s. (DEI; HNHM); “Latif Munela” / “Latif Alban. spt.”, 1 s. (HNHM); “Latif Orosi” / “latifianus typus! Apfelb.” / “typus”, 1 f. syntype (NMW); “Latif Orosi”, 2 syntypes (NMW); “Merdita Mal Shêit” / “latifianus Apf. cotype det. Apflb.”, 1 m. syntype (MNHUB); “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (HNHM); “Merdita M. Scheit” / “Coll. Apfelbeck”, 12 syntypes (HNHM); “Merdita M. Scheit”, 4 s. probably syntypes (NMW); “Merdita Munela”, 8 syntypes (NMW; MNHUB); “Merdita Munela” / “Apfelb. Albanien”, 1 syntype (MNHUB); “Merdita Zebia”, 4 syntypes (MNHUB); “Merdita Zebia” / “Wgth. 1906 Merdita”, 10 s. (NMW); “Merdita Oroschi”, 10 s. (HNHM; NMW); “Derzë” / “Albania leg. Bischoff 1935”, 67 s. (MNHUB). Notes. Jedlička (1939: 52) stated that his P. malissorum ab. munella is identical with P. latifianus, based on revision of type material, but subsequent authors (Schatzmayr, 1943a; Lorenz, 1998: 261) overlooked his opinion. 91.20.2. Pterostichus latifianus strupii J. Müller, 1935 References. Pterostichus latifianus strupii: Müller, 1935: 156 (Qaf e Stames, northeast of Kruja, loc. typ.); Schatzmayr, 1943a: 109 (Qaf e Stames); Bousquet, 2003f: 485. Notes. The ssp. struppi may be a distinct species since its type locality (Qaf e Stames Pass) is situated in the geographically isolated massif of Skënderbe Mt. 91.21. Pterostichus lumensis Apfelbeck, 1905 91.21.1. Pterostichus lumensis lumensis Apfelbeck, 1905 References. Pterostichus lumensis: Apfelbeck, 1905: 242 (Albania / Kosovo border, Koritnik Mt., loc. typ.); Apfelbeck, 1907: 637 (Koritnik Mt., loc. typ.); Apfelbeck, 1917: 225, note 1 (Koritnik Mt.); Apfelbeck, 1930: 169, note 1 (Koritnik Mt.); Schatzmayr, 1930: 255; Jedlička, 1936: 167 (Koritnik Mt.; Korab Mt.); Schatzmayr, 1943a: 108 (Koritnik Mt.). Pterostichus latifianus ssp.?: Schatzmayr, 1930: 256 (Korab Mt.). Pterostichus lumensis lumensis: Drovenik & Peks, 1994: 67 (Koritnik Mt.); Drovenik & Peks, 1999: 78 (Koritnik Mt.); Bousquet, 2003f: 515. Material studied. “Latif Koritnik”, 2 syntypes (NMW); “Latif Alban. spt.”, 1 s. probably syntype (HNHM); “Latif Alban. spt.” / “P. lumensis Apfelb. Typen”, 1 s. probably syntype (HNHM); “Alban. Exp.’18 Gjalica Ljums 17-26.VI.”, 3 s. (NMW); “Alban. Exp. Korab 1918 24.-31.VII.”, 16 s. (NMW); “Korab Albania Bischoff ”, 1 s. (NMW); “Bjeshka Matroxh”, 11 s. COLEOPTERA: CARABIDAE: HARPALINAE

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(MNHUB); “Korab” / “Albania leg. Bischoff 1935”, more than 120 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, more than 120 s. (MNHUB). Notes. Apfelbeck published two descriptions of this taxon (Apfelbeck, 1905; Apfelbeck, 1907b), and the older one is correctly treated as valid. Two specimens (1 m., 1 f.) labelled “Latif Alban. sept.” and identified as Pterostichus djumanus Apfelbeck were found in DEI. They are put provisionally here, since both specimens resemble P. lumensis. Pterostichus djumanus Apfelbeck is nomen nudum. 91.21.2. Pterostichus lumensis litae Csiki, 1940 References. Pterostichus lumensis litae: Csiki, 1940: 216 (Gjalicë Mt., loc. typ.; Korab Mt., loc. typ.); Schatzmayr, 1943a: 108 (Gjalicë Mt.; Korab Mt.); Bousquet, 2003f: 515. Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 m. syntype (HNHM); “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 4 syntypes (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 21 syntypes (HNHM). ? Pterostichus lumensis ljubetensis Apfelbeck, 1908 [= chalybaeus Apfelbeck, 1908] References. Pterostichus lumensis ljubetensis: Schatzmayr, 1930: 255; Bousquet, 2003f: 515. Notes. Apfelbeck published three descriptions of this taxon (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), and the oldest one is correctly treated as valid. All data in the literature and collections about ssp. ljubetensis come from the subalpine and alpine regions of the Shar Planina Mts. that do not extend into Albania. Here, P. lumensis ljubetensis is treated as doubtful for the fauna of Albania. 91.22. Pterostichus malissorum Apfelbeck, 1905 [= nigripes Jedlièka, 1936] References. Pterostichus malissorum: Apfelbeck, 1905: 241-242 (Çukall Mt., loc. typ.); Apfelbeck, 1907a: 497 (Maranajt Mt.: top region, 1500-1576 m); Apfelbeck, 1907b: 637 (Çukall Mt., loc. typ.); Schatzmayr, 1930: 255; Jedlička, 1936: 169 (Malissora; Çukall Mt.; Cafa Valbones; Cafa Pejs; Maja Pejs); Schatzmayr, 1943a: 108 (Çukall Mt.; Malissora; Cafa Valbones; Cafa Pejs; Maja Pejs); Bousquet, 2003f: 515. Pterostichus latifianus ab. nigripes: Jedlička, 1936: 168 (North Albanian Alps: Quf e Valbone, loc. typ.); Jedlička, 1939: 53. Pterostichus latifianus strupii (= nigripes): Schatzmayr, 1943a: 109 (Quf e Valbone). Material studied. “Latif Alban. sept.”, 4 s. (DEI; HNHM); “Latif Cukali” / “Coll. Apfelbeck”, 1 m. syntype (HNHM); “Latif Cukali”, 2 syntypes (NMW); “Maranai Apfelb.”, 1 s. (HNHM); “Maranai Apf.”, 1 s. (HNHM); “Apfelb. Albanien”, 1 m. syntype (MNHUB); “Apfelb. Albanien” / “Alban. spt. Maranaj”, 1 f. probably syntype (MNHUB); “Apfelb. Alban spt” / “Maranaj Geb.”, 2 s. (HNHM); “Apfelb. Alban. spt. IV.V.’05” / “Maranaj”, 6 s. (NMW); “Kunor Vukai Zukali Albanien Lona Giugno 1914”, 6 s. (NMW; MNHUB); “Cukale Albania Bischoff ”, 12 s. (MNHUB); “Cukale-Geb.” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Çafa Stamës” / “leg. Strupi Albania 1934” / “Cotype” / “latifianus a. nigripes n. ab. det. Ing. Jedlička”, 1 f. designated as syntype of Pterostichus latifianus ab. nigripes Jedlička, 1936 (MNHUB); “Cukale” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Çafa Valbonës” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja Pejs” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “M. Mulecifit” / “Albania leg. Bischoff 1936”, 14 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 23 s. (MNHUB); “Boge ober D. Quelle” / “Albania leg. Mueller 158


1939”, 1 s. (MNHUB); “M. Raba Massiv” / “Albania leg. Mueller 1939”, 10 s. (MNHUB); “Maja e Dragomirit” / “Albania leg. Mueller 1939”, 4 s. (MNHUB); North Albanian Alps, Maj’e Redohimes, 1900-2200 m, 28.V.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Maj’e Redohimes, 2200-2550 m, 29.V.1993, 2 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 5 s., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron & B. Petrov leg. (NMNHS). Notes. Apfelbeck published two descriptions of this species (Apfelbeck, 1905; Apfelbeck, 1907b), and the older one is correctly treated as valid. Jedlička (1939: 52) stated that his P. latifianus ab. nigripes Jedlička, 1936 is identical with P. malissorum Apfelbeck, 1905. Later, Schatzmayr (1943a), however, synonymized P. latifianus ab. nigripes Jedlička, 1936 with P. latifianus strupii J. Müller, 1935. I support the view of Jedlička because the type locality of the former is quite close to the known findings of P. malisstorum, whereas P. latifianus strupii lives further in the south: Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. of Pterostichus (Pterostichus) malissorum Apfelbeck, 1905. 91.23. Pterostichus pentheri Apfelbeck, 1918 References. Pterostichus pentheri: Apfelbeck, 1918c: 161, 166 (North Albanian Alps, loc. typ.); Schatzmayr, 1930: 255; Jedlička, 1936: 168 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë Mt.); Schatzmayr, 1943a: 108 (North Albanian Alps; Shkëlzen Mt.; Maja e Jezercë Mt.); Bousquet, 2003f: 515. Material studied. “Albania coll. Lichtn.”, 2 s. (HNHM); “Prokletija Albanien Dr. Penther”, 2 syntypes (NMW); “Prokletija” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes (DEI; NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 5 syntypes (NMW); “Maja e Jezercë” / “Albania leg. Bischoff 1931”, 7 s. (MNHUB); “Maja e Jezercë Çafa Pejs-Çafa Gstars” / “Albania leg. Bischoff 1931”, 2 s. (MNHUB); “Çafa e Markofçes” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Maja e Jezercë” / “Albania leg. Bischoff 1935”, 12 s. (MNHUB); more than 90 s. from locality “Shkëlzën” (MNHUB); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 3 s., P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, Kollatës Mt., Qafa-eRubës, 2000 m, 8.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 2 m., 2 f., L. Latella leg. (cAVT); Valbona, Kollatës Mt., 2000 m, 14.VII.1993, 2 m., A. Gobetti leg. (cAVT). 91.24. Pterostichus reiseri Ganglbauer, 1889 References. Pterostichus reiseri: Apfelbeck, 1918c: 160, 166 (North Albanian Alps, Vermosh, Grebeni, 1100-1700 m); Jedlička, 1939: 52; Schatzmayr, 1943a: 108 (Grebeni). Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 2 s. (NMW); “Vermoša Albanien Dr. Penther”, 2 s. (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW). Notes. Bousquet (2003f: 516) has omitted P. reiseri for Albania. Here the species is confirmed for the country. The record “Albanien: Žljeb plan.” (Jedlička, 1936: 168) refers to the massif shared among Montenegro and the Kosovo Region of Serbia.


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Species insertae sedis ? Pterostichus epiroticus Csiki, 1930 [= corax Ganglbauer, 1889] References. Pterostichus epiroticus: Schatzmayr, 1943a: 107. Notes. The record “Peristerigebirge bei Jannina (Albanien)” (Schatzmayr, 1943a) is based on an older record of Apfelbeck (1904: 267 sub P. corax). Currently this record concerns the territory in Northwestern Greece. In NMW, besides specimens from Central Greece (Oeta Mt.; Korax Mt.), I found five recently collected specimens from the region near to Joannina and Metsovon, which is very close to the Albanian territory. Therefore, P. epiroticus is treated here as likely to be found in Albania.

92. Genus Speluncarius Reitter, 1886 Subgenus Hypogium Tschitschérine, 1900 [= Hypogeobium Tschitschérine, 1903] 92.1. Speluncarius albanicus (Tschitschérine, 1900) References. Platysma albanicum: Tschitschérine, 1900: 49-51 (“Albanie”, loc. typ.). Pterostichus albanicus: Apfelbeck, 1904: 236-237; Schatzmayr, 1930: 329. Tapinopterus albanicus: Reitter, 1914: 262; Müller, 1937: 134; Schatzmayr, 1943a: 131; Jeannel, 1950: 157, 159-160. Hypogeobium albanicum: Jeannel, 1950: 160, 162; Jeannel, 1953: 11. Speluncarius albanicus: Sciaky, 1982: 28; Casale & Giachino, 1991: 215-216; 219; Bousquet, 2003f: 516. Material studied. “Albania” / G.C.Champion Coll. B.M.1927-409” / “Platysma (Hypogium) albanicum m. Typ. Tschitscherin det” / “syntype” (round blue coloured label), 1 f. (BMNH). Notes. The type locality of S. albanicus remains not yet specified. Subgenus Speluncarius s.str. 92.2. Speluncarius minutulus J. Müller, 1937 References. Tapinopterus minutulus: Müller, 1937: 132-123 (Shkëlzen Mt., loc. typ.); Schatzmayr, 1943a: 128 (Shkëlzen Mt.); Turin, 1981: 127. Speluncarius minutulus: Jeanne, 1982: 41-42 (Shkëlzen Mt.); Sciaky, 1982: 20; Drovenik & Peks, 1994: 68; Drovenik & Peks, 1999: 79; Bousquet, 2003f: 516. Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 f. designated as paralectotype by P.M. Giachino (MNHUB). Notes. Drovenik & Peks (1994) and Drovenik & Peks (1999) noted S. minutulus for Montenegro by mistake because it is a local endemic species for Northeast Albania.

93. Genus Stomis Clairville, 1806 Subgenus Stomis s.str. 93.1. Stomis pumicatus (Panzer, 1796) 93.1.1. Stomis pumicatus pumicatus (Panzer, 1796) References. Stomis pumicatus: Apfelbeck, 1904: 213 (Velipoja); Hieke & Wrase, 1988: 68; Bousquet, 2003f: 517. 160


Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB). 93.2. Stomis rostratus (Duftschmid, 1812) 93.2.1. Stomis rostratus rostratus (Duftschmid, 1812) References. Stomis rostratus: Apfelbeck, 1918c: 166 (North Albanian Alps, Grebeni near Vermosh, 1100-1800 m); Magistretti, 1965: 362; Bousquet, 2003f: 518. Material studied. “Grebeni 16.VI.” / “Alban. montenegr. Grenze, Penther’14”, 3 s. (NMW); “Alban. Vermosa Penther”, 2 s. (NMW); “Vermosa 1200 m 3.VI.-5.VII.” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Shkëlzën” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).

94. Genus Tapinopterus Schaum, 1858 Subgenus Tapinopterus s.str. Notes. The species from this subgenus are placed here in three groups of species in agreement with a forthcoming arrangement proposed by P. M. Giachino and L. Picciau (personal communication). I. South Aegean groups. Species group “extensus” # Tapinopterus extensus (Dejean, 1831) # Tapinopterus extensus abreshensis (Lutshnik, 1916) [= convexiusculus Apfelbeck, 1904] References. Pterostichus extensus convexiusculus: Apfelbeck, 1904: 251 (“Turcia – Abresch”, loc. typ.); Schatzmayr, 1943a: 127. Notes. According to Schatzmayr (1943a) the type locality of this race (“Abresch”) must be somewhere between Albania and Palestine. Bousquet (2003f: 519) has noted the subspecies for Greece. Based on Apfelbeck’ short description (Apfelbeck, 1904: 251 sub Pterostichus extensus convexiusculus), I suppose that ssp. abreshensis is not subordinate to T. extensus s.l., taking into consideration also the fact that Apfelbeck (1904) compared the former with T. kaufmanni Ganglbauer, 1897. Dealing with the taxa of Tapinopteris extensus s.l., Mařan (1939b: 298, 303) without hesitation separated T. extensus abreshensis from the remaining races of the species. Only a revision of Apfelbeck’s type (probably in Museum Sarajevo) could establish the systematic position of the taxon in question. Presently, both the species and subspecies are removed from the list of the Albanian Carabidae. 94.1. Tapinopterus extensoides Straneo, 1936 [= extensoides Jedlièka, 1936] References. Tapinopterus extensoides Straneo: Straneo, 1936: 155-156 (Llogora, typ. loc.); Müller, 1937: 132, note 1 (Llogora; cave “Paftali” near Berat; Shëndelli Mt.); Schatzmayr, 1943a: 127 (Llogora; cave “Paftali” near Berat; Shëndelli Mt.). Tapinopterus extensoides Jedlička: Jedlička, 1936: 169 (Llogora, loc. typ.); Turin, 1981: 127. Tapinopterus extensus extensoides “Straneo et COLEOPTERA: CARABIDAE: HARPALINAE

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Jedl.”: Mařan, 1939b: 295-299, 302-303 (Llogora). Tapinopterus extensus extensoides Jedlička: Bousquet, 2003f: 519. Material studied. “Logara Juni” / “Albania leg. Bischoff 1932”, 1 m. syntype (NMW); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Reza e Kanalit Logara, Alban. m.” / “Albania leg. Bischoff 1932”, 1 f. syntype (NMW); “Logara juni” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 7 s. (MNHUB); “Llogora, Alb. Mer.” / “Albania leg. Bischoff 1935”, 1 f. syntype of extensoides Jedlička, 1936 (MNHUB); “Llogora” / “leg. Bischoff Albania 1936”, 17 s. (MNHUB); “Shen Ili” / “leg. Bischoff Albania 1936”, 1 s. (MNHUB); “Borshe” / “Albania leg. Bischoff 1940”, 1 s. (MNHUB); “Shpella Buz i Lugit Mal i Piqs” / “Borshe” / “Albania leg. Bischoff 1940”, 2 s. (MNHUB); “Albania, Periferi Tepelenë, a gorge 1.5 km W of Progonat,” / “785 m, 9.X.2004, Z. Fehér, J. Kontschán & D. Murányi (No. 133)”, 2 f. (HNHM). Notes. In 1936 Jedlička (1936) and Straneo (1936) described T. extensoides having at disposal identical specimens collected by Bischoff from the region of Llogora. Later, Müller (1937: 132, note 1) stated that the Straneo’s description has priority over that of Jedlička. This opinion is correct in spite of Mařan’s statement that both are authors of this taxon (Mařan, 1939b: 302). It is Mařan who first regarded extensoises as subspecies of T. extensus. However, in my view T. extensoides Straneo is clearly distinct from all the other subspecies of T. extensus by an evident apomorphic character – 2 (instead 1) posterior supraorbital setiferous pores (see also Straneo, 1936). The check of all specimens at hand demonstrated that this character is constant (as only a single female from HNHM has a single left and two right posterior supraorbital pores) so that it may define a separate species: Tapinopterus (Tapinopterus) extensoides (Straneo, 1936), stat. rest. from Tapinopterus (Tapinopterus) extensus extensoides (Straneo, 1936). Species group “protensus” 94.2. Tapinopterus protensus (Schaum, 1857) Notes. Bousquet (2003f: 520) noted that T. protensus aetolicus Ganglbauer, 1889 inhabits Montenegro, and not Albania. In spite of that Schatzmayr (1943: 122) cited it for the studied country (see below). Here, the species records and reference data from Albania are referred to two defined and two undefined subspecies. The occurrence of T. protensus in Montenegro needs serious confirmations. 94.2.1. Tapinopterus protensus peristericus Apfelbeck, 1901 References. Tapinopterus aetolicus peristericus: Müller, 1935: 153 (Shëndelli Mt. near Tepelenë). Material studied. “Maja e Papingut” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 2 s. (MNHUB); “Maja e Policanit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 6 s. (MNHUB); “Shen Ili” / “leg. Bischoff Albania 1936”, 3 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”, 31 s. (MNHUB). Notes. This subspecies is confined to the area between the valleys of Drina River and Vijosë River and lives also in Northwestern Greece. Müller (1935: 153-154) noted the identity of syntypes of T. peristericus Apfelbeck, 1901 with specimens from the Shëndelli Mt. In MNHUB, I found specimens from Shëndelli Mt. labelled “aetolicus sbsp. peristericus det. J. Müller”, and others - “aetolicus subsp. nova m. det. J. Müller”.

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94.2.2. Tapinopterus protensus tomoricensis J. Müller, 1935 References. Tapinopterus aetolicus tomoricensis: Müller, 1935: 153-154 (Tomor Mt., loc. typ.). Tapinopterus thessalicus tomoricensis: Mařan, 1940b: 59 (Tomor Mt.). Tapinopterus protensus tomoricensis: Schatzmayr, 1943a: 122 (Tomor Mt.); Bousquet, 2003f: 520. Material studied. “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 2 syntypes (NMW); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 8 syntypes (NMW); 40 s. from Tomor Mt. (MNHUB). ! Tapinopterus protensus ssp.1 Material studied. “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 1 m. (MNHUB). Notes. Josef Müller has noted on the label that the specimen from “Mal i Dajtit” belongs to an undescribed taxon. This specimen has gigantic body dimensions (21.5 mm) in comparison with the other races from Albania, but more material is needed for the correct classification of this local population. ! Tapinopterus protensus ssp.2 References. Tapinopterus aetolicus: Müller, 1935: 154 (Reza e Kanalit Mt. near Llogora). Tapinopterus protensus aetolicus: Schatzmayr, 1943a: 122 (Reza e Kanalit Mt. near Llogora). Material studied. “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Maja e Tartarit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 7 s. (MNHUB). Notes. This still not defined race inhabits the mountains west of the valleys of Vijosë River and Drina River. Müller has identified the specimens from “Llogara” as ssp. aetolicus, and that from “Maja e Tartarit” as “aetolicus sbsp. valonensis m. det. J. Müller”. Both populations from Albania may not belong to the ssp. aetolicus (as Müller stated, 1935), if considering the geographical factor, since the type locality of ssp. aetolicus is the Korax Mt. (Central Greece), situated nearly 270 km south of the area of Llogora. II. North Aegean groups. Species group “balcanicus” 94.3. Tapinopterus bischoffi J. Müller, 1936 References. Tapinopterus bischoffi: Müller, 1936: 106-107 (Mal i Dajtit Mt., loc. typ.); Schatzmayr, 1943a: 127 (Mal i Dajtit Mt.); Turin, 1981: 127; Bousquet, 2003f: 519. Material studied. “Mal i Dajtit Tiranë” / “Albania leg. Bischoff 1931”, 1 s. (MNHUB); “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 6 s. (MNHUB); “Mal i Drini” / “leg. Bischoff Albania 1934” / “cotype” / “Tapinopterus bischoffi m. det. J. Müller”, 1 m. (NMW); “Rushkull” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 5 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1939”, 7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); “AL (Kruje) Paß ca. 5 km ne. Kruje 700-900 m, 1.V.1993 leg. Heinz & Sonja Peks”, 1 f. (MNHUB); Mal i Dajtit Mt., 20 km E Tiranë, 1000 m, 9.V.1995, 2 m., 2 f., L. Penev leg. (NMNHS).


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# Tapinopterus comita Jedlička, 1935 References. Tapinopterus eomita sic!: Turin, 1981: 127. Tapinopterus comita: Bousquet, 2003f: 519. Notes. I do not find any exact records in the literature or material in the collection of this species from Albania, thus the indications of Turin (1981) and Bousquet (2003f) for the country may be treated as incorrect. T. comita inhabits only the Pelister Mt. (Jedlička, 1935; Drovenik & Peks, 1994: 68), a massif shared between Macedonia and Greece. The species is removed from the list of the Albanian Carabidae. 94.4. Tapinopterus detonii Schatzmayr, 1943 References. Tapinopterus detonii: Schatzmayr, 1943b: 110 (Mal i Krujes Mt., 1000 m, loc. typ.); Turin, 1981: 127; Bousquet, 2003f: 519. Material studied. “Albania”, 1 s. (DEI). Notes. Further taxonomic work is needed to demonstrate the differentiation of T. detonii from T. bischoffi J. Müller, 1936, since the former seems not well defined and may be a subspecies or synonym of the latter. 94.5. Tapinopterus dochii (Apfelbeck, 1906) References. Pterostichus dochii: Apfelbeck, 1906: 1663-1664 (Merdita: Mal i Shenjt Mt., loc. typ.; Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 499, 502 (Mal i Shenjt Mt.: top region, 1500 m). Tapinopterus dochii: Jedlička, 1935: 186 (Korab Mt.); Müller, 1936: 107 (Korab Mt.); Schatzmayr, 1943a: 126 (Mal i Shenjt Mt.; Zepë Mt.; Munella Mt.); Hieke, 1981: 81; Bousquet, 2003f: 519. Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05” / “Tapinopterus dochii Apf.”, 1 f. syntype (NMW); more than 100 s. from localities “Berishe”, “Cucale”, “Maja e Shpatit”, “Cafa e Sharit”, “Maja Shales Vogel Iballe”, “Korab”, “Munela”, “Derzë”, “Taja”, “M. Mulecifit”, “Merdita M. Sheit” (MNHUB). [Tapinopterus miridita (Apfelbeck, 1904)] Notes. The only records for T. miridita s.l. from Albania are those of Hieke (1981: 81) and Bousquet (2003f: 520). However, the present author cannot find any exact data in the literature or collections to prove the species occurrence in Albania. The type locality “Sveti Troice” (Apfelbeck, 1904: 245), or more correct “Sveta Trojica”, concerns a monastery near the village of Mušutište, a place situated in the northern slopes of the Shar Planina Mt. (in the Kosovo Region of Serbia) (see also Jedlička, 1935: 186). Oddly enough, the “loc. typ.” and known records of the species are quite removed from the region of Mirdita (situated in the south part of North Albania, south of the Drin River), after which the species was named. T. miridita is treated here as likely to be found in the country. [Tapinopterus miridita miridita (Apfelbeck, 1904)] Notes. The occurrence of this race in the marginal northeastern regions of Albania (the Koritnik Mt.; the Gjalicë Mt.) is possible. [Tapinopterus miridita jablanicensis Mařan, 1935] Notes. This race was described from the Macedonian part of the Jablaniča Mt. (shared between Albania and Macedonia), but so far the subspecies is not properly cited for Albania. 164


# Tapinopterus miridita jacupicensis Jedlička, 1935 Notes. Bousquet (2003f: 520) cited this subspecies for Albania. However, this taxon is a local endemic for the massif of the Jakupiča Mt. situated in the central part of Republic of Macedonia. Probably the author in question has had in mind the spp. jablanicensis when he mentioned ssp. jacupicensis for Albania. The latter subspecies is here excluded from the list of the Albanian Carabidae. 94.6. Tapinopterus monastirensis Reitter, 1913 94.6.1. Tapinopterus monastirensis gestroi J. Müller, 1923 References. Tapinopterus gestroi: Müller, 1923: 110-111 (Tomor Mt.: Toik, loc. typ.). Tapinopterus monastirensis gestroi: Müller, 1936: 107; Mařan, 1939e: 82, 84 (Tomor Mt.: Toik); Schatzmayr, 1943a: 126 (Tomor Mt.); Bousquet, 2003f: 520. Material studied. “Albania”, 1 s. (DEI); “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona”, 1 s. (NMW); “Tomor: Kulmak Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Tomor Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 4 s. (NMW); “TomorTojk juli 1932” / “Albania leg. Bischoff 1932”, 32 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 17 s. (MNHUB).

Tribe SPHODRINI Laporte, 1834 Subtribe ATRANOPSINA Baehr, 1982 95. Genus Platyderus Stephens, 1827 Notes. The present knowledge about the Balkan representatives of this genus poorly corresponds to the actual status of the species. Subgenus Platyderus s.str. 95.1. Platyderus dalmatinus L. Miller, 1881 95.1.1. Platyderus dalmatinus dalmatinus L. Miller, 1881 References. Platyderus dalmatinus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m). Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze, Penther’14”, 1 m. (NMW). Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Here, it is confirmed for the fauna of Albania. 95.2. Platyderus graecus (Reiche & Saulcy, 1855) References. Platyderus graecus: Apfelbeck, 1909: 297 (Vlorë). Notes. Hovorka & Sciaky (2003a: 522) have omitted this species for Albania. Further study is needed to confirm its occurrence in the country.


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95.3. Platyderus minutus (Reiche & Saulcy, 1855) 95.3.1. Platyderus minutus obtusangulus J. Müller, 1923 References. Platyderus minutus: Apfelbeck, 1907a: 498, 501 (Mal i Shenjt Mt.: top region, 1500 m). Platyderus minutus obtusangulus: Müller, 1923: 112 (Tomor Mt.: Kulmak, loc. typ.); Hovorka & Sciaky, 2003a: 523. Material studied. “Albania”, 1 f. (BMNH). 95.4. Platyderus rufus (Duftschmid, 1812) 95.4.1. Platyderus rufus rufus (Duftschmid, 1812) References. Platyderus rufus: Apfelbeck, 1904: 279 (Velipoja near Lezhë); Hieke & Wrase, 1988: 89. Platyderus rufus rufus: Hovorka & Sciaky, 2003a: 523. Material studied. Nearly 50 s. from different localities (MNHUB); Rojan, 6.III.1961, 1 f. (MNST).

Subtribe CALATHINA Laporte, 1834 96. Genus Calathus Bonelli, 1810 Subgenus Bedelinus Ragusa, 1885 96.1. Calathus circumseptus Germar, 1824 References. Calathus circumseptus: Hovorka & Sciaky, 2003b: 524. Material studied. “Albania”, 1 s. (HNHM); “Durazzo”, 1 s. (HNHM); “Apfelb. 1893 Durazzo”, 1 s. (HNHM); “Berat” / “Albania leg. Bischoff 1931”, 4 s. (MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Lum i Erzen” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Tiranë” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1938”, 1 s. (MNHUB); Saranda District, Qeparo, 4-5.V.1994, 2 s., P. Stoev leg. (NMNHS); Ljukove, 100 m, 5.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Kruja Castle, 16.XI.1997, 2 s., R. Ward leg. (CMNH). Notes. First detailed records for Albania. Subgenus Calathus s.str. 96.2. Calathus bosnicus Ganglbauer, 1891 References. Calathus bosnicus: Hovorka & Sciaky, 2003b: 524. Material studied. “Prokletija 1300-2000 m” / “Alban.-montenegr. Grence, Penther’14”, 2 s. (NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 1 s. (NMW); “Gjalica Ljums 17-26.VI.”, 1 s. (NMW); “Llogora…”, 1 s. (NMW); 60 s. from localities “Shkëlzën”, “Çafa Markofçes”, “Maja Pejs”, “Korab”, “Maja e Jezercoë”, “Valbona Çerem” (MNHUB); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS). Notes. First detailed records for Albania. The record “Llogora…” either concerns another congener or is “patria falsa”, since the locality is quite removed in the south from the range of the species. 166


[Calathus distinguendus Chaudoir, 1846] Notes. According to Battoni & Vereschagina (1984: 141) this species may live in Albania, but so far this statement is not proved. 96.3. Calathus ellipticus Reitter, 1889 96.3.1. Calathus ellipticus ellipticus Reitter, 1889 Material studied. “Çafa e Dhembellit”, 2 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 26 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 6 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB). Notes. New species for the fauna of Albania. After a comparison of type material with specimens from Albania, D.W. Wrase (personal communication) concluded that they are conspecific, being in agreement with specimens from Western Greece. A revision of all related taxa involved (including also C. giganteus and C. korax) is needed to clarify the status of the Albanian population. 96.4. Calathus fuscipes (Goeze, 1777) 96.4.1. Calathus fuscipes graecus Dejean, 1831 [= latus (Audinet-Serville, 1821) nec Linnaeus, 1758] References. Calathus fuscipes: Apfelbeck, 1904: 284 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë; Berat); Csiki, 1940: 217 (Shkëlzen Mt.; Kula Ljums); Csiki, 1943: 214; Schatzmayr, 1943b: 110 (“Poggio Boschetto”; Kruja; Dukati); Battoni & Vereschagina, 1984: 152; Hieke et al., 1986: 189 (Tiranë; Tomor Mt.: Abbas Ali Monastery; Bizë near Shëngjergji; Lurë e Vjetër near Lura Mt.); Hieke & Wrase, 1988: 89. Calathus fuscipes var. graecus: Wohlberedt, 1909: 127 (around Liqen i Shkodrës). Calathus fuscipes graecus: Schatzmayr, 1937: 27 (Gjirokastër); Hovorka & Sciaky, 2003b: 525. Calathus fuscipes var. latus: Csiki, 1940: 217 (Gjalicë Mt.; Korab Mt.). Calathus fuscipes fuscipes: Hovorka & Sciaky, 2003b: 525. Material studied. More than 180 s. from different localities (MNHUB); L. Selitës, 8-9.VII.1961, 5 s. (MNST); Lurë, 18-25.VII.1961, 2 s. (MNST); Theth, 1-8.VIII.1961, 2 f. (MNST); Ibë, 14.VII.1962, 1 f. (MNST); Shkodër, 15.V.1963, 1 m.; Mal i Dajtit Mt., 10-11.V.1965, 1 m., 1 f. (MNST); 2 km N Durrës, 24.V.1993, 1 m., 1 f., P. Stoev & D. Zaprjanova leg. (NMNHS); North Albanian Alps, Maj’e Redohimes, 2400-2550 m, 29.V.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 f., P. Beron & B. Petrov leg. (NMNHS); Reshen, 11.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS); Cafasan, 1230 m, 27.VII.1993, 3 m., P. Stoev & D. Zaprjanova leg. (NMNHS); Tiranë, botanical garden, 8.V.1995, 1 m, 1 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Vlorë District, Dukati Mt., 450 m, 11.V.1995, 2 f., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Llogora Pass, 1025 m, 11.V.1995, 2 m., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 8.II.1998, 5 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., 3 f., A. Paparistos leg. (cAVT). Notes. According to A. Vigna Taglianti (personal communication) the populations from Albania belong to ssp. graecus. Besides the material studied above, in NMW I have seen more specimens, which remained unregistered.


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96.5. Calathus giganteus Dejean, 1828 References. Calathus giganteus: Apfelbeck, 1904: 282 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Schatzmayr, 1937: 2, 6; 21 (Vlorë; Sopoti near Gjirokastër); Hovorka & Sciaky, 2003b: 525. Material studied. “Llogora juni” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Tepelene juni 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 6 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Vojusa Tepelene” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Llogora Pass, 1025 m, 11.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). Additional material included (studied S. Hristovski and D.W. Wrase). “Logara” / “Albania, leg Bischoff 1932”, 1 s. (MNHUB); Hotel “Edva” at Radhime (Vlorë District), 8-15.VII.2006, 1 s., leg. M. Komnenov (cSH). ? Calathus glabricollis Dejean, 1828 Material studied. “coll. L.W. Schaufuß” / “Calathus glabricollis Albanien verus”, 1 m. (MNHUB). Notes. It is not sure if the record above concerns the present territory of Albania, so the species is treated here as doubtful for the country. 96.6. Calathus korax Reitter, 1889 References. Calathus corax: Schatzmayr, 1937: 2, 7; 21 (Gjirokastër). Notes. Hovorka & Sciaky (2003b: 525) have omitted C. korax for Albania. Further collections are needed to confirm its occurrence in the country. 96.7. Calathus ravasinii J. Müller, 1935 96.7.1. Calathus ravasinii ravasinii J. Müller, 1935 References. Calathus ravasinii: Müller, 1935: 152-153 (Tomor Mt.: Kulmak, loc. typ.); Schatzmayr, 1937: 2, 7, 22 (Tomor Mt.); Turin, 1981: 136; Hovorka & Sciaky, 2003b: 526. Material studied. “M. Toik Alban. mer.”, 1 s. probably syntype (NMW); “Tomorica Kulmak” / “Albania 1922 Ravasini, Lona”, 3 syntypes (NMW); “Tomor” / “Albania leg. Bischoff 1932”, 22 s. (MNHUB). Notes. Schatzmayr (1937: 22) treated C. ravasinii as separate species, but hinted that it resembles C. macedonicus. Based on the material studied, the present author considers the two taxa are only well-defined subspecies of one species: Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat. nov. for Calathus (Calathus) macedonicus Mařan, 1935. Müller (1935) published his description in Bollettino della Società Entomologica Italiana, volume 67; this issue was printed on 26.II.1935. On the other hand, Mařan (1935) described C. macedonicus in Sborník Entomologického Odděleni Národního Musea v Praze, volume 13; this issue was printed on 1.XII.1935. The nominotypical taxon is easily recognized from ssp. macedonicus by the anteriorly narrowed sides of pronotum. 96.7.2. Calathus ravasinii macedonicus Mařan, 1935 References. Calathus macedonicus: Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 525. Material studied. “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); “Bjeshka Matroxh” / 168


“Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 23 s. (MNHUB); “Teja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Notes. First detailed records for Albania. Subgenus Neocalathus Ball & Nègre, 1972 96.8. Calathus albanicus Apfelbeck, 1906 References. Calathus albanicus: Apfelbeck, 1906: 1666 (Merdita: Zepë Mt., loc. typ.); Apfelbeck, 1907a: 500, 503; Schatzmayr, 1937: 2, 8; 36 (Zepë Mt.); Csiki, 1940: 218 (Gjalicë Mt.; Korab Mt.); Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 526. Material studied. More than 160 s. with three kinds of labeling: “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918” (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); more than 150 s. from the massifs of Shar Planina Mt., Merdita, Munella Mt., Korab Mt., Gjalica Ljums and others (NMW); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 9 s. (MNHUB); “Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1935”, 22 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1935”, 8 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Fusha Korabit” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 12 s. (MNHUB); “Berishe” / “Albania leg. Mueller 1939”, 2 s. (MNHUB); “Maja Shalles i Vogel-Ibale” / “Albania leg. Mueller 1939”, 1 s. (MNHUB); “Kunora Drunit Iballe” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB). Notes. This species inhabits the mountain chain Shar Planina Mt. - Korabi Mt. and massifs in the area of Mirdita, but not the North Albanian Alps, where C. albanicus most probably is replaced by C. metallicus. 96.9. Calathus ambiguus (Paykull, 1790) 96.9.1. Calathus ambiguus ambiguus (Paykull, 1790) Material studied. “AL (Berat) Paß ca. 5 km s. Gllavë-600 m 27.IV.1993 leg. Heinz u. S. Peks”, 3 s. (MNHUB). Notes. The mention of Hieke & Wrase (1988: 90) for Albania is apparently result of following the record “Prisren” (Apfelbeck, 1904: 285). Currently this locality is situated in the Kosovo Region of Serbia. Hovorka & Sciaky (2003b: 526) have most likely followed one of these sources too. Since there is no previous reliable record from Albania, C. ambiguus is treated here as new species for the fauna of the country. 96.10. Calathus cinctus Motschulsky, 1850 [= erythroderus Gemminger & Harold, 1868] References. Calathus mollis f. erythroderus: Schatzmayr, 1943b: 111 (Lushnje). Calathus cinctus: Aukema, 1990: 138. Material studied. “Mal i Daitit” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Okol Shkalla” / “Albania leg. Bischoff 1935”, 7 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Mal i Daitit” COLEOPTERA: CARABIDAE: HARPALINAE

169

/ “Albania 1936 leg. Ramadan coll. Bischoff ”, 4 s. (MNHUB); Kruja Castle, 16.XI.1997, 1 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Rapsh, 15.V.2002, 1 m., A. Paparistos leg. (cAVT). Notes. Hovorka & Sciaky (2003b: 527) have omitted C. cinctus for Albania. Here the species is confirmed for the country. 96.11. Calathus erratus (C.R. Sahlberg, 1827) 96.11.1. Calathus erratus erratus (C.R. Sahlberg, 1827) References. Calathus erratus: Hieke & Wrase, 1988: 90. Calathus erratus erratus: Hovorka & Sciaky, 2003b: 527. Material studied. “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 m., L. Latella leg. (cAVT). Notes. First detailed records for Albania. In addition to the material studied above, the present author has seen several further specimens in NMW, which remained unregistered. 96.12. Calathus melanocephalus (Linnaeus, 1758) 96.12.1. Calathus melanocephalus melanocephalus (Linnaeus, 1758) References. Calathus melanocephalus: Csiki, 1940: 218 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke et al., 1986: 189 (Bizë near Shëngjergji; Lurë e Vjetër near Lura Mt.); Aukema, 1990: 138. Calathus melanocephalus melanocephalus: Hovorka & Sciaky, 2003b: 527. Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 5 s. (HNHM); 30 s. from localities “Shkëlzën”, “Bjeshka Matroxh”, “Tomor”, “Korab”, “Koritnik”, “Gjallica”, “Mal i Shenjit”, “Boge”, “Cucale”, “Taja”, “Llogora”, “Cafa e Sherit Iballe”, “Çafa Markofçes” (MNHUB); Tomor Mt., 21.IX.1961, 1 m., 1 f. (MNST). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona River, 900 m, 5.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. The mention of Hieke & Wrase (1988: 91) for Albania refers apparently to the record “Prisren” (Apfelbeck, 1904: 286). Actually this locality is presently situated in the Kosovo Region of Serbia. Besides the specimens above, I have seen several further specimens in NMW, which remained unregistered. 96.13. Calathus metallicus Dejean, 1828 96.13.1. Calathus metallicus prokletijensis Mařan, 1938 References. Calathus metallicus: Hieke & Wrase, 1988: 90. Calathus metallicus metallicus: Hovorka & Sciaky, 2003b: 527. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 15 s. (NMW); nearly 100 s. from localities “Shkëlzën”, “Çafa Markofçes”, “Maja e Jezerze” (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 m., 1 f., P. Beron & B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, 1000-1100 m, 5-9.VI.1993, 1 m., P. Beron & B. Petrov leg. (NMNHS). 170


Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 2 m., 1 f., L. Latella leg. (cAVT). Notes. First detailed records for Albania. The absence of C. metallicus in the massifs of Shar Planina and Korab Mt. may be a result of its replacement by C. albanicus there. 96.14. Calathus mollis (Marsham, 1802) 96.14.1. Calathus mollis mollis (Marsham, 1802) References.Calathus mollis: Apfelbeck, 1904: 285 (Vlorë; Berat); Apfelbeck, 1909: 297 (Vlorë; Berat); Hieke & Wrase, 1988: 91. Material studied. “Albania Avlona v. Oertzen”, 1 s. (MNHUB). Notes. Hovorka & Sciaky (2003b: 528) have omitted C. mollis for Albania. Here the species is confirmed for the country. C. mollis is exclusively a littoral species and most probably the record from Berat (Apfelbeck, 1904; 1909) concerns C. cinctus (A. Vigna Taglianti, personal communication).

97. Genus Synuchidius Apfelbeck, 1908 97.1. Synuchidius ganglbaueri Apfelbeck, 1908 References. Synuchidius ganglbaueri: Csiki, 1940: 218 (Gjalicë Mt.; Korab Mt.); Hieke, 1981: 82; Hovorka & Sciaky, 2003b: 529. Material studied. 66 s. from three kinds of labeling: “Albania Korab” / “Akad. Balk. Exp. Csiki 1917” (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918” (HNHM); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918” (HNHM); “Alban. Exped. Gjalica Ljums 17-26.VI.”, 12 s. (NMW); “Alban. Exped. Korab 1918 24-31..VII.”, 19 s. (NMW); “Jablanica coll. Lichtneckert”, 2 s. (HNHM); “Bjeshka Matroxh” / “Albania leg. Bischoff 1935”, 4 s. (MNHUB); “Shkalla Harushe” / “Korab” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB). Notes. Apfelbeck published three consecutive descriptions for the subgenus and species (Apfelbeck, 1908; Apfelbeck, 1917; Apfelbeck, 1930), and the oldest ones are correctly treated as valid.

Subtribe SPHODRINA Laporte, 1834 98. Genus Laemostenus Bonelli, 1810 Subgenus Antisphodrus Schaufuss, 1865 98.1. Laemostenus cavicola (Schaum, 1858) 98.1.1. Laemostenus cavicola albanicus J. Müller, 1914 References. Laemostenus cavicola albanicus: Müller, 1914: 1005-1006 (Çukall Mt.: cave “Scpedha n’ kunor Vukai”, loc. typ.; Çukall Mt.: cave “Scpedha Nre Gegs n’ gur t’barzodhs”, loc. typ.); Pretner, 1978: 135, 148 (caves in Çukall Mt.; near Shkodër); Genest & Juberthie, 1994: 593 (Çukall Mt.: cave “n’Kunor Vukai”, cave “Nre Gegs n’gur t’barzodhs”); Casale, 1988: 708, 716, 720 (Çukall Mt.); Casale, 2003: 534. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 7001400 m). Laemosthenes cavicola: Jeannel, 1937b: 89; Magistretti, 1965: 439. Laemostenus cavicola modestus: Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë). COLEOPTERA: CARABIDAE: HARPALINAE

171

Material studied. “Albanie”, 1 s. (MNHUB); “Albanien”, 1 s. (MNHUB); “Scpedha n’Kunor Zukali Albanien Lona Giuno 1914”, 1 syntype (DEI); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze Penther’14”, 12 s. (NMW); “Rapsa 10-18.V.” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. montenegr. Grenze Penther’14”, 3 s. (NMW); “Vermosa 20-21.VII.” / “Alban. montenegr. Grenze Penther’14”, 2 s. (NMW); “Albania Rapsa Dr. Grabowski”, 6 s. (HNHM); “Shpella Kodra Çarit” / “Cukale”, 3 s. (MNHUB); “Brigie Hot” / “leg. Bischoff Albania 1934” / “Shpella e Lugstit”, 1 s. (MNHUB). # Laemostenus cavicola modestus (Schaufuss, 1862) References. Laemostenus cavicola var. modestus: Apfelbeck, 1918c: 168 (Rapshë, 700-1400 m). Laemostenus cavicola modestus: Pretner, 1978: 135, 148, 168 (Rapshë); Casale, 1988: 719 (Rapshë). Notes. I follow Casale (2003: 534) who noted ssp. modestus only for Croatia and Bosnia Herzegovina. 98.2. Laemostenus elongatus (Dejean, 1828) 98.2.1. Laemostenus elongatus pentheri J. Müller, 1924 References. Laemostenus elongatus: Apfelbeck, 1907a: 496 (Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m). Laemostenus elongatus pentheri: Müller, 1924: 70-71 (North Albanian Alps: state attribution of loc. typ. in original description not specified; Rikavac; Vermosh; Rapshë); Pretner, 1978: 136, 148, 168 (Rikavac; Vermosh; Rapshë); Casale, 1988: 700, 704 (Okol near Theth: designated as loc. typ.; Maranaj; Vermosh, Rapshë). Material studied. 49 s. from localities “Maranaj”, “Shkëlzën”, “Maja e Jezercoë”, “Çafa Peis”, “Cucale”, “Boge”, “Maja e Dragomirit”, “Maja Rogamit” (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 4 s., P. Beron & B. Petrov leg. (NMNHS); Reshen District, Merkuth, 920-980 m, 11.VI.1933, 1 m., grassland, P. Beron & B. Petrov leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., Mali i Thatë Mt., 18.VII.1993, 1 f., L. Latella leg. (cAVT). Notes. Casale (2003: 534) has omitted the species for Albania. Here it is confirmed for the country. Subgenus Laemostenus s.str. 98.3. Laemostenus dalmatinus (Dejean, 1828) Material studied. “Rapsa 10-18.V.” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW). Notes. New species for the fauna of Albania. 98.4. Laemiostenus janthinus (Duftschmid, 1812) 98.4.1. Laemiostenus janthinus janthinus (Duftschmid, 1812) References. Laemiostenus janthinus: Csiki, 1940: 218 (Gjalicë Mt.). Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Notes. Casale (2003: 536) has omitted the species for Albania. Here it is confirms for the country. 98.5. Laemostenus venustus (Dejean, 1828) References. Laemostenus venustus: Apfelbeck, 1904: 275 (Velipoja); Casale, 1988: 476 (Velipoja); Hieke & Wrase, 1988: 93; Casale, 2003: 536. 172


Material studied. “Mamurash” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Mamurash” / “Albania leg. Bischoff 1933”, 2 s. (MNHUB); “Gjormi” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Pyla Shullazit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Sukthi bis Gurse” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB). Subgenus Pristonychus Dejean, 1828 98.6. Laemostenus cimmerius (Fischer von Waldheim, 1823) 98.6.1. Laemostenus cimmerius cimmerius (Fischer von Waldheim, 1823) References. Sphodrus cimmerius: Anonymous, 1907: 285. Prustonychus cimmerius: Magistretti, 1965: 433. Laemostenus cimmerius: Wrase, 1991: 11. Laemostenus cimmerius cimmerius: Casale, 1988: 809 (Durrës); Casale, 2003: 537. Material studied. Durrës, amphitheater, 12.VI.1993, 1 f., B. Petrov leg. (NMNHS); Tiranë Vore, 27.VI.1995, 1 m., S. Abadjiev leg. (NMNHS). 98.6.2. Laemostenus cimmerius weiratheri J. Müller, 1932 References. Laemostenus cimmerius weiratheri: Casale, 1988: 809 (M. Sopot); Sakalian & Guéorguiev, 1997: 48; Casale, 2003: 537. 98.7. Laemostenus terricola (Herbst, 1784) 98.7.1. Laemostenus terricola punctatus (Dejean, 1828) References. Pristonychus terricola: Hieke et al., 1986: 189 (Mali me Gropë). Laemostenus punctatus: Hieke & Wrase, 1988: 93. Laemostenus terricola punctatus: Casale, 1988: 787 (Mali me Grope, Dolinen-Gez); Casale, 2003: 539. Material studied. “Vermosa’14 Penther” / “Alban. montenegr. Grenze Penther’14”, 1 s. (NMW); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Tomor-Tojk juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”, 3 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 8 s. (MNHUB); “Vukaj” / “Cucale” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Maja e Policanit” / “Albania leg. Bischoff 1937”, 2 s. (MNHUB); “Mal i That” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); “Albanien – Exp. DEI Mali me Gropë, Dolinengebiet 1350 m, 6.VII.1961”, 1 s. (DEI).

Subtribe SYNUCHINA Lindroth, 1956 99. Genus Synuchus Gyllenhal, 1810 Subgenus Synuchus s.str. 99.1. Synuchus vivalis (Illiger, 1798) 99.1.1. Synuchus vivalis vivalis (Illiger, 1798) [= nivalis (Panzer, 1796)] References. Synuchus nivalis: Hieke & Wrase, 1988: 88. Synuchus vivalis vivalis: Hovorka & Sciaky, 2003c: 546.


173

Material studied. “Shkëlzën” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM). Notes. First detailed records for Albania.

Tribe ZABRINI Bonelli, 1810 Subtribe AMARINA C. Zimmermann, 1831 100. Genus Amara Bonelli, 1810 Subgenus Amara s.str. 100.1. Amara aenea (DeGeer, 1774) References. Amara aenea: Apfelbeck, 1904: 300 (Vlorë; Berat; Velipoja); Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Apfelbeck, 1909: 297 (Vlorë); Csiki, 1940: 215 (Shkëlzen Mt.; Kula Ljums; Ura i Lopez; “Stičen” near Gjalicë Mt.; Nangë; Gostilë); Schatzmayr, 1943b: 105 (Dukati; Tërpan); Hieke et al., 1986: 189 (Tomor Mt.: Ali Abbas Monastery; North Albanian Alps near Theth); Hieke & Wrase, 1988: 100; Hieke, 2003: 547. Material studied. Tiranë, 7.IV.1960, 1 s. (MNST); Ibë, 18.V.1960, 1 s. (MNST); Rodeni, 222.VII.1962, 4 s. (MNST); Zooteknica, 16.V.1963, 1 s. (MNST); Theth, 800-1200 m, 28.V.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS); North Albanian Alps, Bogë, cave No 25, 23.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS); Saranda Dhermi, 2.V.1994, 1 s., P. Stoev leg. (NMNHS); Saranda District, Ljukove, 3-4.VI.1994, 2 s., G. Blagoev leg. (NMNHS); 5 km Leskovic, 2.VII.1994, 1 s., P. Tenchev leg. (NMNHS); Petrela, 15 km NE Tiranë, 300 m, 9.V.1995, 2 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS); 5 km Saranda, 3.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Butrinti, 4.VI.1995, 1 s., G. Blagoev leg. (NMNHS); Lyukove, 5.VI.1995, 1 s., V. Sakalian leg. (NMNHS); Tiranë Park, 1-9.XI.1997 / 12-14.II.1998 / 16-21.II.1998 / 2328.II.1998, 56 s., R. Ward leg. (CMNH). Additional material included (studied A. Vigna Taglianti). Tropoia, valley of Valbona, 900 m, 5.VII.1993, 1 m., 1 f., L. Latella leg. (cAVT); Durrës, Plepa, 15.VIII.1995, 1 f., D. Antonelli leg. (cAVT); Rapsh, 15.V.2002, 2 m., 1 f., A. Paparistos leg. (cAVT). Notes. In addition to the material studied above, the present author has seen numerous further specimens in HNHM, NMW and MNHUB, which remained unregistered. 100.2. Amara anthobia A. Villa & G.B. Villa, 1833 References. Amara anthobia: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke & Wrase, 1988: 102; Hieke, 2003: 547. Material studied. “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Mal i Dajtit” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Mal i Krujes” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Cafa e Dhembellit” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); Vlorë District, Llogora Pass, 1050 m, 11.V.1995, 1 s., L. Penev leg. (NMNHS); Dhermi, 100 m, 11.V.1995, 1 s., S. Golovach, B. Petrov, P. Stoev leg. (NMNHS). 174


100.3. Amara convexior Stephens, 1828 References. Amara convexior: Hieke & Wrase, 1988: 97; Hieke, 2003: 548. Material studied. “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Notes. First detailed record for Albania. 100.4. Amara curta Dejean, 1828 References. Amara curta: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 215 (Ura i Lopez; Gjalicë Mt.; Korab Mt.); Hieke & Wrase, 1988: 98; Hieke, 2003: 548. Material studied. “Apfelb. Alban. spt.. IV.V.’05”, 3 s. (NMW); “Prokletjia 1300-2000” / “Alban. Montenegr. Grenze, Penther’14”, 2 s. (NMW); “Ricavac 1300 m 25.VI.-2.VII.” / “Alban. Montenegr. Grenze, Penther’14”, 1 s. (NMW); “Vermosa’14 Penther”, 2 s. (NMW); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 5 s. (MNHUB); “Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maranaj” / “Albania leg. Bischoff 1937”, 5 s. (MNHUB); North Albanian Alps, Bogë, 1800-1900 m, 20-23.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS); Shkoder, 12.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS). 100.5. Amara eurynota (Panzer, 1796) References. Amara eurynota: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke & Wrase, 1988: 101; Hieke, 2003: 549. Material studied. “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB); “Maja e Konispol Seraqin” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Murthi” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); Saranda District, Qeparo, 4-5.V.1994, 1 s., P. Stoev leg. (NMNHS). ? Amara famelica C. Zimmermann, 1832 Material studied. “Alban. Exp. 1918 Gjalica Ljums 17-26.VI.”, 7 s. (NMW); “Alban. Exp. 1918 Korab, 23-31.VII.”, 1 s. (NMW). Notes. F. Hieke (personal communication) expressed doubt about the correct identification of the specimens in NMW. 100.6. Amara familiaris (Duftschmid, 1812) References. Amara familiaris: Apfelbeck, 1904: 301 (Vlorë); Apfelbeck, 1909: 297 (Vlorë); Hieke & Wrase, 1988: 101; Hieke, 2003: 549. Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 2 s. (NMW); “Logara” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1936”, 2 s. (MNHUB).


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100.7. Amara lucida (Duftschmid, 1812) References. Amara lucida: Apfelbeck, 1904: 301 (Berat); Apfelbeck, 1909: 297 (Berat); Csiki, 1940: 215 (Ura i Lopez; Korab Mt.); Hieke & Wrase, 1988: 102; Hieke, 2003: 549. Material studied. “Merdita M. Scheit” / “Apfelb. Alban. spt. IV.V.’05”, 3 s. (NMW); “Albania Korab” / “Akad. Balk. Exp. Csiki 1917”, 1 s. (HNHM); “Albania Ura i Lopez” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Krabe” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1932”, 3 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 2 s. (MNHUB); “Tomor-Burany juli 1932”, 3 s. (MNHUB); “Berat-Bargulash” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Mal i Dajtit” / “leg. Bischoff Albania 1934”, 4 s. (MNHUB); “Mal i Shejit” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB); “Tomor” / “leg. Bischoff Albania 1934”, 2 s. (MNHUB); “Logara” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Bojana Shkoder” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB). 100.8. Amara morio Ménétriés, 1832 100.8.1. Amara morio nivium Tschitschérine, 1900 References. Amara morio nivium: Hieke, 1995: 288 (Gjalicë Ljums); Hieke, 2003: 550. Material studied. “Albania Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 2 s. (HNHM; MNHUB). 100.9. Amara ovata (Fabricius, 1792) References. Amara ovata: Hieke, 2003: 550. Material studied. “Tomor” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Llogora” / “leg. Bischoff Albania 1936”, 2 s. (MNHUB). Additional material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (MCG); “Lona, Albania”, 1 s. (MSNM). Notes. The mention of Hieke & Wrase (1988: 96) for Albania refers apparently to the record “Prisren” (Apfelbeck, 1904: 299). However, this locality is situated in the Kosovo Region of Serbia. Besides the material studied above, I have seen more specimens in NMW, which remained unregistered. First detailed records for Albania. 100.10. Amara proxima Putzeys, 1866 References. Amara proxima: Hieke & Wrase, 1988: 98; Hieke, 2003: 551. Material studied. “Shendeli Alban. Mer.” / “Winkler. Lona, Bischoff, V.1931”, 6 s. (NMW; MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1936”, 3 s. (MNHUB); “Maja e Tartarit juni 1937” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Additional material included (studied F. Hieke). “Argiro Kastron, S.Alban”, 2 s. (MCG; MTKD); “Ismen Ili, Albania”, 1 s. (NMPC). Notes. First detailed records for Albania.

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100.11. Amara saphyrea Dejean, 1828 References. Amara saphyrea: Hieke & Wrase, 1988: 96; Hieke, 2003: 551. Material included (studied F. Hieke). “Tomor, Kulmak, Alban. mer.”, 1 s. (NMW). Notes. First detailed record for Albania. 100.12. Amara similata (Gyllenhal, 1810) Material studied. Tiranë Park, 1.III.1998, 1 s., shaded lake edge under leaves and wet, rotten wood, R. Ward leg. (CMNH). Notes. New species for the fauna of Albania. # Amara spreta Dejean, 1831 References. Amara spreta: Csiki, 1940: 215 (Gjalicë Mt.). Notes. Basing on the most recent information about the species distribution (Hieke, 2003: 551), the Csiki’s identification is most probably incorrect and that the material in question concerns another congener. F. Hieke (personal communication) expressed similar opinion, so A. spreta is removed from the list of the Albanian Carabidae. 100.13. Amara tibialis (Paykull, 1798) References. Amara tibialis: Csiki, 1940: 215 (Korab Mt.); Hieke & Wrase, 1988: 103; Hieke, 2003: 551. Material studied. “Albania Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM). Subgenus Amarocelia Motschulsky, 1862 100.14. Amara erratica (Duftschmid, 1812) References. Amara erratica: Csiki, 1940: 216 (Gjalicë Mt.); Hieke & Wrase, 1988: 103; Hieke, 2003: 556. Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 1 s. (HNHM); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Skëlzën” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB). Subgenus Bradytus Stephens, 1827 100.15. Amara apricaria (Paykull, 1790) References. Amara apricaria: Hieke et al., 1986: 190 (Borshi near Vlorë; Bizë near Shëngjergji); Hieke & Wrase, 1988: 106; Hieke, 2003: 553. Material studied. “Vermosa’14 Penther”, 1 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Alban. Exp.’18 Pashtrik, 4-14.7.”, 3 s. (NMW); 28 s. from localities “Shkalle Harushe-Korab”, “Korab”, “Shkëlzën”, “Mal i Sheniit”, “Mal i Tartarit”, “Taja”, “Giallica” (MNHUB); “Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 2 s. (HNHM).


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100.16. Amara consularis (Duftschmid, 1812) Material studied. Mal i Dajtit Mt.: Shkall Priskë, 26-30.VI.1961, 1 s. (MNST). Notes. New species for the fauna of Albania. Subgenus Celia C. Zimmermann, 1832 100.17. Amara bifrons (Gyllenhal, 1810) References. Amara bifrons: Csiki, 1940: 216 (Korab Mt.); Hieke & Wrase, 1988: 105; Hieke, 2003: 555. Material studied. “Ricavac 1300 25.VI.-2.VII.” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 4 s. (HNHM); “Alban. Exp. 1918 Korab 23-31.VII.”, 4 s. (NMW); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Derzë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Albania Theth” / “23.VII.1960 leg. Xh. Murraj”, 1 s. (HNHM). Additional material included (studied A. Vigna Taglianti). Tropoia, Valbona, Kollatës Mt., 2250 m, 11.VII.1993, 4 f., L. Latella leg. (cAVT). 100.18. Amara hannemanni Hieke, 1991 References. Amara hannemanni: Hieke, 1991: 1-3 (Shkëlzen Mt., loc. typ.); Hieke, 2003: 556. Material studied. “Skëlzën” / “Albania leg. Bischoff 1935”, 1 paratype (MNHUB); “Skëlzën” / “Albania leg. Bischoff 1936”, 6 paratypes (MNHUB). 100.19. Amara montana Dejean, 1828 References. Amara montana: Hieke, 2003: 557. Material studied. “Skutari, Alb. 4.X.18 Dr. H. Priesner”, 1 s. (NMW); “Albania Shkoder” / “25.V.1958 leg. Xh. Murraj”, 2 s. (HNHM); Vlorë, 1.V.1994, 1 s., P. Stoev leg. (NMNHS); Tiranë Park, 1-9.XI.1997, 1 s., R. Ward leg. (CMNH). Notes. First detailed records for Albania. ? Amara praetermissa (C.R. Sahlberg, 1827) References. Amara praetermissa: Hieke & Wrase, 1988: 105. Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 2 s. (HNHM); “Latif Koritnik”, 3 s. (NMW). Notes. Hieke (2003: 557) has omitted the species for Albania in spite of his previous data (Hieke & Wrase, 1988: 105). He stated me (personal communication) that the specimens above might belong to A. hannemanni, so that A. praetermissa is treated here as doubtful for Albania. Subenus Curtonotus Stephens, 1827 100.20. Amara aulica (Panzer, 1796) References. Amara aulica: Hieke & Wrase, 1988: 109; Hieke, 2003: 559. 178


Material studied. “Tomor Albanien”, 1 s. (MNHUB); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tomor-Buranj juli 1932” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB). Notes. First detailed records for Albania. Subgenus Paracelia Bedel, 1899 100.21. Amara dalmatina Dejean, 1828 References. Amara dalmatina: Hieke (in press). Material included (studied F. Hieke). “SE of Medua, Albania”, 1 s. (MNHUB); “Shendeli, Albania mer.”, 1 s. (MNHUB). Notes. First detailed records for Albania. 100.22. Amara quenseli (Schönherr, 1806) 100.22.1. Amara quenseli quenseli (Schönherr, 1806) References. Amara quenseli: Apfelbeck, 1907a: 497, 499 (Maranajt Mt.: top region, 1500-1576 m; Mal i Shenjt Mt.: top region, 1500 m); Csiki, 1940: 216 (Korab Mt.); Holdhaus & Lindroth, 1940: 146-147, 250 (Koritnik Mt.; Korab Mt.); Hieke & Wrase, 1988: 106. Amara quenseli quenseli: Hieke, 2003: 564. Material studied. “Latif Koritnik” / “Coll. Apfelbeck”, 2 s. (HNHM); “Latif Koritnik”, 8 s. (NMW); “Merdita Munela” / “Apfelb. Alban. spt. IV.V.’05”, 1 s. (NMW); “Prokletija 13002000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 1 s. (NMW); “Albania Galica Lums” / “Akad. Balk. Exp. Csiki 1917”, 2 s. (HNHM); “Albania, Mts. Gyalica Lums” / “Akad. Balk. Exp. Csiki 1918”, 3 s. (HNHM); “Korab” / “Albania leg. Bischoff 1935”, 16 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1936”, 19 s. (MNHUB); “Shkëlzën” / “Albania leg. Bischoff 1936”, 23 s. (MNHUB). Subgenus Parapercosia Tschitschérine, 1899 100.23. Amara taurica (Motschulsky, 1844) [= fallaciosa (J. Müller, 1946)] References. Zabrus fallaciosus: Müller, 1946: 107-110 (South Albania: Shëndelli Mt. near Tepelenë, loc. typ.); Turin, 1981: 148. Amara taurica: Hieke, 2003: 565. Notes. Hieke (1995: 35-36) synonymized Zabrus fallaciosus Müller, 1946 with Amara taurica, noting the type locality of the former as “Patria falsa!”, but later the same has cited A. taurica for Albania (Hieke, 2003: 565). According to F. Hieke (personal communication) A. taurica is known from different parts of Turkey and from two localities in Greece, so that the type locality of Zabrus fallaciosus in Albania should not be considered as doubtful (Hieke, 2005). Subgenus Percosia C. Zimmermann, 1832 100.24. Amara equestris (Duftschmid, 1812) 100.24.1. Amara equestris equestris (Duftschmid, 1812) [= tomoricensis J. Müller, 1923] COLEOPTERA: CARABIDAE: HARPALINAE

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References. Amara tomoricensis: Müller, 1923: 108-109 (Tomor Mt.: Kulmak, loc. typ.). Amara equestris: Csiki, 1940: 216 (Korab Mt.); Hieke, 1978: 229, 257 (Tomor Mt.: Kulmak; Koritnik Mt.; Korab Mt.); Hieke & Wrase, 1988: 108. Amara equestris equestris: Hieke, 2003: 565. Material studied. “Albania Mts. Korab” / “Akad. Balk. Exp. Csiki 1918”, 1 s. (HNHM); “Shen Ili” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Korab” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Koritnik” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Albania Zalle-Dardha” / “24.VII.1958 leg. Xh. Murraj”, 1 s. (HNHM). Subgenus Xenocelia Hieke, 2001 100.25. Amara bischoffi Jedlička, 1946 References. Amara municipalis: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.); Jedlička, 1946: 7 Peshtrik Mt. (loc. typ.); Hieke & Wrase, 1988: 104. Amara bischoffi: Turin, 1981: 145; Hieke, 1983: 328-329, 334 (Peshtrik Mt.; Taja); Hieke, 2001: 33 (two localities). Amara municipalis bischoffi: Hieke, 2003: 557. Material studied. “Taja” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Bashtrik” / “Albania leg. Bischoff 1936”, 1 syntype (MNHUB). Notes. According to F. Hieke (personal communication) the records of Csiki (1940) are problematic since they may concern A. bischoffi, as well as A. messae. 100.26. Amara cursitans C. Zimmermann, 1832 [= properans C. Zimmermann, 1832] References. Amara properans: Csiki, 1940: 215 (Gjalicë Mt.; Korab Mt.). Amara cursitans: Hieke & Wrase, 1988: 104; Hieke, 2001: 50 (five localities); Hieke, 2003: 556. Material studied. “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Skëlzën” / “Albania leg. Bischoff 1935”, 2 s. (MNHUB); “Gjallica” / “Albania leg. Bischoff 1936”, 5 s. (MNHUB); “Llogora” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); “Maja e Dragomit” / “Albania leg. Bischoff 1939”, 1 s. (MNHUB). 100.27. Amara fusca Dejean, 1828 [= complanata Dejean, 1828] References. Amara complanata: Csiki, 1940: 215 (Gjalicë Mt.). Amara fusca: Hieke & Wrase, 1988: 104; Hieke, 2001: 42 (“Koritza-Ebene”, 1 s. in MNHN); Hieke, 2003: 556. 100.28. Amara messae Baliani, 1924 References. Amara messae: Hieke & Wrase, 1988: 104; Hieke, 2001: 65; Hieke, 2003: 557. Material studied. “Giallica” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Maja e Jezercoë” / “Albania leg. Bischoff 1935”, 1 s. (MNHUB); “Cotype” / “Bashtrik” / “Albania leg. Bischoff 1936”, 1 s. (MNHUB); North Albanian Alps, Bogë, 1400-1800 m, 1.VI.1993, 1 s., P. Beron, B. Petrov leg. (NMNHS). Notes. First detailed records for Albania.

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Subgenus Zezea Csiki, 1929 100.29. Amara chaudoiri Schaum, 1858 100.29.1. Amara chaudoiri incognita Fassati, 1946 References. Amara chaudoiri incognita: Hieke, 1970: 138 (Skrofotino near Vlorë); Hieke, 2003: 567. Amara chaudoiri: Hieke & Wrase, 1988: 94. # Amara erythrocnema Dejean, 1828 References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina). Notes. At present the species is treated as local endemic for the island of Corfu (Hieke, 2003: 567). F. Hieke (personal communication) expressed the opinion that the record for A. erythrocnema from Albania most probably concerns A. reflexicollis. Therefore, the former species is removed from the list of Albanian Carabidae. 100.30. Amara fulvipes (Audinet-Serville, 1821) References. Amara fulvipes: Apfelbeck, 1904: 298 (Velipoja); Hieke, 1970: 157 (Velipoja; Lum; Tiranës; Lum i Tiranës; Mal i Dajtit Mt.); Hieke & Wrase, 1988: 94; Hieke, 2003: 567. Material studied. “Lum i Tiranes” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB); “Tirane” / “leg. Bischoff Albania 1934”, 6 s. (MNHUB). 100.31. Amara reflexicollis Motschulsky, 1844 100.31.1. Amara reflexicollis reflexicollis Motschulsky, 1844 References. Amara erythrocnema: Schatzmayr, 1943b: 105 (Skrofotina). Amara reflexicollis reflexicollis: Hieke, 2003: 568. 100.32. Amara tricuspidata Dejean, 1831 References. Amara tricuspidata: Hieke & Wrase, 1988: 94; Hieke, 2003: 568. Material studied. “Museum Paris Albanie Starova (env. de Koritza) Caporal Vuillaume 1918” / “Albanie Armée d’Orient 1916-1918” / “Avril” / “Mai”, 1 s. (MNHUB). Additional material included (studied F. Hieke). 1 s. labelled as that up above (MNHP). Notes. First detailed record for Albania.

Subtribe ZABRINA Bonelli, 1810 101. Genus Zabrus Clairville, 1806 Subgenus Pelor Bonelli, 1810 Notes. The species from this subgenus are arranged here in three groups of species with the aim to delineate the related lineages.


181

Species group “aetolus” 101.1. Zabrus aetolus Schaum, 1864 [Zabrus aetolus kodymi Mařan, 1940] Notes. This race has been described from the Montenegrin part of the North Albanian Alps (Mařan, 1940a: 33), basing on the label data of the material collected by O. Kodym and C. Purkynì in 1937. The ssp. kodymi may inhabit the Albanian part of the mountain, too. 101.1.1. Zabrus aetolus winkleri J. Müller, 1946 References. Zabrus winkleri: Müller, 1946: 110-111 (Maja e Tartarit Mt., south of Vlorë, loc. typ.; Maja e Djembellit Mt., east of Gjirokastër; Maja e Policanit Mt., east of Gjirokastër); Turin, 1981: 148. Zabrus aetolus winkleri: Freude, 1989: 83, 95 (Maja e Tartarit Mt.; Maja e Dhembellit Mt.; Maja e Policanit Mt.); Serrano & Andújar, 2003: 570. Material studied. “Argirokastron Sud-Albania”, 1 s. (MNHUB); “Maja e Tartarit” / “Albania leg. Bischoff 1932”, 10 s. (MNHUB). 101.2. Zabrus albanicus Apfelbeck, 1904 101.2.1. Zabrus albanicus albanicus Apfelbeck, 1904 References. Zabrus albanicus latifianus: Ganglbauer, 1931: 39 (Koritnik Mt.); Mařan, 1939d: 145-146 (Koritnik Mt.); Schatzmayr, 1943b: 108 (Koritnik Mt.); Freude, 1989: 83, 88, 97 (Koritnik Mt.); Drovenik & Peks, 1994: 96 (Koritnik Mt.); Drovenik & Peks, 1999: 111 (Koritnik Mt.). Zabrus albanicus: Csiki, 1940: 221 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Schatzmayr, 1943b: 108 (Shkëlzen Mt.; Gjalicë Mt.; Korab Mt.); Hieke, 1981: 83. Zabrus albanicus albanicus: Serrano & Andújar, 2003: 570. Material studied. “Latif Koritnik” / “albanicus typ. Apfelb.”, 12 s. (NMW); “Latif Alban. spt.”, 3 s. (DEI); more than 140 s. from localities “Albania Galica Lums” (or “Gyalica Lums”), “Skölsen” and “Korab” (HNHM); “Prokletija 1300-2000 m” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Alban. Exped. Pashtrik 1918 29.V.-4.VI.”, 2 s. (NMW); “Alban. Exped. 1918 Gjalica Ljums 17-26.VI.”, 20 s. (NMW); “Alban. Exped. Korab 1918 24-31.VII.”, 71 s. (NMW). Notes. The populations from the Koritnik Mt., as well as the records for these (Ganglbauer, 1931; Mařan, 1939d; Schatzmayr, 1943b; Freude, 1989; Drovenik & Peks, 1994; 1999), are assigned provisionally to the nominotypical subspecies rather than to the ssp. latifianus, since the Zepë Mt. (an unmarked, but potential “loc. typ.” of the latter taxon) has an more southern and well-isolated position from the other massifs in North Albania. Besides the material studied above, in MNHUB the present author has seen other more 140 s. of Z. albanicus s.l. from different localities, which remained unregistered. [Zabrus albanicus jablanicensis Mařan, 1939] References. Zabrus albanicus jablanicensis: Freude, 1989: 88 (“Mittel-Ost-Albanien an der jugoslavischen Grenze”). Notes. Serrano & Andújar (2003: 570) indicated Bosnia Herzegovina as region of the distribution of ssp. jablanicensis by mistake. In fact, this race was described from the Jablanitsa Mt. shared between Albania and Macedonia and so far is cited only from the Macedonian part of the mountain. Most likely the record of Freude (1989) is not based on material coming 182


from collections, and may bases of supposition. The subspecies most probably inhabits the Albanian part of the Jablanitsa Mt., too. 101.2.2. Zabrus albanicus latifianus Ganglbauer, 1915 References. Zabrus albanicus: Apfelbeck, 1907a: 503 (Zepë Mt.). Zabrus albanicus latifianus: Ganglbauer, 1931: 39 (Zepë Mt.); Mařan, 1939d: 145-146 (Zepë Mt.); Schatzmayr, 1943b: 108 (Zepë Mt.); Freude, 1989: 83, 88, 97 (Zepë Mt.); Serrano & Andújar, 2003: 570. Material studied. “Merdita Munela”, 1 s. (NMW); “Merdita Zebia”, 2 s. (NMW). Notes. In my opinion, only the populations from the region of Merdita have to be considered as belonging to spp. latifianus. ! Zabrus albanicus ssp. Zabrus albanicus: Schatzmayr, 1943b: 108 (Tomor Mt.: Abbas Ali and Toik). Material studied. “Tomorica Abbas Ali” / “Albania 1922 Ravasini, Lona”, 4 s. (NMW; MNHUB); “Maja e Dhembelit” / “Albania 1936 leg. Ramadan coll. Bischoff ”, 8 s. (MNHUB); “Albania 1936 leg. Ramadan coll. Bischoff ” / “Maja e Papingut”, 21 s. (MNHUB). Notes. The specimens from the massifs of Tomor Mt., Maja e Dhembelit Mt. and Maja e Papingut Mt. (all in South Albania) may concern another subspecies, since the latter are far removed in the south from the massifs of the Zepë Mt. and Koritnik Mt. # Zabrus apfelbecki Ganglbauer, 1915 References. Zabrus apfelbecki: Magistretti, 1965: 360. Notes. Magistretti (1965) referred older records of Z. ignavus Csiki, 1907 to Z. apfelbecki, but I failed to find any records in the literature. The latter species lives only in Bosnia Herzegovina (Serrano & Andújar, 2003: 570), so that it is removed from the list of the Albanian ground beetles. 101.3. Zabrus ganglbaueri Apfelbeck, 1906 101.3.1. Zabrus ganglbaueri ganglbaueri Apfelbeck, 1906 References. Zabrus ganglbaueri: Apfelbeck, 1906: 1667 (Merdita: Mal i Shenjt Mt. near Oroshi, loc. typ.); Apfelbeck, 1907a: 499-500, 503 (Mal i Shenjt Mt.: top region, 1500 m); Müller, 1936: 108 (Mal i Shenjt Mt. near Oroshi); Schatzmayr, 1943b: 108 (Mal i Shenjt Mt. near Oroshi); Hieke, 1981: 75. Zabrus ganglbaueri ganglbaueri: Freude, 1989: 82, 110 (Mal i Shenjt Mt. near Oroshi; Mal i That Mt. near Gjirokastër); Serrano & Andújar, 2003: 570. Material studied. “Merdita M. Scheit”, 1 s. (NMW); “Merdita 18.19. n. 20/5 1934”, 2 s. (NMW). Notes. Freude (1989) stated that the populations from the Mal i Shenjt Mt. (Merdita District) and the Mal i That Mt. (Gjirokastër District) belong to the nominotypical race. In fact both populations inhabit opposite outskirts of the country, so the specimens from the latter massif may belong either to the following subspecies or to a separate geographical race. 101.3.2. Zabrus ganglbaueri lonae J. Müller, 1923 References. Zabrus lonae: Müller, 1923: 107-108 (Tomor Mt.: Toik, loc. typ.); Müller, 1935: 154 (Tomor Mt.: Toik); Mařan, 1940a: 30-31, 33, 35. Zabrus lonai: Schatzmayr, 1943b: 108 (Tomor Mt.: Toik). Zabrus ganglbaueri lonae: Freude, 1989: 83, 111 (Tomor Mt.: Toik); Serrano & Andújar, 2003: 570. COLEOPTERA: CARABIDAE: HARPALINAE

183

Material studied. “Albania 1922 Ravasini, Lona” / “Tomor Monte Toik”, 1 s. (DEI); “Tomor: Toik Alban. mer.” / “Winkler, Lona, Bischoff, V.1931”, 1 s. (NMW); “Tomor: Tojk” / “Albania leg. Bischoff 1932”, 9 s. (MNHUB). Species group “incrassatus” 101.4. Zabrus bischoffi J. Müller, 1936 References. Zabrus bischoffi: Müller, 1936: 107-108 (Mal i Dajtit Mt. near Tiranë, loc. typ.; Llogora); Schatzmayr, 1943b: 106, 109 (Kruja; Mal i Dajtit Mt.; Llogora); Turin, 1981: 148. Zabrus incrassatus bischoffi: Freude, 1989: 86, 120 (Mal i Dajtit Mt. near Tiranë; Llogara, south of Vlorë); Serrano & Andújar, 2003: 571. Material studied. 19 s. from “Mal i Dajtit” (MNHUB); 5 s. from “Mal i Krues” (MNHUB); 3 s. from “Mal i Shenjit” (MNHUB); 37 s. from “Llogora” (MNHUB); Llogora Pass, 1025 m, 6.6.1995, 1 m., G. Blagoev leg. (NMNHS). Notes. This species has a quite different apex of the median lobe and sides of pronotum narrower anteriorly, which is in contrast with the respective characteristics in Z. incrassatus s.l. In my opinion, the former is separate species not subspecies of the latter: Zabrus (Pelor) bischoffi J. Müller, 1936, stat. rest. of Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936. 101.5. Zabrus boldorii Schatzmayr, 1943 References. Zabrus boldori: Schatzmayr, 1943: 106, 108 (Dukati, south of Vlorë, loc. typ.); Serrano & Andújar, 2003: 570. Zabrus baldori sic!: Turin, 1981: 148. Zabrus incrassatus boldorii: Freude, 1989: 86, 119 (Dukati near Vlorë). Material studied. “Südalbanien Kopstein” / “coll. Prof. F. Werner” / “Pašaliman”, 1 m. (NMW); “Südalbanien Kopstein” / “Valona”, 2 f. (NMW). 101.6. Zabrus incrassatus (Ahrens, 1814) 101.6.1. Zabrus incrassatus incrassatus (Ahrens, 1814) References. Zabrus incrassatus: Apfelbeck, 1904: 310 (Berat; Vlorë); Anonymous, 1907: 285; Apfelbeck, 1907a: 495-496 (northeast of Shkodër; Maranajt Mt.: between Vorfai Postme, 300 m, and Vorfai Siperme, 700 m); Apfelbeck, 1909: 297 (Vlorë; Berat); Müller, 1936: 108 (Shëndelli Mt.; Tepelenë; Maja e Tartarit Mt.; Tiranë; Gjirokastër; Berat; Vlorë); Schatzmayr, 1943b: 106, 108 (“Poggio Boschetto”; Lushnje; Berat; Vlorë; Shen Ili; Tepelenë; Maja e Tartarit Mt.; Tiranë; Shkodër); Hieke & Wrase, 1988: 110. Material studied. “Südalbanien Kopstein”, 2 s. (NMW); “Albanien Ochrida”, 1 s. (NMW); “Hani Grabom Mai” / “Alban. montenegr. Grenze, Penther’14”, 2 s. (NMW); “Rapsa 1018.V.” / “Alban. montenegr. Grenze, Penther’14”, 4 s. (NMW); “Alban. Exp. 1918 Bicaj 14.15.VI.”, 3 s. (NMW); Ibë, 2-11.VII. 1962, 1 m. (MNST); Shkodër, 15.V.1963, 3 m., 1 f. (MNST); 2 km N Durrës, 24.V.1993, 1 m., P. Stoev leg. (NMNHS); Vlorë, 1.V.1994, 2 m., 3 f., P. Stoev leg. (NMNHS); Saranda District, Himarë, 3-4.V.1994, 1 f., P. Stoev leg. (NMNHS); Saranda District, Borsh, 5.V.1994, 1 f., P. Stoev leg. (NMNHS); Saranda District, Ljukova, 34.VI.1995, 1 f., G. Blagoev leg. (NMNHS); Vlorë District, Llogora Mt., 1027 m, 4.VI.1995, 1 f., G. Blagoev leg. (NMNHS). 184


Notes. Serrano & Andújar (2003: 571) have omitted the species for Albania, so it is here confirmed for the country. Besides the material studied above, in MNHUB I have seen other more than 100 s. from different localities (“Shen Ili”, “Tirane”, “Murthi”, “Tepelene”, “Munela”, “Burelle”, “Vlorë”, “Mal i Gjer”, “Myzege”, “Bose”, “Maja e Dhembellit”, “Tarabosh”, “Mal i Tartarit”, etc.). Species group “spinipes” 101.7. Zabrus spinipes (Fabricius, 1798) 101.7.1. Zabrus spinipes insignis J. Müller, 1932 References. Zabrus spinipes: Burmeister, 1939: 127; Hieke et al., 1986: 190 (Tomor Mt.: Abbas Ali Monastery); Hieke & Wrase, 1988: 110; Freude, 1989: 147. Material studied. “Shen-Ili Juni” / “Albania leg. Bischoff 1932”, 11 s. (MNHUB); “Mal i That” / “Albania leg. Bischoff 1937”, 22 s. (MNHUB). Notes. Serrano & Andújar (2003: 572) have omitted the species for Albania, so it is here confirmed for the country. Subgenus Zabrus s.str. 101.8. Zabrus tenebrioides (Goeze, 1777) 101.8.1. Zabrus tenebrioides tenebrioides (Goeze, 1777) References. Zabrus tenebrioides: Schatzmayr, 1943b: 105-106 (Dukati; Lushnje). Zabrus tenebrioides tenebrioides: Serrano & Andújar, 2003: 573. Material studied. “Myzeqe” / “Albania leg. Bischoff 1937”, 1 s. (MNHUB); 11 km west of Tiranë, 20.VI.1994, 1 s., B. Guéorguiev leg. (NMNHS); Radona, Shkiela, 21.VI.1994, 5 s., B. Guéorguiev leg. (NMNHS). Additional material included (studied A. Vigna Taglianti). Durrës, Currila, 8.V.2002, 1 f., A. Paparistos leg. (cAVT); Qesarat near Memaliaj (Tepelenë District), valley of Vijosë River, 300360 m, 18.IV.2003, 1 f., P. Crucitti & C. Cavalieri leg. (cAVT).

Tribe ZUPHIINI Bonelli, 1810 Subtribe ZUPHIINA Bonelli, 1810 102. Genus Polystichus Bonelli, 1810 102.1. Polystichus connexus (Geoffroy, 1785) References. Polystichus connexus: Hieke & Wrase, 1988: 163. Material studied. “Liqen i Butrintes Vivari” / “leg. Bischoff Albania 1934”, 1 s. (MNHUB). Notes. Baehr (2003e: 574) has omitted the species for Albania. Here it is confirmed for the country as first detailed record is given.


185

103. Genus Zuphium Latreille, 1806 Subgenus Zuphium s.str. 103.1. Zuphium olens (P. Rossi, 1790) 103.1.1. Zuphium olens olens (P. Rossi, 1790) References. Zuphium olens: Schatzmayr, 1943b: 112 (Stan-Karbunarë); Hieke & Wrase, 1988: 162. Zuphium olens olens: Baehr, 2003e: 574. Material studied. “Durres” / “Albania leg. Bischoff 1932”, 1 s. (MNHUB).

186


Results When omitting doubtful species and taxa still not well defined, the Albanian Carabidae listed in this catalogue represent 543 species. They are distributed among 11 subfamilies (Nebriinae – 25, Cicindelinae – 10, Carabinae – 20, Elaphrinae – 1, Omophroninae – 1, Brachininae – 17, Siagoninae – 1, Scaritinae – 28, Apotominae – 2, Trechinae – 115, and Harpalinae – 323), 31 tribes and 103 genera (Table 1). One tribe, 10 genera, 11 subgenera and 55 species are new for the fauna of Albania, and 1 subgenus and 3 species are new for the fauna of the Balkan Peninsula (Appendix A.1-2). From these, 1 tribe, 1 subtribe, 7 genera, 20 subgenera and 72 species are reported firstly with detailed record/s for the country (Appendix A.3). Altogether 119 species, omitted in the recent Palaearctic Catalogue (Löbl & Smetana, 2003), are here confirmed for Albania (Appendix A.4). Twelve taxa from the species group remain still not well defined, as likely most of them represent new for the science species or subspecies (Appendix A.5). Two subgenera and 22 species and subspecies are doubtful for the country (Appendix A.6); generally the published records about them concern adjacent regions in Montenegro, the Kosovo Region of Serbia, Macedonia or Northwestern Greece. Most of the species and subspecies from the last category may inhabit Albania, but until now we do not have reliable data to confirm this. Two subgenera and 29 species and subspecies are deleted from the list (Appendix A.7). The systematic list includes also some endemics described from massifs shared between Albania and neighbour countries (but not recorded yet from the former), as well as several more widespread species found from localities very close to Albania. A moderate taxonomic contribution is also achieved (Appendix B). Principally, it is based on study of types, topotypes or more extensive series of previously problematic species and subspecies from Albania and neighbouring countries. In result of that 38 various taxonomic innovations (new synonyms, new status or revalidations) are introduced. Further, nine taxa from the species group (5 valid species and 4 synonyms), all described by V. Apfelbeck, have been found to have assigned an incorrect year of description. Accordingly, the relevant earlier years of their descriptions are fixed. Finally, eight nomina nuda were also traced out, as one of them may be a result of mislabeling, and one is supplied with year of description in the literature.

RESULTS

187

Conclusions Undoubtedly the Balkan Peninsula contains the richest ground beetle diversity compared to other diverse European regions like the Iberian Peninsula, Italy or the Caucasus Major (Turin, 1981; Kryzhanovskij et al., 1995; Serrano, 2003; Vigna Taglianti, 1993; 2004; 2005a; 2005b). Currently an impressive total of at least 1487 species of Carabidae, following the classification used in Löbl & Smetana, 2003, is reported from this territory (unpublished data). This figure is provisional while the systematics, diversity and distribution of the Carabidae in the Balkan Peninsula are better known. Until now, the carabid fauna of Bulgaria is probably the best investigated one in the region (Hieke & Wrase, 1988; Guéorguiev & Guéorguiev, 1995). The species composition of no other Balkan regions has been examined in details; only that of ex-Yugoslavia was subject of uncritical registration (Drovenik & Peks, 1994; Drovenik & Peks, 1999). Therefore, the Carabidae of Slovenia, Croatia, Bosnia Herzegovina, Montenegro, Serbia, Macedonia, Romania, Greece and European Turkey need scrutiny and writing of up-to-date checklists. Bearing in mind the surface of Albania, the number of the carabid species and subspecies there seems relatively high (543), if compared it with that of other Balkan or Mediterranean regions. In Bulgaria there are known more than 720 species (Guéorguiev & Guéorguiev, 1995), from Macedonia there are reported about 420 species (Guéorguiev, 1998; Hristovski, personal communication), and for ex-Yugoslavia more than 1500 species and subspecies have been cited (Drovenik & Peks, 1999). Both the Iberian Peninsula and Italy, having territories from 10 to 20 times bigger than Albania, have a number of species only 2.13-2.42 times higher than is that in the last country (Serrano, 2003; Vigna Taglianti, 2005a; 2005b).

188


Taxonomic structure of Albanian Carabidae (Table 1) Subfamily

Tribe

Nebriinae

Nebriini

Cicindelinae

Carabinae

Subtribe

Apotominae Trechinae

Nr. species

Eurynebria Leistus

1 7

Nebria

9

Notiophilus Calomera Cicindela Cylindera

8 2 4 3

Myriochila Calosoma

1 4

Carabus

15

Cychrus Elaphrus Omophron Aptinus Brachinus

1 1 1 3 14

Clivinina Reicheiina Dyschiriini

Siagona Clivina Reicheadella Dyschirius

1 3 1 21

Scaritini

Scaritina

Scarites

3

Apotomini Bembidiini

Bembidiina

Apotomus Asaphidion Bembidion

2 4 63

Notiophilini Cicindelini

Carabini

Cychrini Elaphrinae Elaphrini Omophroninae Omophronini Brachininae Brachinini

Siagoninae Scaritinae

Genus

Siagonini Clivinini

Subgenus

Nr. species

Leistus Pogonophorus Alpaeus Boreonebria Eunebria Nebria

2 5 5 1 1 2

Cicindela Cylindera Eugrapha Myriochila Calosoma Microcallisthenes Archicarabus Carabus Chaetocarabus Megodontus Oreocarabus Pachystus Procerus Procrustes Tachypus Tomocarabus

4 1 2 1 2 2 1 1 1 3 3 2 1 1 1 1

Neoelaphrus Omophron Aptinus Brachinus Brachynidius Brachynolomus Cnecostolus insertae sedis

1 1 3 5 4 1 3 1

Reicheadella Chiridysus Dyschiriodes Dyschirius Eudyschirius Paradyschirius Parallelomorphus Scallophorites

1 1 14 1 3 2 2 1

Bembidion

1

TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE

189

Subfamily

Tribe

Subtribe

Tachyina

Patrobini Pogonini Trechini

Harpalinae

Chlaeniini

Cyclosomini Dryptini

190

Perileptina Trechina

Trechodina Callistina Chlaeniina

Masoreina

Genus

Nr. species

Ocys Sinechostictus

1 3

Lymnastis Paratachys Polyderis Sphaerotachys Tachys Tachyta Tachyura Deltomerus

1 3 1 1 1 1 2 2

Pogonistes Pogonus Perileptus Albanotrechus Duvalius

2 2 1 1 10

Neotrechus Trechus Thalassophilus Callistus Chlaenius

3 12 1 1 13

Masoreus Drypta

1 1

Subgenus

Nr. species

Bembidionetolitzkya Chlorodium Diplocampa Emphanes Euperyphus Metallina Microserrullula Nepha Notaphemphanes Notaphus Ocydromus Ocyturanes Odontium Omoperyphus Peryphanes Peryphus Philochthus Phyla Princidium Talanes Terminophanes Testedium Trepanes

9 1 2 5 3 2 1 3 1 1 3 6 2 1 4 4 6 1 1 1 1 1 3

Pseudolimnaeum Sinechostictus

1 2

Tachyta Tachyura Deltomerus Paradeltomerus

1 2 1 1

Pogonus Perileptus

2 1

Duvalius Euduvalius

9 1

Trechus

12

Chlaeniellus Chlaenites Chlaenius Dinodes Epomis Trichochlaenius Masoreus Drypta

6 1 1 2 2 1 1 1


Subfamily

Tribe

Subtribe

Harpalini

Amblystomina Amblystomus Anisodactylina Anisodactylus

Harpalus

32

Ophonus

20

Parophonus

6

Trichotichnus Acupalpus Anthracus Bradycellus Dicheirotrichus Loxoncus Stenolophus

1 10 2 4 1 1 7

Cymindidina

Cymindis

9

Dromiina

Dromius Microlestes Paradromius Philorhizus Lebia

1 7 1 2 5

Apristus Lionychus Syntomus Badister

2 2 4 4

Licinus

4

Odacantha Oodes Omphreus Panagaeus Agonum

1 2 3 1 15

Stenolophina

Lebiina Lionychina

Odacanthini Oodini Omphreini Panagaeini Platynini

3 4 1 1 3 2 3 1 1 4

Harpalina

Licinini

Nr. species

Diachromus Gynandromorphus Carterus Ditomus Dixus Graniger Odontocarus Acinopus

Ditomina

Lebiini

Genus

Licinina

Subgenus

Nr. species

Anisodactylus Pseudodichirius

3 1

Carterus

3

Acinopus Oedematicus Osimus Cryptophonus Harpalus Pseudophonus Brachyophonus Hesperophonus Incisophonus Metophonus Ophonus Ophonomimus Parophonus Trichotichnus Acupalpus

2 1 1 1 29 2 1 4 1 10 4 1 5 1 10

Bradycellus Dicheirotrichus

4 1

Egadroma Stenolophus Cymindis Menas Tarulus Dromius

1 6 7 1 1 1

Manodromius Philorhizus Lamprias Lebia

1 2 1 4

Lionychus

2

Badister Baudia Licinus Neorescius Odacantha Oodes Omphreus Panadaeus Agonum Europhilus

2 2 3 1 1 2 3 1 13 2

TAXONOMIC STRUCTURE OF ALBANIAN CARABIDAE

191

Subfamily

Tribe

Subtribe

Zabrini

Zuphiini 11 subfamilies 31 tribes

192

Nr. species

Anchomenus Olisthopus Paranchus Platynus

1 2 1 2

Abax

2

Molops

15

Myas Pedius Poecilus

1 2 8

Pterostichus

24

Speluncarius

2

Atranopsina Calathina

Stomis Tapinopterus Platyderus Calathus

2 6 4 14

Sphodrina

Synuchidius Laemostenus

1 7

Synuchina Amarina

Synuchus Amara

1 32

Zabrina

Zabrus

8

Pterostichini

Sphodrini

Genus

Polystichus Zuphium 103 genera

Subgenus

Nr. species

Anchomenus

1

Batenus Platynus Abacopercus Abax Molops Stenochoromus Myas

1 1 1 1 14 1 1

Angoleus Poecilus Argutor Bothriopterus Cheporus Feronidius Melanius Morphnosoma Parahaptoderus Phonias Platypterinus Platysma Pseudomaseus Pterostichus Hypogium Speluncarius Stomis Tapinopterus Platyderus Bedelinus Calathus Neocalathus

1 7 1 1 1 1 2 1 1 3 1 1 4 7 1 1 2 6 4 1 6 7

Antisphodrus Laemostenus Pristonychus Synuchus Amara Amarocelia Bradytus Celia Curtonotus Paracelia Parapercosia Percosia Xenocelia Zezea Pelor Zabrus

2 3 2 1 13 1 2 3 1 2 1 1 4 4 7 1

1 1 Zuphium 543 species


1

Appendix A. Faunistic contributions A.1. New taxa for the fauna of the Balkan Peninsula Subgenera 1. Terminophanes Müller-Motzfeld, 1998 Species 1. Bembidion (Terminophanes) terminale Heer, 1841 2. Cymindis (Cymindis) angularis Gyllenhal, 1810 3. Lionychus (Lionychus) maritimus Fairmaire, 1862 A.2. Taxa new for the fauna of Albania Tribes 1. ODACANTHINI Laporte, 1834 Genera 1. Eurynebria Ganglbauer, 1891 2. Myriochila Motschulsky, 1858 3. Lymnastis Motschulsky, 1862 4. Polyderis Motschulsky, 1862 5. Pogonistes Chaudoir, 1871 6. Odontocarus Solier, 1835 7. Dromius Bonelli, 1810 8. Lionychus Wissmann, 1846 9. Odacantha Paykull, 1798 10. Paranchus Lindroth, 1974 Subgenera 1. Myriochila Motschulsky, 1858 s.str. 2. Dyschirius Bonelli, 1810 s.str. 3. Talanes Motschulsky, 1864 4. Terminophanes Müller-Motzfld, 1998 5. Pseudolimnaeum Kraatz, 1888 6. Menas Motschulsky, 1864 7. Dromius Bonelli, 1810 s.str. 8. Lamprias Bonelli, 1810 9. Lionychus Wissmann, 1846 s.str. 10. Odacanta Paykull, 1798 s.str. 11. Parahaptoderus Jeanne, 1969 Species (subspecies not considered) 1. Eurynebria complanata (Linnaeus, 1767) 2. Leistus (Pogonophorus) apfelbecki Ganglbauer, 1891 APPENDIX

193

3. Nebria (Alpaeus) germarii Heer, 1837 4. Myriochila (Myriochila) melancholica (Fabricius, 1798) 5. Aptinus (Aptinus) acutangulus Chaudoir, 1876 6. Dyschirius (Dyschiriodes) luticola Chaudoir, 1850 7. Dyschirius (Dyschirius) latipennis Seidlitz, 1867 8. Scarites (Parallelomorphus) terricola Bonelli, 1813 9. Asaphidion stierlini (Heyden, 1880) 10. Bembidion (Bembidion) quadripustulatum Audinet-Serville, 1821 11. Bembidion (Euperyphus) testaceum (Duftschmid, 1812) 12. Bembidion (Peryphanes) deletum Audinet-Serville, 1821 13. Bembidion (Peryphus) femoratum Sturm, 1825 14. Bembidion (Philochthus) mannerhaimii C.R. Sahlberg, 1827 15. Bembidion (Talanes) subfasciatum Chaudoir, 1850 16. Bembidion (Terminophanes) terminale Heer, 1841 17. Bembidion (Trepanes) maculatum Dejean, 1831 18. Sinechostictus (Pseudolimnaeum) doderoi (Ganglbauer, 1891) 19. Sinechostictus (Sinechostictus) millerianus (Heyden, 1883) 20. Lymnastis galilaeus Piochard de la Brûlerie, 1876 21. Polyderis cardioderus (Chaudoir, 1850) 22. Pogonistes convexicollis Chaudoir, 1871 23. Pogonistes gracilis (Dejean, 1828) 24. Carterus (Carterus) rotundicollis Rambur, 1837 25. Odontocarus robustus (Dejean, 1831) 26. Harpalus (Harpalus) euchlorus Ménétriés, 1839 27. Harpalus (Harpalus) flavescens (Piller et Mitterpacher, 1783) 28. Harpalus (Harpalus) hospes Sturm, 1818 29. Ophonus (Hesperophonus) similis (Dejean, 1829) 30. Ophonus (Metophonus) melletii (Heer, 1837) 31. Ophonus (Ophonus) stictus Stephens, 1828 32. Acupalpus (Acupalpus) brunnipes (Sturm, 1825) 33. Stenolophus (Stenolophus) abdominalis Gené, 1836 34. Cymindis (Cymindis) angularis Gyllenhal, 1810 35. Cymindis (Cymindis) coadunata Dejean, 1825 36. Cymindis (Menas) miliaris (Fabricius, 1801) 37. Dromius (Dromius) quadrimaculatus (Linnaeus, 1758) 38. Lebia (Lamprias) cyanocephala (Linnaeus, 1758) 39. Lebia (Lebia) trimaculata (Villers, 1789) 40. Apristus reticulatus Schaum, 1857 41. Lionychus (Lionychus) maritimus Fairmaire, 1862 42. Lionychus (Lionychus) quadrillum (Duftschmid, 1812) 43. Badister (Badister) bullatus (Schrank, 1798) 44. Odacantha (Odacantha) melanura (Linnaeus, 1767) 45. Agonum (Agonum) afrum (Duftschmid, 1812) 46. Agonum (Agonum) muelleri (Herbst, 1784) 47. Paranchus albipes (Fabricius, 1796) 48. Poecilus (Poecilus) versicolor (Sturm, 1824) 194


49. Pterostichus (Parahaptoderus) brevis (Duftschmid, 1812) 50. Pterostichus (Pseudomaseus) nigrita (Paykull, 1790) 51. Calathus (Calathus) ellipticus Reitter, 1889 52. Calathus (Neocalathus) ambiguus (Paykull, 1790) 53. Laemostenus (Laemostenus) dalmatinus (Dejean, 1828) 54. Amara (Amara) similata (Gyllenhal, 1810) 55. Amara (Bradytus) consularis (Duftschmid, 1812) A.3. Taxa firstly reported with detailed record/s for Albania Tribes 1. CYCLOSOMINI Laporte, 1834 Subtribes 1. MASOREINA Chaudoir, 1870 Genera 1. Tachys Dejean, 1821 2. Masoreus Dejean, 1821 3. Diachromus Erichson, 1837 4. Paradromius Fowler, 1887 5. Myas Sturm, 1826 6. Synuchus Gyllenhal, 1810 7. Polystichus Bonelli, 1810 Subgenera 1. Boreonebria Jeannel, 1937 2. Eunebria Jeannel, 1937 3. Procerus Dejean, 1821 4. Notaphemphanes Netolitzky, 1920 5. Omoperyphus Netolitzky, 1931 6. Phyla Motschulsky, 1844 7. Masoreus Dejean, 1821 s.str. 8. Pseudodichirius Lutshnik, 1921 9. Brachyophonus Sciaky, 1987 10. Incisophonus Sciaky, 1987 11. Tarulus Bedel, 1906 12. Manodromius Reitter, 1905 13. Batenus Motschulsky, 1865 14. Myas Sturm, 1826 s.str. 15. Bothriopterus Chaudoir, 1835 16. Morphnosoma Lutshnik, 1915 17. Platypterinus Reitter, 1908 18. Bedelinus Ragusa, 1885 19. Synuchus Gyllenhal, 1810 s.str. 20. Curtonotus Stephens, 1827 Species (subspecies not considered) 1. Leistus (Leistus) nitidus (Duftschmid, 1812) 2. Leistus (Pogonophorus) parvicollis Chaudoir, 1869 APPENDIX

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3. Nebria (Boreonebria) rufescens (Strøm, 1768) 4. Nebria (Eunebria) jockischi Sturm, 1815 5. Notiophilus aquaticus (Linnaeus, 1758) 6. Notiophilus germinyi Fauvel, 1863 7. Calosoma (Calosoma) inquisitor (Linnaeus, 1758) 8. Carabus (Procerus) gigas Creutzer, 1799 9. Clivina collaris (Herbst, 1784) 10. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837 11. Bembidion (Emphanes) axillare (Motschulsky, 1844) 12. Bembidion (Notaphemphanes) ephippium (Marsham, 1802) 13. Bembidion (Odontium) foraminosum Sturm, 1825 14. Bembidion (Omoperyphus) hypocrita Dejean, 1831 15. Bembidion (Philochthus) guttula (Fabricius, 1792) 16. Bembidion (Phyla) tethys Netolitzky, 1926 17. Paratachys micros (Fischer von Waldheim, 1828) 18. Tachys scutellaris Stephens, 1828 19. Tachyura (Tachyura) parvula (Dejean, 1831) 20. Chlaenius (Chlaeniellus) nitidulus (Schrank, 1781) 21. Chlaenius (Chlaeniellus) tristis (Schaller, 1783) 22. Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829 23. Chlaenius (Epomis) circumscriptus (Duftschmid, 1812) 24. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813) 25. Anisodactylus (Pseudodichirius) intermedius Dejean, 1829 26. Diachromus germanus (Linnaeus, 1758) 27. Harpalus (Harpalus) laevipes Zetterstedt, 1828 28. Harpalus (Harplaus) pumilus Sturm, 1818 29. Harpalus (Harpalus) subcylindricus Dejean, 1829 30. Harpalus (Harpalus) latus (Linnaeus, 1758) 31. Harpalus (Harpalus) solitaris Dejean, 1829 32. Harpalus (Harpalus) autumnalis (Duftschmid, 1812) 33. Ophonus (Brachyophonus) krueperi Apfelbeck, 1902 34. Ophonus (Incisophonus) incisus (Dejean, 1829) 35. Ophonus (Metophonus) rufibarbis (Fabricius, 1792) 36. Ophonus (Metophonus) schaubergerianus (Puel, 1937) 37. Ophonus (Ophonus) ardosiacus (Lutshnik, 1922) 38. Acupalpus (Acupalpus) elegans (Dejean, 1829) 39. Acupalpus (Acupalpus) luteatus (Duftschmid, 1812) 40. Acupalpus (Acupalpus) meridianus (Linnaeus, 1761) 41. Acupalpus (Acupalpus) suturalis Dejean, 1829 42. Bradycellus (Bradycellus) distinctus (Dejean, 1829) 43. Bradycellus (Bradycellus) verbasci (Duftschmid, 1812) 44. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758) 45. Paradromius (Manodromius) linearis (Olivier, 1795) 46. Philorhizus (Philorhizus) notatus (Stephens, 1827) 47. Lebia (Lebia) humeralis Dejean, 1825 48. Lebia (Lebia) scapularis (Geoffroy, 1785) 196


49. Licinus (Licinus) cassideus (Fabricius, 1792) 50. Agonum (Agonum) marginatum (Linnaeus, 1758) 51. Platynus (Batenus) scrobiculatus (Fabricius, 1801) 52. Abax (Abax) ovalis (Duftschmid, 1812) 53. Myas (Myas) chalybaeus (Palliardi, 1825) 54. Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787) 55. Pterostichus (Morphnosoma) melanarius (Illiger, 1798) 56. Pterostichus (Phonias) diligens (Sturm, 1824) 57. Pterostichus (Platypterinus) ottomanus Apfelbeck, 1908 58. Calathus (Bedelinus) circumseptus Germar, 1824 59. Calathus (Calathus) bosnicus Ganglbauer, 1891 60. Calathus (Neocalathus) erratus (C.R. Sahlberg, 1827) 61. Calathus (Neocalathus) metallicus Dejean, 1828 62. Synuchus (Synuchus) vivalis (Illiger, 1798) 63. Amara (Amara) convexior Stephens, 1828 64. Amara (Amara) ovata (Fabricius, 1792) 65. Amara (Amara) proxima Putzeys, 1866 66. Amara (Amara) saphyrea Dejean, 1828 67. Amara (Celia) montana Dejean, 1828 68. Amara (Curtonotus) aulica (Panzer, 1796) 69. Amara (Paracelia) dalmatina Dejean, 1828 70. Amara (Xenocelia) messae Baliani, 1924 71. Amara (Zezea) tricuspidata Dejean, 1831 72. Polystichus connexus (Geoffroy, 1785) A.4. Species omitted for Albania in Löbl & Smetana (2003); here their presence in the country is confirmed 1. Leistus (Leistus) fulvibarbis Dejean, 1826 2. Leistus (Pogonophorus) magnicollis Motschulsky, 1866 3. Nebria (Eunebria) jockischi Sturm, 1815 4. Notiophilus aestuans Dejean, 1826 5. Notiophilus aquaticus (Linnaeus, 1758) 6. Notiophilus biguttatus (Fabricius, 1779) 7. Notiophilus danieli Reitter, 1897 8. Notiophilus germinyi Fauvel, 1863 9. Notiophilus palustris (Duftschmid, 1812) 10. Notiophilus rufipes Curtis, 1829 11. Notiophilus substriatus C.R. Waterhouse, 1833 12. Calomera fischeri (M.F. Adams, 1817) 13. Carabus (Oreocarabus) preslii Dejean, 1830 14. Elaphrus (Neoelaphrus) uliginosus Fabricius, 1792 15. Brachinus (Brachinus) elegans Chaudoir, 1842 16. Brachinus (Brachinus) plagiatus Reiche, 1868 17. Brachinus (Cnecostolus) bayardi Dejean, 1831 18. Dyschirius (Dyschiriodes) nitidus (Dejean, 1825) APPENDIX

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19. Dyschirius (Dyschiriodes) cylindricus (Dejean, 1825) 20. Dyschirius (Dyschiriodes) laeviusculus Putzeys, 1846 21. Dyschirius (Eudyschirius) gracilis (Heer, 1837) 22. Scarites (Scallophorites) buparius (Forster, 1771) 23. Asaphidion flavipes (Linnaeus, 1761) 24. Bembidion (Bembidionetolitzkya) conforme (Dejean, 1831) 25. Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837 26. Bembidion (Bembidionetolitzkya) rhodopense Apfelbeck, 1902 27. Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812) 28. Bembidion (Bembidionetolitzkya) varicolor (Fabricius, 1803) 29. Bembidion (Chlorodium) splendidum Sturm, 1825 30. Bembidion (Diplocampa) assimile Gyllenhal, 1810 31. Bembidion (Emphanes) axillare (Motschulsky, 1844) 32. Bembidion (Emphanes) latiplaga Chaudoir, 1850 33. Bembidion (Emphanes) tenellum Erichson, 1837 34. Bembidion (Euperyphus) scapulare Dejean, 1831 35. Bembidion (Metallina) properans (Stephens, 1828) 36. Bembidion (Microserrullula) quadricolle (Motschulsky, 1844) 37. Bembidion (Nepha) callosum Küster, 1847 38. Bembidion (Nepha) caucasicum (Motschulsky, 1844) 39. Bembidion (Notaphemphanes) ephippium (Marsham, 1802) 40. Bembidion (Ocyturanes) pindicum Apfelbeck, 1901 41. Bembidion (Odontium) foraminosum Sturm, 1825 42. Bembidion (Odontium) striatum (Fabricius, 1792) 43. Bembidion (Omoperyphus) hypocrita Dejean, 1831 44. Bembidion (Peryphanes) brunnicorne Dejean, 1831 45. Bembidion (Peryphanes) grandipenne Schaum, 1862 46. Bembidion (Peryphanes) tetracolum Say, 1823 47. Bembidion (Philochthus) guttula (Fabricius, 1792) 48. Bembidion (Philochthus) inoptatum Schaum, 1857 49. Bembidion (Philochthus) lunulatum (Geoffroy, 1785) 50. Bembidion (Phyla) tethys Netolitzky, 1926 51. Bembidion (Princidium) punctulatum Drapiez, 1820 52. Bembidion (Testedium) bipunctatum (Linnaeus, 1761) 53. Bembidion (Trepanes) octomaculatum (Goeze, 1777) 54. Ocys harpaloides (Audinet-Serville, 1821) 55. Sinechostictus (Sinechostictus) tarsicus (Peyron, 1858) 56. Paratachys bistriatus (Duftschmid, 1812) 57. Paratachys fulvicollis (Dejean, 1831) 58. Paratachys micros (Fischer von Waldheim, 1828) 59. Sphaerotachys haemorrhoidalis (Ponza, 1805) 60. Tachys scutellaris Stephens, 1828 61. Tachyta (Tachyta) nana (Gyllenhal, 1810) 62. Tachyura (Tachyura) diabrachys (Kolenati, 1845) 63. Tachyura (Tachyura) parvula (Dejean, 1831) 64. Pogonus (Pogonus) littoralis (Duftschmid, 1812) 198


65. Chlaenius (Chlaeniellus) terminatus Dejean, 1826 66. Chlaenius (Epomis) dejeani (Dejean, 1831) 67. Masoreus (Masoreus) wetterhallii (Gyllenhal, 1813) 68. Harpalus (Harpalus) cupreus Dejean, 1829 69. Harpalus (Pseudophonus) griseus (Panzer, 1796) 70. Ophonus (Hesperophonus) subquadratus (Dejean, 1829) 71. Parophonus (Ophonomimus) hirsutulus (Dejean, 1829) 72. Acupalpus (Acupalpus) parvulus (Sturm, 1825) 73. Bradycellus (Bradycellus) caucasicus (Chaudoir, 1846) 74. Cymindis (Cymindis) axillaris (Fabricius, 1794) 75. Cymindis (Cymindis) humeralis (Geoffroy, 1785) 76. Cymindis (Cymindis) lineola L. Dufour, 1820 77. Cymindis (Tarulus) vaporariorum (Linnaeus, 1758) 78. Microlestes luctuosus Holdhaus, 1904 79. Microlestes minutulus (Goeze, 1777) 80. Microlestes negrita (Wollaston, 1854) 81. Paradromius (Manodromius) linearis (Olivier, 1795) 82. Philorhizus (Philorhizus) notatus (Stephens, 1827) 83. Lebia (Lebia) cruxminor (Linnaeus, 1758) 84. Lebia (Lebia) humeralis Dejean, 1825 85. Lebia (Lebia) scapularis (Geoffroy, 1785) 86. Apristus subaeneus Chaudoir, 1846 87. Syntomus impressus (Dejean, 1825) 88. Syntomus pallipes (Dejean, 1825) 89. Syntomus truncatellus (Linnaeus, 1761) 90. Omphreus (Omphreus) morio Dejean, 1828 91. Agonum (Agonum) extensum Ménétriés, 1849 92. Agonum (Agonum) hypocrita (Apfelbeck, 1904) 93. Agonum (Agonum) marginatum (Linnaeus, 1758) 94. Agonum (Agonum) sexpunctatum (Linnaeus, 1758) 95. Agonum (Agonum) sordidum Dejean, 1828 96. Agonum (Agonum) viridicupreum (Goeze, 1777) 97. Agonum (Europhilus) antennarium (Duftschmid, 1812) 98. Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763) 99. Olisthopus fuscatus Dejean, 1828 100. Olisthopus glabricollis (Germar, 1817) 101. Platynus (Batenus) scrobiculatus (Fabricius, 1801) 102. Platynus (Platynus) assimilis (Paykull, 1790) 103. Molops (Molops) alpestris (Dejean, 1828) 104. Molops (Molops) apfelbecki Ganglbauer, 1891 (considering former Molops reseri Apfelbeck, 1904) 105. Molops (Molops) curtulus Ganglbauer, 1891 106. Molops (Molops) holdhausi J. Müller, 1917 107. Molops (Molops) plurisetosus J. Müller, 1918 108. Molops (Molops) piceus (Panzer, 1793) 109. Pterostichus (Cheporus) burmeisteri Heer, 1838 110. Pterostichus (Platysma) niger (Schaller, 1783) APPENDIX

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111. Pterostichus (Pterostichus) reiseri Ganglbauer, 1889 112. Platyderus (Platyderus) dalmatinus L. Miller, 1881 113. Calathus (Neocalathus) cinctus Motschulsky, 1850 114. Calathus (Neocalathus) mollis (Marsham, 1802) 115. Laemostenus (Antisphodrus) elongatus (Dejean, 1828) 116. Laemiostenus (Laemostenus) janthinus (Duftschmid, 1812) 117. Zabrus (Pelor) incrassatus (Ahrens, 1814) 118. Zabrus (Pelor) spinipes (Fabricius, 1798) 119. Polystichus connexus (Geoffroy, 1785) A.5. Taxa still not well defined 1. Nebria (Alpaeus) germarii ssp. 2. Dalmatoreicheia sp. 3. Reicheadella (Reicheadella) sp. 4. Bembidion (Bembidionetolitzkya) sp. (aff. complanatum Heer, 1937) 5. Bembidion (Bembidionetolitzkya) sp. (ex. gruppo “fasciolatum”) 6. Bembidion (Terminophanes) terminale ssp. 7. Adriaphaenops sp. 8. Trechini gen. & sp. indet. (? aff. Pheggomisetes Knirsch, 1923) 9. Omphreus (Omphreus) ravasinii ssp. 10. Tapinopterus (Tapinopterus) protensus ssp.1 11. Tapinopterus (Tapinopterus) protensus ssp.2 12. Zabrus (Pelor) albanicus ssp. A.6. Taxa (cited, mapped or found in collections) with doubtful presence in Albania Subgenera 1. Platycarabus A. Morawitz, 1886 2. Trachycarabus Géhin, 1876 Species and subspecies 1. Nebria (Alpaeus) bosnica tresnjevikensis Horvatovich, 1974 2. Nebria (Alpaeus) dahli (Duftschmid, 1812) (considering ssp. montenegrina Apfelbeck, 1904) 3. Carabus (Platycarabus) irregularis Fabricius, 1792 (considering ssp. ramanus Sokolář, 1909) 4. Carabus (Trachycarabus) scabriusculus Olivier, 1795 (considering ssp. bulgarus Lapouge, 1908) 5. Siagona europaea oberleitneri Dejean, 1830 6. Dyschirius (Eudyschirius) gracilis (Heer, 1837) (considering nominotypical subspecies) 7. Asaphidion cyanicorne (Pandellé, 1867) (considering nominotypical subspecies) 8. Bembidion (Bembidion) humerale Sturm, 1825 9. Bembidion (Bembidion) quadrimaculatum (Linnaeus, 1761) (considering nominotypical subspecies) 10. Bembidion (Bembidionetolitzkya) relictum Apfelbeck, 1904 11. Deltomerus (Deltomerus) malissorum Apfelbeck, 1918 12. Deltomerus (Paradeltomerus) paradoxus paradoxus Apfelbeck, 1908 13. Pogonus (Pogonus) luridipennis (Germar, 1823) 14. Trechus (Trechus) irenis Csiki, 1912 15. Cymindis (Cymindis) pindicola Apfelbeck, 1901 200


16. Olisthopus rotundatus (Paykull, 1790) (considering nominotypical subspecies) 17. Molops (Molops) spartanus parnassicola Kraatz, 1875 18. Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908 19. Pterostichus (?) epiroticus Csiki, 1930 20. Calathus (Calathus) glabricollis Dejean, 1828 21. Amara (Amara) famelica C. Zimmermann, 1832 22. Amara (Celia) praetermissa (C.R. Sahlberg, 1827) A.7. Taxa deleted from the list of Albanian Carabidae Subgenera 1. Eucarabus Géhin, 1876 2. Oreoplatysma Jakobson, 1907 Species and subspecies 1. Nebria (Nebria) salina Fairmaire & Laboulbène, 1854 2. Cicindela (Cicindela) hybrida Linnaeus, 1758 3. Carabus (Eucarabus) catenulatus Scopoli, 1763 4. Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812 5. Cychrus semigranosus semigranosus Palliardi, 1825 6. Cychrus semigranosus balcanicus Hopffgarten, 1881 7. Cychrus semigranosus montanus Winkler, 1939 8. Brachinus (Brachinus) alexandri F. Battoni, 1984 9. Dyschirius (Dyschiriodes) punctatus (Dejean, 1825) 10. Bembidion (Bembidionetolitzkya) ascendens K. Daniel, 1902 11. Bembidion (Bembidionetolitzkya) fasciolatum (Duftschmid, 1812) 12. Bembidion (Peryphanes) milleri Jacquelin du Val, 1852 13. Tachyura (Tachyura) sexstriata (Duftschmid, 1812) 14. Duvalius (Bicharotrechus) maglicensis Winkler, 1933 15. Trechus (Trechus) obtusiusculus Ganglbauer, 1889 16. Acinopus (Acinopus) subquadratus Brullé, 1832 17. Harpalus (Harpalus) anatolicus Tschitschérine, 1898 18. Harpalus (Harpalus) danieli Reitter, 1900 19. Molops (Molops) weiratheri J. Müller, 1930 20. Molops (Molops) simplex simplex Chaudoir, 1868 21. Molops (Molops) rufipes klisuranus Apfelbeck, 1902 22. Pterostichus (Oreoplatysma) lodosi Heinz, 1977 23. Tapinopterus (Tapinopterus) comita Jedlička, 1935 24. Tapinopterus (Tapinopterus) extensus (Dejean, 1831) 25. Tapinopterus (Tapinopterus) miridita jacupicensis Jedlička, 1935 26. Laemostenus (Antisphodrus) cavicola modestus (Schaufuss, 1862) 27. Amara (Amara) spreta Dejean, 1831 28. Amara (Zezea) erythrocnema Dejean, 1828 29. Zabrus (Pelor) apfelbecki Ganglbauer, 1915

APPENDIX

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B. Taxonomic contributions B.1. Nomenclatorial acts Nebriini Nebria (Alpaeus) macedonica Mařan, 1938, stat. rest. for Nebria (Alpaeus) aetolica macedonica Mařan, 1938 Nebria (Alpaeus) macedonica galicica Horvatovich, 1974a, new assignment of Nebria (Alpaeus) aetolica galicica Horvatovich, 1974a Nebria (Alpaeus) macedonica papingutica Horvatovich, 1974a, new assignment of Nebria (Alpaeus) aetolica papingutica Horvatovich, 1974a Nebria (Alpaeus) macedonica rambouseki Mařan, 1938, new assignment of Nebria (Alpaeus) aetolica rambouseki Mařan, 1938 Carabini Carabus (Oreocarabus) neumeyeri Schaum, 1856, stat. nov. for Carabus (Oreocarabus) hortensis neumeyeri Schaum, 1856 Carabus (Oreocarabus) preslii Dejean, 1830, stat. rest. for Carabus (Oreocarabus) hortensis preslii Dejean, 1830 Cychrini Cychrus semigranosus albanicus Hopp, 1929, syn. rest. of Cychrus semigranosus montenegrinus Apfelbeck, 1904 Cychrus semigranosus dinaricus Mandl, 1989, syn. nov. of Cychrus semigranosus montenegrinus Apfelbeck, 1904 Licinini Licinus (Licinus) submarginatus J.R. Sahlberg, 1903, syn. nov. of Licinus (Licinus) graecus Apfelbeck, 1901 Platynini Platynus (Batenus) scrobiculatus serbicus Csiki, 1904, stat. nov. for Platynus (Batenus) serbicus Csiki, 1904 Platynus (Batenus) scrobiculatus turcicus Apfelbeck, 1904, stat. rest. for Platynus (Batenus) turcicus Apfelbeck, 1904 Pterostichini Molops (Molops) maderi J. Müller, 1933, syn. nov. of Molops (Molops) malshentianus Apfelbeck, 1917 Molops (Molops) apfelbecki reiseri Apfelbeck, 1904, new assignment of Molops (Molops) reiseri reiseri Apfelbeck, 1904 Molops (Molops) apfelbecki corallipes J. Müller, 1936, new assignment of Molops (Molops) reiseri corallipes J. Müller, 1936 Molops (Molops) apfelbecki pseudoalpestris Mlynář, 1977, new assignment of Molops (Molops) reiseri pseudoalpestris Mlynář, 1977 Molops (Molops) apfelbecki victoris J. Müller, 1918, new assignment of Molops (Molops) reiseri victoris J. Müller, 1918 Molops (Molops) holdhausi J. Müller, 1917, stat. rest. for Molops (Molops) reiseri holdhausi J. Müller, 1917 Aphaon Heyden, 1906, syn. nov. of Pterostichus s. str. Bonelli, 1810 Pterostichus (Pterostichus) bischoffianus Jedlička, 1936, new assignment of Pterostichus (Aphaon) bischoffianus Jedlička, 1936 202


Pterostichus (Pterostichus) brucki Schaum, 1859, new assignment of Pterostichus (Aphaon) brucki Schaum, 1859 Pterostichus (Pterostichus) latifianus latifianus Apfelbeck, 1906, new assignment of Pterostichus (Aphaon) latifianus latifianus Apfelbeck, 1906 Pterostichus (Pterostichus) latifianus strupii J. Müller, 1935, new assignment of Pterostichus (Aphaon) latifianus strupii J. Müller, 1935 Pterostichus (Pterostichus) lumensis lumensis Apfelbeck, 1905, new assignment of Pterostichus (Aphaon) lumensis lumensis Apfelbeck, 1905 Pterostichus (Pterostichus) lumensis litae Csiki, 1940, new assignment of Pterostichus (Aphaon) lumensis litae Csiki, 1940 Pterostichus (Pterostichus) lumensis ljubetensis Apfelbeck, 1908, new assignment of Pterostichus (Aphaon) lumensis ljubetensis Apfelbeck, 1908 Pterostichus (Pterostichus) malissorum Apfelbeck, 1905, new assignment of Pterostichus (Aphaon) malissorum Apfelbeck, 1905 Pterostichus (Pterostichus) latifianus ab. nigripes Jedlička, 1936, syn. rest. for Pterostichus (Pterostichus) malissorum Apfelbeck, 1905 Pterostichus (Pterostichus) meisteri Reitter, 1885, new assignment of Pterostichus (Aphaon) meisteri Reitter, 1885 Pterostichus (Pterostichus) merklii (J. Frivaldszky, 1879), new assignment of Pterostichus (Aphaon) merklii (J. Frivaldszky, 1879) Pterostichus (Pterostichus) pentheri Apfelbeck, 1918, new assignment of Pterostichus (Aphaon) pentheri Apfelbeck, 1918 Pterostichus (Pterostichus) reiseri Ganglbauer, 1889, new assignment of Pterostichus (Aphaon) reiseri Ganglbauer, 1889 Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874, new assignment of Pterostichus (Aphaon) rhilensis rhilensis Rottenberg, 1874 Pterostichus (Pterostichus) rhilensis kourili Mařan, 1933, new assignment of Pterostichus (Aphaon) rhilensis kourili Mařan, 1933 Pterostichus (Pterostichus) rhilensis vitosensis Mařan, 1933, new assignment of Pterostichus (Aphaon) rhilensis vitosensis Mařan, 1933 Pterostichus (Pterostichus) walteri Reitter, 1883, new assignment of Pterostichus (Aphaon) walteri Reitter, 1883 Tapinopterus (Tapinopterus) extensoides (Straneo, 1936), stat. rest. for Tapinopterus (Tapinopterus) extensus extensoides (Straneo, 1936). Sphodrini Calathus (Calathus) ravasinii macedonicus Mařan, 1935, stat. nov. for Calathus (Calathus) macedonicus Mařan, 1935 Zabrini Zabrus (Pelor) bischoffi J. Müller, 1936, stat. rest. for Zabrus (Pelor) incrassatus bischoffi J. Müller, 1936. B.2. Year of description The following taxa (valid or synonyms) have long time been accredited to have a correct year of description. However, it was found out that this is incorrect. Below, the relevant earlier APPENDIX

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years of publication dates of the original descriptions are given. All the descriptions considered fulfil the conditions of the ICZN (1999). Carabus croaticus ljubetensis Apfelbeck, 1917 (synonym of Carabus croaticus durmitorensis Apfelbeck, 1904) instead Carabus croaticus ljubetensis Apfelbeck, 1918a Carabus violaceus merditanus Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857) instead Carabus violaceus merditanus Apfelbeck, 1918a Carabus violaceus shardagensis Apfelbeck, 1917 (synonym of Carabus violaceus dryas Gistl, 1857) instead Carabus violaceus shardagensis Apfelbeck, 1918a Aptinus merditanus Apfelbeck, 1917 (valid species) instead Aptinus merditanus Apfelbeck, 1918b Aptinus merditanus var. granulipennis Apfelbeck, 1917 (synonym of Aptinus merditanus Apfelbeck, 1917) instead Aptinus merditanus var. granulipennis Apfelbeck, 1918b Trechus albanicus Apfelbeck, 1905 (valid species) instead Trechus albanicus Apfelbeck, 1907b Ophonus krueperi Apfelbeck, 1902 (valid species) instead Ophonus krueperi (Apfelbeck, 1904) Licinus graecus Apfelbeck, 1901 (valid species) instead Licinus cassideus graecus Apfelbeck, 1904 Molops malshentianus Apfelbeck, 1917 (valid species) instead Molops malshentianus Apfelbeck, 1918b B.3. Nomina nuda found in both collections and literature Eurynebria complanata orientalis Apfelbeck Eurynebria complanata albanica Apfelbeck Eurynebria complanata var. albanica Apfelbeck ? Nebria ganglbaueri ditana Apfelbeck (or incorrect spelling of Nebria ganglbaueri merditana Apfelbeck, 1906) Aptinus lugubris rufipes Apfelbeck (according to Apfelbeck, 1907a: 499, 501) Bembidion merditanum Apfelbeck Harpalus bischoffi Schauberger Molops transfuga Apfelbeck (according to Müller, 1930: 124; Mlynář, 1977: 120) Molops apfelbecki bischoffi J. Müller Pterostichus djumanus Apfelbeck

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References General work on the names, the systematics and the distribution of the Carabidae from the Balkan Peninsula (the separate parts of the first work are cited in the following section) 1. 2.

Löbl I., A. Smetana. 2003. Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata Myxophaga - Adephaga. Stenstrup: Apollo Books, 819 pp. Lorenz W. 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Tutzing: W. Lorenz, iv + 937 pp.

Faunistic and taxonomic references dealing with Albanian Carabidae (sources containing data but not found are in brackets) 1.

Absolon K., J. Mařan. 1947. Příspěvek k poznání geografického rozšíření, systematiku a vzniku druhů rodu Neotrechus J. Müll. – Příroda, 39: 1-6. 2. Anonymous. 1907. Collections du Musée d’Histoire Naturelle de son altesse Royale Ferdinand I Prince de Bulgarie. Sofia, xiv + 484 p. 3. Apfelbeck V. 1901a. Izvestaj o entomoloskom istrazivackom putovanju u Tursku i Grcko u godini 1900. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 13: 405-438. 4. Apfelbeck V. 1901b. Zwei neue Rassen der Gattung Carabus aus Albanien. – Societas Entomologica, 16: 121-122. 5. Apfelbeck V. 1902a. Zur Kenntnis der palaearktischen Carabiden. Synonymische und zoogeographische Beiträge. – Münchener Koleopterologische Zeitschrift, 1 [1902-1903]: 95-101. 6. Apfelbeck V. 1903. Ophonus suturifer Reitt. Ist = O. fallax Peyr. = suturalis Chaud. – Wiener Entomologische Zeitung, 22: 53-54. 7. Apfelbeck V. 1904. Die Käferfauna der Balkanhalbinsel, mit Berücksichtigung KleinAsiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea. Berlin: R. Friedländer und Sohn, IX + 422 pp. 8. Apfelbeck V. 1905. Paeninsulae balcanicae coleopterorum species novae - Nove vrste koleoptera s balkanskog poluostrva. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 17: 239-252. 9. Apfelbeck V. 1906. Neue Koleopteren, gesammelt während einer in Jahre 1905 mit Subvention der kaiserlichen Akademie der Wissenschaften in Wien durchgeführten zoologischen Forschungsreise nach Albanien und Montenegro (I. Serie). – Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-naturwissenschaftliche Klasse (Abt.I), 115: 1661-1674. 10. Apfelbeck V. 1907a. Koleopterologische Ergebnisse der mit Subvention der Kaiserlichen Akademie der Wissenschaften in Wien im Frühjahr 1905 ausgeführten Forschungsreise nach Albanien und Montenegro. – Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-naturwissenschaftliche Klasse (Abt.I), 116: 493-506. REFERENCES

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11. Apfelbeck V. 1907b. Neue Koleopteren von der Balkanhalbinsel. – Wissenschaftliche Mitteilungen aus Bosnien und der Hercegovina, 10: 635-644. 12. Apfelbeck V. 1909. Prinosi poznavanju faune koleoptera u Albanii. I. Valona i Berat. Koleoptera sabrana po gosp. Dru Karlu Pacu (Patsch) u okoloci Valone i Berata godine 1900. prigodom arheoloskog putovanja. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 21: 297-301. 13. Apfelbeck V. 1911. Contributiones ad coleopterorum faunam paeninsulae balcanicae. II. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 23: 209-223. 14. Apfelbeck V. 1916. Fauna insectorum balcanica. VI. 1. Beschreibung von 27 neuen Coleopteren der Balkanhalbinsel. – Wissenschaftliche Mitteilungen aus Bosnien und der Hercegovina, 13: 340-341. 15. Apfelbeck V. 1917. Dopune opisa i primjedbe o novim koleopterima Balkanskog Polusotrova. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 29: 219-243. 16. Apfelbeck V. 1918a. Diagnosen neuer Koleopteren von der Balkanhalbinsel. – Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematischnaturwissenschaftlische Klasse, 55: 75-77. 17. Apfelbeck V. 1918b. Diagnosen neuer Koleopteren von der Balkanhalbinsel. – Anzeiger der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematischnaturwissenschaftlische Klasse, 55: 88-90. 18. Apfelbeck V. 1918c. Koleopteren aus dem nordalbanisch-montenegrinischen Grenzgebiete (Ergebnisse einer von der Kaiserl. Akademie der Wissenschaft in Wien veranlaâten naturwissenschaftlichen Forschungsreise in Nordalbanien). –Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-naturwissenschaftliche Klasse (Abt.I), 127: 159-168. 19. Apfelbeck V. 1919a. Poznavanju balkanske faune koleoptera. I. Visokoalpinske pasmine Carabus croaticus-a i C. caelatus-a uz opis novih oblika. – Glasnik Zemaljskog Museja u Bosni i Hercegovini, 31: 265-272. 20. Apfelbeck V. 1919b. Zur Kenntnis der Balkanfauna (Coleoptera). I. Die hochalpinen Rassen von Carabus croaticus und caelatus, nebst Beschreibungen neuer Formen. – Koleopterologische Rundschau, 8: 44-47. 21. Apfelbeck V. 1930. Fauna insectorum balcanica. VII. 2. Teil. – Neue Beiträge zur systematischen Insektenkunde, 4: 161-176. 22. Arnold W. 1939. Bestimmung europäischer Käfer. (4. Stück.) II. Fam. Carabidae. Subfam. Carabinae. Tribus Carabini. 1. Calosoma. - 2. Carabus. - 3. Cychrus (Arten Mittel- und Nordeuropas.). – Koleopterologische Rundschau, 25: 145-200. 23. Aukema B. 1990. Taxonomy, life history and distribution of three closely related species of the genus Calathus (Coleoptera: Carabidae). – Tijdschrift voor Entomologie, 133: 121-141. 24. Baehr M. 2003a. Subfamily Siagoninae Bonelli, 1813. Pp. 218. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 25. Baehr M. 2003b. Tribe Dryptini Bonelli, 1810. Pp. 358-359. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 26. Baehr M. 2003c. Tribe Licinini Bonelli, 1810. Pp. 439-443. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 206


27. Baehr M. 2003d. Tribe Panagaeini Bonelli, 1810. Pp. 446-448. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 28. Baehr M. 2003e. Tribe Zuphiini Bonelli, 1810. Pp. 573-574. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 29. Balkenohl M. 2003. Subfamily Scaritinae Bonelli, 1810. Pp. 219-234. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata Myxophaga - Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 30. Bänninger M. 1925. Neunter Beitrag zur Kenntnis der Carabinae: die Nebriini. – Entomologische Mitteilungen, 14 (3-4): 256-281. 31. Bernau G. 1913. Über die Rassen von Carabus cancellatus Illig. Ein Beitrag zu deren Kenntnis. – Wiener Entomologische Zeitung, 32: 191-210. 32. Bousquet Y. 2003a. Tribe Pogonini Laporte, 1834. Pp. 286-288. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 33. Bousquet Y. 2003b. Tribe Cyclosomini Laporte, 1834. Pp. 356-358. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 34. Bousquet Y. 2003c. Tribe Oodini LaFerté-Sénectére, 1851. Pp. 444-445. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata Myxophaga - Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 35. Bousquet Y. 2003d. Tribe Omphreini Ganglbauer, 1891. Pp. 445-446. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata Myxophaga - Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 36. Bousquet Y. 2003e. Tribe Platynini Bonelli, 1810. Pp. 449-469. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 37. Bousquet Y. 2003f. Tribe Pterostichini Bonelli, 1810. Pp. 469-521. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 38. Bousquet Y., A. Barševskis. 2003. Tribe Notiophilini Motschulsky, 1850. Pp. 96-98. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 39. Bousquet Y., B. Březina, A. Davies, J. Farkač, A. Smetana. 2003. Tribe Carabini Latreille, 1802. Pp. 118-201. In: Löbl, L. & Smetana, A. (eds.): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, Denmark, 819 pp. 40. Breuning S. 1928a. Beitrag zur Carabenfauna von Bulgarien (Carabidae, Col.) mit Berücksichtigung der angrenzenden Gebiete. – Mitteilungen aus dem Königlichen Narurwissenschaftichen Instituten Sofia, 1: 108-117. {Breuning S. 1928b. Monographie der Gattung Calosoma Web. (Carab.). III. Teil. – Koleopterologische Rundschau, 14: 43-101.} 41. Breuning S. 1932. Monographie der Gattung Carabus L. Bestimmungs-Tabellen der europäischen Coleopteren. 104. Heft. Troppau: Emmerich Reitter, 288 pp. 42. Breuning S. 1933. Monographie der Gattung Carabus L. (IV. Teil). Bestimmungs-Tabellen der europäischen Coleopteren. 107. Heft. Troppau: Emmerich Reitter, pp. 707-912. REFERENCES

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Index (valid taxa are given in bold) Abacopercus 138 Abax 138 Abax s.str. 139 abdominalis (Stenolophus) 119 abdominalis persicus (Stenolophus) 119 Acinopus 97 Acinopus s.str. 97 Acupalpus 12, 115 Acupalpus s.str. 115 acutangulus (Aptinus) 39 Adriaphaenops 79 aenea (Amara) 174 aeneocephalus (Chlaenius) 92 aeneocephalus aeneocephalus (Chlaenius) 92 aeneus (Dyschirius) 46 aeneus (Harpalus) 107 aeneus aeneus (Dyschirius) 46 aestuans (Notiophilus) 23 aetolica (Nebria) 18 aetolica galicica (Nebria) 18, 21 aetolica korabica (Nebria) 21 aetolica macedonica (Nebria) 18 aetolica papingutica (Nebria) 18, 21, 22 aetolica rambouseki (Nebria) 18 aetolica tomoricensis (Nebria) 21, 22 aetolicum (Bembidion) 63 aetolicum aetolicum (Bembidion) 63 aetolicus (Carabus) 30 aetolicus (Tapinopterus) 163 aetolicus peristericus (Tapinopterus) 162 aetolicus tomoricensis (Tapinopterus) 163 aetolus (Zabrus) 182 aetolus kodymi (Zabrus) 182 aetolus winkleri (Zabrus) 182 affinis (Harpalus) 107 affinis ab. limbopunctatus (Harpalus) 107 affinis ab. viridulus (Harpalus) 107 afrum (Agonum) 133 agnatus (Dyschirius) 46 224

Agonum 133 Agonum s.str. 133 albanensis (Carabus) 31 albanensis (Harpalus) 111 albanensis (Ophonus) 111 albanicola (Carabus) 34 albanicum (Bembidion) 68 albanicum (Hypogeobium) 160 albanicum (Platysma) 160 albanicus (Calathus) 169, 171 albanicus (Carabus) 35 albanicus (Cychrus) 37 albanicus (Harpalus) 103 albanicus (Molops) 142 albanicus (Omphreus) 132 albanicus (Ophonus) 111 albanicus (Pterostichus) 160 albanicus (Speluncarius) 160 albanicus (Tapinopterus) 160 albanicus (Trechus) 87 albanicus (Zabrus) 182, 183 albanicus albanicus (Zabrus) 182 albanicus latifianus (Zabrus) 182, 183 albanicus jablanicensis (Zabrus) 182 Albanotrechus 79 albipes (Paranchus) 137 alexandri (Brachinus) 40 alinae (Molops) 140 Alpaeus 18 alpestre (Agonum) 133 alpestris (Molops) 141 alpestris imitator (Molops) 141 Amara 12, 174 Amara s.str. 174 AMARINA 174 Amarocelia 177 ambiguus (Calathus) 169 ambiguus (Chlaenius) 91 ambiguus ambiguus (Calathus) 169


AMBLYSTOMINA 93 Amblystomus 93 ammophilus (Acinopus) 98 anatolicus (Harpalus) 104 anatolicus anatolicus (Harpalus) 104 Ancholeus 149 Anchomenus 136 Anchomenus s.str. 136 andreae (Bembidion) 68 andreae albanicum (Bembidion) 68 andreae bualei (Bembidion) 68 angularis (Cymindis) 121 angustatum (Agonum) 134 ANISODACTYLINA 94 Anisodactylus 12, 94 Anisodactylus s.str. 94 antennarium (Agonum) 136 antennarius (Platynus) 136 anthobia (Amara) 174 anthracinus (Pterostichus) 154 anthracinus anthracinus (Pterostichus) 154 anthracinus biimpressus (Pterostichus) 155 Anthracus 117 Antisphodrus 171 anxius (Harpalus) 102 apfelbecki (Leistus) 17 apfelbecki (Molops) 143 apfelbecki (Nebria) 19 apfelbecki (Pterostichus) 153 apfelbecki (Zabrus) 183 apfelbecki apfelbecki (Molops) 143 apfelbecki bischoffi (Molops) 143 apfelbecki corallipes (Molops) 142 apfelbecki kulti (Nebria) 19 apfelbecki plurisetosus (Molops) 145 apfelbecki pseudoalpestris (Molops) 142 apfelbecki reiseri (Molops) 142, 143 apfelbecki victoris (Molops) 142, 143 Aphaenopsis 79 Aphaon 155 apicalis (Dyschirius) 47 APOTOMINAE 51, 187 APOTOMINI 51 Apotomus 51 apricaria (Amara) 177 Apristus 127

apterus (Microlestes) 124 Aptinus 39 Aptinus s.str. 39 aquaticus (Notiophilus) 24 Archicarabus 30 ardosiacus (Ophonus) 112 arenaria (Cicindela) 28 arenaria (Cylindera) 28 arenaria viennensis (Cylindera) 28 areolatus (Perileptus) 79 areolatus areolatus (Perileptus) 79 Argutor 151 articulatum (Bembidion) 72 Asaphidion 52 ascendens (Bembidion) 56, 57 assimile (Bembidion) 58 assimilis (Platynus) 138 ATRANOPSINA 165 atratus (Harpalus) 100 atratus var. subsinuatus (Harpalus) 100 atrocaeruleum (Bembidion) 56 attemsi (Nebria) 19 attenuatus (Harpalus) 99 atterimus (Pterostichus) 152 atterimus atterimus (Pterostichus) 152 aulica (Amara) 178 auriculatus (Dyschirius) 47 austriacus (Molops) 147 autumnalis (Harpalus) 104 axillare (Bembidion) 58 axillare occiduum (Bembidion) 58 axillaris (Cymindis) 121 axillaris axillaris (Cymindis) 121 axillaris adriatica (Cymindis) 121 axillaris lineola (Cymindis) 122 azurescens (Bembidion) 59 azurescens azurescens (Bembidion) 59 azureus (Chlaenius) 91 azureus (Harpalus) 109 azureus (Ophonus) 109, 110 azureus oberthuri (Harpalus) 109 babinjensis (Carabus) 32 babinjensis (Trechus) 85 bacillus (Dyschirius) 47 bacillus arbensis (Dyschiriodes) 47 bacillus arbensis (Dyschirius) 47 INDEX

225

bacillus bacillus (Dyschirius) 47 bacillus lucanus (Dyschirius) 47 Badister 128 Badister s.str. 128 balcanicum (Bembidion) 64 balcanicum balcanicum (Bembidion) 64, 65, 66 baldori sic! (Zabrus) 184 Batenus 138 Baudia 129 baudii (Acinopus) 97 baudii hellenicus (Acinopus) 97, 98 bayardi (Brachinus) 43 Bedelinus 166 BEMBIDIINA 52 BEMBIDIINI 12, 52 Bembidion 12, 53 Bembidion s.str. 53 Bembidionetolitzkya 54 beroni (Albanotrechus) 12, 79 Bicharotrechus 79 bifrons (Amara) 178 biguttatus (Notiophilus) 24 binaghii (Agonum) 133 binotatus (Anisodactylus) 94 binotatus ab. spurcaticornis (Anisodactylus) 94 bipunctatum (Bembidion) 71 bipunctatum nivale (Bembidion) 71 bipunctatum var. pyritosum (Bembidion) 71 bipustulatus (Brachinus) 43 bisbimaculatus (Tachys) 76 bischoffi (Amara) 180 bischoffi (Duvalius) 80 bischoffi (Harpalus) 105 bischoffi (Neotrechus) 83 bischoffi (Reicheadella) 45 bischoffi (Reicheia) 45 bischoffi (Tapinopterus) 163, 164 bischoffi (Zabrus) 184 bischoffianus (Pterostichus) 155, 156 bistriatus (Paratachys) 74 bistriatus (Tachys) 74 bodemeyeri (Brachinus) 41 boldorii (Zabrus) 184 bombarda (Aptinus) 39 Boreonebria 22 bosnica (Nebria) 19 226

bosnica apfelbecki (Nebria) 19 bosnica jezercensis (Nebria) 19 bosnica kulti (Nebria) 19 bosnica prokletiensis (Nebria) 19 bosnica sturanyi (Nebria) 19 bosnica tresnjevikensis (Nebria) 20 bosnicus (Calathus) 166 Bothriopterus 151 Brachynidius 41 BRACHININAE 7, 39, 187 BRACHININI 39 Brachynolomus 42 Brachinus 40 Brachinus s.str. 40 Brachyophonus 109 Bracteon 57 Bradycellus 12, 118 Bradycellus s.str. 118 Bradytus 177 brevicollis (Brachinus) 42 brevicollis (Nebria) 22, 23 brevicollis (Ophonus) 111 brevicollis var. salina (Nebria) 22, 23 brevis (Pterostichus) 153 brevis brevis (Pterostichus) 153 brucki (Pterostichus) 155, 156 brunnicorne (Bembidion) 66 brunnicorne brunnicorne (Bembidion) 66 brunnipes (Acupalpus) 115 brunoi (Bembidion) 56, 57 bualei (Bembidion) 68 bualei bualei (Bembidion) 68 bucephalus (Carabus) 35 bullatus (Badister) 128 buparius (Scarites) 51 burmeisteri (Pterostichus) 151 burmeisteri burmeisteri (Pterostichus) 151 caelatus (Carabus) 31 caelatus caelatus (Carabus) 31 caelatus caelatus m. malissorum (Carabus) 31 caelatus caelatus n. metalkanus (Carabus) 31 caelatus dalmatinus (Carabus) 32 caelatus dalmatinus n. procerus (Carabus) 32 caelatus malissorum (Carabus) 31 caelatus metalkanus (Carabus) 31 caelatus metalkanus v. malissorum (Carabus) 31


caelatus sarajevensis sic! (Carabus) 31 caelatus sarajevoensis (Carabus) 31 caelatus var. procerus (Carabus) 32 CALATHINA 166 Calathus 166 Calathus s.str. 166 CALLISTINA 89 Callistus 89 callosum (Bembidion) 61 callosum callosum (Bembidion) 61 Calomera 25 Calosoma 29 Calosoma s.str. 29 calydonius (Ditomus) 95 calydonius calydonius (Ditomus) 95 campestris (Cicindela) 26, 27 campestris campestris (Cicindela) 26 campestris olivieria (Cicindela) 26, 27 campestris pontica (Cicindela) 26 cancellatus (Carabus) 36 cancellatus alessiensis (Carabus) 36 cancellatus emarginatus (Carabus) 36 cancellatus subgraniger var. alessiensis (Carabus) 36 cancellatus var. alessiensis (Carabus) 36 CARABINAE 7, 29, 187 CARABINI 29 caraboides (Asaphidion) 52, 53 caraboides balcanicus (Asaphidion) 52 caraboides var. balcanicus (Asaphidion) 52 Carabus 29 Carabus s.str. 30 carbonarium (Agonum) 133 carbonarium carbonarium (Agonum) 133 cardioderus (Polyderis) 74 cardioderus balcanicus (Trechus) 86 carinatus (Abax) 138 carinatus carinatus (Abax) 138 carinatus var. latus (Abax) 138 Carterus 95 Carterus s.str. 95 cassideus (Licinus) 129 cassideus cassideus (Licinus) 129 cassideus dohrni (Licinus) 129 cassideus graecus (Licinus) 129, 130 cassideus latus (Licinus) 129 catenatus (Carabus) 31

catenatus var. albanensis (Carabus) 31 catenulatus (Carabus) 31 catenulatus catenulatus (Carabus) 31 caucasicum (Bembidion) 62 caucasicus (Bradycellus) 118 cavernosus (Carabus) 34 cavernosus cavernosus (Carabus) 34 cavernosus n. cavernosus (Carabus) 34 cavernosus var. sterbai (Carabus) 34 cavicola (Laemostenus) 171 cavicola albanicus (Laemostenus) 171 cavicola modestus (Laemostenus) 171, 172 cavicola var. modestus (Laemostenus) 171, 172 Celia 178 centralis (Harpalus) 98 Cephalota 26 Chaetocarabus 30 chalybaeus (Myas) 148 chalybaeus (Pterostichus) 158 chalybeus (Dyschirius) 49 chalybeus gibbifrons (Dyschirius) 49 chaudoiri (Amara) 181 chaudoiri incognita (Amara) 181 Cheporus 151 chionobatus (Carabus) 30 chionophilus (Carabus) 36 Chiridysus 46 Chlaeniellus 89 CHLAENIINA 89 CHLAENIINI 89 Chlaenites 90 Chlaenius 89 Chlaenius s.str. 91 Chlorodium 58 Cicindela 26 Cicindela s.str. 26 CICINDELINAE 7, 12, 25, 187 CICINDELINI 25 cimmerius (Laemostenus) 173 cimmerius (Pristonychus) 173 cimmerius (Sphodrus) 173 cimmerius cimmerius (Laemostenus) 173 cimmerius weiratheri (Laemostenus) 173 cinctus (Calathus) 169, 170, 171 circumdata (Cephalota) 26 circumdata circumdata (Cephalota) 26 INDEX

227

circumscriptus (Chlaenius) 92 circumseptus (Calathus) 166 clathratus (Carabus) 31 clathratus auraniensis (Carabus) 31 Clivina 44 CLIVININA 44 CLIVININI 44 clypeatus (Aristus) 96 clypeatus (Ditomus) 96 clypeatus (Dixus) 96 Cnecostolus 43 coadunata (Cymindis) 121 coadunata coadunata (Cymindis) 121 coerulescens (Poecilus) 151 coerulescens (Pterostichus) 151 coeruleum (Bembidion) 57 coeruleum concoeruleum (Bembidion) 57 collaris (Bradycellus) 118 collaris (Clivina) 44 columbinus (Ophonus) 113 combustum (Bembidion) 60, 78 combustum combustum (Bembidion) 60 comita (Tapinopterus) 164 complanata (Amara) 180 complanata (Eurynebria) 16 complanata albanica (Eurynebria) 16 complanata orientalis (Eurynebria) 16 complanata var. albanica (Eurynebria) 16 complanatum (Bembidion) 55 concoeruleum (Bembidion) 57 concoeruleus (Ocydromus) 57 conforme (Bembidion) 55 connexus (Polystichus) 185 consularis (Amara) 178 convexicollis (Pogonistes) 78 convexior (Amara) 175 convexiusculus (Argutor) 153 convexiusculus (Pterostichus) 153 convexiusculus (Tapinopterus) 161 convexus (Carabus) 36 convexus dilatatus (Carabus) 36 convexus dilatatus n. miliusi (Carabus) 36 convexus dilatatus var. miliusi (Carabus) 36 convexus miliusi (Carabus) 36 convexus var. chionophilus (Carabus) 36 corax (Pterostichus) 160 228

corax sic! (Calathus) 168 cordatus (Ophonus) 111 coriaceus (Carabus) 35 coriaceus cerisyi (Carabus) 35 coriaceus cerisyi m. weiratheriana (Carabus) 35 coriaceus excavatus (Carabus) 35 coriaceus excavatus dalmaticus (Carabus) 36 coriaceus excavatus m. dalmatica (Carabus) 35 coriaceus excavatus n. joniensis (Carabus) 35 coriaceus joniensis (Carabus) 36 coriaceus joniensis var. albanicus (Carabus) 36 coriaceus mediterraneus (Carabus) 36 coriaceus rugosus (Carabus) 35 coriaceus rugosus n. joniensis m. albanicus (Carabus) 35 coriaceus subrugosus (Carabus) 35 coriaceus subrugosus v. weiratherianus (Carabus) 35 coriaceus var. hopffgarteni (Carabus) 35 corticalis (Microlestes) 123 corticalis escorialensis (Microlestes) 123 crepitans (Brachinus) 40 crepitans ab. fallax (Brachinus) 40 cribricollis (Harpalus) 110 cribricollis (Ophonus) 110 croaticus (Carabus) 32 croaticus babinjensis (Carabus) 32 croaticus durmitorensis (Carabus) 32 croaticus durmitorensis var. babinjensis (Carabus) 32 croaticus durmitorensis var. ljubetensis (Carabus) 32 croaticus ljubetensis (Carabus) 32 croaticus ljubetensis m. durmitorensis (Carabus) 32 croaticus ljubetensis n. ljubetensis (Carabus) 32 cruciatum (Bembidion) 68 cruciatum albanicum (Bembidion) 68 cruralis (Chlaenius) 91 cruxmajor (Panagaeus) 133 cruxmajor ab. schaumi (Panagaeus) 133 cruxminor (Lebia) 126 cruxminor cruxminor (Lebia) 126 Cryptophonus 98 cupreus (Harpalus) 105 cupreus (Poecilus) 149 cupreus (Pterostichus) 149 cupreus cupreus (Poecilus) 149 cupreus fastuosus (Harpalus) 105 cursitans (Amara) 180 cursor (Pterostichus) 151


cursorius (Poecilus) 150 cursorius (Pterostichus) 150 cursorius cursorius (Poecilus) 150 curta (Amara) 175 Curtonotus 178 curtulus (Molops) 144 curtulus cakorensis (Molops) 144 curtulus winklerianus (Molops) 144 cyaneomicans (Bembidion) 61 cyanicorne (Asaphidion) 52 cyanicorne cyanicorne (Asaphidion) 52 cyanocephala (Lebia) 125, 126 cyanocephala cyanocephala (Lebia) 125 CYCHRINI 37 Cychrus 37 CYCLOSOMINI 92 Cylindera 28 Cylindera s.str. 28 cylindricus (Dyschirius) 47 cylindricus hauseri (Dyschirius) 47 CYMINDIDINA 121 Cymindis 121 Cymindis s.str. 121 dahli (Nebria) 20 dahli montenegrina (Nebria) 20 dahli velebitica (Nebria) 20 dalmaticum sic! (Bembidion) 67 dalmaticus (Carabus) 35 dalmatina (Amara) 179 dalmatinum (Bembidion) 67, 90 dalmatinum dalmatinum (Bembidion) 67 dalmatinus (Laemostenus) 172 dalmatinus (Platyderus) 165 dalmatinus dalmatinus (Platyderus) 165 dalmatinus malissorum (Neotrechus) 84 dalmatinus pentheri (Neotrechus) 84 dalmatinus procerus (Carabus) 32 dalmatinus suturalis var. pentheri (Neotrechus) 84 Dalmatoreicheia 45 dama (Carterus) 95 dama (Ditomus) 95 danieli (Harpalus) 104 danieli (Notiophilus) 24 decipiens (Chlaenius) 91 decipiens laticollis (Chlaenius) 91 decipiens var. ambiguus (Chlaenius) 91

decolor (Bembidion) 69 decorum (Bembidion) 63 decorum decorum (Bembidion) 63 dejeani (Chlaenius) 92 dejeani (Epomis) 92 dejeani (Parophonus) 114 deletum (Bembidion) 67 deletum deletum (Bembidion) 67 Deltomerus 76 Deltomerus s.str. 76 dentata (Drypta) 93 depressa (Siagona) 44 detonii (Tapinopterus) 164 diabrachys (Tachys) 76 diabrachys (Tachyura) 76 Diachromus 94 Dicheirotrichus 118 Dicheirotrichus s.str. 118 didonnai (Duvalius) 80 diffinis (Harpalus) 113 diffinis (Ophonus) 113 dilatatus (Badister) 129 diligens (Pterostichus) 153 dimidiatus (Harpalus) 105 dinaricus (Cychrus) 37 Dinodes 91 Diplocampa 58 discophorus (Acupalpus) 119 discophorus (Stenolophus) 119 distinctissimus (Trechus) 87 distinctus (Bradycellus) 118 distinguendus (Calathus) 167 distinguendus (Harpalus) 107 distinguendus distinguendus (Harpalus) 107 DITOMINA 95 Ditomus 95 Dixus 96 djumanus (Pterostichus) 158 dochii (Pterostichus) 164 dochii (Tapinopterus) 164 doderoi (Sinechostictus) 73 dorsale (Platynus) 136 dorsalis (Acupalpus) 117 dorsalis (Anchomenus) 136 dorsalis (Platynus) 136

INDEX

229

DROMIINA 123 Dromius 123 Dromius s.str. 123 droveniki (Duvalius) 79 Drypta 93 Drypta s.str. 93 DRYPTINI 93 duftschmidi (Agonum) 133 Duvalius 79 Duvalius s.str. 80 dyrrhachii (Bembidion) 63 DYSCHIRIINI 46 Dyschiriodes 46 Dyschirius 12, 46 Dyschirius s.str. 49 effluviorus (Sinechostictus) 73 Egadroma 119 egregius (Trechus) 87 ejaculans (Brachinus) 41 ELAPHRINAE 38, 187 ELAPHRINI 38 Elaphrus 38 elegans (Acupalpus) 115 elegans (Bembidion) 52, 53, 57, 58 elegans (Brachinus) 41 ellipticus (Calathus) 167 ellipticus ellipticus (Calathus) 167 elongatum (Bembidion) 73 elongatum tarsicum (Bembidion) 73 elongatum var. effluviorum (Bembidion) 73 elongatus (Laemostenus) 172 elongatus (Pterostichus) 152 elongatus (Sinechostictus) 73 elongatus pentheri (Laemostenus) 172 emarginatus (Acinopus) 97 Emphanes 58 eomita sic! (Tapinopterus) 164 ephippium (Bembidion) 62 epirensis (Carabus) 31 epiroticus (Pterostichus) 160 Epomis 92 equestris (Amara) 179, 180 equestris equestris (Amara) 179, 180 eremita (Ditomus) 96 eremita (Dixus) 96 erratica (Amara) 177 230

erratus (Calathus) 170 erratus erratus (Calathus) 170 erythrocnema (Amara) 181 erythroderus (Calathus) 169 escorialensis (Microlestes) 123 etruscus (Gynandromorphus) 95 etruscus etruscus (Gynandromorphus) 95 Eucarabus 31 euchlorus (Harpalus) 105 Euduvalius 82 Eudyschirius 49 Eugrapha 28 Eunebria 22 Euperyphus 60 europaea (Siagona) 44 europaea europaea (Siagona) 44 europaea oberleitneri (Siagona) 44 europaea var. oberleitneri (Siagona) 44 Europhilus 136 Eurynebria 16 eurynota (Amara) 175 exhalans (Brachinus) 43 exilis (Microlestes) 124 exilis luctuosus (Microlestes) 124 explodens (Brachinus) 42 explodens ab. glabratus (Brachinus) 42 explodens ab. strepens (Brachinus) 42 explodens sichemita (Brachinus) 42 extensoides Jedlička, 1936 (Tapinopterus) 161, 162 extensoides Straneo, 1936 (Tapinopterus) 161, 162 extensum (Agonum) 133 extensus (Tapinopterus) 161, 162 extensus abreshensis (Tapinopterus) 161 extensus convexiusculus (Pterostichus) 161 extensus extensoides (Tapinopterus) 161, 162 fallaciosa (Amara) 179 fallaciosus (Zabrus) 179 fallax (Brachinus) 40 fallax (Parophonus) 114 fallax (Tachyophonus) 114 famelica (Amara) 175 familiaris (Amara) 175 fasciatopunctatus (Pterostichus) 155 fasciolatum (Bembidion) 56, 57 femoralis (Carterophonus) 96 femoralis (Graniger) 96


femoratum (Bembidion) 68 femoratum femoratum (Bembidion) 68 Feronidius 152 festivus (Chlaenius) 91 festivus festivus (Chlaenius) 91 fischeri (Calomera) 25 fischeri (Cicindela) 25 fischeri fischeri (Calomera) 25 fissuralis (Microlestes) 123 flavescens (Harpalus) 103 flavicornis (Harpalus) 101 flavicornis flavicornis (Harpalus) 101 flavipes (Asaphidion) 52, 53 flavipes (Chlaenius) 89 fodori (Duvalius) 82 foraminosum (Bembidion) 66 fossor (Clivina) 45 fossor fossor (Clivina) 45 franziniorum (Ophonus) 113 fuchsi (Duvalius) 81 fugax (Harpalus) 108 fulvibarbis (Leistus) 16 fulvibarbis fulvibarbis (Leistus) 16 fulvicollis (Paratachys) 74 fulvicollis (Tachys) 74 fulvipes (Amara) 181 fumigatum (Bembidion) 58 fusca (Amara) 180 fuscatus (Olisthopus) 137 fuscipes (Calathus) 167 fuscipes fuscipes (Calathus) 167 fuscipes graecus (Calathus) 167 fuscipes var. graecus (Calathus) 167 fuscipes var. latus (Calathus) 167 galilaeus (Lymnastis) 73 ganglbaueri (Brachinus) 41 ganglbaueri (Bradycellus) 118 ganglbaueri (Nebria) 20 ganglbaueri (Synuchidius) 171 ganglbaueri (Zabrus) 183 ganglbaueri ditana (Nebria) 21 ganglbaueri ganglbaueri (Nebria) 20 ganglbaueri ganglbaueri (Zabrus) 183 ganglbaueri gyalicaensis (Nebria) 20 ganglbaueri korabensis (Nebria) 20 ganglbaueri lonae (Zabrus) 183

ganglbaueri merditana (Nebria) 20, 21 ganglbaueri zebiaensis (Nebria) 21 genei illigeri (Bembidion) 62 geniculatum (Bembidion) 55 geniculatum geniculatum (Bembidion) 55 georgii (Duvalius) 80 germanica (Cicindela) 28 germanica (Cylindera) 28 germanica germanica (Cylindera) 28 germanica muelleri (Cylindera) 28 germanus (Diachromus) 94 germarii (Nebria) 21 germarii durmitorensis (Nebria) 21 germinyi (Notiophilus) 24 gestroi (Tapinopterus) 165 gibbifrons (Dyschirius) 49 giganteus (Calathus) 167, 168 gigas (Carabus) 35 gigas gigas (Carabus) 35 gigas parnassicus (Carabus) 35 gilvipes (Carterus) 95 gisellae (Agonum) 134 glabratus (Brachinus) 42 glabricollis (Calathus) 168 glabricollis (Olisthopus) 137 globosus (Dyschirius) 49 gracilis (Dyschirius) 49, 50 gracilis (Oodes) 131 gracilis (Pogonistes) 78 gracilis (Pterostichus) 155 gracilis gracilis (Dyschirius) 49 gracilis gracilis (Pterostichus) 155 graecus (Carabus) 35 graecus (Licinus) 129, 130 graecus (Platyderus) 165 graecus graecus (Carabus) 35 graecus morio (Carabus) 35 graecus morio var. neowinkleri (Carabus) 35 graecus morio var. winkleri (Carabus) 35 graecus n. winkleri (Carabus) 35 graecus winkleri (Carabus) 35 grandipenne (Bembidion) 67 grandipenne grandipenne (Bembidion) 67 Graniger 96 granulatus (Carabus) 30 granulatus aetolicus (Carabus) 30 INDEX

231

granulatus granulatus m. merdita sic! (Carabus) 30 granulatus granulatus n. interstitialis (Carabus) 30 granulatus interstitialis (Carabus) 30 granulatus interstitialis v. miridita (Carabus) 30 granulatus miridita (Carabus) 30 granulatus m. miridita (Carabus) 30 granulatus var. miridita (Carabus) 30 granulipennis (Aptinus) 39 griseus (Harpalus) 108 gudenzii (Bembidion) 64 guttula (Bembidion) 69 guttula guttula (Bembidion) 69 gyllenhali (Nebria) 22 Gynandromorphus 95 haemorrhoidalis (Sphaerotachys) 75 haemorrhoidalis (Tachys) 75 hannemanni (Amara) 178 HARPALINA 97 HARPALINAE 7, 89, 187 HARPALINI 12, 93 harpaloides (Ocys) 72 Harpalus 12, 98 Harpalus s.str. 99 helopioides (Oodes) 131 helopioides helopioides (Oodes) 131 herzegowinensis (Carabus) 34 Hesperophonus 109 hilleri (Carabus) 34 hirsutulus (Harpalus) 113 hirsutulus (Ophonomimus) 113 hirsutulus (Ophonus) 113 hirsutulus (Parophonus) 113 holdhausi (Agonum) 134 holdhausi (Molops) 142, 143, 144, 145 holdhausi (Platynus) 134 honestus (Harpalus) 99 honestus honestus (Harpalus) 99 hopffgarteni (Carabus) 35 hortensis (Carabus) 33, 34 hortensis hortensis (Carabus) 34 hortensis neumeyeri (Carabus) 33, 34 hortensis neumeyeri n. herzegowinensis (Carabus) 34 hortensis neumeyeri n. neumeyeri (Carabus) 34 hortensis neumeyeri var. albanicola (Carabus) 34 hortensis neumeyeri var. hilleri (Carabus) 34 hortensis neumeyeri var. pindicus (Carabus) 34 232

hortensis preslii (Carabus) 33 hortensis preslii m. jonica (Carabus) 34 hortensis rhodopensis (Carabus) 34 hospes (Harpalus) 106 hospes hospes (Harpalus) 106 humerale (Bembidion) 53 humeralis (Cymindis) 121 humeralis (Lebia) 126 hybrida (Cicindela) 27 hybrida albanica (Cicindela) 27 hybrida riparia (Cicindela) 27 Hygrocarabus 31 hypocrita (Agonum) 134 hypocrita (Bembidion) 66 hypocrita (Notiophilus) 24 hypocrita illyricum (Bembidion) 66 Hypogeobium 160 Hypogium 160 hypsobius (Carabus) 36 ignavus (Zabrus) 183 illigeri (Bembidion) 62 immaculicornis (Brachinus) 42 immaculicornis immaculicornis (Brachinus) 42 immundus (Acupalpus) 116 importunus (Dyschirius) 50 importunus importunus (Dyschirius) 50 impressum sic! (Syntomus) 127 impressus (Metabletus) 127 impressus (Syntomus) 127 impressus impressus (Syntomus) 127 inaequalis (Tachys) 76 incertus (Brachinus) 43 Incisophonus 111 incisus (Ophonus) 111 incrassatus (Zabrus) 184 incrassatus bischoffi (Zabrus) 184 incrassatus boldorii (Zabrus) 184 incrassatus incrassatus (Zabrus) 184 inoptatum (Bembidion) 70 inquinatus (Pedius) 149 inquinatus (Pterostichus) 149 inquisitor (Calosoma) 29 inquisitor inquisitor (Calosoma) 29 inserticeps (Bembidion) 61 intermedius (Anisodactylus) 94 intermedius (Dyschirius) 46


intricatus (Carabus) 30 intricatus chinobatus (Carabus) 30 intricatus intricatus (Carabus) 30 intricatus intricatus m. montenegrinus (Carabus) 30 intricatus intricatus n. weiratheri (Carabus) 30 intricatus intricatus var. weiratheri (Carabus) 30 intricatus montenegrinus (Carabus) 30 intricatus var. montenegrinus (Carabus) 30 irenis (Trechus) 86 iricolor (Bembidion) 70 irregularis (Carabus) 35 irregularis bucephalus (Carabus) 35 irregularis ramanus (Carabus) 35 janthinus (Laemostenus) 172 janthinus janthinus (Laemostenus) 172 javurkovae (Bembidion) 68 jockischi (Nebria) 22 jockischi jockischi (Nebria) 22 jonicus (Carabus) 34 joniensis (Carabus) 35 justinae (Bembidion) 55 kaufmanni (Tapinopterus) 161 klisuranus (Molops) 144, 146 kobingeri (Trechus) 87 kobingeri bulgaricus (Trechus) 88 kobingeri kobingeri (Trechus) 87, 88 kobingeri pawlowskianus (Trechus) 88 korabensis (Carabus) 33 korabica (Nebria) 21 korax (Calathus) 167, 168 koyi (Poecilus) 150 koyi (Pterostichus) 150 koyi dinaricus (Pterostichus) 150 koyi koyi (Poecilus) 150 kratteri (Nebria) 23 kratteri kratteri (Nebria) 23 kratteri valonensis (Nebria) 23 krueperi (Harpalus) 109 krueperi (Ophonus) 109 Laemostenus 171 Laemostenus s.str. 172 laeviceps (Parophonus) 114, 115 laevicollis (Trichotichnus) 115 laevicollis laevicollis (Trichotichnus) 115 laevifrons (Clivina) 45 laevigatus (Scarites) 50

laevipes (Harpalus) 101 laeviusculus (Dyschirius) 47 lafertei (Dyschirius) 49 Lamprias 125 lampros (Bembidion) 60 lampros var. properans (Bembidion) 61 laterale (Bembidion) 61 laterale pseudolaterale (Bembidion) 61 laticollis (Chlaenius) 91 laticollis (Ophonus) 111 latifianus (Pterostichus) 157 latifianus ab. nigripes (Pterostichus) 158, 159 latifianus latifianus (Pterostichus) 156, 157 latifianus strupii (Pterostichus) 156, 157, 158, 159 latipennis (Dyschirius) 49 latiplaga (Bembidion) 59 latiplaga latiplaga (Bembidion) 59 latus (Abax) 138 latus (Calathus) 167 latus (Harpalus) 103 latus (Licinus) 129 Lebia 125 Lebia s.str. 126 LEBIINA 125 LEBIINI 121 Leistus 16 Leistus s.str. 16 leonhardi stenocephalus (Duvalius) 81 leonhardi sturanyi (Duvalius) 81, 82 lepidus (Poecilus) 150 lepidus (Pterostichus) 150 lepidus lepidus (Poecilus) 150 levantinus (Amblystomus) 93 levantinus levantinus (Amblystomus) 93 LICININA 128 LICININI 128 Licinus 129 Licinus s.str. 129 limbatum (Omophron) 38 limbopunctatus (Harpalus) 107 Limnocarabus 31 linearis (Dromius) 125 linearis (Paradromius) 125 linearis linearis (Paradromius) 125 lineata (Cymindis) 122 INDEX

233

lineola (Cymindis) 122 LIONYCHINA 127 Lionychus 127 Lionychus s.str. 127 literata sic! (Cicindela) 28 litorale (Bembidion) 57, 58 litoralis sic! (Pogonus) 78 litterata (Cicindela) 28 littoralis (Calomera) 26 littoralis (Pogonus) 78 littoralis nemoralis (Calomera) 26 ljubetensis (Carabus) 32 lodosi (Pterostichus) 153 lonae (Neotrechus) 82 lonae (Trechus) 82 lonae atavus (Neotrechus) 83 lonae lonae (Neotrechus) 82, 83 lonae shkodrensis (Neotrechus) 83 lonai (Omphreus) 131 lonai sic! (Neotrechus) 82 lonai sic! atavus (Neotrechus) 83 lonai sic! lonai sic! (Neotrechus) 82 lonai sic! shkodrensis (Neotrechus) 83 longicollis (Pedius) 149 longicorne (Agonum) 134 longicornis (Anthracus) 117 longicornis (Thalassophilus) 88 Loxoncus 119 lucida (Amara) 176 lucidus (Dyschirius) 46 luctuosus (Microlestes) 124 luctuosus luctuosus (Microlestes) 124 lugubris rufipes (Aptinus) 39, 40 lumensis (Pterostichus) 157, 158 lumensis litae (Pterostichus) 156, 158 lumensis ljubetensis (Pterostichus) 156, 158 lumensis lumensis (Pterostichus) 156, 157 lunatus (Callistus) 89 lunatus lunatus (Callistus) 89 lunulata (Cicindela) 26 lunulata var. nemoralis (Cicindela) 26 lunulatum (Bembidion) 70 luridipennis (Pogonus) 78 luteatus (Acupalpus) 115 luticola (Dyschirius) 47 luticola luticola (Dyschirius) 47 234

Lymnastis 73 macedonica (Nebria) 18, 21 macedonica galicica (Nebria) 18, 21 macedonica papingutica (Nebria) 18, 21 macedonica rambouseki (Nebria) 18, 22 macedonicus (Calathus) 168 macedonicus (Pachycarus) 97 macroderus (Dyschirius) 48 macroderus macroderus (Dyschirius) 48 maculatum (Bembidion) 72 maculatum maculatum (Bembidion) 72 maculatus (Acupalpus) 116, 117 maculicornis (Harpalus) 114 maculicornis (Ophonus) 114 maculicornis (Parophonus) 114 maculicornis (Trichotichnus) 114 maderi (Molops) 139 maglicensis (Duvalius) 79 magnicollis (Leistus) 17 malissorum (Carabus) 31, 32 malissorum (Deltomerus) 76, 77 malissorum (Neotrechus) 83, 84 malissorum (Molops) 145 malissorum (Pterostichus) 156, 158, 159 malissorum (Trechus) 84 malissorum ab. munella (Pterostichus) 157 malshentianus (Molops) 139 mannerheimii (Bembidion) 70 Manodromius 125 marginata (Egadroma) 119 marginatum (Agonum) 134 marginatus (Stenolophus) 119 maritimus (Lionychus) 127 MASOREINA 92 Masoreus 92 Masoreus s.str. 92 maurus (Microlestes) 124 maurus maurus (Microlestes) 124 megacephalus (Acinopus) 98 Megodontus 29, 31 meisteri (Pterostichus) 156 melanarius (Pterostichus) 153 melanarius melanarius (Pterostichus) 153 melancholica (Myriochila) 28 melancholica melancholica (Myriochila) 28 Melanius 152


melanocephalus (Calathus) 170 melanocephalus melanocephalus (Calathus) 170 melanura (Odacantha) 131 melas (Pterostichus) 152 melas depressus (Pterostichus) 152 melletii (Ophonus) 111 Menas 122 mendax (Harpalus) 114 mendax (Ophonus) 114 mendax (Parophonus) 114 mendax (Tachyophonus) 114 merditana (Nebria) 20, 21 merditanum (Bembidion) 57 merditanus (Aptinus) 39 merditanus (Carabus) 33 merditanus (Molops) 141, 142 merditanus (Trechus) 86 merditanus intermedius (Molops) 142 merditanus merditanus (Aptinus) 39, 40 merditanus merditanus (Molops) 141 merditanus var. granulipennis (Aptinus) 40 meridianus (Acupalpus) 116 meridionale (Bembidion) 59 meridionalis (Ophonus) 110 merklii (Pterostichus) 156 meschniggi (Bembidion) 56 messae (Amara) 180 metalkanus (Carabus) 31 metallescens (Amblystomus) 93 metallicus (Calathus) 169, 170, 171 metallicus (Pterostichus) 151 metallicus metallicus (Calathus) 170 metallicus prokletijensis (Calathus) 170 Metallina 60 Metophonus 111 Microcallisthenes 29 Microlestes 12, 123 micros (Paratachys) 74 micros (Tachys) 74 Microserrullula 61 miliaris (Cymindis) 123 miliusi (Carabus) 36 milleri (Bembidion) 67, 68 milleri milleri (Bembidion) 68 millerianus (Sinechostictus) 73 minor (Pterostichus) 155

minor minor (Pterostichus) 155 minutulus (Blechrus) 124 minutulus (Microlestes) 124 minutulus (Speluncarius) 160 minutulus (Tapinopterus) 160 minutulus apfelbecki (Microlestes) 124 minutus (Dyschirius) 48 minutus (Platyderus) 166 minutus albanicus (Dyschirius) 48 minutus obtusangulus (Platyderus) 166 miridita (Tapinopterus) 164 miridita jablanicensis (Tapinopterus) 164, 165 miridita jacupicensis (Tapinopterus) 165 miridita miridita (Tapinopterus) 164 miriditus (Carabus) 30 mixtus (Stenolophus) 120 moestum (Agonum) 133 moestus (Platynus) 133 mollis (Calathus) 171 mollis f. erythroderus (Calathus) 169 mollis mollis (Calathus) 171 Molops 139 Molops s.str. 139 monachum var. binaghii (Agonum) 133 monastirensis (Tapinopterus) 165 monastirensis gestroi (Tapinopterus) 165 montana (Amara) 178 montenegrinus (Carabus) 30 montenegrinus (Molops) 147, 148 montenegrinus cavernicola (Molops) 147 monticola (Cicindela) 27 monticola albanica (Cicindela) 27 montivagus (Carabus) 30 montivagus m. sutomorensis (Carabus) 30 montivagus vellepiticus (Carabus) 30 morio (Amara) 176 morio (Dyschirius) 48 morio (Omphreus) 132 morio albanicus (Omphreus) 132 morio malissorum (Omphreus) 132 morio nivium (Amara) 176 Morphnosoma 153 muelleri (Agonum) 134, 135 muelleri (Duvalius) 80 muelleri (Platynus) 134 munella (Pterostichus) 157 INDEX

235

municipalis (Amara) 180 municipalis bischoffi (Amara) 180 Myas 148 Myas s.str. 148 Myriochila 28 Myriochila s.str. 28 Mystropterus 97 nana (Tachyta) 75 Nebria 12, 18 Nebria s.str. 22 NEBRIINAE 7, 16, 187 NEBRIINI 16 nebulosum (Asaphidion) 52, 53 nebulosum balcanicum (Asaphidion) 52, 58 negrita (Microlestes) 124 negrita apfelbecki (Microlestes) 124 nemoralis nemoralis (Lophyridia) 26 nemorivagus (Anisodactylus) 94 Neocalathus 169 Neoelaphrus 38 Neorescius 130 Neotrechus 82 neowinkleri (Carabus) 35 Nepha 61 neresheimeri (Bembidion) 70 neumeyeri (Carabus) 33, 34 niger (Amblystomus) 93 niger (Pterostichus) 154 niger niger (Pterostichus) 154 nigricornis (Brachinus) 43 nigricornis (Chlaenius) 67, 89, 90 nigrinus (Trechus) 87 nigripes (Pterostichus) 158 nigrita (Pterostichus) 155 nitidulus (Chlaenius) 90 nitidus (Dyschirius) 46 nitidus (Leistus) 17 nitidus nitidus (Dyschirius) 46 nivalis (Synuchus) 173 nodulosus (Carabus) 31 normannum (Bembidion) 59 normannum apfelbecki (Bembidion) 59 normannum mediterraneum (Bembidion) 59 normannum meridionale (Bembidion) 59 Notaphemphanes 62 Notaphus 63 236

notatus (Acupalpus) 116 notatus (Dromius) 125 notatus (Philorhizus) 125 NOTIOPHILINI 23 Notiophilus 12, 23 oberthuri (Ophonus) 109 oblitus (Harpalus) 108 oblitus oblitus (Harpalus) 108 oblongopunctatus (Pterostichus) 151 oblongopunctatus oblongopunctatus (Pterostichus) 151 oblongum (Bembidion) 60 obscuroguttatus (Metabletus) 128 obscuroguttatus (Syntomus) 128 obscurus (Aristus) 96 obscurus (Ditomus) 96 obscurus (Dixus) 96 obsoletus (Dicheirotrichus) 118 obtusangulus intermedius (Molops) 142 obtusiusculus (Trechus) 88 obtusus (Trechus) 85 obtusus obtusus (Trechus) 85 octomaculatum (Bembidion) 72 Ocydromus 63 Ocys 72 Ocyturanes 64 Odacantha 130 Odacanta s.str. 131 ODACANTHINI 130 Odontium 66 Odontocarus 97 Oedematicus 98 oertzeni (Licinus) 130 oertzeni albanicus (Licinus) 130 oertzeni oertzeni (Licinus) 130 olens (Zuphium) 186 olens olens (Zuphium) 186 Olisthopus 12, 137 Omoperyphus 66 Omophron 38 Omophron s.str. 38 OMOPHRONINAE 7, 38, 187 OMOPHRONINI 38 OMPHREINI 131 Omphreus 131 Omphreus s.str. 131


Oodes 131 OODINI 131 Ophonomimus 113 Ophonus 12, 109 Ophonus s.str. 112 Oreocarabus 33 Oreoplatysma 153 Osimus 98 osmanilis (Molops) 145 osmanilis osmanilis (Molops) 145, 146 osmanilis tomoricensis (Molops) 145, 146 ottomanus (Pterostichus) 154 ottomanus ottomanus (Pterostichus) 154 ovalis (Abax) 139 ovata (Amara) 176 Pachycarus 97 Pachystus 34 pallipes (Metabletus) 128 pallipes (Syntomus) 128 paludicola (Acupalpus) 116, 117 palustris (Notiophilus) 24 PANAGAEINI 133 Panagaeus 133 Panagaeus s.str. 133 Paracelia 179 Paradeltomerus 77 paradoxus (Deltomerus) 77 paradoxus korabensis (Deltomerus) 77 paradoxus paradoxus (Deltomerus) 77 Paradromius 125 Paradyschirius 50 Parahaptoderus 153 Parallelomorphus 50 parallelus (Dyschirius) 50 parallelus (Ophonus) 111 parallelus albanensis (Harpalus) 111 parallelus albanicus (Ophonus) 111 parallelus ruficornis (Dyschiriodes) 50 parallelus ruficornis (Dyschirius) 50 Paranchus 137 Parapercosia 179 Paratachys 74 parnassicola (Molops) 140 parnassicola alinae (Molops) 140 parnassicola valonensis (Molops) 140 parnassicus (Carabus) 35

Parophonus 12, 113 Parophonus s.str. 114 parreyssi (Molops) 141 parreyssi lonae (Molops) 141 parumpunctatus (Deltomerus) 77 parvicollis (Leistus) 17 parvula (Tachyura) 76 parvulus (Acupalpus) 117 parvulus (Tachys) 76 PATROBINI 76 Pedius 149 Pelor 181 peltatus (Badister) 129 peltatus peltatus (Badister) 129 pentheri (Calosoma) 29 pentheri (Pterostichus) 156, 159 pentheroides (Calosoma) 29 Percosia 179 peregrinus (Brachinus) 42 PERILEPTINA 79 Perileptus 79 Perileptus s.str. 79 peristericus (Tapinopterus) 162 permoestum (Agonum) 135 Peryphanes 66 Peryphus 68 Phaon 155 Pheggomisetes 88 Philochthus 69 Philorhizus 125 Philorhizus s.str. 125 Phonias 153 Phyla 70 picenus peristericus (Carabus) 33 piceus (Molops) 147 piceus balcanicus (Molops) 147 picipennis (Harpalus) 102 picipes (Acinopus) 97 pindicola (Cymindis) 122 pindicum (Bembidion) 64, 65 pindicus (Carabus) 34 plagiatus (Brachinus) 40, 41 plagiatus (Microlestes) 124 plagiatus escorialensis (Microlestes) 123 planicollis (Acupalpus) 117 planicollis Apfelbeck (Harpalus) 114 INDEX

237

planicollis Apfelbeck (Ophonus) 114 planicollis Apfelbeck (Parophonus) 114, 115 planicollis Dejean (Harpalus) 114 planicollis Dejean (Ophonus) 114 planicollis Dejean (Parophonus) 114 Platycarabus 35 Platyderus 165 Platyderus s.str. 165 PLATYNINI 133 Platynus 138 Platynus s.str. 138 Platypterinus 154 Platysma 154 plurisetosus (Molops) 145 Poecilus 149 Poecilus s.str. 149 POGONINI 78 Pogonistes 78 Pogonophorus 17 Pogonus 78 Pogonus s.str. 78 Polyderis 74 Polystichus 185 pominii (Duvalius) 81 praetermissa (Amara) 178 praeustum (Bembidion) 65, 74 praeustum praeustum (Bembidion) 65 praeustum viridifluum (Bembidion) 65 preslii (Carabus) 33, 34 preslii neumeyeri (Carabus) 34 priapus (Trechus) 86 priapus priapus (Trechus) 86 Princidium 71 Pristonychus 173 Procerus 29, 35 procerus (Anoplogenius) 119 procerus (Carabus) 32 procerus (Loxoncus) 119 Procrustes 29, 35 properans (Amara) 180 properans (Bembidion) 61 protensus (Tapinopterus) 162 protensus aetolicus (Tapinopterus) 162, 163 protensus peristericus (Tapinopterus) 162 protensus tomoricensis (Tapinopterus) 163 proxima (Amara) 176 238

proximus (Stenolophus) 120 Pseudodichirius 94 pseudolaterale (Bembidion) 61 Pseudolimnaeum 73 Pseudomaseus 154 Pseudophonus 108 psophia (Brachinus) 41 PTEROSTICHINI 138 Pterostichus 151 Pterostichus s.str. 155 pubescens (Harpalus) 108 pubescens (Ophonus) 108 pueli (Anisodactylus) 94 pueli pueli (Anisodactylus) 94 pumicatus (Stomis) 160 pumicatus pumicatus (Stomis) 160 pumilus (Harpalus) 102 punctatostriatus (Harpalus) 106 punctatulus (Harpalus) 111 punctatulus (Ophonus) 111 punctatus (Dyschirius) 48 punctatus (Laemostenus) 173 punctatus (Leistus) 18 punctatus albanicus (Dyschirius) 48 puncticeps (Harpalus) 112 puncticeps (Ophonus) 112 puncticollis Coquerel (Acupalpus) 117 puncticollis Auct. nec Coquerel (Acupalpus) 116 puncticollis (Harpalus) 112 puncticollis (Ophonus) 112 puncticollis (Poecilus) 149 puncticollis (Pterostichus) 149 punctulatum (Bembidion) 71 punctulatum punctulatum (Bembidion) 71 punctulatus sic! (Ophonus) 111 pusillus (Dyschirius) 48 pusillus (Notiophilus) 23 pygmaeus (Harpalus) 106 quadricolle (Bembidion) 61 quadriguttatum (Fabricius) (Bembidion) 53 quadriguttatum (Olivier) (Bembidion) 54 quadrillum (Lionychus) 127 quadrimaculatum (Bembidion) 53, 54 quadrimaculatum quadrimaculatum (Bembidion) 53 quadrimaculatus (Dromius) 123


quadripunctatus (Harpalus) 101 quadripustulatum (Bembidion) 54 quadripustulatum quadripustulatum (Bembidion) 54 quadristriatus (Trechus) 85 quarnerensis (Anthracus) 117 quenseli (Amara) 179 quenseli quenseli (Amara) 179 ravasinianus (Trechus) 88 ravasinii (Calathus) 168 ravasinii (Omphreus) 132 ravasinii (Trechus) 88 ravasinii macedonicus (Calathus) 168 ravasinii ravasinii (Calathus) 168 rebeli (Poecilus) 150 rebeli (Pterostichus) 150 reflexicollis (Amara) 181 reflexicollis reflexicollis (Amara) 181 regiszogui (Duvalius) 81 Reicheadella 45 Reicheadella s.str. 45 REICHEINA 45 reiseri (Bembidion) 66 reiseri (Molops) 143, 144 reiseri (Pterostichus) 156, 159 reiseri corallipes (Molops) 142 reiseri holdhausi (Molops) 142, 143, 144, 145 reiseri pseudoalpestris (Molops) 142 reiseri reiseri (Bembidion) 65 reiseri reiseri (Molops) 142, 143 reiseri victoris (Molops) 142, 143, 144 reiseri vranense (Bembidion) 65 relictum (Bembidion) 55 relictum (Calosoma) 29 relictum var. pentheroides (Calosoma) 29 reticulatus (Apristus) 127 rhilensis (Pterostichus) 155 rhilensis kourili (Pterostichus) 156 rhilensis rhilensis (Pterostichus) 156 rhilensis vitosensis (Pterostichus) 156 rhodopense (Bembidion) 54 rhodopensis (Carabus) 34 rilvensis (Carabus) 32 riparia (Cicindela) 27 riparius (Pogonus) 78 rivulare (Bembidion) 58

robustus (Odontocarus) 97 rossii (Asaphidion) 53 rostratus (Stomis) 161 rostratus rostratus (Stomis) 161 rotundatus (Olisthopus) 137 rotundatus rotundatus (Olisthopus) 137 rotundicollis (Carterus) 95 rubripes (Harpalus) 99 rufescens (Nebria) 22 rufescens rufescens (Nebria) 22 rufibarbis (Harpalus) 112 rufibarbis (Ophonus) 112 ruficornis (Dyschirius) 50 rufipalpis (Harpalus) 100 rufipalpis rufipalpis (Harpalus) 100 rufipes (Harpalus) 108 rufipes (Molops) 146 rufipes (Notiophilus) 25 rufipes (Pseudophonus) 108 rufipes klisuranus (Molops) 146 rufipes munelensis (Molops) 146 rufipes pseudoosmanilis (Molops) 146 rufipes rufipes (Molops) 146 rufipes steindachneri (Molops) 147 rufipes sturanyi (Molops) 147 rufitarsis (Harpalus) 100 rufithorax (Apotomus) 51 rufomarginatus (Leistus) 18 rufus (Apotomus) 51 rufus (Platyderus) 166 rufus rufus (Platyderus) 166 rugosicollis (Myas) 148 rugosus (Carabus) 35 rumelicum (Bembidion) 63 rupicola (Harpalus) 112 rupicola (Ophonus) 112 sabulicola (Harpalus) 113 sabulicola (Ophonus) 113 sabulicola columbinus (Harpalus) 113 sabulicola columbinus (Ophonus) 113 sahlbergi albanica (Cicindela) 27 salina (Nebria) 23 salinus (Dyschirius) 49 salinus striatopunctatus (Dyschirius) 49 saphyrea (Amara) 177 sarajevoensis (Carabus) 31 INDEX

239

saxicola (Harpalus) 108 scabriusculus (Carabus) 37 scabriusculus bulgarus (Carabus) 37 Scallophorites 51 scapulare (Bembidion) 60 scapulare oblongum (Bembidion) 60 scapularis (Cymindis) 122 scapularis (Lebia) 126 scapularis scapularis (Cymindis) 122 scapularis scapularis (Lebia) 126 Scarites 50 SCARITINA 50 SCARITINAE 7, 12, 44, 187 SCARITINI 50 schaubergerianus (Ophonus) 112 schaumi (Panagaeus) 133 scrobiculatus (Platynus) 138 scrobiculatus serbicus (Platynus) 138 scrobiculatus turcicus (Platynus) 138 scutellaris (Tachys) 75 semigranosus (Cychrus) 37 semigranosus albanicus (Cychrus) 37, 38 semigranosus balcanicus (Cychrus) 37 semigranosus dinaricus (Cychrus) 37, 38 semigranosus jakupicensis (Cychrus) 38 semigranosus montanus (Cychrus) 37 semigranosus montenegrinus (Cychrus) 37, 38 semigranosus n. balcanicus (Cychrus) 37 semigranosus n. montenegrinus (Cychrus) 37 semigranosus semigranosus (Cychrus) 37 semigranosus travnikanus (Cychrus) 38 serbicus (Platynus) 138 serripes (Harpalus) 101 serripes ab. subchalybaeus (Harpalus) 101 serripes serripes (Harpalus) 101 sexpunctatum (Agonum) 135 sexstriata (Tachyura) 76 sexstriatus (Tachys) 76 sexstriatus var. bisbimaculatus (Tachys) 76 shardagensis (Carabus) 33 shljebensis (Carabus) 32 Siagona 44 SIAGONINAE 7, 44, 187 SIAGONINI 44 sichemita (Brachinus) 42 siculum (Bembidion) 64 240

siculum smyrnense (Bembidion) 64 sigma (Dromius) 125 sigma (Philorhizus) 125 signatipenne (Bembidion) 65 silphoides (Licinus) 130 similata (Amara) 177 similis (Ophonus) 110 simplex (Molops) 140, 141 simplex meridionalis (Molops) 140, 141 simplex simplex (Molops) 140 simulator (Dyschirius) 48 Sinechostictus 10, 73 Sinechostictus s.str. 73 skrimshiranus (Stenolophus) 120 skutariensis (Harpalus) 104 smaragdinus (Harpalus) 104 solitaris (Harpalus) 103 sordidum (Agonum) 135 sordidum sordidum (Agonum) 135 sordidus (Platynus) 135 spartanus (Molops) 140 spartanus parnassicola (Molops) 140 spartanus valonensis (Molops) 140 Speluncarius 12, 160 Speluncarius s.str. 160 Sphaerotachys 75 SPHODRINA 171 SPHODRINI 165 spinibarbis (Leistus) 17, 18 spinibarbis punctatus (Leistus) 18 spinibarbis rufipes (Leistus) 18 spinipes (Zabrus) 185 spinipes insignis (Zabrus) 185 splendidum (Bembidion) 58 splendidum pincum (Bembidion) 58 spoliatus (Chlaenius) 90 spoliatus spoliatus (Chlaenius) 90 spreta (Amara) 177 spurcaticornis (Anisodactylus) 94 steindachneri (Molops) 147 stenocephalus (Duvalius) 82 stenocephalus (Trechus) 81 Stenochoromus 147 STENOLOPHINA 12, 115 Stenolophus 119 Stenolophus s.str. 119


sterbai (Carabus) 34 stictus (Ophonus) 113 stierlini (Asaphidion) 53 stolfai (Bembidion) 65, 66 Stomis 160 Stomis s.str. 160 strenuus (Argutor) 154 strenuus (Pterostichus) 154 strepens (Brachinus) 42 striatopunctatus (Poecilus) 150 striatul sic! (Bembidion) 66 striatum (Bembidion) 66 strumosus (Dyschirius) 46 strupii (Duvalius) 82 sturanyi (Duvalius) 81, 82 sturanyi (Molops) 147 sturanyi (Nebria) 19 sturanyi matejkai (Duvalius) 82 sturanyi munelensis (Molops) 146 sturanyi stenocephalus (Duvalius) 81, 82 sturanyi sturanyi (Duvalius) 82 subaeneus (Apristus) 127 subchalybaeus (Harpalus) 101 subcostatum (Bembidion) 68 subcostatum javurkovae (Bembidion) 68, 69 subcostatum spartanum (Bembidion) 69 subcostatum vau (Bembidion) 68, 69 subcylindricus (Harpalus) 102 subfasciatum (Bembidion) 71 submarginatus (Licinus) 129, 130 subnotatus (Trechus) 85 subnotatus ljubetensis (Trechus) 86 subnotatus subnotatus (Trechus) 85 subquadratus (Acinopus) 97, 98 subquadratus (Ophonus) 110 subrugosus (Carabus) 35 subsinuatus (Harpalus) 100 substriatum (Bembidion) 62 substriatus (Dyschiriodes) 50 substriatus (Dyschirius) 50 substriatus (Notiophilus) 25 substriatus priscus (Dyschirius) 50 sulphuripes (Harpalus) 100 sulphuripes sulphuripes (Harpalus) 100 sutomorensis (Carabus) 30 suturalis (Acupalpus) 117

suturalis (Neotrechus) 84 suturalis (Trechus) 84 suturalis pentheri (Neotrechus) 84 suturalis pretneri (Neotrechus) 83 suturalis suturalis (Acupalpus) 117 sycophanta (Calosoma) 29 sylvatica (Cicindela) 27 sylvatica sylvatica (Cicindela) 27 Syntomus 12, 127 Synuchidius 171 SYNUCHINA 173 Synuchus 173 Synuchus s.str. 173 TACHYINA 73 TACHYINI 10 Tachypus 36 Tachys 75 Tachyta 75 Tachyta s.str. 75 Tachyura 76 Tachyura s.str. 76 Taenidia 26 Talanes 71 Tapinopterus 12, 161 Tapinopterus s.str. 161 tardus (Harpalus) 103 tarsicus (Sinechostictus) 73 Tarulus 123 taurica (Amara) 179 tenebrioides (Zabrus) 185 tenebrioides tenebrioides (Zabrus) 185 tenebrosus (Harpalus) 98 tenebrosus var. centralis (Harpalus) 98 tenellum (Bembidion) 59 tenellum tenellum (Bembidion) 59 terminale (Bembidion) 71 terminatus (Chlaenius) 90 Terminophanes 71 terricola (Laemostenus) 173 terricola (Pristonychus) 173 terricola (Scarites) 51 terricola punctatus (Laemostenus) 173 terricola terricola (Scarites) 51 testaceum (Bembidion) 60 testaceum testaceum (Bembidion) 60 Testedium 71 INDEX

241

tethys (Bembidion) 70 tetracolum (Bembidion) 69 tetracolum tetracolum (Bembidion) 69 teutonus (Acupalpus) 120 teutonus (Stenolophus) 120 Thalassophilus 88 thessalicus tomoricensis (Tapinopterus) 163 thoreyi (Agonum) 136 thoreyi (Platynus) 136 thoreyi thoreyi (Agonum) 136 tibiale (Bembidion) 54 tibialis (Amara) 177 Tomocarabus 36 tomoricensis (Amara) 179, 180 tomoricensis (Molops) 145 tomoricensis (Nebria) 21 Trachycarabus 37 transfuga (Molops) 143 transversalis (Cicindela) 27 transversalis transversalis (Cicindela) 27 TRECHINA 79 TRECHINAE 7, 52, 187 TRECHINI 79 TRECHINITAE 10 TRECHODINA 88 Trechus 85 Trechus s.str. 85 Trepanes 72 Trichochlaenius 92 Trichotichnus 12, 115 Trichotichnus s.str. 115 tricolor (Bembidion) 56 tricuspidata (Amara) 181 tricuspidatus (Carterus) 95 tricuspidatus (Ditomus) 95 trimaculata (Lebia) 126 triseriatus (Harpalus) 105 trisignata (Cylindera) 28 trisignata trisignata (Cylindera) 28 tristis (Chlaenius) 90 tristis (Harpalus) 104 tristis (Trechus) 87 tristis tristis (Chlaenius) 90 truncatellus (Metabletus) 128 truncatellus (Syntomus) 128 turcicus (Platynus) 138 242

uliginosus (Elaphrus) 38 unipustulatus (Badister) 128 ustulatum (Bembidion) 69 valbonensis (Trechus) 85 valonensis (Nebria) 23 vaporariorum (Cymindis) 123 varicolor (Bembidion) 56 varicolor varicolor (Bembidion) 56 variolosus (Carabus) 31 variolosus nodulosus (Carabus) 31 varium (Bembidion) 63 varium dyrrhachii (Bembidion) 63 velutinus (Chlaenius) 91 venustus (Laemostenus) 172 verbasci (Bradycellus) 118 vernalis (Harpalus) 102 vernalis cursor (Pterostichus) 151 versicolor (Poecilus) 151 vestitus (Chlaenius) 90 victoris (Molops) 143, 144 viduus moestus (Platynus) 133 viduus var. moestus (Platynus) 133 vignai (Duvalius) 82 violaceus (Carabus) 32 violaceus azurescens (Carabus) 32 violaceus azurescens m. merditana (Carabus) 33 violaceus azurescens m. merditanus (Carabus) 33 violaceus azurescens m. zabljakensis (Carabus) 32 violaceus azurescens n. zabljakensis (Carabus) 32 violaceus azurescens var. shljebensis (Carabus) 32 violaceus dryas (Carabus) 33 violaceus dryas var. korabensis (Carabus) 33 violaceus dryas var. merditanus (Carabus) 33 violaceus exasperatus (Carabus) 33 violaceus merditanus (Carabus) 33 violaceus shardagensis (Carabus) 33 violaceus var. korabensis (Carabus) 33 violaceus var. shardagensis (Carabus) 33 violaceus vlasuljensis (Carabus) 33 violaceus zabljakensis v. merditanus (Carabus) 33 viridicupreum (Agonum) 135 viridicupreum viridicupreum (Agonum) 135 viridicupreus (Platynus) 135 viridulus (Harpalus) 107 vivalis (Synuchus) 173 vivalis vivalis (Synuchus) 173


vlasuljensis (Carabus) 33 vseteckai (Bembidion) 62 vseteckai dissimile (Bembidion) 62 walteri (Pterostichus) 156 weiratheri (Carabus) 30 weiratheri (Molops) 140 weiratherianus (Carabus) 35 wetterhallii (Masoreus) 92 wetterhallii wetterhallii (Masoreus) 92 winkleri (Carabus) 35 winkleri (Zabrus) 182 winklerianus (Molops) 144

Xenocelia 180 zabljakensis (Carabus) 32 ZABRINA 181 ZABRINI 174 Zabrus 181 Zabrus s.str. 185 Zezea 181 zhalovi (Duvalius) 82 ZUPHIINA 185 ZUPHIINI 185 Zuphium 186 Zuphium s.str. 186

INDEX

243

244


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