A revision of the Dryinidae (Hymenoptera) Part 2

D r y i n i n ae 947 N E O T R O P I C SPEC I ES O F TRI DRYJVU5 T w e l v e spec i es are known . K E Y TO THE S P ...

Author: Massimo Olmi

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D r y i n i n ae

947

N E O T R O P I C SPEC I ES O F TRI DRYJVU5

T w e l v e spec i es are known .

K E Y TO THE S P ECI E S

F E MALES

2

H ead u n u su ally e x cav a t e d , w i t h ey es v e r y b u l gin g ( F ig . 6 4 3 A , ..... ... 2 B) . • . • • . . . . . . • • . • • • . • . . • • H ead c on v e x o r f l a t o r n o t v e r y ex cav ated, w i th ey es n o rma l l y b u lg i n g ( F i g . 6 4 3 C ) . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . .4 H ead mo r e e x cavated ( F i g . 6 4 3 A I , ey es mo r e b u lg i n g ( F i g . 6 4 3 A I............................ 1. ma xiexop h t h a lmi c u s n . sp . Head less ex cav a t ed

3

4 5

6

7

8

( F i g . 643 . . . . BI . . . . . . . . . . . . . . . . . . H ead with P O L sligh t ly l o n g e r a c e I I us . . . . . . . . . . . . . . . . . . • . . .

BI ey es less b u l g i n g ( F i g . 6 4 3 .3 . . . . . . . . . . . . . . . . . . . . . . . . . . . t h an b r ead t h o f each pos t e r i o r . 8 . exophthalmicus n . sp .

H ead w i t h P O L sh o rt e r t han b r ead t h o f post e r i o r o c e l l i . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 . que c h u a nus n . sp . .5 F o r e tarsi w i th s e gme n t 1 p r o d u c e d i n t o a h o o k . . . . .6 Fo r e tarsi with segmen ts 2 an d 3 p r o d u c e d i n t o h o oks . E n l a r ged claw with su b ap i cal t o o t h lo ca t e d v e r y n ea r apex ( F i g . 6 4 5 1; c o c o o n c i r c u la r . . . . . . . . . 2. rufic auda ( R i c h a r d s ) E n la r ged claw w i t h subap i cal to o t h mo r e d i stan t f r o m apex ( F ig . 6 4 7 1; c o c o o n ov 0 i da I . . . . . . . . . . 3 . poecilooterae ( R i c h a rd s ) H ead w i t h OL n ea rly t h r e e t i mes o r mo r e than t h r e e t i mes as l on g as P O L . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 H ead w i t h O L s l i gh tly mo r e o r less l o n g e r than P O L . . . . . . . . . . . 9 Rad i al v e i n weakly c u r v i l i n ea r ( F i g . 6 4 8 A ) ; p r on o t u m almost f u l l y b l a ck . . . . . . . . . • . . . . . . . . . 4 . maximus n . sp . Rad i al v e i n s t r o n gly S - sh ap e d ( F i g . 6 4 8 B); p r on o t u m f u lly r e d d i sh - t est a c e o u s . . . . . 8 H ead ex cavate d , b l aCk , w i th c l y pe u s and man d i b l e s testac e o u s .

..... ..... ............ .5. stri a ticer)s ( K i e f f e r : - H ead almost f l at , r e d d ish . . . . . . 6 . ruficeps (Came r o n ) 9 Segmen t o f f r o n t ta r su s mo r e th an t w i c e as l o n g as segme n t 4

.

. 1 0

S e gmen t

of

f r o n t ta rsus less th an t w i c e as l o n g as segme n t 4 .

948

Mem. Amer.

•

10 Head black,

with

·

.

.

.

•

•

.

.

.

•

.

.

.

.

.

.

•

f ro n t

some t imes o rb i t s

Ent.

•

•

•

•

part

Inst.,

•

t e staceous; •

.

.

.

.

.

.

.

.

.

.

.

.

of

.

11

.

.

.

.

Disc

.

.

.

of

.

.

.

.

.

.

.

.

•

•

.

.

.

.

.

.

p r o n o t um du l l ,

.

.

•

.

•

of

1 1 .

.

•

•

•

•

•

•

•

•

•

.

.

•

.

•

clypeus,

a n dinus

n.

•

•

•

•

•

•

•

•

•

•

•

•

11

mandi b l e s and smal l ....

sp.

scutum with areolae b road......

gib b o s u s

12.

black,

p ronotum sh i ny ,

•

scutum with areolae .

n.

sp.

granulat ed and with

p icescens

around disc..................10. Disc

•

37, 1984

v e r t ex ,

Head f ully r eddish-t estaceou s ; •

•

no.

t e staceous,

not

n.

granulat ed,

n igrofla vus

st riae around disc............ 7.

some

st riae

sp.

n.

with

some

sp.

MALES

No t auli c e s visib l e and comp l e t e ; not

v i si b l e b eh i nd t h e

w i t h a median 2

ke e l

t emp l e s;

Notau l i c e s i nv i sib l e ;

occipital vei n

of

radial

f ro n s w i t h a median wi thout •

•

•

•

•

•

•

LJi stal

a median •

st rong •

•

•

•

•

•

•

•

part

wi thout

·

•

•

•

of

.

•

•

•

.

•

•

•

.

.

•

•

•

•

•

•

.

post e ri o r

•

•

•

•

v ei n

ke e l ;

keel ; •

•

.

•

•

•

.

•

•

.

•

surface

than

of

•

2.

•

.

•

•

2

part

propodeum

proximal

surface

circular .

complet e..........

l o ng as p roximal

.

p oecilopterae ( R i chards)

3.

•

l o ng e r

.

propodeum

(Ki eff e r )

ovoidal ....................

post eri o r

cocoon •

carina

nearly as

cocoon

•

radial

a median median

•

•

ke e l ;

ke e l ;

carina i n comp l e t e , surface o f

s tria ticeps

. . . . . . . . . . . . 5.

Di stal

part

o c c i p i t al post e r i o r

•

.

•

•

.

of •

•

•

part ;

frons

propodeum wi t h a •

•

•

•

•

•

•

•

•

•

•

•

•

•

•

•

•

.

rufic a u da ( R i chards)

The males of other species are u nknow n.

1.

Ft:MALE:

Tridryin u s ma xiexoph th a lmicus

fully winged;

testaceous,

with sides testaceous; abdomen black; hind coxae bulging,

shiny,

pronoturn dull,

8,75

mm;

brown;

mesothorax,

head

(Fig.

643 A)

without sculpture, invisible;

OL

sp.

head testaceous;

pro pectus black;

metathorax

abdomen black-brown;

black;

occipital carina hairy;

le ngth

with segments 2-5

n.

anten nae

pro notum

legs testaceous,

with

unusually excavated,

with

black,

and propodeum black; partly mid and with eyes very

numerous weak longitudinal keels;

more than three times as lo ng as POL;

with sides without sculpture and disc with irregular keels,

anterior transversal im pressio n and anterior collar very weak;

humped,

raised into

a carina;

posterior collar invisible;

disc

posterior transversal irnpressio n and

pro notal tubercles not reaching teguJaej

scutellum and metanotum reticulate rugose,

shiny;

scutum,

notaulices invisible;

949

Dryininae

Fig. 644

-

Chela of Tridryin us exoph t h a lmic u s n. sp.

(type)

propodeum reticulate rugose, with 2 longitudinal keels on poste"i",' "urface; fore-wing with 2 brown transversal bands and with distal part of radial vein longer' '.hem proximal part ( 2 1: 18); segment 1 of front tarsus nearly as long as segment 4; segment" 2 and 3 of front tarsus produced

Mem . A m er .

950

Fig.

645

-

Che l a

of

En!.

I n s!. ,

no.

37, 1984

Tridryinus l'uficauda

( R i chards ) ( t y p e )

951

D r y i n i n ae

i n to h oo k s ;

en l a rged c l a w

segme n t:; of front tars u s

644) w i th sub a p ic a l tooth a n d 18 l a m e l l ae;

( F i g. ( F i g.

644 )

w i th 2 r o w s of a t l e a s t 60 lame l l ae

exte n c! i n g co n ti n u o u s ly to the a p ex; MAL E :

ti b i a l

s p u r s 1 , 1 ,2 .

u n kn ow n

L O C U S TYPI C US :

Moe n ga t a p oe-Vii a v>i a

TYP I CAL IvIAT ERIAL : H OSTS:

h o l oty p e F !

( Su rin a"lf·)

i n LE .

unknown

D I STRltlU T I ON:

,'Iee>\"ur:;i c r e gi o n :

2.

SU RINAME :

Tridryin u s r u fic a u da

Moe n ga ta poe-lV i a w i a ,

( R i c h a rcJs)

/vlesodryin u s r u fic a u da R i c h a r d s 195 3 :

F E MAL E :

f u l l y w i n ge d 4, 25-6, 87 m m ;

m a n d i b les red d i s h - y e ! l ow ; black ;

h e a c b l a ck ,

abd orT,en f u lly b row n or b r ow n w i th

( F i g . 643 C )

du l l ,

h a i l'y ,

s h orter th a n P O L ; hairy,

sh i n y ,

w i th

r e a c h i n g tegu l a e ; s c u l p tu re d a s

retic u late r u gcse ;

a n terio r c o l l a r i n v i s i b le ;

scutum,

head;

i n vi s i b le,

i n c l in e d ;

ve i n

as lo n g as p ro x i ma l

f u lly

w i n ged;

tu b ercles n ot h a i ry ,

segm e n t 1

ba n d s a n d " i th d i stal

as w i th

p a r t of

p rod u c e d

i n to a h o ok ;

en I a rged c l a w

n e a r a p e x a n d w i th 20 lam e l l ae ( F i g .

( Fig.

645 ) w i th 2 row s o f a t l e a s t 80

l a m e l l ae exten d i n g c o n t i n u ou s ly to th e a p e x ; MAL E :

p o s te r i or c o l l a r very

p r on ota l

segme n t 1 of f r o n t ta r s u s s l i ghtl y

part;

located very

segme n t 5 of f r o n t ta r s u s

OL

pron otu n l

p os te r i o r s u rface w i th 2 lo n g i tu d i n a l

fore- w i n g w i th 2 b row n tr a n s ver sa l

tooth

head

p r o p o d e u m retic u l a te r u gose ,

rad i a l

s h orter th a n segm e n t 4 ;

cari na;

s c u te l l u m a n d rnetan otUrTi s h i n y ,

n ot a u l i ces

l egs red d i s h ,

occ i p i ta l c a r i n a comp l e te ;

to u c h i ng occ i p i ta l

kee l s;

6 45 ) ;

apex redd i sh;

n u mero u s str i a e arou n d d i sc ;

poste r i or s u r f a c e very

w ith s u b a p i c a l

w i th c l y p e u s a n d

thorax a n d p ro p od e u rT'

mid a n cJ h i n d c l u b s of feclOra b r ow n ;

p o s terior oce l l i

reduced a n d s h ort;

n . comb .

59 .

a n te n n a e tes taceOU5j

w i th m i d a n d h i n d c ox a e b l ack ,

LE!

l e n gth 4 m m ;

b l ack ;

ti b i a l s p u r s 1 , 1 , 2 .

m a n d i b l e s,

c l y peus a n d partl y

gen ae testaceou s; a n te n n a e testaceO U5j l egs te staceOU5, w i th h i n d cox a e p a r tly b ro w n ; a n te n n a e n o t d i s ta l ly th i c k e n e d j a n te n n a l segm e n ts i n fo l l o w i n g p ro p o r t i o n s :

9:

5:

21 :

1 3:

1 2:

1 2:

f u l l y retic u l a te r u gose ; fro n s w i th ou t med i a n c a r i n a com p l e te ; POL = 8; OL = 3; O OL = 5; s c u te l l u m a n d meta n ot u m d u ll, p ro p od e u m reti c u la te r u go s e ; med i a n

l on g i tu d i n a l

k ee l ;

1 0:

1 0:

and UPL

9:

1 0, 5;

head du l l ,

l a te r a l k ee l s ; occ i p ital = 1 ; TL = 0; s c u torTl,

f u l l y reti c u l a te r u gose ;

n o ta u l i c es i n vi s i o l e ;

p o s te r i o r s u rf a c e of p rop o d e u m w i th

b a nd b e n e a th the p terosti gm a ;

d i s ta l

p a rt of r a d i a l

ve i n

l on ger th a n

p rox i m a l p a rt ( 1 4 : 1 1 ) ; gen i ta l i a : f i g . 646; t i b i a l s p u rs 1, 1 , 2 . L O C U S T YP I C U S : S a n Ped r o d e iv\o n te s d e Oca ( C osta R i c a ) T Y P I CAL MATER I AL : HOS T S:

a stro n g

fore-w i n g h y a l i n e o r w ith a w e a k tr a n sve r s a l

h o l oty p e F !

in Trin id a d Orrnenis s p .

i n IVA. on C a c a o .

952

Mem . Amer .

F i g . 646

En!.

I n s! . ,

no.

37,

1984

M a l e ge nitalia of Tridryinus ruficauda (Richards) from S ! . Au g u s t i n e (T r i n i d a d ) (r i g h t h a l f remove d )

-

U I S T R I B U T I ON :

Neo t r o p ic reg i o n : COSTA R I CA : S a n P e d r o d e Mon t e s d e Oca,

S ! . Augu st i n e , OL! B RAZ I L : iVlanaus ( Am a zo n a s ) , BM! 50° 3D' W 1 2° 3�' 5 (Vi l a Vera), T W ! NOTES : t h e cocoon of Tr. ruficauda is a l m o s t ci rcu l a r a n d com posed of o n e o n l y lining. WA!

T R I N I DAD :

3.

il!esodryinus

Tridryinus poecilopterae ( R i ch a rd s )

p oecilop ter ae

r�

i ch a r d s 1 947:

Mesodryinus poecilopterae Ri ch a rd s : Mesodryinus sp . :

Teran

1 980 :

289 .

n . comb.

868.

Ri ch a r d s 1 9 5 3 :

59 .

D r y i n i n ae

953

F i g . 647 - C h e l a of Tridryinus p oec ii op te ra e ( R i c h a rds ) from 51. A u g u s t i n e ( Tr i n idad )

F E MA L E : f u l l y w i n g ed ; l e n g t h 6 , 0-7 , 5 m m ; head [,iceck, w i th c l yp""" errH ", a n cJ i b l e s red d i sh- tes t aceo u s ; a n t e n n a e b row n , w i t h segme n t s 1 - 2 redd i sh ; t horax a n d propodeum b l ac k ; a b domen b l ac k ; l e g s b ro w n , w i t h t i b ia e , femora a n d p a r t l y t roc h a n ters of f o r e l e g s redd i sh ; h e a d h a i r y , d u l l , ret i c u l a te ru gose, occ i p i t a l c a r i n a comp l e t e ; po s t e r i or oce l l i touch i n g occ i p i t a l ca r i n a ; O L shorter t h a n POL ; pronot u m h a i r y , sh i n y , w i t h n u merous s t r i ae a ro u n d d i sc ; pos t e r i o r co l l a r very reduced a n d s h o r t ; a n t e r i o r co l l a r i n v i s i b l e ; p ron o t a l t u b erc l e s not rea c h i n g te g u l ae ; scu t u m , sc u te l l u m a n d m e t a n o t u m sh i n y , h a i r y a n d r e t i c u l a t e ru gose ; n o t a u l i ces i n v i s i b l e ; p ropode u m re t i c u l a t e ru gose, w i t h pos t e r i or s u rf a c e v ery i n c l i n ed

954

Mem . A me r . E n ! .

I ns ! . ,

no.

37 ,

1 984

� �� . . .. . .

F i g . 648 - R a d i a l v e i n s of fema l e s of Tridryinus maXlmus n . s p . a n d striaticeps ( K i ef f e r ) ( B ) f rom Su r i n a me

( t y pe ) ( A )

a n d w i t h 2 l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h 2 brown t r a n s v e r s a l b a n d s a n d w i t h d i s t a l p a r t o f r a d i a l v e i n a s l o n g a s p rox i ma l p a r t ; segme n t 1 of f ro n t t a r s u s n e a r l y as l o n g as segmen t 4 ; segmen t 1 p ro d u ced i n to a hook ; e n l a r g e d c l a w w i t h a su b a p i c a l too t h a n d 24 l a me l l ae ( F i g . 647 ) ; s e g me n t 5 of f r o n t t a rs u s ( F i g . 647 ) w i t h 2 rows of a t l e a s t 80 l a me l l ae e x t en d i n g con t i n u o u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M A L E ( F i g s 634, 635 ) : f u l l y w i n g e d ; l e n g t h 4 , 0-4 , 7 m m ; h e a d b l ack , w i t h m a n d i b l e s r e d d i s h ; a n te n n a e brow n , w i t h seg men t s 1 -2 a n d 1 0 redd i s h ; some t i mes o n l y seg m e n t s 1 -2 a n d p a r t l y 3 red d i sh ; t h o r a x a n d p ropodeum b l ack ; a b domen b ro w n ; l e g s redd i s h - t e s t aceo u s , w i t h mi d a n d h i n d cox a e b row n ; some t i mes o n l y h i n d cox a e p a r t l y bro w n ; h e a d d u l l , re t i c u l a t e r u gose ; occ i p i t a l c a r i n a comp l e te ; a n t e n n a e n o t d i s t a l l y t h i c k ened ; a n t e n n a I seg men t s i n fo l l o w i n g p ropo r t i o n s : 1 2 : 9 : 2 9 : 1 5 : 1 8 : 1 5 : 1 4 : 1 2 : 1 1 : 1 4 ; f ro n t a l l i n e comp l e t e ; P O L = 1 0 ; Ol = 5 ; O O l = 8 ; O P l = 2 , 5 ; Tl = 2 ; scu t u m , sc u te l l u m , met a n o t u m a n d p ropodeum d u l l , ret i c u l a t e ru gose ; n o t a u l i c e s i n v i s i b l e ; pos t e r i o r s u rface of p ropodeum v e r y i n c l i n ed , w i t h o u t l o n g i t u d i n a l k ee l s ; fore-w i n g w i t ho u t d a rk t r a n sv e rs a l b a n d s o r w i t h a w e a k d a r k b a n d b e n e a t h t h e p t e r os t i g m a ; d i s t a l p a r t of rad i a l v e i n n e ar l y a s l o n g a s p ro x i m a l p a r t ( 1 6 : 1 5) ; t i b i a l s p u r s 1 , 1 , 2 . lOCUS TYP I C U S : S I . A u g u s t i n e ( T r i n i d ad ) T Y P I CAL MAT E R I A L : ho l o t y pe F ! a n d 1 p a ra t y p e Ivi ! i n BM . H O STS : i n T r i n i d a d Poekilloptera (=Poeciloptera) phalaenoides ( l . ) (Fl a tidae) ( R i c h a r d s 1947 ) ; i n E I Sa l v a d o r Poekilloptera completa Jac . ( R i c h a rd s 1 95 3 ) ; i n Venez u e l a Poekilloptera phalaenoides ( l . ) ( Te r a n 1 980 ) . D I ST R I B U T I O N : Neotrop i c reg i o n : T R I N I DA D : S! . Au g u s t i n e , BM ! E l SALVA D O R : S a n S a l v a d o r , O l ! PANAMA : Ba l boa ( C a n a l Zon e ) , AM ! V E N E Z U E L A : C a racas ( D . F.), WA ! O l !

4 . Tridryinus maximus n . s p .

D r y i n i n ae

955

�.",

,

\\� F i g . 649

-

.

Ch e l a of Tridryillus maxil1lus n . sp .

( type)

\\

\J

956

M e m . Amer . E n ! .

I nst . ,

no.

37 ,

1 984

F E M A L E ( F i g s 632, 63 3 ) : f u l l y w i n ged ; l e n g t h 9 , 37 m m ; head b l ac k , w i t h c l y pe u s w h i t i s h except f o r a med i a n reg i o n b l ac k ; e y e s su rrou n d e d b y a n a r row w h i t i s h s t r i p e ; a n tennae b l a c k , w i t h segme n t s 6- 1 0 l i g h t ; m ax i l l a r y p al p i w h i t i s h ; propec t u s b l ack ; p ro no t u rn b l ac k , w i t h s i d es p a r t l y red d i sh ; meso t h o r a x , m e t a t h o r a x a n d propodeum b l ac k ; a b domen b l a c k - b row n : l eg s b row n , w i t h cox a e b l ack ; dorsa I reg i o n o f s t a I k s o f fore femora a n d dorsa I reg i o n of fore t i b i ae w h i t i s h ; h e a d d u l l , g r a n u l a ted ; oce l l a r reg i o n c o n v e x ; vertex exc a v a t e d ; oce l l a r reg i o n a n d su rrou n d i n g reg i o n w i t h n u merous l o n g i t u d i n a l k ee l s ; occ i p i t a l c a r i n a comp l e t e ; t em p l es i n v i s i b l e ; poster i o r oce l l i touch i n g occ i p i t a l c a r i n a ; O L n e a r l y t h ree t i mes a s l o n g a s POL ; p ro n o t u m s h i n y , w i t h n u merous t r a n s v e rs a l k ee l s on d i sc a n d w i t h p u nc t u res a n d some k ee l s or areo l ae on s i de s a n d on a n te r i o r c o l l a r ; a n t e r i o r t r a n s v e rs a l i m p ress ion s t ron g ; a n t e r i o r co l l a r d i s t i n c t ; pos ter i or t r a n sver s a l i m press i o n s t r o n g a n d l o c a ted n e a r pos t e r i o r m a r g i n of p ro n o t u m ; pos t e r i o r co l l a r s h or t , d i s t i n c t ; d i sc h a i r y ; p rono t a l t u berc l es n o t reach i n g t e g u l a e ; scu t u m , s c u te l l u m a n d rne t an o t Unl ret i c u l a t e ru gose ; no t a u l i c e s i n v i s i b l e ; porpod e u m ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s o n pos t e r i o r s u rf ace ; fore-w i n g w i t h 2 b row n t r asv ers a l b a n d s a n d w i t h r a d i a l v e i n n o t S- s h a ped ( F i g . 648 A ) ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p rox i m a l p ae t ( 24 : 1 7 ) ; seg m e n t 1 of f ro n t t a r s u s n ea r l y a s l o n g a s seg men t 4 ; seg men t 3 of f ro n t t a rs u s produ ced i n to a h ook ; en l a r ged c l aw ( F i g . 649 ) w i t h s u b a p i c a l too t h a n d 30 l a me l l a e ; segme n t 5 of fron t t a r s u s ( F i g . 649 ) w i t h 2 row s of a t l e a s t 80 l a me l l ae e x t e n d i n g c on t i n u o u s l y to the a p e x ; t i b i a l s p u rs 1 , 1 , 2 . IiIAL E : u n k now n LOCUS TYP I C U S : Z a n d e r i j ( Su r i n a m e ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n LE . HOSTS : u n k n o w n D I ST R I B UT I O N : Neo t ro p i c reg i o n : S U R I N AM E : Z a n de r i j , L E !

5.

Tridryinus striaticeps

( K i effer )

Prodryinus striaticeps K i effer 1 9 0 9 :

( K i effer ) : Tridryinus striaticeps ( K i effer ) : Tridryinus striaticeps

334 . K i effer 1 9 1 3c : 325 . K i effer 1 9 1 4b : 29 .

F E M A L E : fu l l y w i n g e d ; l e n g t h 5 , 00-7 , 50 m m ; h e a d b l ac k , w i t h c l y p e u s a n d m a n d i b l e s y e l l o w ; a n te n n a e b row n ; a n t e n n a e b row n , w i t h seg men t s 6- 1 0 t e s t aceo u s ; p ropec t u s a n d pro n o t u m redd i s h ; meso t hora x , m e t a t h orax a n d propod e u m b l a c k , abdomen b l ac k -y e l l o w ; l eg s t e s t aceou s , w i t h s t a l k s of h i n d femora a n d h i n d t i b i ae b row n ; he a d ex c a v a te d , sh i n y , w i t h n u mero u s l o n g i t u d i n a l k e e l s ; oc i p i t a l c a r i n a comp l e t e ; poste r i o r oce l l i t o u c h i n g occ i p i t a l c a r i n a ; OL more t h a n t h ree t i mes as l o n g as POL ; p ro n o t u m sh i n y ,

hairy,

w i t h n u merous w e a k s t r i ae a ro u n d d i sc ;

a n t e r i o r t r a n sv ersa l

957

Dryininae

Fig.

650

-

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of

Tridryinus striatice}ls

(Kieffer)

from

Suriname

958

Mem . Amer .

En t .

I nst . ,

no.

37,

1 984

i mpres s i on and a n te r i o r co l l a r very w e a k ; pos t e r i o r t r a n s v e r s a l i mp r e s s i o n s t rong a n d l oc a ted v e r y n e a r pos ter'ior rnal'S:j i n of p ro no t u m j p o s t e r i o r co l l a r i n v i s i b l e ; p ro no t a l t u b erc l e s n o t reach i n g t e g u l a e ; scu t u m , scu te l l u m a n d m e t a n o t u m re t i c u l a t e r u g ose ; n o t a u l i ce s i n v i s i b l e ; p ropodeum ret i c u l a t e r u g os e , w i t h 2 l on g i t u d i n a l k ee l s o n pos t e r i o r s u rface ; fore-w i n g w i t h 2 b ro w n t r a n sv e r s a l b n a d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n ne a r l y a s l on g a s prox i m a l p a r t ; d i s t a l p a r t of ra d i a l v e i n s t ron g l y S-s h a ped ( F i g . 648 B ) ; segme n t 1 o f f r o n t t a r s u s l on g e r t h a n seg m e n t 4 ( 1 8 : 1 5 ) ; segme n t s 2 a n d 3 of f r o n t t a r s u s p ro d u ced i n t o hook s ; en l a rged c l a w ( F i g . 650 ) w i t h s u b a p i c a l tooth a n d 1 3 l a me l l ae ; segme n t 5 of f ro n t t a rsus ( F i g . 650 ) w i t h 2 rows of 40 l ame l l a e ex tencJ i r's; con t i n u o u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : f u l l y w i n g e d ; l e n g t h 4 , 75-4 , 87 m m ; b l ac k ; m a n d i b l e s a n d c l y pe u s t e s t ac eo u s ; a n t e n n a e b row n , w i t h segmen t s 1 - 2 test aceo u s ; l eg s t e s t aceO U 5 , w i t h m i d a n d h i n d cox a e , h i n d femora a n d h i n d t i b i ae p a r t l y b l ack o r b row n ; a n t e n n a e n o t d i s t a l l y t h i c kened ; a n t en n a l segme n t s i n fo l l ow i n g p ropo r t i o n s : 8 : 7 : 23 : 1 3 : 1 4 : 1 4 : 1 2 : 1 0 : 9 : 1 1 ; h e a d d u l l , f l a t , g r a n u l a ted a n d s t r on g l y re t i c u l a t e r u gose ; f r o n s w i t h o u t med i a n a n d l a t e r a l k ee l s ; occ i p i t a l c a r i n a i ncomp l e t e , n o t v i s i b lf> bE-h i n d t h e t em p l es ; POL ; 8 ; OL ; 4 ; OOL ; 4 ; OPL ; 3 ; TL ; 0 ; scu t u m , scu t e l l u m a n d met a n o t u m d u l l , g r a n u l a ted a n d s t r on g l y re t i c u l a te r u g o se ; n o t a u l i ces comp l e t e , p o s te r i or l y sep a ra t ed ; p ropodeum re t i c u l a t e rugose, w i t h o u t t r a n s v e rs a l k ee l s ; p o s te r i o r s u rf a ce w i t h a med i a n s t r o n g l o n g i t u d i n a l k e e l ; fore-w i n g w i t h a d a r k t r a n s v ersa l b a n d b e n e a t h t h e p te ros t i g m a ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ( 1 8 : 1 5 ) ; t i b i a l s p u r s 1 , 1 , 2. L O C U S TYP I C U S : P a r a ( B r az i l ) TYP I CA L M A T E R I AL : ho l o t y p e F ! i n CA . H OS T S : u n k n o w n D I STR I B UT I ON : Neot rop i c reg i o n : SUR I N AME : M a R e t r a i te p l a n t a t i o n ( P a r a m a r i bo ) , L E ! BOL I V I A : At Mou t h of R i o B a u re s ( R i o I te n e z , De p t . B en i ) , AM ! B R AZ I L : P a r a , CA l Teodoro S am p a i o ( S ao P a u l o ) , TW ! OL! J a t a i ( Go i a s ) , T W !

6.

Tridryinus ruficeps ( C a m eron )

n . com b .

Dryin u s ruficeps C a meron 1 88 8 : 445 . Prodryin u s ruficeps ( C ameron ) : K i ef f e r i n K i ef f e r et M a rs h a l l 1 906 : 498 . Prodryin u s ruficeps ( C ameron ) : K i effer 1 9 1 4b : 5 1 . Les todryinus dichrous F e n t o n 1 927 : 1 ( n . sy n . ) ; locus typ ic u s C h a p a d a ( Br a z i I ) •

F E M AL E :

f u l l y w i nged ;

l e n g t h 5 , 0-7 , 1 8 mm ;

head,

a n te n n a e ,

p ropec t u s a n d

Dry i n i na e

651

F i g.

p ro n o t u m r e d d i s h ;

- C h e la of Tridryinus ruficeps

rneso t h orax ,

meta thorax

b ro w n - red d i s h o r redd i s h ;

l egs redd i sh ;

gra n u l a t e d ,

l ongi t u d i n a l

w i t h n u merous

occ i p i t a l c a rin a ve r tex ;

inc o m p l e te ,

OL more t h a n

occ i p i t a l c a r i na ;

9:'9

vis i b l e on l y

t h ree t i rnes as

p r o n o t u rn h a i r y ,

and p ro p o de u rn b lack ;

head sh i n y ,

ab d o m e n

a l m o s t fla t ,

weakl y

k ee l s on fro n s an d vertex ; beh i n d t h e oce l l i

l o n g as POL ;

sh i n y ,

( C amero n )

w i th

a n d on s i des of

p o s t e r i o r oce l l i touch i n g

n u mero u s kee l s a r o u n d d i sc ,

960

lvie m . Amer . E n t .

I n st. ,

no.

37,

1 984

A

B

Fig .

652 - Pronotum of fema l es of Tridry in u s n igrofl a vu s n . sp . from Nova Teuto n i a ( 5r a zi l ) ( A )

a n d picescen s n . sp .

( ty p e ) ( B )

with a posterior tr a n sversa l impression n e a r posterior mar gin ( th e poste r i o r co l l a r so is invis i b le ) ; t h e a n terior tra n sversa l imp ression is w e a k ; the p r o n otum so h a s a sw o l l e n disc a n d a not ve ry visible a n terior c o l l a r ; p ro n o t a l tuberc l es not r e a c h i n g tegu l a e ; scutum d u l l , reticu l a te r u gose; notau l ices i n visib l e; inste a d of t h e notau l ices there a re 2 l on gitudin a l k ee l s reachin g a p p roximate l y 0 , 5 l e n gth of scutum; scute l l um a n d met a n o tum retic u l ate rugose; p ro p odeum ret i c u l ate rugose ; posterior surface of p ro p odeum very inc l in e d , with 2 l o n gitudina l k ee l s ; fore- w i n g with a distinct b r o w n tr a n sversa l b a n d ben eath the pterostigma a n d with 2 t r a c k s of p ro x im a l b ro w n tra nsversa l b ands; dista l p a rt of r a d ial ve i n a s l on g a s p roximal p a rt ; seg me n t 1 of front ta rsus n e a r l y a s l ong a s segme n t 4; se g me n ts 2 - 3 p ro duced i n t o a hOOK; enalrged c l a w with sub a pic a l tooth and 1 3 l a me l l ae ( Fig . 6 5 1 ) ; segment 5 of front t a rsus ( F i g . 651) with 2 rows of at l e ast 50 l ame l l a e e x te n d i n g c o n tin uous l y to the a p e x ; tib i a l spurs 1 , 1 , 2 . MAL E : u n k n o w n LOC U S TYP I C U S : Bug a b a ( Pa n anla ) TYP I CAL MAT E R I AL: ho l o t y p e F! of D. ruficep s in 81'-1; ho l oty p e F! a n d 1 p a r a t y p e F! of L. dichrous i n WA . HOSTS: u n k n o w n O I STRIBU T I ON : Neotropic region: PAI�AMA: Bug a b a , BM! SUR I NAME: K w atta , L E ! P a r amaribo, TW! COLOMBIA: M a r t a ( Rio D o n Die g o , I� Sierra Neva d a d e S . ) , B M ! V E N E Z U E LA: San Esta b a n ( Puerto C a b e110), TW! E C UADOR :

Dry i n i nae

961

F i g . 653 - C h e l a o f Tridry in u s n igrofl a v u s n . s p .

( ty pe )

L i moncoc h a ( N a po ) , O L ! O T ! P E R U : T i n g o M a r i a ( Monzon V a l l ey ) , C A ! Q u i n c em i l ( M a r c a p a t a ) , UL ! TW ! BRAZ I L : N o v a T e u t o n i a ( S . t a C a t a r i n a ) , U L ! B M ! Cha p a d a ( R i o G r a n d e do Su i ) , VIA ! S a n tos ( Sao P a u l o ) , ZO ! S a n t a rem ( P a r a ) , 'ivA ! J a t a i ( Go i a s ) , OT ! Represa do R i o G r a n d e ( G u a n ab a r a ) , TW ! 7 1 ° 38 ' W 4 ° 33' S ( A mazon a s ) , TW ! OL ! ARG ENT I NA : E l do r a d o ( M i s i o n e s ) ,

AM !

7.

F E MA L E :

f u l l y w i n g ed : ;

Tridryin u s nigrofla v u s n . s p .

l e n g t h 5 , 50-8 , 62 m m ;

h e a d b l ac k ,

w i t h f ro n t p a r t

962

of ver tex

Mem . Amer .

( more a l o n g orb i t s ) ,

En! .

I n s! . ,

no.

37,

1 984

c l y p e u s a n d m a n d i b l e s y e l l ow ;

a n te n n a e

t e s t aceo u s , w i t h segmen t s 2-5 a n d 8- 1 0 b row n ; propec t u s b l ack , w i t h a med i a n dors a l reg i o n y e l l ow ; p ro n o t u rn b l ack , w i t h p os t e r i or co l l a r a n d p a r t l y pos t e r i o r l obes y e l l ow ; rnesotho ra x , rne t a t h o r a x a n d p ropodeurn b l ac k ; a b dornen b ro w n -red d i s h ; l eg s t e s t aceou s ; h e a d d u l l , ret i c u l a t e ru gose ; occ i p i t a l c a r i n a i n comp l e te , o n l y v i s i b l e beh i n d t h e oce l l i a n d o n s i des of vertex ; pos t e r i o r oce l l i n o t touch i n g occ i p i t a l c a r i n a ; O P L shorter than P O L ; O L s l i g h t l y l o n g e r t h a n P O L ; p ro n o t u rn ( F i g . 652 A) sh i n y , w i t h n urnerou s l o n g i t u d i n a I a n d t r a n sversa I k e e l s , crossed by a t r a n sversa I a n t e r i o r i rnp ress i o n ; pos t e r i o r t r a n s versa l i rn p ress i o n a n d p o s t e r i o r co l l a r i n v i s i b l ej p ron o t a l t u b e rc l e s n o t reach i n g t eg u l ae ; sc u t u rn sh i n y , re t i cu l a te ru gose ; n ot a u I i ces i nv i s i b I e ; scu te I I urn sh i n y , w i t h n u rnerous l o n g i t u d i n a I k ee l s ; m e t a n o t u rn ret i c u l a t e r u g ose ; p ropodeurn ret i c u l a t e rugo se , w i t h 2 l o n g i t u d i n a l k ee l s on pos t e r i or s u rf a ce ; fore-w i n g w i t h 2 b ro w n t r a n sversal b a n d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n s h o r t t h a n prox i m a l p a r t ( 9 : 1 2 ) ; segmen t 1 of f ro n t t a rs u s nea r l y as l o n g as segmen t 4 ; segmen ts 3 a n d 2 of fron t t a rs u s p ro d uced i n t o hook s ; en l a r ged c l a w ( F i g . 653 ) w i t h s u b a p i c a l tooth a n d 8 l a me l l ae ; segmen t 5 of fro n t t a r s u s ( F i g . 653 ) w i t h 2 row s of at l e a s t 60 l a m e l l ae ex t en d i n g con t i n uo u s l y to t h e a pex ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n L O C U S TYP I C U S : N o v a T e u t o n i a ( B r az i l ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n B M . HOSTS : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i on : BRAZ I L : Nov a T e u t o n i a ( S . t a C a t a r i n a ) , BM !

8.

Tridryi n u s exophtha lmicus n . sp .

F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 93 mm ; h e a d red d i s h - te s t aceo u s , w i t h a b ro w n s p o t on oce l l a r reg i o n : a n te n n a e t e s t aceo u s , w i t h segmen t s 1 -5 d a r k e n e d ; p ro p ec t u s b l ack ; pro n o t u m b l ack , w i t h s i des y e l l ow ; meso t h orax , m e t a t h o r a x a n d propode u m b l ac k ; a b d o m e n b row n-b l ack ; l e g s t e s t aceou s , w i t h h i n d cox a e p a r t l y b l ac k ; h e a d ( F i g . 643 B ) u n u s u a l l y exc a v a t e d , w i t h e y e s very b u l g i n g , w i t h n u merou s l o n g i t u d i n a l keels on frons; occ i p i t a l c a r i n a i n v i s i b l e ; P O L s l i g h t l y l o n g e r t h a n b r e a d t h of each poster i o r oce l l u s ; OL more than t h ree t i mes a s l o n g a s PO L ; p ron o t u rn d u l l , h a i ry , w i t h n u merous kee l s a ro u n d d i sc ; a n t e r i o r t r a n sver s a l i m p ress i o n a n d a n t er i or co l l a r weak ; pos t e r i o r t r a n s v e rs a l i m p r es s i o n a n d pos t e r i or co l l a r i n v i s i b l e ; d i sc h u m p e d , r a i sed i n to a c a r i n a ; scu t u m , scu t el l u m a n d m e t a n o t u m re t i c u l a t e ru gose ; n o t a u l i ce s i n v i s i b l e ; rne t a n o t u m r a i sed i n to a med i a n c a r i n a ; propodeu m ret i c u l a t e r u g ose w i t h 2 l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t o f ra d i a l v e i n l o n g e r t h a n p ro x i m a l p ar t ( 1 8- 1 2 ) ; segmen t 1 of fro n t t a r s u s n ea r l y a s l o n g a s segmen t 4 ;

segmen t s 2 a n d 3 o f fron t t a rs u s p rod uced i n t o

D r y i n i n ae

F i g . 654

-

C he l a o f Tr idryinus exo,ohthalmicus n . sp .

963

( type)

hook s ; e n l a r g e d c l aw ( F i g . 654) w i t h su b a p i c a l too th a n d 11 l a me l l ae ; segmen t 5 of f r o n t t a rsus ( F i g . 654) w i t h 2 rows o f 44 l a m e l l a e ex ten d i n g con t i nuou s l y t o t h e apex ; t i b i a l s p u rs 1,1,2.

964

Mem . Amer .

En!.

I ns ! . ,

no.

37,

1 984

F i g . 655 - C h e l a of Tridryin us quech u a n u s n . sp .

( ty p e )

D ry i n i n ae

MAL E : u n k n ow n LOCUS TYP I C U S : N o v a Teu t on i a ( B r az i l ) TYP I CAL MA T E R I A L : ho l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I B U T I O N : Neo t rop i c reg i o n : BRAZ I L : BM !

9.

965

N o v a Teu t on i a

Tridryin u s quech u a n u s

n.

( S . t a Catarina ) ,

sp .

FEMAL E : f u l l y w i n g ed ; l e n g t h 6 , 56 m m ; h e a d f u l l y t e s t a ceou s ; a n t e n n a e b row n , w i t h se g men t 1 t e s t aceo u s ; t ho r a x a n d p ro p o d e u m b l ac k ; abdomen b l a ck , w i t h some reg i o n s t e s t aceo u s ; l e g s b l ac k , w i t h t a rs i , t roc h a n t e r s a n d f o r e t i b i a e p a r t l y tes t ac eo u s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l segmen t s i n fo l l o w i n g p ro p or t i on s : 1 6 : 8 : 3 7 : 1 5 : 1 3 : 1 1 : 9 : 9 : 7 , 5 : 1 0 ; h e a d d u l l , v e r y exca v a t e d , w i t h eyes very b u l g i n g ; f ro n s a n d v e r tex s c u l p t u red b y weak l o n g i t u d i n a l s t r i a e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; OOL = 1 6 ; PUL shorter t h a n b read t h of each p o s t e r i o r oce l l u s ; p ro n o t u m h u mped , w i t h a s t ro n g pos t e r i o r t r a n s v e rs a l i mp r e s s i o n a n d w i t h a w e a k a n te r i o r t r a n s v e rs a l i m p re s s i o n ; po s t e r i o r co l l a r i n v i s i b l e ; p ron o t a l t u berc l e s n o t rea ch i n g t e g u l a e ; sc u t u m d u l l , f u l l y re t i c u l a t e r u gose ; n o t a u l i ces i n v i s i b l e ; sc u t e l l u m f u l l y re t i c u l a t e r u g os e ; met a n o t u m ra i sed i n to a med i a n c a r i n a , ret i c u l a t e r u g ose n e a r a n terior m a r g i n and smoo t h n e a r p o s t e r i o r m a r g i n ; propodeum ret i c u l a t e r u g o s e , w i t h o u t a t r a n s v ersa l k ee l b e t w een dor s a l a n d p o s te r i o r s u rface ; p o s t e r i o r s u rface w i t h t w o l o n g i t u d i n a l k ee l s ; fore-w i ng w i t h t w o d a r k t r a n sv e r s a l b a n d s , o n e o n t h e b a s a l c e l l s a n d a n o t h er b e n e a t h t h e p t eros t i g m a ; d i s t a l p a r t of ra d i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 7 : 1 4 ) ; fore t a rs a l seg men t s i n fo l l ow i n g propor t i o n s : 24 : 4 : 8 : 1 8 : 32 ; segmen t s 2 a n d 3 o f f ro n t t a r s u s produ ced i n to hook s ; en l a rged c l a w ( F i g . 655 ) w i t h a s u b a p i c a l tooth and 1 5 l a me l l a e ; se g m e n t 5 of f ro n t t a r s u s ( F i g . 655 ) w i t h 2 row s of n e a r l y 47 l a me l l ae ex t e n d i n g con t i n u ou s l y to t h e a p e x ; t i b i a l s p u r s 1 , 1 , 2 . MAL E : u n k n o w n L O C U S TYP I C U S : P u y o ( E c u ador ) TYP I CA L M A T E R I AL : ho l o t y p e F! i n O T . HOST S : u n k n o w n D I S T R I B U T I O N : N eo t rop i c reg i on : ECUADOR : P u y o ( P a s t a z a ) , O T !

10.

Tridryin u s picescens n . s p .

FEMA L E : f u l l y w i n g ed ; l e n g t h 5 , 0 mm ; h e a d b l a ck , w i t h f ro n t p a r t of v e r t e x , c l y p e u s a n d rna n d i b l e s t e s t a ceou s ; a n t e n n a e b row n , w i t h seg m e n t t e s t a c eo u s ; occ i p u t redd i s h ; tem p l es w i t h a sm a l l red d i s h s po t ; t h orax a n d p ropodeurn b l ac k , a b domen b ro w n - re d d i sh ; l e g s t e s t a ceou s , w i t h rn i d a n d

966

Mem . Am er . E n ! .

I n s! . ,

n o . 37,

1 9 84

F i g . 656 - C h e l a of Tridryinus p icescens n . sp .

( type)

h i n d cox a e b l ac k ; h e a d d u l l , h a i ry , ret i c u l a t e ru gose ; occ i p i t a l c a r i n a comp l e t e ; pos t e r i or oce l l i n o t touch i n g occ i p i t a l c a r i n a ; O P L shorter t h a n O L ; OL s h o r t e r t h a n PO L ; prono t u m ( F i g . 652 B ) d u l l , h a i ry , g r a n u l a t e d , w i t h n u m e ro u s s t r i a e a ro u n d d i sc ; a n te r i o r t r a n sver s a l i mpres s i o n a n d a n te r i o r co l l a r weak ; pos t e rior t r a nsver s a l i mp ress i o n s t ro n g a n d l o c a t e d

D r y i n i n ae

F i g . 657 - C h e l a of Tridryinus a n dinus n . sp .

967

( type)

n e a r po s t e r i o r m a r g i n o f prono t u m j pos t e r i o r co l l a r i n v i s i b l e j p ro n a t a l t u b erc l e s not reac h i n g t eg u l ae ; scu t u m , s c u t e l l u m a n d m e t a n o t u m ret i c u l a te r u gose ; not a u l i ce s i n v i s i b l e ; propode u m ret i c u l a t e r u g o s e , w i t h 2 l on g i t u d i n a l k ee l s on pos t e r i o r s u rf a c e ; fore- w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t of r a d i a l ve i n l o n g e r t h a n p rox i mo l p a r t ( 1 5 : 1 0 ) ; s e g m e n t 1 o f fron t t a r s u s l es s t h a n t w i ce a s l o n g a s seg men t 4 ( 1 9 : 1 2 ) ; s e g m e n t 3 of f ro n t t a r s u s p ro d u c ed i n to a hook ; e n l a r g ed c l aw ( F i g . 656 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l a e ; s e g m e n t 5 of f ron t t a r s u s ( F i g . 656 ) w i t h 1 row o f 1 3 lame l l ae ; a pex w i t h a g ro u p o f 1 3 l a m e l l a e ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n ow n LOCUS TYP I C US : Las Cej a s ( A r g e n t i n a ) TYP I CAL MAT E R I A L : h o l o t y p e F! i n T U . H O ST S : u n know n DISTR I B U T I O N : I�eo t rop i c reg i o n : A R G E N T I NA : L a s Cej a s ( T u c u m a n ) , TU !

11.

Tridry in u s andinus n . s p .

F E MAL E : f u l l y w i n g ed ; l e n g t h 5 , 00-5 , 62 m m ; h e a d b l ack, w i t h f ron t p a r t of ver tex , cly p e u s and m a n d i b l e s t e s t aceo u sj som e t i me s a l so orb i t s t e s t aceou sj occ i p u t b l ac k or t e s t aceou s ; an t e n n a e , p ro no t u m a n d p ropec t u s redd i s h t e s t aceou s ; mesot h o r a x , m e t a t h o r a x a n d p ropodeum b l ack ; a b domen a n d l e g s redd i s h - t e s t aceou s ; h e a d d u l l , re t i c u l a t e r u gose ; occ i p i t a l c a r i n a com p l e t e ; pos t e r i o r oce l l i n o t tou c h i n g occ i p i t a l c a r i n a ; O P L s h o r t e r t h a n OL ; O L s h o r t e r t h a n P O L ; p rono t u m d u l l , h a i r y , g r a n u l a ted , w i t h n u merous s t r i ae a rou n d d i sc ; a n te r i or t r a n sv e rs a l i m p ress i o n a n d a n t e r i o r co l l a r w e a k ; pos t e r i o r t r a n s v e rs a l i m p ress i on s t ro n g a n d l oc a ted on pos t e r i o r m a r g i n of p ro n o t u m j pos t e r i o r co l l a r i n v i s i b l e j pra na t a l t u berc l e s not reach i n g

968

Mem . Amer . E n ! .

I ns! . ,

no.

37 ,

1 984

F i g . 658 - C h e l a of Tridryl'n u s gib b o s u s n . s p .

( type)

t e g u l ae ; scu t u m , s c u t e l l u m a n d m e t a n o t u m d u l l , re t i c u l a t e r u g ose ; n o t a u l i ces i n v i s i b l e ; p ropodeum ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s o n pos t e r i o r s u r f ac e ; fore-w i n g w i t h 2 b rown t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t of r ad i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 3 : 1 1 ) ; s e g m e n t 1 of fron t t a rs u s more t h a n t w i ce as l o n g as seg m e n t 4 ( 2 1 : 8 ) ; seg m e n t 3 of fron t t a rs u s p roduced i n t o a hook ; en l a r ged c l aw ( F i g . 657 ) w i t h s u b a p i c a l too t h a n d 5 l a m e l l a e ; segmen t 5 of f ron t t a rs u s ( F i g . 657 ) w i t h 1 row o f 1 3 l ame l l ae ; a p e x w i t h a g r o u p o f 1 2 l a me l l ae ; t i b i a l s p u r s 1 , 1 , 2. MALE : u n k n o w n L O C U S T Y P I C U S : Yacoch u y a ( A rge n t i n a ) TYP I CA L MA T E R I AL : h o l o t y p e F ! i n T U ; 1 p a r a t y p e F ! i n G V ; 1 p a r a t y p e F! in OL . HOST S : u n k n o w n

969

D ryininae

D I ST R I B U T I O N : T U!

Ol!

Neotropic region :

Cordo b a ,

ARGEI�TINA:

Yacoc h u ya ( C afayate ,

Sa l ta ) ,

GV!

1 2 . Tridryin u s gil) b o s u s n . s p .

F E r.lAlE : fu l l y wing e d : l ength 4,00-6 , 2 5 mm; head r e d d is h -testac eo u s ; sometimes oce l l ar region brown; antennae b r o w n , with seg ments 1 -2 testac eo u s; p ro p ectu s b l ac k , with dorsal re g i on testaceou s; p ro p o d e u m b l ack w i th sides l ig ht o r fu l l y r e d dish; occasiona l l y black with margins and a median stripe l i g ht or r e d dish with a b rown spot on sides; mesothorax , metathorax an d p ro p o d e u m b l ack; ab d omen reddish-b rown; l e g s testac eo u s , with m i d and hind coxae part l y b l ac k ; c l u b s o f femora p artly brown; head d u l l , retic u l ate r u g o se; occip i ta l carina comp l ete; posterior oce l l i not to u c hing occipital carina; OPl s h orter than Ol; Ol sh orter than P O L; p ronotulTI d u l l , h a i ry, w i th n u m e ro u s weak striae around disc; anterior transversa l impre ssion and ante rior co l lar weak; p osterior transversa l impres s i on strong and l ocated on posterior m ar g i n of pronotu m; p osterior co l l ar invisib l e; scutu m d u l l , h a i ry , retic u l ate r u gose; notau l ices inv i sib l e; metanotum d u l l , very s h ort and r e d u c e d ; p ro p o d e u m retic u l ate r u g os e , with 2 l ong i t u dinal k ee l s on posterior s u rface; fore-wing with 2 b rown transve rsal bands and with distal p art of radia l vein s h orter th an p roxilTIal part ( 1 2 : 1 5 ) ; s e g m ent 1 of front tar s u s more than twice as l on g as s e g m ent 4 ( 28 : 1 2 ) ; s e g m ent 3 o f front tar s u s p ro d u ce d into a h o o k ; e n l arged c l aw ( Fig . 658 ) with s ubapical tooth and 5 l am el l ae; segment 5 of fore tarsi ( Fig . 6:'8 ) with 2 r o w s of 14 l am e l l ae; apex with a g r o u p of 1 5 l ame l l ae; tibial s p u rs 1 , 1,2. MAL E : unknown lOC U S TYP I C U S : St . A u g u stine ( T rini d ad ) paraty p e TYP I CAL MAT E R I AL : h o l otype F! in BI'-I; 1 para type F! i n Ltc; p aratype F ! in F ! in WA ; 1 p araty p e F ! in C A ; 1 paratype F! i n AM ; Ol. HOSTS: unknown D I ST R I B U T IO N : f,eotro pic re gion : TR I N I D AD : St . A u g u stine ( St . Georg e ) , 81\;i! COSTA R I '=A : E s p arta, WA! SUR I N AM E : Kre p i ( C h ar l esbu r g , Paramaribo ) , lE! PER U : Tingo Maria ( Monson Va l l ey ) , CAl BOL I V I A : At m o u th of Rio Baures ( Rio I tenez, D e pt . Beni ) , AM! Rio I tenez ( Oept. [leni ) , Ol!

4 . j\;egadryin u s Richards

Mega dryin u s Ric h a r d s 1 9 5 3 :

55 .

TYPE S P E C I E S : Megadryin u s magnificus Richards 1 9 5 3 ,

orig .

d esig .

970

Mem . Am e r . E n ! .

I n st . ,

no.

37,

1 984

F i g . 659 - Fema l e of /o,egadryinu5 magnificus R i ch a rds ( t y p e )

FEMALE ( F i g s 659 , 660 ) : f u l l y w i n ged ; m ax i l l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segme n t s ; en l a rged c l a w ( F i g s 659 , 660 , 661 ) more t h a n t h ree t i mes as l o n g a s fro n t t i b i a , m u c h l o n g e r t h a n a ro l i u m , w i t hou t s U b a p i c a l too t h ; no t a u l i c e s i nv i s i b l e; pron o t a l t u b e rc l e s not reac h i n g t e g u lae ; occ i p i t a l c a r i n a corn p l e te; t i b i a l s p u rs 1 , 1 , 2 . MAL E : u nKnow n DIST R I BU T I OI;J : Neot rop i c HOS T S : Flatidae SP E C I ES: 1

D r y i n i n ae

971

F i g . 6 6 0 - F em a l e of Mega dryin u s m a gn ific u s R i c h a r d s ( t y p e )

N E O T R O P I C S P E C I ES OF MEGADRYINUS

O n e spec i es i s k no w n e

IY/ega dryin u s m a gn ificus R i c h a r d s

Megadryin u s m a gn ificus R i c h a r d s 1 95 3 : 55 .

F E MA L E ( F i g s 6 5 9 , 6 60 ) : f u l l y w i n g ed; l en g t h 1 0 , 93 m m ; h e a d b l ac k , w i t h f ro n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow-red d i s h ; a n t e n n a e w i t h s e g me n t s 1 -3 red d i s h - t e s t aceou s , 4- 1 0 b row n ; p ropec t u s b l ac k ; p ro n o t u m b l a c k , w i t h a p e x e s o f t e g u l a e a n d l a tera l m a r g i n s r e d d i s h ; s c u t u m ,

972


I ns t . ,

n o . 37,

1 984

F i g . 6 6 1 - C h e l a of Nega dryln u s magnlhcus R i c h a r d s ( t y p e )

s c u te l l u m , p ropode u m , mesop l e u r a , m e t a p l e u r a , mesos t e r n u m a n d m e t a s t e r n u m b l ack; pet i o l e b l ack; a b d omen b row n - b l ack; fore l eg s w i t h coxae a n d troc h a n ters p a r t l y b l a ck , femora a n d t i b i ae redd i sh , t a r s i b row n i s h , en l a r ged c l a w l i g h t; m i d a n d h i n d l e g s b row n -b l ack; te g u l ae

973

D ry i n i n ae

y e l l o w ; a n t e n n a l seg men t s in fo l l ow i n g p rop o r t i on s : 22: 6: 67 : 20: 15: 9 : 7 : 7 : 6 : 1 0 ; h e a d sh i n y , h a i r y , f u l l y re t i c u l a te ru gose ; occ i p i t a l c a r i n a comp l e t e ; OPL s h o r t e r t h a n b r ea d t h o f each oce l l u s ; p ro n o t u m sh i n y , c o n v ex , w i t ho u t t r a n sversa I i m p ress i o n ; p ron o t u m w i t h n u m ero u s weak l on g i t u d i n a l k ee l s , w i t h s i des w i t h ou t scu l p t u r e ; scu t u m h a i r y , f u l l y r e t i c u l a t e ru gose ; n o t a u l i c es i n v i s i b l e ; sc u t e l l u m a n d met a n o t u m ret i c u l a te r u gose ; p ro p o d e u m f u l l y ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s on p o s t e r i o r s u rf a ce ; poste r i o r s u rf a ce of p ro n o t u m n e a r l y as l o n g as a n t e r i o r s u rf ac e ; fore-w i n g w i t h 2 b rown t r a n sversa l b a n d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n l o n ger than p rox i m a l p a r t ; en l a r ged c l a w ( F i g . 6bl) n e a r l y t h ree t i mes a s l o n g a s fore t i b i a e , w i t h ou t s u b a p i c a l too th , s p a t u l a t e a t a p e x , w i t h 1 row of nea r l y 80 p eg - l i k e h a i r s ; seg men t 5 o f fro n t t a r s u s ( F i g . 6 6 1 ) w i t h 2 row s of near l y 1 70 l a me l l a e ; a pex w i t h a g ro u p of n e a r l y 26 l a m e l l ae ; seg m e n t 1 of f r o n t t a rsu s s h o r t e r t h a n seg men t 4 ( 1 8 : 28 ) ; 1 , 1 , 2.

seg m e n t s 2 a n d 3 of fro n t t a r s u s w i t h o u t hook s ;

t i b i a l s p u rs

MALE : u n k n o w n LOC US TYP I C U S : Ar i m a ( T r i n i d ad ) TYP I C AL ''i,AT EI''AL : h o l o t y p e F ! i n WA ; 1 p a r a t y p e F ! i n B M . HOSTS : i n T r i n i d a d Ormenis s p . (Fla tidae) ( R i c h a rd s 1953 ) D I STR I B U T I ON : N eo t rop i c reg i o n : TR I N I DA D : A r i m a , \VA ! S I . A u g u s t i n e ( R i c h a rd s 1 9 53 ) ;

D i ego M a r t i n ,

tl M !

5 . A lph a dryin us n .

gen .

T Y P E S P E C I E S : A lph a dryin u s a sia ticus n. s p . F E M A L E ( F i g s 662, 663 ) : f u l l y w i n ged ; rnax i l l a ry p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segme n t s ; en l a r g e d c l a w ( F i g . 666 ) a s l o n g a s or s h o r t e r t h a n front t i b i a , much l o n g e r t h a n a ro l i u m , w i t h o u t s u b a p i ca l too t h ; n o t a u l i ces com p l e t e o r i ncomp l e t e ; pron o t a l t u berc l es reac h i n g t e g u l ae ; occas i o n a l l y n o t reach i n g t eg u l a e ; occ i p i t a l c a r i n a com p l e t e or i n com p l e te ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n D I STR I B U T I ON : P a l a e a rc t i c ( on l y fos s i l spec i e s ) , Neot rop i c , A u s t r a l i a n HOSTS : u n k n o w n SPEC I ES : 1 4

E t h i op i a n ,

PALAEARCT I C S P E C I E S CJF A LPHADRYINUS

T w o foss i l

spec i e s a r e k n ow n .

Urien t a l ,

974


Fig.

662

I ns! . ,

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37,

1 984

F e m a l e of A lph a dryin u s lamellatus n . sp .

-

( ty p e )

�EY T O THE SP E C I E S

F EMAL E S

P r on o t u n1 h u mp ed , w i t h a d i s t i n c t d i s c a n d t w o t r a n s v e r s a l i mp r e s s i o n s ( F i g . 6 6 4 B ) ; an t e r i o r a n d p o s t e r i o r c o l l a r d i s t i n c t .

.

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975

i mp r e s s i on b e t w e e n a n t e r i o r and p o s t e r i o r c o l l a r . . . • . . . . . • • . . . . . . . • . . . • • . . 2 . b altic u s n . sp . •

(F i g .

665

A)

•

•

MALES

U n k nown

Fig.

663

-

F e rn a l e of A lphadryin u s l a m el l a t u s

n .

sp .

( t y pe )

1 . A lp h a dryin u s bruesi n . s p .

F E M A L E ( F i g . 664 ) : f u l l y w i n g e d ; l e n g t h 6 , 7 rn rn ; h e a d b row n - l i g h t , w i t h t e e t h of rn a n d i b l es b row n - d a rk a n d ternp l e s b l ac k ; a n t e n n a e b l a c k , w i t h segrnen t 1 t e s t aceou s ; p ro n o t u rn black , w i t h t r a n sv e r s a l i rn press i o n a p p a ren t l y t e s t aceou s ; sc u t u m b l ac k ; s c u te l l u m a p p a r e n t l y t e s t aceou s ; rn e t a n o t u rn i n v i s i b l e ; p ropodeurn b l ac k , w i t h po s t e r i o r s u rface a p p a ren t l y

•

976

M e m . Amer .

En!.

I ns! . ,

no.

37,

F i g . 664 - F e m a l e o f A lp h a dryin u s bruesi n . s p . B : p ron o t u m . '

1 984

( t y p e )( A ) ( p a r t l y v i s i b l e ) ;

t e s t aceo u s ; ven t r a l s i de of t h o r a x b l a c k ; a b domen b l ac k ; l e g s t e s t ac e o u s , w i t h cox a e , c l u b s of femora a n d p a r t l y t i b i ae b l a c k ; a n t e n n ae 1 0seg men t e d , shor t , weak l y s l e n d e r , d i s t a l l y w e a k l y t h i ck e n e d ; d e n s e l y h a i ry ; a n t e n n a l h a i r s v e r y s h or t , m u c h s h o r t e r t h a n brea d t h o f segmen t s ; a n t en n a l seg men t s i n fo l l o w i n g propor t i o n s : 1 1 : 6 : 32 : 1 9 : 1 5 : 1 5 : 7 : 7 : 8 : 1 0 ; a n t e n n a e l es s t h a n t h ree t i me s a s l o n g a s h e a d ( l en g t h o f h e a d dorsa l l y m e a s u red from occ i p i t a l c ar i n a beh i n d t h e oce l l i to a pex of m a n d i b l e s ) : 1 22 : 50 ; c l y p e u s and a n te n n a I soc k e t s n o t v i s i b l e ; m a n d i b l es w i t h 4 teeth p r o g ress i n g l a r g e r f ro m a n t e r i o r o n e to poster i o r ; eyes a p p a re n t l y b a re ; m a l ar s p ac e w i t h a su bocu l a r su l c u s ; occ i p i t a l c a r i n a comp l e t e ; tem p l e s p rom i n en t ; eyes norma l l y b u lg i n g ; P O L � 3 , 5 ; OL � 2 ;

Dryininae

DOL

=

8;

OPL

=

3,5;

TL

=

977

6; frontal line complete; vertex and frons dull,

granulated, rugose mostly on vertex and behind the ocelli; pronoturn

(Fig.

664 B) crossed by an anterior weak transversal impression between anterior collar and disc;

disc weakly humped;

pronotum also crossed '='y a

weak posterior transversal impression between disc and posterior collar; posterior collar very short, rnuch shorter than anterior collar ( 1 ,5:5); pronotum dull, granulated, slightly shorter than head (19:26); pronotal tubercles apparently reaching or almost reaching tegulae; sides of pronotum contracted where they meet the scutum;

posterior transversal

impression and posterior collar without a distinct dorsal rnedian carina; scutum shorter than pronotum (13: 1 9), du I I,

weak I y granu I ated;

notau Iices

complete, broad, posteriorly separated, composed of numerous strong punctures; scutellum dull, weakly granulated, shorter than scutum ( 1 0: 1 3), metanotum invisible; shape of scuturr" Dryininae;

scutellum and pronotum usual for

propodeum not easily distinct;

propodeum

(dorsally viewed)

invisible because hidden under the wings; laterally viewed visible, showing numerous areolae on sides and on posterior surface; posterior surface of propodeurn apparently with two incomplete longitudinal keels; posterior surface of propodeum very step; fore-wing apparently hyaline, without dark transversal bands, with the usual venation of Dryininae; pterostigma narrow, much longer than broad open;

(38:5); radial cell apparently

distal part of radial vein much longer than proximal part (22:1 0);

radial ve',n not s-shaped;

fore wings with the usual three oasal cells

clearly enclosed (costal, median and submedian cells); hind wings apparently hyaline, without dark bands, not easily distinct;

fore-wing

very slender, with shape usual for Dryininae; front leg segments in following proportions: 1 2 (coxae): 1 4 (trochanters): 22 (femora): 2 1 (tibiae): 8 (tarsal segment 1 ): 3 (tarsal segment 2): 5 (tarsal segment 3): 6 (tarsal segment 4): 23

(tarsal segment 5):

16 (enlarged claw); fore

trochanters less than four times as long as broad: 1 4: 4;

fore trochanters

with a long proximal slender stalk, broadened after the half-way; segments 2 and 3 of front tarsus produced into hooks; rudimentary cia" not distinct;

enlarged claw much longer than arolium (22:1 0); enlarsed

claw apparently without subapical tooth with some long lanlellae;

(but apex weakly distinct) and

apex of enlarged cia" not pointed, but spatulate;

segment 5 of front tarsus with some median lame! lae and

\II,. ith

nUfT'erOU5

apical lamellae; number of lamellae on enlarged claw and on tarsal segment 5 not distinct;

mid leg segments in following proportions: 8

(coxae): 5 (trochanters): 6

15 (femora):

1 4 (tibiae):

13 (tarsal segments 1 ):

(tarsal segment 2): 4,5 (tarsal segment 3): 3 (tarsal segment 4): 4,5

,tarsal segment 5);

hind leg segments in following proportions: 1 7 (coxae):

5 (trochanters): 33

(femora):

segment 2): 5):;

7

32 (tibiae):

1 8 (tarsal segment 1 ): 8 (tarsal

(tarsal segment 3): 3 (tarsal segment 4):

tibial spurs 1 , 1 ,2;

5 (tarsal segment

abdomen without a distinct petiole;

max'lllary

Jalpi apparently with 6 segments; labial palpi apparently with 3 segments; last labial segment very long and slender; shape, length and Jreadth of wings usual for Dryininae; shape and

978

Mem . Amer.

Ent .

I n st . ,

no.

37,

1 984

morp h o l o g y of body usua l for Dryininae. M/,LE: u n k n oV\n L OC U S T YP I CUS : German a m b e r T YP I CAL �iAT E f, I AL :

h o l o t y p e F!

in CO .

H OSTS : u n k n ow n o I STR I BUT I O N : on I y k n o w n from G e rm a n a rll b e r . N O T ES : t h i s spec i e s seems show morph olog i c c h a r a c t e r i st i cs of l i v i n g ,11phaclryinus. N o spec i es o f Alphaclryinus i s l i v i n g a c t ua l l y i n Pa l a e a rc t i c r eg i o n . T h e spec i es i s n amed i n h o n o r of the l a t e spec i al i st Mr . C . T . B rues .

2 . Alphadryinus balticus n . sp .

FEI\:ALE ( F i g . 665 ) : fu l l y w i n g ed ; l e n g t h 1 , 80 mOl ; a p p a re n tly b l ac k , w i t h a n t e n n a e a n d l e gs t e s t a ceous ; a bdomen a n d a p p a re n tly ve n tr a l side o f t h o r a x b r o w n - t est a ceousj a n t e n n a e lO-se g me n t ed, s h o r t , d i st a l l y th i c k e n ed ; dense l y h a i r y ; a n t e n n a l h a i rs short, sh o r t e r t h a n b re a d t h o f segme n t s; a n t e n n a l se g m e n ts i n f ol l ow i n g p ro p o r t i o ns: 7 : 5 : 5 : 4 : 3 , 5: 3 , 5 : 3 : 3 : 3 : 6 ; a n t en n ae l e ss t h a n tw i ce a s l o n g a s h e a d ( l e n g t h o f h e a d dorsa l l y measured f rom occ i p i t a l c a r i n a be h i n d t h e oce l l i to a p e x o f mand i b l es ) : 4 2 : 25 ; clyp eus a p p a re n tly w i t h ven t r a l m a r g i n b i den t a t e ; m a n d i b l es quadr i de n t a t e , w i th tee t h p ro g re ss i n g l a r g e r f rom a n te r i o r o n e to post e r i o r ; e y es a p p a re n t l y b a r e ; ma l a r space n o t v i s i b l e ; occ i p i t a l c a r i n a comp l ete ; temp l es p romi n e n t; e y es norma l l y b u l g i n g ; POL � 5; OL � 4 ; OOL � 7 ; O PL � 5 ; : TL � 6 ; f r o n s a p p a r e n t l y w i th out med i a n k ee l ; ve r t e x a n d f ro n s du l l , g r a n u l a t ed, a p p a re n tly n o t rugose ; p ro p odeum very sh o r t , w i t h o u t d i sc, n o t humped, b u t w i th a t r a nsve rsa l i m p ress i o n b e t w ee n a n te r i o r a n d post e r i o r co l l a r ; p r o n o t a l tuberc l es a p p a re n t l y r e a c h i n g t e g u l a e; p ro n o tum much s h o r t e r t h a n h e a d ( 5 : 2 5 ) ; scutum l o n g e r t h a n p ro n o tum ( 1 3 : 5 ) ; poste r i o r co l l a r o f p r o n o tum a p p a r e n t l y ver y short; scutum d u l l , g ra n u l a ted, a p p a re n t l y n o t rugose ; n o t au l i ces comp l e t e , poster i o r l y sep a r a t ed ; scut e l l um w e a k l y v i s i b l e , a p p a r e n t l y a s sc u l p t ured a s scutum ; met a n o t um not v i s i b l e ; p ropodeum o n l y l a te ra l l y v i s i b l e , a p p a r en t l y re t i cu l a t e rug ose , w i t h dorsa l surf ace n e a r l y a s l o n g as post e r i o r sur f a c e ( 1 0 : 1 2 ) ; post e r i or surf a c e n o t v i s i b l e f rom t n e dorsa l s i de ; sh a p e of scutum, scut e l lum a n d pro podeurn usua l f o r iJryininae; s h a pe of p r o n o turn u n u sua l ; fore-w i n g h y a l i n e , w i t h out dark t r a nsversa l b a nds, w i t h t h e usua l ven a t i o n of Dryininae; p t e rost i gma n a r row, much l o n g e r t h a n b road ( 1 0 : 4 ) ; r a d i a l c e l l not f u l l y v i s i b l e ; d i st a l p a r t of r ad i a l ve i n much l o n g e r t h a n p roxim a l p a r t ( but r ad i a l p a r t o n l y p a r t l y v i s i b l e) ; r a d i a l ve i n n o t s-sh a p e d ; f o re w i n g s w i t h t h e usua l t h re e b a s a l c e l l s c l e a r l y e n c l osed ( cost a l , med i a n a n d submed i a n c e l l s ) ; h i n d w i n g s a p p a rently h y a l i n e, w i t h o u t d a r k b a n ds ; f o re-w i n g s l ende r , w i t h sh a pe usua l f o r Dryininae ; f r o n t l e g segme n t s i n fo l l ow i n g p ro p o r t i o n s: 1 1 ( co x a e ) : 5 ( t r oc h a n t e rs ) : 2 1 ( femora ) : 1 6 ( t i b i ae ) : 1 0 ( t a rsa l se gment 1 ) :

D r y i n i n ae

979

A

:" i g .

665

-

Fe m a l e of Alph a dryin u s o a l ticus n . s p . ( t y p e )( B ) ; A : p ron o t u m .

2

head and

( t a r s a l s e g m e n t 2 ) : 2 ( t a rs a l s e g m e n t 3 ) : 1 3 ( t a r s a l s e g m e n t 4 ) : 1 5 : a r s a l seg m e n t 5 ) : 1 4 ( e n l a r g e d c l a w ) ; fore t roc h a n t ers more t h a n t w i c e 2S lo n g as b ro a d ( 5 : 2 ) , w i t h a short prox i m a l s t a l k ; e n l a r ged c l a w m u c h ""lger t h a n a ro l i u m ( 1 4 : 4 ) ; en l a r g e d c l a w s p a t u l a te , w i t h apex n o t =ci n ted ; a pex o f e n l a r g e d c l a w w i t h a row of t e e t h ( 5 t e e t h v i s i b l e ) a n d "itn n u m e r o u s b r i s t l es ; s e g m e n t 5 of f ro n t t a r s u s weak l y v i s i b l e , s:o=aren t l y w i t h some med i a l l a me l l ae ( 3 l a m e l l ae v i s i b le ) ; apex w i t h a ,;-cJp of l a m e l l ae ; r u d i me n t a r y c l a w d i s t i n c t ; s e g m e n t s 2 a n d 3 of fron t :,,- s u s p roduced i n to hook s ; m i d l e g s e g m e n t s i n fo l l ow i n g p rop o r t i on s : 6 coxae ) : 2 ( troc h a n t e r s ) : 1 3 ( femora ) : 1 7 ( t i b i ae ) : 7 ( t a r s a l s e g m e n t 1 ) : 3 :srsa l se g m e n t 2 ) : 3 ( t a r sa l s e g m e n t 3 ) : 2 , 5 ( t a r s a l s e g m e n t 4 ) : 5 ( t a r s a l "",,"len t 5 ) ; h i n d l e g seg me n t s i n fo l l ow i n g p roport i o n s : 9 ( cox a e ) : 4 :-xh a n t ers ) : 1 8 ( t i b i ae ) : 7 ( t a r s a l seg men t 1 ) : 4 ( t a r s a l s e g m e n t 2 ) : 4 :5-sa l se g m e n t 3 ) : 4 ( t a r s a l s e g m e n t 4 ) : 6 ( t a r s a l seg m e n t 5 ) ; t i b i a l "=_rs 1 , 1 , 2 ;

a bdomen w i t h o u t a d i s t i n c t pe t i o l e ;

m ax i l l a ry p a l p i

980


I ns t . ,

no.

37,

1 984

a p p a re n t l y w i t h 6 s e g me n t s ; l a b i a l p a l p i a p p aren t l y w i t h 3 s e g me n t s ; l a s t l a b i a l seg men t not v i s i b l e ; s h a p e , l e n g t h a n d b re a d t h o f t h e w i n g s u s u a l for Dryin in a e ; s h a pe a n d mor p h o l o g y o f bod y u su a l for Dryin in a e . MALE : u n k nown LOCUS TYP I C U S : Germ a n a m b e r TYP I CAL MAT E R I AL : h o l o t y p e F ! i n CO . HOSTS : u n k n o w n D I ST R I B U T I O N : on l y k n o w n f r o m Germ a n a m b e r . N O T E S : t h i s spec i e s i s d i ffere n t from o t h e r fos s i I a n d I i v i n g spec i es of A lph a dryin u s for t h e u n u su a l s h a pe of p ro n o t u m ( ex c a v a ted , n o t h u m pe d ) . T h· i s exce p t i o n , however, i s k n o w n i n o t h e r g e n e r a ( see Dryinus irregularis n . sp . ) a n d i s not con s i dered a gen e r i c c h a ra c t e r .

E TH I OP I AN S P E C I ES OF ALPHADRYLVUS

O n e spec i es i s k now n .

A lph a dryin u s a b erra n s ( Be n o i t ) n . com b .

Dryin u s a b erra n s Be n o i t 1 95 3 b : 4 1 2 .

F E MAL E : f u l l y w i n g e d ; l e n g t h 5 , 0 m m ; he a d b l ac k , w i t h c l y p e u s a n d m a n d i b l e s t e s t a ceou s ; a n te n n a e tes t aceou s ; p ropec t u s b l ac k ; prono t u m b l ac k , w i t h m a r g i n s a n d pos t e r i o r co l l a r redd i s h ; meso t h orax , m e t a t h o r a x a n d p ropo d e u m b l ac k ; a b domen b row n ; l e g s t e s t a c eo u s , w i t h cox a e a n d c l u b s o f femora b row n ; a n t e n n ae d i s t a l l y w e a k l y t h i cken e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p ropor t i on s : 9 : 4 : 2 5 : 1 3 : 9 : 5 , 5 : 5 : 5 : 4 , 5 : 6 ; h e a d d u l l , g r a n u l a t e d ; occ i p i t a l c a r i n a comp l e te ; pos t e r i o r oce l l i t o u c h i n g occ i p i t a l c a r i n a ; OL � POL ; p ro n o t u m d u l l , g r a n u l a t e d , w i t h a n t e r i o r i m p re s s i o n a n d a n t e r i o r co l l a r weak ; p o s te r i o r t r a n s v e rs a l i m p res s i o n s t ron g ; pos t e r i o r co l l a r s h or t , d i s t i n c t ; pronot a l t u b erc l e s n o t reach i n g te g u l a e ; scu t u m s h i n y , g r a n u l a t e d ; n o t a u l i ces i n c om p l e t e , reac h i n g a p p ro x i m a te l y 0 , 65 l e n g t h of sc u t u m ; s c u t e l l u m a n d m e t a n o t u m d u l l , g r a n u l a t e d ; propod e u m re t i c u l a te r u g o s e , w i t h o u t l o n g i t u d i n a l k ee l s on pos ter i o r su rface ; fore-w i n g w i t h 3 b rown t r a n s v e r s a l b a n d s and w i t h d i s t a l p a r t o f r a d i a l ve i n n e a r l y tw i c e a s l o n g a s p rox i m al p a r t ; en l arged c l a w ( F i g . 666 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 5 l ame l l ae ; segmen t 5 of f ro n t tarsus ( F i g . 666 ) w i t h 2 rows of 1 7 l a m e l l ae ; apex w i t h a g roup of 9 l a me l l ae ; s e g m e n t 1 of f ro n t t a r s u s l o n g er t h a n s e g m e n t 4 ( 1 7 : 1 0 ) ; 3 of f ro n t t a rs u s p r o d u ced i n to a hook ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k nown LOCUS TYP I C SU : Bek i l y ( M a d a g a s c a r )

seg men t

�81

Dry i n i nae

//��-----

I

y/

i��

//

/1

,

V

',,\

'"

�

/\

F i g . 666

-

Chela of Al,)hadryinus a:oerrans

(Benoit) (type)

M em . Amer .

982

T Y P I CAL MAT E R I AL : H O STS :

En!.

holotype F !

I ns ! . ,

no.

37,

1 984

in P .

u n know n

D I STR I B U T I ON :

E t h i o p i a n reg i o n :

MADAGASCAR ;

Bek i l y ,

P!

OR I E N TAL S P E C I E S O F !l � PIIADJ( YINU.C:

T w o spec i e s are know n .

KEY T O THE SPEC I ES

F EMALES

Oc c i p ap i ca Occ i p a p i ca

i t a l c a r i n a c omp l e l p r om i n en c e ( F i g . i t a l c a r i n a i n c omp l p r om i n en c e ( F i g .

t e ; s eg m e n t 5 o f f r on t t a r s u s w i t h ou t 6 6 7 ) . . . . . . 1 . a si a ticus n . s p . l e t e ; s e g me n t 5 o f f r on t t a r su s w i t h 6 6 8 ) . . . . . . 2 . gib b o s u s n . s p .

MALES

U n know n

1 . A lp h a dryin u s a s i a ticus

n.

sp .

F E MA L E : f u l l y w i n g ed : l e n g t h 3, 75-5 , 62 m m ; h e a d b l ack, w i t h c l y p e u s a n d m a n d i b l e s y e l l o w ; some t i mes a l so ma l a r s p a c e redd i s h ; a n t e n n a e bro w n , w i t h se g men t s 1 -2 y e l l ow ; p ropec t u s b l a c k ; p ro n o t u rn b l ack, w i t h a n t e r i or m a rg i n , s i des a n d pos t e r i o r col l a r redd i sh ; rnesot h o ra x , rne t a thorax , p ropod e u m a n d p e t i o l e b l ac k ; a b domen b rown ; l e g s tes t aceou s , w i t h cox a e a n d c l u b s o f f e m o r a b row n ; somet i mes a l so t i b i ae b row n ; h e a d d u l l , g ra n u l a t ed , w i t h 1 -4 l o n g i t u d i n a l kee l s on t h e s i des of t h e oce l l i , from occ i p i t a l c a r i n a to c l y p eu s ; some t i mes a l so i r reg u l a r short kee l s are v i s i b l e on fro n s ; occ i p i t a l c a r i n a comp l e t e ; a n t e n n a e shorter t h a n bod y , w i t h seg men t 3 l o n g er t h a n seg n len t 4 ; p ron o t u m h u m p e d , w i t h a s t ron g t ra n s v e r s a l p o s t e r i o r i mp re s s i o n a n d a weak a n t e r i o r t r a n sv e rs a l i rn p ress i on ; pos t e r i or co l l a r v i s i b l e ; a n ter i or co l l a r weak l y v i s i b l e ; p ronot u m sh i n y , w e ak l y g r a n u l a t e d , w i t h n u mero u s l o n g i t u d i n a l kee l s arou n cJ d i sc , sc u t u m d u l l , g r a n u l a te d , w i t h l a t e r a l reg i o n s ret i c u l a t e r u gose ; not a u l i ces i n comp l e t e , e x t en d i n g t o b e y o n d 0 , 5 l e n g t h of sc u t u m ;

Dry i n i n ae

F i g . 667 - C h e l a of A lph adryinus a si a ticus n . sp .

983

( ty p e )

s c u t e l l u m a n d m e t a n o t u m g r a n u l a t e d ; p ropod e u m re t i c u l a t e ru gose w i t h 2 l o n g i t u d i n a l kee l s on pos t e r i or s u rf ace ; fore-w i n g w i t h 3 b ro w n t r a n s v ers a l o a n d s ( t h e t w o b a s a l b a n d s j o i n ed ) ; d i s t a l p art o f rad i a l v e i n m u c h l o n ge r t h a n p ro x i m al p a r t ; e n l a r g e d c l a w ( F i g . 667 ) not s p a t u l a t e , w i t h o u t s u b a p i c a l too t h a n d l a m e l l ae , w i t h 1 prox i m a l b i g p e g - l i ke h a i r ; se g m e n t

984

Mem . A m er . E n t .

I n st . ,

no.

37,

1 984

F i g . 668 - C h e l a of A lp h a dryin u s gib b o s u s n . s p .

( type)

5 o f f ro n t t a rs u s ( F i g . 667 ) w i t h 3-4 row s o f a t l e a s t 4 5 l ame l l ae e x t en d i n g con t i n uo u s l y to t h e a pex ; en l a rged c l a w m u c h l o n g e r t h a n a ro l i u m ; seg m e n t 1 of fron t t a r s u s s l i g h t l y l o n ger t h a n segmen t 4 ; t i b i a l spurs 1 , 1 , 2 .

Dry j n j nae

t/IAL E :

985

unknown

L O C U S TYP I C U S :

Kal abakan

T Y P I C A L �'IA T E R I A L : H OS T S :

( Sabah ,

ho l o t y p e F !

Ma l a y s i a )

in 8;

1

p a r a t y p e F" !

D I ST R I BU T I ON : V I E T I�AM :

Oriental

8 i an

(Mt.

reg i o n :

La n g ) ,

f',\A LAYS I A :

Kal abakan

F e MA L E :

f u l l y w i n g ed ;

l e n g t h 5 , 0 m rn ;

( m ore a l o n g o rb i t s ) ,

red d i s h ;

p ro p ec t u s b l ac k ;

( Tawau,

Saoah ) ,

8!

OL !

2 . A l�h a dryin u s gi 0 bo s u s n .

v er tex

i n UL .

unknown

sp .

h e a d b l a ck ,

w i t h f r o n t p a r t of

c l y pe u s and ma n d i b l es redd i sh ;

p ro n o t u m b l a c k ,

w i th anterior and

a n t e n n ae l a tera l

m a r g i n s re d d i s h ; pos t e r i o r co l l a r redd i s h ; mesoth o r a x , met a t h o r a x a n d p ro p o d e u m b l a c k , a b domen b r o w n - r e d d i s h ; l e g s t e s t a ceo u s , w i t h c l u b s o f femora a n d h i n d c o x a e b r ow n ; h e a d s w o l l e n , sh i n y , w e a k l y p u n c t a t e ; oc i p i t a l c a r i n a i n comp l e t e ; p o s t e r i or oce l I i n o t t o u c h i n g occ i p i t a l ca r i n a ; U ? L ::: P O L ;

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66SJ - C h e l a of A lph a dryin u s lamellatus n . sp .

( type)

a ro u n d co l l a rs ; p ronot a l t u b erc l e s n o t reach i n g t e g u l a e ; sc u t u m d u l l , g r a n u l a t ed a n d w e a k l y re t i c u l a t e ru gose ; n o t a u l i ces com p l e t e , s t ron g l y con verg i n g , pos t e r i o r l y sep a r a ted ; s c u te l l u m a n d m e t a n o t u m sh i n y , w ea k l y p u nc t a t e ; propode u m ret i c u l a te ru gose, w i t h 2 l on g i t u d i n a l k ee l s on pos t e r i o r s u rf a c e ; fore-w i n g w i t h 2 b ro w n t r a n s vers a l b a n d s and w i t h d i s t a l p a r t of rad i a l v e i n much l o n g e r t h a n p rox i m a l p a r t ( 28 : 1 4 ) ; seg m e n t 1 o f f ro n t t a r s u s shorter t h a n seg men t 4 ( 1 7 : 20 ) ; segme n t s 2 a n d 3

988


Fig.

I ns ! . ,

no.

37,

1 984

670 - C h e l a of A lp h a dryin u s p a n amensis

n .

sp.

( ty p e )

p ro d u c e d i n to hook s ; en l arged c l a w ( F i g . 669 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 1 u n u s u a l v e r y l o n g l a me l l a + 4 s h o r t e r l a me l l ae ; segmen t 5 o f f ron t t a r s u s ( F i g . 669 ) w i t h 1 row of 1 1 l a me l l a e ; a pex w i t h a g roup of 1 5 l ame l l ae ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n

Dry i n i nae

989

LOCUS TYP I C U S : Nova Teu ton i a ( B raz i l ) TYP I CA L MA T E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n OL. HOSTS: u n k nown D I ST R I B U T I O N : Neot rop i c reg i o n : BM ! OL !

BRAZ I L :

i n B fYI ;

1

paratype F !

Nova Teuton i a ( S . t a C a t a r i na ) ,

2 . A lp h a dryi n u s p a n amensis n . s p .

F E M A L E : f u l l y w i n g e d ; l e n g t h 6 , 93 m m ; h e a d b l ac k , w i t h fron t p a r t of ver tex , c l y p e u s , m a n d i b l e s a n d orb i t s y e l l o w ; occ i p u t b row n -y e l l ow ; a n t e n n a e w i t h se g m e n t s 2-5 b row n , seg m e n t 1 p a r t l y y e l l o w , seg men t 6 w i t h d i s t a l p a r t y e l l o w ; seg men t s 7- 1 0 m i s s i n g i n h o l o t y p e ( o n l y k n o w n spec i men ) ; t h o r a x , p ropodeum a n d a b domen b l ac k ; l e g s t e s t a c eo u s j h e a d d u l l , f l a t , w i t h oce l l a r reg ion sw o l l e n , g r a n u l a te d a n d ret i c u l a t e r u g o se, h a i ry ; occ i p i t a l c a r i n a i n comp l e t e , v i s i b l e on l y beh i n d the oce l l i and on t h e s i des of v e r tex ; pos t e r i o r oce l l i not t o u c h i n g occ i p i t a l c a r i n a , O P L shorter t h a n P O L ; O L nea r l y t h ree t i mes a s l o n g as P U L ; p rono t u m sh i n y , h a i r y , w i t h n u m erou s l o n g i t u d i n a l k ee l s on co l l a r a n d w i t h n u ro erous k ee l s a ro u n d d i sc ; a n t e r i o r t r a n s versa l i m p ress ion s t r o n g ; a n ter ior co l l a r d i s t i n c t ; p os t e r i o r t r a n sver s a l i m p ress i o n s t ro n g a n d n e a r pos t e r i or m a r g i n o f pronot u m j pos t e r i o r co ! l a r i n v i s i b l e ; p ranot a l t u b erc l es not reach i n g t e g u l a e ; sc u t u m d u " , h a i r y , ret i c u l a t e ru gose ; not a u l i ces i n comp l e t e , reach i n g 0 , 3 3 l e n g t h o f sc u t u m ; sc u t e l l u m d u l l a n d ret i c u l a te r u g ose ; m e t a n o t u m sh i n y , smoo t h , w i t h o u t scu l p t u re , e x c e p t for some p u n c t u res l oc a t e d n e a r sc u te l l u m ; p ropodeum ret i c u l a t e r u gose , w i t h 2 l o n g i t u d i n a l k ee l s o n po s t e r i o r s u rface ; fore- w i n g w i t h 2 b ro w n t r a n s versa ! b a n d s a n d " i t h d i s t a l p a r t o f ra d i a l v e i n m u c h l o n g e r t h a n prox i m a l p a r t ( 25 : 1 1 ) ; r a d i a l ce l l broade l y open ; seg men t 1 of f ron t t a r s u s n e a r l y as l o n g as seg men t 4; se g m e n t s 2 and 3 of front tarsus p ro d u c e d i n to hook s ; en l a r g e d c l a w ( F i g . 670 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 1 :' l a m e l l ae ; s e g rnen t 5 o f f ron t t a r s u s ( F i g . 670 ) w i t h 1 r o w of 1 0 l a m e l l ae ; a pex w i t h a g ro u p of a t l e a s t 40 l a me l l ae ; t i b i a l s p u r s 1 , 1 , 2 . MAL E : u n k n o w n L O C U S TYP I C U S : G a m boa ( P a n a m a ) TYP I C A L lviA T E R I A L : h o l o t y p e F ! i n LE . .., OSTS : u n k n o w n U I ST R I B U T I O N : Neot rop i c reg ion : PANAMA :

G a nlboa ( P i pe l i n e ['(o a d ) ,

LE !

3 . A l;,h a dryin u s parvus n . s p .

c :: MAL E : f u l l y w i n g ed ; l e n g t h 3 , 75 m m ; t e s t a ceou s , " i t h p e t i o l e b l ack ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a i se g m e n t s i n fo l l ow i n g p roport i o n s : 8 :

990

IVlem . Amer . E n ! .

F i g . 67 1

I ns! . ,

no.

37,

1 984

- C h e l a of A l p h a dryin us p arvus n . sp .

( type )

4 , 5 : 1 9 : 1 1 : 1 0 : 9 : 7 : 6 : 6 : 7 , 5 ; h e a d d u l l , g r a n u l a te d ; frons w i t h o u t a med i a n k ee l ; occ i p i t a l c a r i n a i n com p l e t e , v i s i b l e o n l y on dors a i s i d e of o cc i p u t ; occ i p i t a l c a r i n a n o t v i s i b l e on temp l e s ; P U L � 3 ; OL � 4 ; O O L 5 , 5 ; O P L � 1 ; temp l e s i n v i s i b l e ; pro n o t u m d u l l , crossed b y a n a n t e r i o r t r a n s ve rs a l i m p ress i o n a n d a pos t e r i o r t r a n s v e rs a l i m p ress i o n ; d i sc h u m p e d ; p ro n o t a l t u b erc l e s n o t reac h i n g t eg u l a e ; pos t e i o r c o l l a r very s h or t ; pro n o t u m and scu t u m d u l l , g r a n u l a t e d ; n o t a u l i c e s i n com p l e t e , reac h i n g 0 , 3 3 l e n g t h o f sc u t u m ; sc u te l l url1 a n d m e t a n o t u m sh i n y , w e a k l y g ra n u l a t ed ; p ropodeum ret i c u l a te r u g ose ; pos t e r i o r s u rface w i t h t w o l o n g i t u d i n a l k ee l s ; fore- w i n g w i t h t w o d a r k t r a n s v e rs a l b a n d s , one o n t h e b a s a l c e l l s a n d a n o t h er b e n e a t h t h e p t e ros t i g m a ; d i s t a l p a r t of ra d i a l v e i n l o n ge r t h a n prox i m a l p a r t ( 1 3 : 8 ) ; fore t a rsa l segmen t s i n fo l l o w i n g p roport i o n s : 1 2 : 2 : 3 , 5 : 1 3 : 20 ; en l a r g e d c l a w ( F i g . 6 7 1 ) w i t h o u t s u b a p i c a l toot h , w i t h 1 r o w of 8 l a me l l ae , a m o n g w h i c h 1 l o n g a p i c a l l a me l l a ; seg m e n t 5 o f f ro n t t a rs u s ( F i g . 6 7 1 ) w i t h 2 row s o f 1 2 l a me l l ae ; a p ex w i t h a g roup of 1 0 l am e l l a e ; t i b i a l s p u rs 1 , 1 , 2 . I� ALE : u n k n o w n L O C U S TYP I C U S : C a r u a r u ( B r az i l ) TYP I CA L MAT E R I A L : ho l o t y p e F ! i n T W . H O ST S : u n k n o w n D I ST R I B UT l m, : Neot rop i c reg i o n : BRAZ I L :

C a r u a ru ,

TW!

D r y i n i n ae

991

F i g . 672 - Ch e l a of A l p h a dryin u s pegn a i n . s p .

4 . A lp h a dryin u s pegn a i n .

F E MALE :

f u l l y w i n g ed ;

l e n g t h 3 , 75-4 , 50 m m ;

( type)

sp .

h e a d b l ac k ,

w i t h fron t p a r t of

992


I ns! . ,

no.

37,

1 984

v ertex ( more a l o n g orb i t s ) , c l y p e u s a n d m a n d i b l e s t e s t aceou s ; some t i mes head t e s t aceou s , w i t h occ i p u t and ver tex b l a c k ; some t i mes a l so frons p a r t l y b l a c k ; a n ten n a e t e s t aceou s ; p ropec t u s b l ack ; p rono t u m b l ac k , w i t h p os t e r i o r h a l f or on l y pos t e r i or co l l a r a n d l a t e r a l m a r g i n s red d i sh-brow n ; l e g s t e s t a ceou s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l seg men t s i n fo l l ow i n g p ropor t i o n s : 1 0 : 5 : 20 : 1 2 : 1 1 : 1 0 : 8 : 7 : 6 : 9 ; head sw o l l e n , d u l l , g r a n u l a ted , w i t h some w e a k kee l s o n f ro n s ; occ i p i t a l c a r i n a i n comp l e t e ; P O L = 5 , 5 ; O L = 2 ; O O L = 4 , 5 ; pos t e r i o r oce l l i touch i n g occ i p i t a l ca r i n a ; p rono t u m g r a n u l a t e d , sh i n y ; a n t e r i o r t r a n sv e r s a l i m p r e s s i o n a n d a n t e r i or co l i a r w e a k ; pos t e r i o r t r a n s versa J i m p ress ion s t ron g ; pos t e r i o r co l l a r shor t , d i s t i n c t ; d i sc w e a k l y h u m p e d ; p ron o t a l t u b erc l es n o t reach i n g t e g u l a e ; scu t u m d u l l , g r a n u l a t e d , w i t h i r regu l a r k ee l s or areo l a e n e a r pos t e r i o r m a r g i n a n d on l a t e r a l reg i o n s ; not a u l i ces i n comp l e t e , reac h i n g a p p rox i m a t e l y 0 , 5 l e n g t h of scu t u m ; sc u te l l u m a n d m e t a n o t u m sh i n y , g r a n u l a t ed ; p ropodeum ret i c u l a t e r u gose, w i t h 2 l o n g i t u d i n a l k ee l s on pos t e r i o r s u rface ; fore-w i n g w i t h 1 b r o w n t r a n s v ersa l b a n d a n d w i t h d i s t a l p a r t of r a d i a l ve i n l o n g e r t h a n p rox i m a l p a r t ( 1 8 : 9 ) ; seg me n t 1 o f f ro n t t a rs u s l o n g e r t h a n segmen t 4 ( 1 7 : 1 3 ) ; seg rne n t 3 of fron t t a rs u s p ro d u ced i n to a hook ; e n l a r g e d c l a w ( F i g . 672 ) w i t h o u t s u b a p i c a l t o o t h a n d w i t h 1 0 l a m e l l a e a rn o n g w h i c h 1 a p i c a l v e r y b i g l a rn e l l a ; se g m e n t 5 o f f ro n t t a r s u s ( F i g . 672 ) w i t h 1 row o f 1 6 l a me l l ae ; a pex w i t h a group of 1 0 l a me l l ae ; t i b i a l s p u r s 1 , 1 , 2 . MALE : u nk no w n L O C U S TYP I C U S : Qu i n cem i l ( Peru ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n TW ; 1 p a r a t y p e F ! i n O T ; paratype F ! i n LE ; 2 p a r a t y p es F F ! i n OL . HOSTS : u n k n o w n D I ST R I B U T I O N : Neo t ro p i c reg i o n : PANAMA : B a rro C o l orado I s l a n d , Lt: ! OL ! E C U A D O R : L i moncoc h a ( N a po ) , OL ! O T ! P E R U : Qu i ncem i l ( M a rca p a t a ) , T W ! N O T E : t h e s p ec i es i s n a med i n honor of �I r . L u i s E . P en a .

5 . A lpha dryinus brasilia n u s n .

sp.

F E M A L E : f u I I y w i n g e d ; l e n g t h 8 , 3 1 mrn ; head redd i s h - t e s t aceou s ; a n t e n n a e a n d p rono t u m t e s t aceou s ; p ropec t u s t e s t aceo u s ; s c u t u m r e d d i sh ; sc u t e I I u m , m e t a n o t u m , p ropod e u rn , mesop l e u r a , met a p l e u r a b l a c k ; a b d omen b ro w n t e s t ac eo u s ; l e g s t e s t aceou s , w i t h f o r e t i b i a e , m i d a n d h i n d s t a l k s o f femora bro w n ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l segmen t s i n fo l l o w i n g propor t i on s : 1 5 : 8 : 57 : 33 : 26 : 1 8 : 1 3 : 1 1 : 9 : 1 1 ; h e a d d u l l , re t i c u l a te r u gose ; frons w i t h a com p l e t e med i a n k ee l ; t e m p l e s i n v i s i b l e ; occ i p i t a l c a r i n a comp l e t e ; pos t e r i o r oce l l i touch i n g occ i p i t a l c a r i n a ; P O L = 1 0 ; UL 5 ; OOL = 5 ; p ro no t u m sh i n y , w e a k l y g r a n u l a te d , w i t h a weak a n t e r i o r t r a n s v e rs a l i m p r es s i on a n d w i t h a s t r o n g pos t e r i o r t r a n s v e r s a l i m p r e s s i o n ; d i sc h u m ped ; pos t e r i o r co l l a r i n v i s i b l e ; p ro n o t a l t u berc l e s not reach i n g t e g u l a e ; scu t u m d u l l , w e a k l y ru gose ; n o t a u l i ces i ncomp l e t e reach i n g

993

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c l a w ( F i g . 673 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 1 3 l a me l l a e ; s e g m e n t 5 of f ro n t t a rs u s ( F i g . b73 ) w i t h t w o row s of n e a r l y 45 l a me l l a e e x t en d i n g co n t i n u o u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . Iv:AL E : u n k n o w n L O C U S TYP I C U S : 1 2 ° 3 1 ' S 5 5 ° 3 7 ' VI ( S i n o p , Iv: a t o G rosso, 8 r az i l ) TYP I C AL l,iA T E I� I A L : h o l o t y p e F ! i n T v'i . H O ST S : u n k n o w n U I SH
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T y p e A o f Pseu dogo n a top u s m a lesiae n . sp . o r sara,,'akensis n . sp . ( s e e t ex t ) D o r s a l p r o c e s s o f g on o f o r c e p s v e ry r ed u c ed , a l mo s t i n v i s i b l e . . . . . . . . . . . . . . . . . . . . Dicondyl u s india n u s n . sp . •

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o c c i p u t s t ra i g h t . . . Neodryin u s typh locyb a e ( A s h m e a d ) T emp l e s p r e s e n t o c c i p u t c o n Ca v e An t e n n a I s e gmen t 3 l e s s t h a n t h r e e t i m e s a s l o ng a s b r oad A n t e n n a l s egmen t 3 mo r e t h a n t h r e e t i me s a s l on g a s b r oad . P r o p o d e u m smo o t h , s h i n y , n o t r e t i c u l a t e r u g o s e . . . . . . . . . . . L a beo a l b i t arsis K i e f f e r ( s e e n om i n a d u b i a . . . . P r op od e u m d u l l , r e t i c u l a t e ru g o s e . . . . . . . . . . N o t a u l i c e s c omp l e t e , p o s t e r i o r l y s ep a r a t ed . . . . . . . . . . . . . . . . T e mp I e s a b s en t ;

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N o t a u l i c e s i n c omp l e t e D o r s a l p r o c e s s o f g on o f o r c ep s s l e n d e r ( F i g . 1 1 8 8 ) . . . . . . . Gona top u s herb arum ( R . C . L . P e r k i n s ) D o r s a I p r o c e s s o f g on o f o r c ep s b r o a d en ed ( F i g . 1 0 3 6 ) . . . . . . . . . . . . . E s a gona top u s niger ( F e n t o n ) N o t a u l i c e s r e a c h i n g app r o x i ma t e l y 0 , 8 0 l e n g t h o f s c u t u m ; .

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p r o c e s s o f g o n o f o r c ep s l on g a n d s l e n d e r ( F i g . 1 2 6 5 A ) . . . Embolemus n a s u tu s ( A s h m e a d ) ( s e e n om i n a d u b i a ) .

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N o t a u l i c e s o n l y v i s i b l e o n p r ox i ma l h a l f o f s cu t u m ; d o r sa l p r o c e s s o f g on o f o r c e p s r ed u c ed a n d l ob e s h a p e d ( F i g . 1 0 1 5 ) 7 8

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Tetrodontochelys unicus ( R . C L . P e rk i n s ) P r op od e u m f u l l y smo o t h , s h i n y , n o t r e t i c u l a t e r u g o s e . . . . . . . . 8 P r op od eu m d u l I , r e t i c u l a t e r u g o s e 9 N o t a u l i c e s i n c o mp l e t e , o n l y v i s i b l e on p r ox i ma l h a l f o f s c u t u m ; s c u t u m d u l l , g ra n u l a t ed · Gona top u s cyphonotus B ra d l ey N o t a u l i c e s c omp l e t e , p o s t e r i o r l y s e p a ra t ed ; s c u t u m s h i n y , w i t h ou t s c u l p t u r e . A crodontochelys m im u s ( R . C . L . P e rk i n s ) N o t a u l i c e s c o mp l e t e a n d j o i n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dicondy l u s americ a n u s ( R . C . L . P e rk i n s ) N o t a u l i c e s c o mp l e t e a n d mo r e o r l e s s s ep a r a t ed . . . . . . . . . . . 10 D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g s 860 , 863 , 1 1 95 , 1 264 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 D o r s a l p r o c e s s o f g on o f o r c ep s n ea r l y a s l on g a s g on o f o r c ep s (F i gs 857 , 1 1 80 ) 14 D o r s a l p r o c e s s o f g on o f o r c ep s v e ry r e d u c ed ( F i g s 8 6 0 , 8 6 3 ) 12 D o r s a l p r o c e s s o f g o n o f o r c ep s l on g e r ( F i g s 1 1 9 5 , 1 2 6 4 ) . . . . . . 1 3 T h o ra x a n d p r op od eu m b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogo n a top u s stenocra n i R . C . L . P e rk i n s T h o ra x a n d p r o p od e u m b r own - r e d d i sh Pse u dogona top u s arizonicus R . C . L . P e rk i n s D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex r o u n d ed ( F i g . 1 1 9 5 ) . . . · . . . . . . . . . . Gon a top u s a shmeadi K i e f f e r D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex i r r e g u l a r l y t ru n c a t ed ( F i g . 1 2 6 4 ) . . . . . L a beo longitarsis A s h m e a d ( s e e n om i n a d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h med i a n a n d d i s t a l r eg i o n b r o a d e n ed ( F i g s 8 5 4 , 8 5 7 , 1 2 5 7 , 1 2 5 8 ) . . . . . . . . . . . . . . . . . . . . . . . 1 5 D o r s a l p r o c e s s o f g on o f o r c ep s f u l l y s l e n d e r o r on l y w i t h d i s t a l r eg i on b r o a d e n ed ( F i g s 1 0 1 3 , 1 1 8 0 , 1 1 9 7 , 1 2 7 0 ) . . . . . . . 1 8 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p e x p o i n t ed ( F i g . 8 5 7 ) . · . . . . . . . . . . . . Pseu dogon a top u s sjoestedti ( K i e f f e r ) D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p e x r o u n d ed ( F i g s 8 5 4 , 1 25 7 , 1 25 8 ) 16 D o r sa l p r o c e s s o f g on o f o r c ep s w i t h d i s t a l r e g i on w i t h o u t p a p i l l a e a n d w i t h d i s t a l ma r g i n n o t s e r r a t e ( F i g . 1 2 5 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . Neogona top u s ie n t om' F r ey t a g ( n om i n a d ub i a ) D o r s a I p r o c e s s o f g on o f o r c ep s w i t h d i s t a I r eg i o n c ov e r ed w i t h p ap i l l a e a n d w i t h d i s t a l ma r g i n s e r ra t e ( F i g s 8 5 4 , 1 2 5 8 ) . . . . 1 7 N o t a u l i c e s n ea r e r , a l mo s t j o i n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogon a top u s c an a densis n . sp . N o t au l i ce s l es s n e a r · Neogon a top u s gra n dis F r ey t ag ( n o m i na d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h d i s t a l r eg i on v e r y b r o a d en ed a n d w i t h d i s t a l ma r g i n s l i g h t l y s e r ra t e ( F i g . 1 1 9 7 ) . Gona topus ombrodes ( R . C . L . P e rk i n s ) ·

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D o r s a l p r o c e s s o f g o n o f o r c ep s f u l l y s l e n d e r or a t mo s t w i t h d i s t a l r e g i on s l i g h t l y b road e n e d { F i g s 1 0 1 3 , 1 1 8 0 , 1 2 7 0 ) . . . . 1 9 1 9 R a d i a l v e i n r e g u l a r l y c u r v ed { F i g . 1 1 7 8 ) · . . . . . . . . . . Gon a top u s a gropyrus F en t o n R a d i a l v e i n w i t h d i s t a l a n d p r ox i ma l p a r t s f o rm i n g a n a n g l e {F i g . 1 269 ) 20 2 0 D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p e x p o i n t ed { F i g . 1 2 7 0 ) .

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N E OTROP I C R E G I ON

T emp l e s a b s e n t ; o c c i p u t s t ra i g h t · . . . . . . . . . . Neodry i n u s trin ita tis R i c h a rd s T emp l e s p r e s e n t , m o r e o r l e s s p r o m i n e n t ; o c c i p u t c o n c a v e 2 A n t e n n a l s eg m e n t 3 I e s s t h a n t h r e e t i m e s a s l on g a s b r o a d · . . . . . . . . . L a beo s a n ctivincenti A sh m e a d ( n om i n a d u b i a ) A n t en n a l s eg m e n t 3 m o r e t h a n t h r e e t i me s a s l on g a s b r oad 3 P r o p o d e u m f u l l y r e t i cu l a t e ru g o s e 4 P r op od eu m a t mo s t p a r t l y r e t i c u l a t e ru g o s e , f u l l y o r p a r t l y s m o o t h , s ome t i me s r u g o s e 6 D o r s a l p r o c e s s o f g on o f o r c ep s l on g e r t h a n p en i s { F i g . 1 2 6 6 ) L a berin u s p aranen sis Og I o b l i n ( n om i n a d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c e p s s h o r t e r t h a n p en i s ( F i g s 9 3 9 , 1 25 9 ) 5 D o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p e x s l i g h t I y b r o a d en ed {F i g . 939) . . . . . A p terodryin u s tij u c a n u s ( A r l e ) •

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Gon a top u s bartletti n . s p . P r o p o d e u m w i t h d o r s a I s u r f a c e f u I I Y smo o t h ; p o s t e r i o r s u r f a c e r e t i cu l a t e r u g o s e ; d o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p e x mo r e b r o a d e n ed { F i g . 1 2 6 8 ) · . . . . . . L a beo sim u l a n s A s h m e a d ( s e e n om i n a d u b i a ) D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n p en i s ( F i g . 1 20 6 ) Gona top u s fern a n din a e n . s p . D o r s a l p r o c e s s o f g on o f o r c ep s s l i g h t l y s h o r t e r t h a n p en i s {F i g . 1 20 8 ) 9 L eg s b r o wn - d a rk , w i t h d i s t a l a p e x o f c ox a e , t r o c h a n t e r s a n d f emo ra y e l l ow ; p r ox i ma l a p e x o f t i b i a e a n d t a r s i y e l l ow .

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p r o c e s s o f g o n o f o r c ep s v e r y b r o a d en ed { F i g . 1 2 5 9 ) L a beo gren a den sis A s h m e a d ( s e e n om i n a d u b i a ) N o t a u l i c e s c o mp l e t e 7 N o t a u I i c e s i n c omp l e t e 8 P r o p o d e u m n o t smo o t h , f u l l y weak l y r u g o s e ; d o r sa l p r o c e s s o f g on o f o r c e p s w i t h a p e x s l i g h t I y b r o a d en ed { F i g . 1 2 2 0 ) .

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1 064

M e m . Amer .

En!.

I ns!. ,

n o . 37 ,

1 984

. . . . . . . . . . . . . . . . . . A crodontochelys cubensis R i c h a r d s L e g s t e s t a c e o u s -d a r k , w i t h d i s t a l a p ex o f c ox a e , t r o c h a n t e r s a n d f em o r a s l i g h t I y l i g h t ; p r o x i ma l a p e x o f t i b i a e a n d t a r s i s l i gh t I y l i gh t Gona top u s breviforceps K i e f f e r .

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AUSTRA L I AN REG I ON

T emp l e s a b s en t ; o c c i p u t s t ra i g h t o r a l mo s t s t ra i g h t . . . . . . . . . . . Neodryin u s koebelei R . C . L . P e rk i n s T emp l e s p r e s en t ; o c c i p u t c o n ca v e .

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H ea d w i t h a d i s t i n c t o c c i p i t a l v en t ra l s i d e o f t h e h ea d •

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Head w i shor t , pos t er i No t au l i

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ca r i n a f r om p o s t e r i o r o c e l l i •

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t h ou t o cc i p i t a l ca r i n a o r w i t h o c c i p i t a l c a r i n a v e ry o n l y v i s i b l e b eh i n d o c e l l i o r on t h e s i d e s o f t h e o r o c e l l i , b u t n ev e r r e a c h i n g t emp i e s c e s p os t e r i o r l y j o i n t •

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. . . . . . . . . . Echthrodelp h a x fairch il dii R . C . L . P e rk i n s N o t a u l i c e s p o s t e r i o r l y d i s t i n c t l y s ep a ra t ed . Ech throdelp h a x nigricollis R . C . L . P e rk i n s 4 L a b i a l p a l p i w i t h 1 s eg m e n t . . . . . . . . . . . Haplogona top u s vitie n s is R . C . L . P e rk i n s L a b i a l p a l p i w i t h 2 - 3 s e g men t s 5 5 N o t a u I i c e s i n c omp l e t e 6 N o t a u I i c e s c omp l e t e 8 6 P r op od e u m s mo o t h a n d s h i n y ; p r ox i ma l p a r t o f rad i a l v e i n mu c h s h o r t e r t ha n d i s t a I p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Gona top u s mel a n i a s ( R . C . L . P e rk i n s ) P r op od e u m r e t i c u l a t e ru g o s e , d u l l ; d i s t a l p a r t o f ra d i a l v e i n n ea r l y a s l on g a s p r ox i ma l p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 7 D o r s a l p r o c e s s o f g o n o f o r c ep s n ea r l y a s l o n g a s g o n o f o r c e p s Gona top u s zea l a n dicus n . s p . (F i g . 1 239) D o r s a l p r o c e s s o f g o n o f o r c ep s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g . 1 246 ) 1 . . Gona top u s op t a b ilis ( R . C . L . P e rk i n s ) Gon a top u s decora t u s ( R . C . L . P e rk i n s ) 9 8 N o t a u l i c e s s e p a ra t ed 1 1 N o t au l i c e s j o i n t 9 D o r sa l s u r f a c e o f p r op od e u m smo o t h , n o t r e t i cu l a t e ru g o s e , o n l y w i t h med i a n f u r r ow . . . . . . . . . . . . . . . . Pseudogona topus s a c c h a retorum R . C L . P e rk i n s D o r s a l s u r f a c e o f p r op od eu m f u l l y r e t i c u l a t e ru g o s e 10 1 0 D o r s a l p r o c e s s o f g o n o f o r c ep s l on g , w i t h d i s t a l h a l f .

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1 T h e o n l y k n ow m a l e spec i men c a n b e l o n g t o Gona top u s op ta b ilis or decor a t u s ( see tex t ) .

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Gon a topo d i n a e

b r o a d e n ed , p a r a l l . . . . . . . . . . . . . . . . . Dorsa l p ro c e s s of 960 ) •

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1 065

e l t o t h e p en i s ( F i g . 8 7 8 ) . . . . . . . . . . . . . . . . . . . . Ps e u dogo n a top u s hospes R . C . L . P e rk i n s g on o f o r c ep s s h o r t , s l en d e r , t r a n s v e r s e ( F i g . Dicondylus perkinsi ( A s h m ea d ) •

1 1

P r op od e u m s m o o t h , n o t r e t i c u l a t e ru g o s e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogon a top u s dichromus R . C . L . P e rk i n s P r op o d e u m f u l l y o r a l mo s t f u l l y r e t i c u l a t e r u g o s e . . . . . . . . . . . 1 2 1 2 D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h n u m e r o u s s h o r t , p o i n t ed l am e l l a e ( F i g . 9 6 9 ) ; p r o p o d e u m f u l l y r e t i c u l a t e rug o s e . . . . . . . . . . . . . . . . . . . . . . . Dicondylus alpin u s ( G o u r I a y ) D o r s a l p r o c e s s o f g on o f o r c ep s smo o t h , w i t h ou t s h o r t l a m e l l a e .

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( F i g s 8 7 5 , 8 7 6 ) ; p r o p o d e u m r e t i c u l a t e ru g o s e , b u t w i t h d o r s a l s u r f a c e s h ow i n g t w o l a t e r a l s m o o t h a r e a s . . . . . . . . . . Pseu dogon a top u s n igric a n s R . C . L . P e r k i n s .

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1 . Neodryin u s R .

C.

L Perk i n s

Neodryin u s R . C . L . Perk i ns 1 905 : 50 . Psilodryin us K i effer i n K i effer et M a r s h a l l 1 90 6 : 497 ( s y n . p ro p osed b y R i c h a r d s 1 953 ) ; t y p e spec i e s Psilodryin u s a c u ticollis ( K i effer 1 905 ) , ori g . des i g . Neodryin u s R . C . L . P e r k i n s : K i effer i n K i effer e t M a rsh a l l 1 906 : 497 . Prodry i n u s K i effer i n K i effer et M a rs h a l l 1 906 : 497 ( s y n . p roposed by R i c h a rd s 1 95 3 ) ; t y p e spec i es Prodryin u s bra chycerus ( K i effer 1 904 ) , or i g . des i g . Neodryin u s R . C . L . Perk i n s : K i effer 1 9 1 4b : 2 5 . Prodryin u s partim K i effer : K i effer 1 9 1 4b : 4 9 . Ps ilodryin u s K i effer : K i effer 1 9 1 4b : 53 . Ph a n erodryin us Roep k e 1 9 1 6 : 289 ( s y n . p roposed by R i c h a r d s 1 95 3 ) ; t y pe s p ec i e s Ph a n erodryin us ja v a n u s R o e p k e 1 9 1 6 , or i g . d es i g . Neodryin u s R . C . L . Perk i n s : R i c h a r d s 1 939 : 1 89 . Prodryin u s I< i ef fe r : R i c h a r d s 1 939 : 1 89 . Psilodryin u s K i effe r : R i c h a rd s 1 93 9 : 1 89 . L eptodryin us R i c ha rd s 1 95 3 : 67 ( n . sy n . ) ; t y p e s p ec i e s Leptodryin us leptop u s R i c h a rd s 1 95 3 , m o n o t y p i c . Neodryin u s R . C . L . Perk i n s : D l m i a n d C u rrado 1 97 6 a : 1 84 . Psilodryinus K i ef fe r : D i m i a n d C u rr a d o 1 976a : 1 84 . Prodryin u s K i ef fe r : D l m i a n d C u rrado 1 976a : 1 84 .

T Y P E SP E C I E S : Neodryin u s koebelei R . C . L . Perk i n s 1 905 , o r i g . des i g . F E MA L E ( F i g s 7 1 5 , 7 1 6 ) : f u l l y w i n ged ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; p ro no t u m crossed by a s t ro n g t r a n s v e rs a l

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1 066

Mem . Amer .

Fig.

715

-

Ent.

I ns t . ,

n o . 37 ,

1 984

F em a l e of Neodryin u s nelsoni R . C . L . Perk i n s

i m p ress i o n ( F i g . 7 1 6 ) ; n ot a u l ices i n v i s i b l e ; e n l a r g e d c l a w ( F i g . 7 1 9 ) w i t h s u b a p i c a l too t h a n d l am e l l ae or b r i s t l es or p e g - I i k e h a i rs ; t i b i a l s p u rs 1 , 0 , 2 ( F i g . 758 B ) . MALE ( F i g s 7 1 7 , 7 1 8 ) : f u l l y w i n g e d ; m a x i l l a ry p a l p i w i t h 6 s e g men t s ; l ab i a l p a l p i w i t h 3 segmen ts ; occ i p u t s t ra i g h t ; temp l es i n v i s i b l e ; o c c i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d t h e oce l l i o r o n t h e s i des o f t he pos te r i o r oce l l i , b u t n e v e r reac h i n g t e m p l es ; t i b i a l s p u rs 1 , 1 ,2.

G o n a topod i n ae

Fig. 716

-

1 067

Fem a l e of Neodryinus n e lsoni R . C . L .

D I ST R I BU T I O N : E t h i op i a n , O r i e n t a l , Nearc t i c , HOST S : Fl a tidae, Ric aniidae, Nogodinidae S P E C I E S : 26

Perk i n s

Neotrop i c , A u s t r a l i a n

E T H I O P I AN S P E C I E S O F NEODRYINUS

T w o s p ec i e s a re k n ow n .


F E MALES

O c c i p i t a l c a r i n a i n v i s i b l e ; s cu t u m n o t r e t i c u l a t e ru g o s e , w i t h n u m e r ou s l on g i t u d i n a l k e e l s 1 . a n tiq u u s B en o i t O c c i p i t a l c a r i n a v i s i b l e b eh i n d t h e o c e l l i a n d on t h e s i d e s o f t h e p o s t e r i o r o c e l l i ; s c u t u m f u l l y r e t i c u l a t e ru g o s e . . . . . . . . . . •

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1 068

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I ns!. ,

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1 984

gigas C eb a l l o s

M a l e of Neodryin u s koebelei R . C . L . Perk i n s

MALES

U n k no w n

no. 37,

G o n a topod i n a e

Fig.

1 069

7 1 8 - M a l e of Neodryin u s koebelei R . C . L . Perk i n s

1 . Neodryin u s a n t i q u u s Beno i t

Neodryin u s a n ti q u u s Be n o i t 1 95 3 b : 402 .

F E MALE : f u l l y w i n g e d ; l e n g t h 4 , 37 m m ; h e a d b l ac k , w i t h m a n d i b l e s redd i s h ; a n t e n n ae b ro w n , w i t h seg m e n t 5 , d i s t a l p a r t o f seg m e n t 4 a n d p ro x i m a l p a r t of seg men t 6 ye l l o w ; p ron o t u m b l a c k , w i t h s i des p a r t l y redd i s h ; meso t h o r a x , met a t ho r a x a n d p ro p o d e u m b l a c k ; a b domen b ro w n b l ac k ; l e g s b ro w n -red d i s h ; head w ea k l y exc a v a ted , w i t h n U merous l o n g i t u d i n a l k ee l s o n fron s , v e r t e x and occ i p u t ; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m crossed b y a t r a n s v e rs e i mp r es s i on , d u l l , g r a n u l a ted , w i t h n u merous l o n g i t u d i n a l kee l s on co l l a r a n d d i s c ; pos t e r i o r edges of :) rono t u m ro u n ded , not p ro d u ced i n to l ob e s ; scu t u m du I I , as scu I p t u red as ,::J ro no t u m , w i t h n u m e rous l o n g i t u d i n a l kee l s ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m a s scu I p t u red a s scu t u m ; met a n o t u m v ery reduced a n d sho r t ; p ropode u m f u l l y re t i c u l a t e ru gose , w i t ho u t l o n g i t u d i n a l k ee l s on pos ter i or s u rface ; fore- w i n g w i t h t w o b r o w n t r a n svers a l b a n d s ; d i s t a l p a r t of r a d i a l ve i n m u c h l o n ger t h a n p ro x i m a l p a r t ; fore t ro c h a n ters a p p ro x i m a te l y f o u r t i mes a s l on g a s b ro a d ; segment 1 of f ro n t t a r s u s l o n g e r t h a n seg m e n t 4 ( 20 : 1 0 ) ; e n l a r ged c l a w ( F i g . 7 1 9 ) w i t h a s U b a p i c a l [Oo t h a n d 4 p e g - l i ke h a i rs ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 7 1 9 ) w i t h t w o - o w s of n e a r l y 9 l a m e l l ae ; a p e x w i t h a g r o u p of a t l e a s t 20 l a m e l l a e ; .,-, a x i I l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segm e n ts ; t i b i a l s ;:) u rs 1 , 0 , 2 . '.'ALE : u n k n o w n

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LOCUS T Y P I C U S : M a d a g as c a r T Y P I CA L M AT E R I A L : ho l o t y p e F ! i n P ( b y m i s t a k e t h e ho l o t y p e w a s l ab e l l ed Neodryin u s berla n di n . sp . ) . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n re g i o n : MADAGASCAR , P !

2 . Neodryin u s gigas Ceb a l l os

Neodryin u s gigas C eb a l l os 1 936 : 5 1 . Neodryin u s giga s C eb a l l o s : Beno i t 1 953b : 401 .

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1 072

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- C he l a of Neodryin u s gigas C eb a l l os ( t y p e )

prono t u m ro u n de d , n o t p ro d uced i n to l o b e s ; sc u t u m d u l l , f u l l y ret i c u l a t e r u gose ; n o t a u I i ce s i n v i s i b I e ; sc u t e l l u m d u I I , f u I I y re t i c u I a te r u gose ; m e t a n o t u m re t i c u l a te ru gose , v e r y red u ced a n d s h o r t ; p ropodeum f u l l y

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1 073

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1 075

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1 076


I nst . ,

n o . 37 ,

1 984

1 . Neodryin u s leptopus R i c h a r d s

Neo dryin u s (Leptodryin u s ) leptop u s R i c h a rd s 1 95 3 : 66 .

F E M A L E : f u l l y w i n ged ; l e n g t h 5 , 93 m m ; head b l ac k , w i t h f ro n t p a r t o f v e r t e x ( more a l o n g orb i t s ) , c l y p e u s a n d m a n d i b l es y e l l o w ; a n te n n ae testaceo u s , w i t h segmen ts 3-5 d a r k ; t ho r a x a n d p ropodeum b l ac k , w i t h poste r i o r s u rface o f pron o t u m red d i s h ; a b domen b row n ; l eg s b row n , w i t h t a rs i l i g h t ; hea d d u l l , g r a n u l a t e d , w i t h a s t ro n g med i a n k e e l between t he a n t E! n n a l soc k e t s ; f r o n s w i t h some short t r a n svers a l k ee l s o n t he s i d es of the med i a n k ee l ; oce l l a r t r i a n g l e e q u i l a t e ra l ; a n t e r i o r h a l f o f pronot u m

( F i g . 7 2 2 A ) c rossed b y a w e a k t r a n s v e rs a l i m p ress i on ; poste r i or h a l f w i t h a no t h e r w e a k t r a n sversa l i mp ress i o n ; c o l l a r s h i n y , w i t h o u t s c u l p t u re , s m oo t h ; d i s c s h i n y , w i t h s t r o n g I d n g i t u d i n a l k e e l s ; poster i o r ed ges o f t h e p ron o t u m n o t p ro d u ced i n to l obes ; sc u t u m s h i n y , r u gose , w i t h s t ro n g l o n g i t u d i n a l k e e l s ; n o t a u l i ces i n v i s i b l e ; s c u te l l u m s h i n y , f u l l y re t i c u l a t e r u g ose ; m e t a n o t u m i n v i s i b l e ; p ropodeum f u l l y ret i c u l a te r u gose , w i t h t w o l o n g i t u d i n a l kee l s on poste r i o r s u rf a ce ; fore t ro c h a n ters a p p ro x i m a te l y four t i mes as l o n g as b road ; fore-w i n g w i t h two b ro w n t r a n s v e r s a l b a n d s ; d i s t a l p a r t o f rad i a l ve i n m u c h l on g e r t h a n p ro x i m a l p a r t ; segmen t 1 o f f r o n t t a r s u s w e a k l y s horter t h a n segmen t 4 ( 27 : 30 ) ; e n l a rged c l a w ( F i g . 723 ) w i t h s U b a p i c a l too t h a n d 8 l a m e l l ae ; segme n t 5 of f ro n t t a r s u s ( F i g . 723 ) w i t h t w o rows of 1 8 + 1 1 l ame l l ae ; apex w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 seg men t s ; l ab i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n k n o w n LOCUS T Y P I C US : K uc h i n g ( Borneo , I n dones i a ) T Y P I C A L MAT E R I A L : h o l o t y p e F ! i n BM . HOST S : u n k no w n D I S T R I BU T I O N : O r i e n t a l reg i o n : I N D O N E S I A : K uc h i n g ( Borneo ) , B M ! LAOS : B a n - v a n-eue ( SE of P ho u - k o w - k u e i ) , B !

2 . Neo dryin u s retic u l a t u s ( F o u t s )

Psilodryin u s reticu l a tus F o u t s 1 92 2 : 632 . Neodryin u s retic u l a t u s ( F o u t s ) : B a l t a z a r 1 966 : 204 . F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 25 m m ; head b l ac k , w i t h m a n d i b l es red i s h ; a n te n n ae b row n , w i t h segme n ts 8- 1 0 l i g h t a n d ve n t r a l face of segmen t 1 w h i te ; t ho r a x , p ropodeum a n d a b domen b l ac k ; l eg s b l ack b row n ; head v e r y e x c a v a te d , s h i n y , g r a n u l a t e d , w i t h n u merous l o n g i t u d i n a l k ee l s o n fro n s , vertex and o cc i p u t ; SOme k ee l s a re a l so v i s i b l e amon g the oce l l i ; occ i p i t a l c a r i n a i n v i s i b l e ; p ron o t u m crossed b y

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1 077

C he l a of Neodryin u s reti c u l a t u s ( F o u t s ) ( t y p e )

a t r a n s v e rs a l i m pres s i on , h a i r y , w i t h some l o n g i t u d i n a l k ee l s m a i n l y on the co l l a r ; pos t e r i o r edges of the p ro n o t u m rou n d e d , n o t p ro d u ced i n to

Mem . A m e r .

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Ent.

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no. 37,

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F i g . 725 - C he l a of Neodryin u s chel a tu s n . s p .

( type )

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l obes ; s c u t u m d u l l , g r a n u l a t ed a n d f u l l y r e t i c u l a t e r u gose , w i t h t w o k ee l s i n s t e a d of t h e no t a u l i ces ; n o t a u l i ces i n v i s i b l e ; sc u te l l u m a s scu I p t u red as sc u t u m ; m e t a n o t u m very reduced a n d s ho r t ; p ropodeum fu I I y r e t i c u l a t e r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e pos t e r i o r s u rface ; fore- w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; fore t roc h a n t ers a p p rox i m a te l y f o u r t i me s a s l o n g a s b road ; segmen t 1 of fron t t a r s u s l o n g e r t h a n s e g m e n t 4 ( 30 : 22 ) ; en l a r g e d c l a w ( F i g . 724 ) w i t h s u b a p i c a l t o o t h a n d 5 b r i s t l es ; segme n t 5 of f r o n t t a rs u s ( F i g . 724 ) w i t h 2 row s of 1 7 + 1 1 l am e l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 1 4 l am e l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n known l O C U S T Y P I C US : los B a no s ( P h i I i p p i n es ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n VIA . HOST S : u n k no w n D I ST R I B U T I O N : O r i e n t a l reg i o n : P H I L I P P I N E S : L o s B a nos ( L u zon ) , WA !

3 . Neodryin u s chela tus n . s p .

F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 25-6 , 80 m m ; h e a d b l ac k , w i t h s i des of c l y p e u s and m a n d i b l es y e l l ow ; a n te n n ae b l ac k , w i t h segme n t s 6-1 0 r ed d i s h a n d v e n t r a l f a c e of seg m e n t s 1 -2 y e l l o w ; t horax , p ropodeum a n d p e t i o l e b l ac k , w i t h s i des o f prono t u m p a r t l y redd i s h ; a b domen red d i s h ; fore l e g s b l ac k , w i t h coxae a n d t roc h a n t e r s p a r t l y y e l l ow , c l u b s of femora redd i s h , e x t e r n a l f ace of t i b i ae red d i s h , e n l a r ged c l a w y e l l ow ; m i d a n d h i n d l e g s b l a ck , w i t h coxae p a r t l y red d i s h , t roc h a n t e r s a n d c l u b s o f fe mora f u l l y red d i s h , t a r s i y e l l o w , w i t h p rox i m a l h a l f of s e g m e n t

1 b l ac k ; h e a d e xc a v a t e d , s h i n y , w i t h n u merous l o n g i t u d i n a l k ee l s on frons and v e r t e x ; occ i p i t a l c a r i n a i n comp l e t e , reach i n g a p p rox i m a te l y 0 , 5 l e n g t h of OOl ; p rono t u m ( F i g . 722 B ) crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , a s scu l p t u re d a s h ea d , w i t h n u merous l o n g i t u d i n a l k ee l s ; p o s te r i or ed ges o f t h e p ron o t u m rou n d e d , n o t p roduced i n t o l ob e s ; scu t u m h a i r y , s h i n y , a s s c u l p t u red a s h ea d , w i t h cen t r a l reg i o n ret i c u l a t e r u gose , o n t he s i des w i t h n u merous l o n g i t u d i n a l k ee l s ; no t a u l i ce s i n v i s i b l e ; sc u te l l u m f u l l y ret i c u l a t e r u g os e , a s scu l p t u red a s h e a d ; m e t a n o t u m v ery reduced a n d s hort ; p ropodeum f u I I y re t i c u I a t e r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on pos te r i or s u rface ; fore-w i n g w i t h t wo b r o w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f rad i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; segmen t 1 of f ron t t a rs u s l o n g e r t h a n seg m e n t 4 ( 35 : 2 5 ) ; e n l a r g e d c l a w ( F i g . 725 ) w i t h s u b a p i c a l t oo t h a n d 4 peg- I i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 725 ) w i t h t w o rows of 1 0 + 8 l a me l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 3 0 l ame l l a e ; m a x i l l a r y p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n k n o w n l O C U S T Y P I C U S : B a n V a n E u e ( L aos )

Mem . A m e r . E n ! .

1 080

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-

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n o . 37 ,

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C he l a of Neodryinus rub escens n . s p .

( type)

T Y P I CAL M A T E R I A L : h o l o t y p e F ! i n B ; 1 p a r a t y pe F ! i n CO . HOSTS : u n k n o w n D I STR I BU T I O N : O r i e n t a l reg i o n : LAOS : B a n V a n E u e ( V i en t i a n e ) , P H I L I PP I NE S : D a l a w a n B a y ( Ba l a b ac ) , CO !

B!

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4 . Neodryin u s rubescens n .

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sp .

F E MA L E : f u l l y w i n ge d ; l e n g t h 5 , 00 m m ; h e a d b l ack , w i t h s i de s of c l y p e u s y e l l ow a n d m a n d i b l es redd i s h ; a n t e n n a e b l ac k , w i t h seg m e n t s 7- 1 0 redd i s h a n d v e n tra l face of s e g m e n t 1 y e l l o w ; t ho ra x , propode u m a n d pe t i o l e b l ac k , w i t h redd i s h n u a nces on t h e s i des o f t h e p ronot u m ; a b domen b row n - re d d i s h ; l e g s redd i s h ; head exc a v a ted , s h i n y , g r a n u l a ted , w i t h n u merous l o n g i t u d i n a l k ee l s on f ro n s a n d verte x ; occi p i t a l ca r i n a d i s t i n c t , not e x te n d i n g b e y o n d 0 , 5 l en g t h o f OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p re s s i o n , h a i r y , as scu l p t u red as h e a d , w i t h n u m e ro u s l o n g i t u d i n a l k ee l s on t h e c o l l a r ; poster i or e d g e s of t h e p ro n o t u m rou n ded , not produced i n to l o b e s ; s c u t u m d u l l , as sc u l p t u red as p ronot u m , ret i c u l a t e r u gose , w i t h two k ee l s i n s t e a d of t h e n o t a u l i ces ; no t a u l i ce s i n v i s i b l e ; sc u te l l u m a s scu l p t u red a s sc u t u m ; m e t a n o t u m v e r y reduced a n d s hort ; p ropod e u m f u l l y ret i c u l a t e ru gose , w i t h t w o l o n g i t u d i n a l k ee l s o n poste r i o r s u rf ace ; fore-w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f ra d i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; fore t roc h a n ters ap prox i m a te l y four t i mes a s l o n g a s b road ; s e g m e n t 1 of fron t t a rs u s l o n g e r t h a n s e g m e n t 4 ( 2 3 : 1 5 ) ; en l a rged c l a w ( F i g . 726 ) w i t h s u b a p i c a l tooth a n d 7 b r i s t l es ; s e g m e n t 5 of fro n t t a r s u s ( F i g . 726 ) w i t h t wo rows of 10 palpi

+

1 3 l a m e l l ae ; apex w i t h a g ro u p of a t l ea s t 1 5 l am e l l ae ; m a x i I l ary w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g m e n ts ; t i b i a l s p u rs 1 , 0 , 2 .

M AL E : u n k n o w n L O C U S TYP I C U S : K a n d y ( S r i L a n k a ) T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n B M . HOST S : u n k no w n D I ST R I BU T I O N : O r i e n t a l reg i o n : SR I LANKA : K a n d y ,

BM !

5 . Neodryin u s dolosus n . s p .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 68 m m ; head b l ac k , w i t h s i des of c l y p e u s a n d m a n d i b l es redd i s h ; a n te n n a e b l ac k - b ro w n , w i t h s e g m e n t s 8- 1 0 red d i s h a n d ven t r a l face of s e g m e n t 1 y e l l ow ; t hora x , p ropodeum a n d p e t i o l e b l ac k ; a b do m e n b row n ; l e g s b ro w n - b l ac k ; h e a d exca v a ted , g r a n u l a ted , s h i n y , h a i r y ; fro n s a n d v e r te x w i t h n u merous l on g i t u d i n a l k ee l s ; occ i p i t a l car i n a d i s t i n c t , not exte n d i n g beyon d 0 , 5 l en g t h of OO L ; pron o t u m crossed b y a t r a n s vers a l i m p re s s i o n , s h i n y , h a i r y , a s s c u l p t u red a s head , w i t h S o m e l o n g i t u d i n a l k ee l s on t h e co l l a r ; poste r i or edges of t h e p ro n o t u m ro u n ded , not p rod uced i n to l obes ; s c u t u m h a i r y , a s sc u l p t u red a s p ronot u m , w i t h two i n c i m p l e te k ee l s i n s tead of t h e n o t a u t i ces j no t a u l i c e s i n v i s i b l e ; s c u te l l u m a s scu l p t u red a s sc u t u m ; m e t a n o t u m v e r y reduced a n d short ; p ropod e u m f u l l y ret i c u l a t e ru gose , w i t h two l o n g i t u d i n a l k ee l s on t h e poster i o r s u rface ; fore-w i n g w i t h two b ro w n t r a n s v e rs a l b a n d s ; fore troc h a n ters a p p rox i m a te l y fo u r t i mes a s

1 082

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no.

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F i g . 729 - C he l a of Neodryin us dolosus n . sp .

( ty p e )

l o n g a s b ro a d ; segmen t 1 o f f ro n t t a r s u s l o n ge r t h a n segmen t 4 ( 2 5 : 1 8 ) ; en l a r g e d c l a w ( F i g . 7 2 7 ) w i t h s u b a p i c a l too t h a n d 8 peg - I i ke h a i rs ; segmen t 5 o f f r o n t t a r s u s ( F i g . 727 ) w i t h t w o rows o f 1 3 + 1 4 l a me l l ae ; a p e x w i t h a g r o u p o f a t l ea s t 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . I,j AL E : u n k n o w n L O C U S T Y P I C US : T a i p o k a u ( Ho n g Kon g ) TYP I CA L MAT E R I AL : ho l o t y pe F ! i n 8 . H OST S : u n k n o w n D I ST I, 1 8 U T I O N : O r i e n t a l reg i o n : HOr� G K O N G : T a i po k a u ( Ko w l oon ) , B !

6 . Neodryin u s rob ustus n .

sp .

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728 - C h e l a of Neodryin u s rob u s t u s n . sp .

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F E MA L E : f u l l y w i n g e d ; l e n g t h 5 m m ; h e a d b l ac k , w i t h m a n d i b l es redd i sh ; a n t e n n a e b l ac k , w i t h se g m e n t 1 0 redd i s h a n d v e n t r a l s i d e o f s e g m e n t 1 l i g h t ; thorax , p ropodeu m , p et i o l e , a b domen a n d l e g s b l ac k ; h e a d e x c a v a ted , d u l l , g r a n u l a t e d , w i t h n u me ro u s l o n g i t u d i n a l k ee l s o n fron s a n d verte x ; occ i p i t a l ca r i n a i n com p l et e , not exte n d i n g b e y o n d 0 , 5 l e n g t h o f OOL ; p ro n o t u m d u l l , a s scu l p t u red a s h e a d , crossed b y a t r a n svers a l i m pres s i o n , w i t h n u me ro u s l o n g i t u d i n a l k ee l s ; poster i o r e d g e s o f pron o t u m

1 084

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rounded , not p rod uced i n to l ob e s ; sc u t u m d u I I , a s s c u I p t u re d as p ro no t u m , w ea k l y re t i c u l a te r u g ose , h a i r y , w i t h 2 i n comp l e te kee l s i n s t e a d of t he n o t a u l i ces ; n o t a u l i ces a b s e n t ; s c u te l l u m a s s c u l p t u red a s s c u t u m ; m e t a n o t u m very reduced , s h or t ; p ro p o d e u m f u I I y re t i c u I a te r u g ose , w i t h t w o l o n g i t u d i n a l kee l s on poste r i o r s u rface ; fore troc h a n ters approx i m a te l y fo u r t i mes a s l o n g a s b road ; fore w i n g s w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n m u c h l o n g e r t h a n prox i m a l p a r t ; se g me n t 1 o f f r o n t t a r s u s l o n g e r t h a n seg m e n t 4 ( 2 7 : 1 7 ) ; en l arged c l a w ( F i g . 728 ) w i t h s u b a p i c a l too t h and 8 b r i s t l e s ; seg m e n t 5 of fro n t t a r s u s ( F i g . 728 ) w i t h t w o ro w s o f 2 1 l am e l l a e ; a p e x w i t h a g ro u p o f a t l ea s t 1 2 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n known L O C U S T Y P I C U S : B a n V a n E u e ( L ao s ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n B . HOSTS : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : LAO S : B a n V a n E u e ( V i e n t i a n e ) , B !

7 . Neodryin u s j a va n us ( Roep k e )

Ph anerodryin us ja v a n u s Roepke 1 9 1 6 : 290 . Psilodryin u s thoracicus F o u t s 1 92 2 : 6 3 1 ( n . s y n . ) ; locus typicus P uerto P r i ncesa ( P h i l i p p i n es ) . Neodryin u s l a w a n a e R i c h a r d s 1 953 : 64 ( n . s y n . ) ; locus typicus K u a l a Lumpur ( Malaya ) . Neodryin u s j a v a n u s ( Roep k e ) : R i c h a r d s 1 953 : 6 1 . Neodryin us thoracic us ( F o u t s ) : B a l t az a r 1 966 : 204 .

F E MA L E : f u l l y w i n ged ; l e n g t h 5 , 00-6 , 25 m m ; h e a d b l ac k , w i t h s i de s o f c l y p e u s a n d m a n d i b l e s y e l l o w ; a n te n n ae redd i s h , w i t h v e n t ra l face of s e g m e n t 1 y e l l o w ; t hora x , p ropodeum and pe t i o l e b l ac k ; a b domen redd i s h ; l e g s redd i s h ; h e a d exca v a t e d , h a i r y ; frons a n d v e r t e x w i t h n u merous l o n g i t u d i n a l k ee l s ; occ i p i t a l ca r i n a i n comp l e t e , e x t e n d i n g beyond 0 , 5 l e n g t h o f OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m press i o n , h a i r y , g r a n u l a t e d , w i t h some l o n g i t u d i n a l kee l s on t he s i d es of t he co l l a r ; poste r i or edges o f t h e p ro n o t u m rounded , not p rod uced i n to l ob e s ; s c u t um d u l l , h a i r y , as sc u l p t u red as p ro no t u m , w i t h t w o i n c i m p l e te k ee l s i n s t e a d of t h e not a u l i ce s ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m a s sc u l p t u red a s scu t um ; m e t a n o t u m v e r y reduced a n d s h or t ; p ropodeum f u l l y re t i cu l a te r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e post e r i o r s u rf ace ; fore-w i n g w i t h t w o t r a n s v e r s a I b r o w n b a n d s ; d i s t a I p a r t o f ra d i a I v e i n m u c h l on g e r t h a n p rox i m a l p a r t ; f o r e t roc h a n ters a p p rox i m a t e l y f o u r t i mes a s l on g a s b ro a d j se g m e n t 1 of fron t t a r s u s l o n g e r t h a n seg m e n t 4 ( 3 1 : 20 ) ; en I a r ged c l a w ( F i g . 729 ) w i t h s u b a p i c a l tooth a n d 8 b r i s t l e s ; seg m e n t 5 o f fron t

1 086

Mem . A mer . E n t .

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n o . 37 ,

1 984

F i g . 730 - L a b i a l p a l p u s of fem a l e of !leodryin u s j a va n u s ( Roep k e ) from K ua l a Lumpur ( M a l a y a )

t a rs u s ( F i g . 72 9 ) w i t h t w o ro ws o f 1 4 + 1 5 l a m e l l ae ; apex w i t h a g ro u p o f a t l e a s t 1 5 l a me l l ae ; m a x i l I a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 seg m e n t s ( F i g . 730 ) ; t i b i a l s p u rs 1 , 0 , 2 . M AL E : f u l l y w i n ged ; l e n g t h 3 , 37-3 , 50 m m ; h e a d b l ac k , w i t h m a n d i b les y e l l o w ; t h o r a x and propod e u m b l ack ; a b do m e n b row n ; legs b row n , w i t h t a rs i a n d fore t i b i a e l i g h t ; a n te n n a l seg men t s i n fo l l o w i n g p roport i on s : 6 : 6 : 1 6 : 1 2 : 1 2 : 1 2 : 1 2 : 1 0 : 9 : 1 0 ; h e a d h a i r y , g r a n u l a ted , w i t h a m ed i a n kee l b e t w e e n t he a n t e n n a I sock e t s ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 6; O L = 1 ; OOL = 7; b re a d t h of t h e oce l l i : 5 ; te m p l es i n v i s i b l e ; s c u t u m h a i r y , d u l l , g r a n u l a ted ; n o t a u l i ces d i s t i nc t a n d com p l e t e , poster i o r l y s e p a r a t e d ; m i n i m u m d i s t a n ce b e t ween t h e n o t a u J i ces n e a r l y a s l o n g a s a n te n n a I se g m e n t 2 ( 5 : 6 ) ; sc u te l l u m a s scu l p t u red a s scu t u m ; m e t a n o t u m w i t h a n te r i or t h i rd r u g os e ; m i d a n d pos t e r i o r t h i rd smoot h , s h i n y , w i t h o u t sc u l p t u r e ; p ropodeurn fu l l y ret i c u l a t e r u gose , w i t ho u t l o n g i t u d i n a l k ee l s o n t he poste r i or s u rface ; fore-w i n g h y a l i ne , w i t ho u t d a r k t r a n s ve rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n n ea r l y a s l o n g a s p rox i m a l p a r t ; dors a l p rocess o f gon oforceps ( F i g . 7 3 1 ) s h o r t , s l en d e r , i r reg u l a r a n d po i n ted , l oc a t e d on t h e d i s t a l h a l f of t he gonoforcep s ; m a x i l l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 s e g rn e n t s . LOCUS T Y P I C U S : Ass i n a n ( I n d o n e s i a ) T Y P I CA L �IA T [ R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( hereb y d es i g n a te d ) of Ph . j a va n u s i n B M ; h o l o t y pe F ! of Ps . thora cicus i n W A ; h o l o t y pe F ! a n d 4 p a r a t y p e s ( 2 F F , 2 M M ) ! o f N . l a w a n a e i n B M . HOST S : i n M a l a y a L a wa n a conspersa ( W a l k . ) ( Fl a ti da e ) a n d can dida F . ( Roepke 1 9 1 6 ) . D I STR I B U T I O N : O r i e n t a l reg i o n : I N D O I� E S I A : A s s i n a n ( Sa l a t i g a , J a v a ) , B M ! MALAYS I A : K u a l a L u m p u r ( M a l a y a ) , B M ! P H I L I P P I N E S : P u e r t o P r i ncesa ( P a l a w a n ) , VIA !

8 . Neodry in u s pseu dodiffu s u s n .

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from C h i a n g dao


1 089

red d i s h ; a n t e n n a e b r o w n o r redd i s h , w i t h s e g m e n t s 7 - 1 0 b r ow n ; t ho r a x a n d p ropodeum b l ac k , w i t h s i des a n d pos t e r i o r m a r g i n re d d i s h ; a b domen b l a c k - b r o w n ; l e g s b row n , w i t h t a r s i l i g h t ; h e a d we a k l y e x c a v a ted , s h i n y , w i t h f ro n s a n d v e r t e x f u l l y re t i c u l a t e r u gose ; occ i p u t g r a n u l a t ed , n o t ret i c u l a t e r u gose ; occ i p i t a l c a r i n a i n comp l e te , e x te n d i n g b e y o n d 0 , 5 l e n g t h of OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , h a i ry , w i t h c o l l a r smoo t h , w i t ho u t sc u l p t u re ; d i sc w i t h some l o n g i t u d i n a l kee l s ; poste r i o r edges of t h e p ro n o t u m ro u n d ed , n o t p rod uced i n t o l obes ; sc u t u m h a i r y , f u l l y re t i c u l a t e r u gose ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m ret i c u l a t e r u gose ; met a n o t u m v e r y r e d u c e d a n d s h or t ; p ropodeum f u l l y ret i c u l a te r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e poste r i o r s u rf ace ; fore-w i n g w i t h t w o b r o w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n m u c h l o n g e r t h a n p ro x i m a I p a r t ; fore t r oc h a n t e r s a p p ro x i m a te I y fo u r t i mes a s l o n g as b road ; s e g m e n t 1 of f ro n t t a rs u s l on g e r t h a n 4 ( 24 : 1 9 ) ; e n l a r g e d c l a w ( F i g . 732 ) w i t h s u b a p i c a l too t h a n d 1 5 peg - l i ke h a i r s ; seg men t 5 of f r o n t t a rs u s ( F i g . 732 ) w i t h t w o r o w s o f 3 + 7 l a me l l a e ; a p e x w i t h w g ro u p o f 5 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 segm e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MAL E : u n k n o w n lOCUS T Y P I C U S : C h i a n g d ao ( T h a i I a n d ) TYP I CAL MAT E R I A L : h o l o t y pe F ! i n B ; 2 p a r a t y p es F F ! i n WA ; 1 p a r a t y pe F!

i n OL.

HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : T HA I LAN D : C h i a n g d a o , 8 ! P H I L l P P I ;, E S : W a - w a D a m ( M t . M o n t a l b a n , R i z a l , L u z o n ) , O l ! M a n i l a ( L u zo n ) , WA ! SR I lANKA : P a l a t u p a n a ( H a m . D i s t . ) ( Ponom a r e n ko 1 9 8 1 b , u n der n a me ,Veodryin u s koebelei R . C . l . Perk i n s ) , WA ! U l h i t i y a O y a ( 1 5 m i . �m:=: of M a h i y a n g a n a , B a d . D i s t . ) , WA !

9 . Neodryin u s diffu s u s n . s p .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 68-5 , 62 m m ; h e a d b l ac k , w i t h m a n d i b l es redd i s h ; a n t e n n ae b l a c k - b row n ; t ho r a x , p ropodeum a n d p e t i o l e b l a ck , w i t h s i des of p r o n o t u m re d d i s h ; a b domen b l ac k , w i t h a p e x re d d i s h ; l e g s b ro w n , w i t h t a r s i l i g h t ; h e a d e x c a v a ted , s h i n y , w i t h f r o n s a n d v e r t e x f u l l y re t i c u l a t e r u g ose ; occ i p u t g r a n u l a ted , n o t re t i c u l a te r u gose ; occ i p i t a l c a r i n a i n comp l e t e , e x t e n d i n g b e y o n d 0 , 5 l e n g t h of O O l ; p ro n o t u m c rossed b y a t r a n s vers a l i m p ress i o n , h a i r y , w i t h co l l a r smoot h , w i t ho u t scu l p t u r e ; d i sc w i t h some l o n g i t u d i n a l k ee l s ; poste r i o r ed ges of t h e p ro no t u m rou n de d , n o t p ro d uced i n to l o b e s ; s c u t u m h a i r y , f u l l y re t i c u l a te r u gose ; n o t a u l i ces i n v i s i b l e ; scu t e ! l u m ret i c u l a t e r u gose ; m e t a n o t u m very reduced a n d s h or t ; p ropodeum f u l l y ret i c u l a t e r u gose , w i t h two l o n g i t u d i n a l k ee l s o n the pos t e r i o r s u r f ace ; fore-w i n g w i t h two brown t r a n s v e rs a l b a n d s ; d i s t a l part o f rad i a l v e i n m u c h l o n g er than p ro x i m a l p a r t ; fore t r oc h a n ters a p p ro x i m a t e l y fo u r t i mes a s l o n g a s b ro a d ; seg men t

1 090

Mem . Amer .

Ent.

I ns t . ,

n o . 37 ,

1 984

/1 ��------ ----� ----- �

Fig.

733

-

C he l a of Neodry in u s diffu s u s n .

sp .

( ty p e )

1 of fro n t t a rs u s l o n ger t h a n seg m e n t 4 ( 2 0 : 1 4 ) ; e n l a rged c l a w ( F i g . 733 ) w i t h s u b a p i c a l too t h a n d 1 3 peg- I i ke h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 733 ) w i t h t w o r o w s o f 9 + 9 l a me l l ae ; a p e x w i t h a g ro u p of 1 0 l a me l l a e ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n known L O C U S TYP I C U S : K h aoc h a n g ( T h a i l a n d ) T Y P I CA L �lAT E R I AL : h o l o t y p e F ! i n O L ; 1 p a r a t y p e F ! i n B . HOSTS : u n k n o w n D I ST R I BU T I O N : O r i e n t a l reg i o n : T HA I LAN D : K h aoch a n g ( T ra n g , K a ho p h a p p h a ) , O L ! MALAYS I A : Rej a n g D e l t a ( Sa r i k e i D i s t . , S a r a w a k ) ,

6!


1 09 1

I� E A R C T I C S P E C I E S O F NEODR YINUS

T h ree s p ec i es a re k n ow n .

K E Y TO T H E S P E C I E S

F EMALES

S c u t u m g ra n u l a t ed a n d p a r t l y r e t i c u l a t e ru g o s e , n o t s c u l p t u r ed by l on g i t u d i n a l k e e l s ; p r o t h o rax f u l l y o r p a r t l y b l a c k • 1 . typ h locybae ( A s h mead ) S c u t u m f u l l y s c u l p t u r ed by p a ra l l e l l o n g i t u d i n a l k e e l s ; p r o t h o ra x b l a c k o r f u l l y r ed d i sh 2 P r o t h o rax f u l l y r ed d i sh , w i t h c o l l a r a n d d i s c smoo t h a n d s h i n y , w i t h ou t s c u l p t u r e 2 . balli n . sp . P r o t h o ra x a l mo s t f u l l y b l a ck , w i t h c o l l a r a n d d i s c d u l l , g ra n u l a t e d a n d w i t h k e e l s a r o u n d t h e d i s c 3 . incaic u s n . sp . ( s e e N e o t r o p i c s p e c i e s ) •

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MALES

O n I y t h e ma I e of N.

typh locyb a e i s k n ow n .

1 . Neodryin u s typ h locyb a e ( As h m e a d ) n . com b .

L a beo typ h locyb a e A s h m e a d 1 893 : 89 . Dryin u s ormenidis A s h me a d 1 903 : 1 92 ( s y n . p roposed b y M u esebeck a n d W a l k l e y 1 95 1 ) ; locus typ i c u s S a n d u s k y ( O h i o , U . S. A . ) . Dryin u s n igrellus B r u e s 1 90 4 : 1 1 7 ( sy n . p roposed b y M u esebec k a n d Wa l k l e y 1 95 1 ) ; locus typicus P a r k v i l l e ( N . Y . , U . S . A . ) . Dryin u s ormenidis A s h mea d : R . C . L . Perk i n s 1 906a : 4 9 2 . Neodryin u s arizonic u s R . C . L . Perk i n s 1 90 7 : 38 ( n . s y n . ) ; locus typ ic u s Noga l e s ( A r i z . , U. S. A . ) . Neodryin u s d u b iosus R . C . L . Perk i n s 1 907.: 39 ( n . s y n . ) ; locus typ ic u s Nog a l e s ( A r i z . , U . S. A . ) . Dryinus ormenidis A s h mead : K i effer 1 90 7 b : 7 . Psilodryin u s n igrell u s ( B r u es ) : K i effer 1 907b : 8 . L a b eo typ h locyb a e A s h m e a d : K i effer 1 907b : 1 6 .

1 092

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Ent.

I n st . ,

no . 3 7 ,

1 984

F i g . 734 - C h e l a of Neodryin u s typ h locyb a e ( As h mead ) from Sa n d u sk y (Ohio)

Gon a topod i n ae

1 093

--- --------------

F i g . 735 - Ma l e gen i t a l i a of Neodryin u s typh locyb a e ( A shmead } ( l ec t o t y p e )

Prodryin u s ormenidis ( As h m e a d ) : K i effer 1 9 1 4b : 5 1 . Psilodryinus arizon icus ( R . C . L . Perk i n s ) : K i effer 1 9 1 4b : 5 5 . Psilodryinus dubiosus ( R . C . L . Perk i n s ) : K i effer 1 9 1 4b : 55 . L a b erius typhlocyb a e ( As h me a d ) : K i effer 1 9 1 4b : 65 . Gon a top u s typ h locy b a e ( As h me a d ) : B r u e s 1 9 1 6 : 6 1 4 . Psilodryin u s carin a t u s Fou t s 1 924 : 1 60 ( n . s y n . ) ; locus typicus C a r l i s l e ( P a . , U. S. A . ) . Psilodryin u s arizon icus ( R . C . L . Perk i n s ) : M u e sebeck a n d Wa l k l ey 1 95 1 : 1 03 5 . Psilodryinus carin a tus Fou t s : M u esebeck a n d Wa l k l e y 1 95 1 : 1 03 5 .

1 094


I nst . ,

n o . 37,

1 984

Psilodryin u s dubiosus ( R . C . l . P e rk i n s ) : Mu esebec k and W a l k l ey 1 9 5 1 : 1 03 5 . Psilodryin u s typ h locyb a e ( As h m ea d ) : M u e sebeck a n d W a I k l ey 1 95 1 : 1 0 35 . Neodryin u s ormenidis ( As h m e a d ) : R i c h a rd s 1 95 3 : 68 . Neodryin u s (Prodryin u s ) arizon ic u s R . C . l . Perk i n s : Krombe i n 1 97 9 : 1 244 . Neodryin u s (Prodryin u s ) c a rin a t u s ( Fou t s ) : K romb e i n 1 979 : 1 244 . Neodryin u s (Prodryin u s ) du b io s u s R . C . l . Perk i n s : Krom b e i n 1 979 : 1 244 . Neodryin u s (Prodryin u s ) typ h locyb a e ( As h me a d ) : Krom b e i n 1 97 9 : 1 24 4 .

F E M A L E : fu l l y w i n g e d ; l e n g t h 4 , 00-5 , 1 2 m m ; h e a d b l ac k o r b row n , w i t h c l y p e u s a n d m a n d i b l es y e l l o w ; a n te n n a e u s u a l l y y e l l o w , w i t h segmen t s 6-8 b row n , occas i o n a l l y fu l l y y e l l ow ; p r o n o t u m b l ac k , w i t h s i de s a n d poster i o r m a r g i n red d i sh ; mesot h o ra x , m e t a t h orax a n d propodeum b l ac k ; abdomen b ro w n - b l ac k ; l e g s u s u a l l y t e s t a ceou s , occ a s i on a l l y w i t h cox a e , c l u b s o f femora a n d t i b i a e d a r k e n e d o r b l ac k i sh ; h e a d a l mo s t f l a t , s h i n y , g r a n u l a t e d a n d w e a k l y rug ose ; occ i p i t a l c a r i n a w e a k l y v i s i b l e on t he s i des of the oce l l i , not e x t e n d i n g b e y o n d 0 , 5 l e n g t h o f OOl ; p r o n o t u m c rossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , g r a n u l a t e d , w e a k l y s t r i a t e ; pos t e r i o r e d g e s of p ro n o t u m rou n ded , n o t p rodu ced i n t o l obes ; sc u t u m s h i n y , g r a n u l a t e d , w i t h t r a c k s o f a reo l ae , m a i n l y a l o n g i n v i s i b l e n o t a u l i ces ; sc u te l l u m sh i n y , w i t h a n t e r i or h a l f g r a n u l a ted a n d poster i or h a l f smooth a n d w i t h o u t sc u l p t u re ; m e t a n o t u m very reduced a n d short ; p ropodeum fu l l y ret i c u l a t e r u gose , w i t h l o n g i t u d i n a l kee l s on poster i o r s u rf a ce ; i n s o m e spec i me n s a t r a n svers a l f o l d i s v i s i b l e b e t ween a n t er i or a n d p o s t e r i o r s u rf a c e of p ropodeu m ; fore-w i n g w i t h two b ro w n t r a n s versa l b a n d s ; d i s t a l p a r t of ra d i a l v e i n as l o n g a s , or s l i g h t l y l o n ger t h a n prox i m a l p a r t ; fore troch a n ters a p p rox i m a te l y f o u r t i mes as l o n g as b road ; seg m e n t 1 of fro n t tarsus l on g e r than segme n t 4 ( 20 : 1 1 ) ; en l arged c l a w ( F i g . 734 ) w i t h s u b a p i c a l tooth a n d 4 b r i s t l e s ; seg men t 5 of fron t t a rs u s ( F i g . 7 3 4 ) w i t h 1 row of 23 l a m e l l a e ; a p ex w i t h a group of n ea r l y 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MAL E : f u l l y w i n g e d ; l e n g t h 2 , 8 1 m m ; head b l a c k - b row n , w i t h m a n d i b l es y e l l o w ; a n ten n ae b ro w n ; t h o r a x a n d p ropodeum b l ac k ; a b domen b ro w n t e s t a ceo u s ; l e g s t e s t aceou s ; a n t e n n a l seg men t s i n f o l l ow i n g proport i on s : 7 : 6 : 1 0 : 8 : 8 : 8 : 7 : 7 : 7 : 1 0 ; head sh i n y , g ra n u l a t ed , w i t h f r o n s a n d vertex re t i c u l a t e ru gose , w i t h a med i a n kee l b e t w een t jo e a n te n n a l sock e t s ; POL = 8 ; O L = 3 ; OOl = 4 ; b read th o f t h e oce l l i : 4 ; occ i p i t a l c a r i n a i n v i s i b l e ; te m p l es i n v i s i b l e ; oce l l a r t r i a n g l e w i t h b a s e v ery l o n g ; s c u t u m d u l l , g ra n u l a t e d ; n o t a u l i ces i n comp l e t e , reach i n g a p p rox i ma te l y 0 , 75 l e n g t h o f scu t u m ; scu te l l u m a n d m e t a n o t u m g r a n u l a t e d ; p ropodeum ret i c u l a te rugose, w i t hou t l o n g i t u d i n a l k ee l s o n pos ter i or

1 09 5

Gona topod i n a e

s u rf ac e ; fore-w i n g h y a l i n e ,

w i t h o u t d a rk t r a n s v ers a l b a n d s ;

d i s t a l part

o f ra d i a l v e i n n e a r l y as l o n g as p rox i m a l p a r t ; dor s a l p rocess of gonoforceps ( F i g . 735 ) shor t , n a rrow a n d po i n ted ; t i b ; a I s p u rs 1 , 1 , 2 . L O C U S T Y P I C US : Wa s h i n g ton ( D . C . , U . s . A . ) T Y P I CAL M A T E R I A L : l ec t o t y p e M ! a n d 1 p a r a l ec to t y p e M ! ( h ereby des i g n a ted ) of L. typ h locyb a e in WA ; l ec to t y p e F ! and 2 para l ec t o t y pes F F ! ( h e r e b y des i g n a t ed ) of D . ormenidis i n WA ; l ec t o t y pe F ! ( d es i g n a ted b y O l m ; 1 982 ) of N. arizon icus in B ; h o l o t y p e F ! of N. dubiosus in B ; h o l o t y pe F ! of Ps . carin a t u s i n WA ; h o l o t y p e F ! of D . n igrellus i n A M . H O S T S : i n U . S . A . Metca lfa (� Ormenis) pruinosa ( Sa y ) a n d Ormenis septentrionalis ( Sp i n . ) ; Typ hlocy b a s p . a I so accord i ng to A s h m e a d 1 893 ( i nform a t i o n p ro b a b l y w ron g :

the

l ec t o t y p e i n fact

is l a b e l l ed as f o l l ow s :

" P a r : o n Homop t e r a o n C e l t i s + E l m , J u l y 23 . 83" ; n o h o s t i s g l u ed toge t h er w i t h the t y pe spec i men ; the i n for m a t i on so is o n l y in o r i g i n a l desc r i p t i on a n d not o n l a b e l s ) . PARAS I T E S : i n U . S . A . Cheiloneurus swezey; A s h m e a d ( Sw ez e y 1 903 ; R . C . L . Perk i n s 1 906b ) . D I S T R I B U T I O N : Nearc t i c reg i o n : CANADA : V i n e l a n d ( O n t . ) , a T ! S h e t l a n d ( On t . ) , O T ! U . S . A . : Nog a l es ( Ar i z . ) , B ! K i r k w ood ( Mo n t . ) , WA ! Koppe ' s B r i d g e ( We l l bo r n , Brazos Co . , T ex . ) , T E ! H e a r n e ( Rober t so n Co . , T ex . ) , OL ! Mercedes ( Tex . ) , WA ! C M ! W i c h i t a ( K a n s . ) , OL ! A n n Arbor ( M i ch . ) , BM ! I ow a ( M uesebeck a n d W a l k l ey 1 95 1 ) ; U r b a n a ( I I I . ) , WA ! Terrace P a r k ( Oh i o ) , B ! C o l u m b u s ( O h i o ) , B ! S a n d u s k y ( Oh i o ) , WA ! B ! Ben t on v i l l e ( Ark . ) , WA ! A i k en ( S . C . ) , O l ! a T ! Wash i n g t on ( D . C . ) , WA ! C a r l i s l e ( P a . ) , WA ! N . C u m b er l a n d ( P a . ) , Ol ! Moores t o w n ( N . J . ) , T W ! I n d l ow v i l l e ( N . Y . ) , I T ! N e w b u r g h ( N . Y . ) , ZO ! S i x M i l e C reek ( I t h a c a , N . Y . ) , O T ! Pou g h k e� p s i e ( N . Y . ) , O L ! T W ! P a r k v i l l e ( lo n g I s l a n d , N . Y . ) , A M ! Sou t h H a ven ( lo n g I s l a n d , N . Y . ) , OL ! H a rbor H i l l ( N . Y . ) , C M !

2 . Neodryin u s b a l l i n .

sp .

F E M A L E : f u l l y w i n ged ; l en g t h 4 m m ; h e a d b l ac k , w i t h c l y p e u s a n d m a n d i b l es t e s t aceo u s ; a n t e n n ae t e s t aceou s ; p ro t horax redd i s h ; meso t h o r a x , m e t a t h orax a n d p ropodeu m b l ac k ; a b domen b l ac k ; fore l e g s t e s t a ceou s ; m i d a n d h i n d l e g s b row n , w i t h t a r s i l i g h t ; a n te n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I segmen t s i n fo l l ow i n g p roport i on s : 1 4 : 6 : 24 : 1 1 : 1 0 : 7 : ( l a s t four a n t en n a l

seg m e n t s m i s s i n g

i n t h e on l y k n o w n spec i men ) ;

head f l a t ,

w i t h oce l l a r reg i o n s w o l l en , d u l l , g r a n u l a t e d a n d f u l l y ret i c u l a t e rugose ; frons w i t h a m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 4 ; OL = 2 ; OOL = 1 0 , 5 ; p ro n o t u m sh i n y , smoo t h , w i t ho u t p o s t e r i or l obes d i rected t o w a r d s the t eg u l ae , w i t h a s t r o n g a n t e r i o r t r a n sv e rs a l i m p res s i on ; d i sc h u mped ; p ron o t u m w i t h o u t scu l p t u re , w i t h on l y w e a k s t r i ae a r o u n d t h e d i sc ; scu t u m d u l l , f u l l y sc u l p t u red b y l o n g i t u d i n a l p a ra l l e l k ee l s ; n ot a u l i ces i n v i s i b l e ; sc u te l l u m sh i n y , w i t h p os t e r i o r h a l f smoo t h a n d w i t hou t scu l p t u re ; a n t e r i o r h a l f p a r t l y ret i c u l a t e ru gose a n d w ea k l y

1 096

Mem . A m e r . E n t .

I n st . ,

n o . 37,

1 984

F i g . 736 - C h e l a of Neodryinus b a ll i n . sp .

( ty p e )

1 097


g r a n u l a ted ; m e t a n o t u m very reduced and f u l l y re t i c u l a t e ru gose ; p ropodeum f u l l y ret i c u l a t e ru gose a n d w i t h t w o i n comp l e t e l o n g i t u d i n a l kee l s o n poster i or s u rf ace ; fore-w i n g w i t h t w o d a r k t ra n sversa I b a n d s ; d i s t a l p a r t o f ra d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ; fore t a r s a l segmen t s i n fo l l ow i n g p roport i on s : 1 8 : 3 : 8 : 1 7 : 3 2 ; en l arged c l a w ( F i g . 7 3 6 ) w i t h s u b a p i c a l t o o t h a n d 6 b r i s t l es ; segmen t 5 o f f r o n t t a r s u s ( F i g . 7 3 6 ) w i t h 1 ro w of n e a r l y 2 2 l am e l l a e ; apex w i t h a g rou p of a t l ea s t 30 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n k nown L O C U S T Y P I C U S : T i n aj a s A l t a s ( Ar i z . , U . S . A . ) TYP I CA L M A T E R I AL : h o l o t y p e F ! i n WA . HOSTS : u n k n o w n D I ST R I B U T I ON : Nearc t i c reg i on : U . S . A . : T i n a j a s A l t a s ( Ar i z . ) , NOT E S : t h e s p ec i es i s n a med i n h onor of M r . E . D . B a l l .

WA !

N E O T R O P I C S P E C I E S OF NEODR YINUS

N i n e s p ec i es a re k n o w n .


F E I\IA L E S

S eg m e n t

2

3

4

5

5 of

f r on t

t a r s u s a l mo s t

t w i c e as

l on g a s

en l a rg ed

c I a w ( F i g . 7 3 7 ) • . . . • 1 . a c u t i co 11 i s ( K i e f f e r ) S eg m e n t 5 o f f r on t t a r s u s s l i g h t l y l o n g e r t h a n en l a rg ed c l a w ( F i g s 739 , 740 , 742 , 743, 744, 745, 746 , 747 ) . . . . . . . . . . . . . . . . 2 F o r e -w i n g w i t h ra d i a l v e i n s t r o n g l y S - s h a p ed ( F i g . 7 3 8 B , D , E ; F i g . 741 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .3 F o r e - w i n g w i t h ra d i a l v e i n n o t o r w e a k l y S - s h a p e d ( F i g . 7 3 8 A , C) • • • • • . • • • • • • • • • • • • • • • • • . • • • • . • . • . . • • • • . . . . . • . . • • • • • . • . • . • • • 7 D i s t a l p a r t o f ra d i a l v e i n l on g e r t h a n p r o x i ma l p a r t ( F i g . 7 3 8 D) • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • . • • . • • • • • . • • .4 D i s t a l p a r t o f ra d i a l v e i n a s l on g a s o r sh o r t e r t h a n p r ox i ma l pa r t ( F i g . 738 B , E ; F i g . 7 4 1 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 H ea d a n d s c u t u m g ra n u l a t ed , n o t r e t i c u l a t e r u g o s e ; b od y r ed d i sh - t e s t a c e ou s , w i t h i r r e g u l a r b r own s p o t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . dom inicd )l I J ,-, ( E v a n s ) H ea d a n d s c u t u m g r a n u l a t ed a n d r e t i c u l a t e r u g o s e ; b od y b l a c k . . . . . . . . . . . . . . . . . . . . . . 3 . albosign a tu s n . sp . D i s t a l p a r t o f ra d i a l v e i n a s l on g a s p r ox i ma l pa r t ( F i g . 7 3 8

Mem . A m e r . E n ! .

1 098

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(Fig.

747 )

( F i gs 744 , . .8 745 ) S c u t u m f u l l y w e a k l y r e t i c u l a t e r u g o s e , w i t h ou t l o n g i t u d i n a l 6 . m a c u licornis ( C a m e r o n ) . k ee I s S c u t u m w i t h n u me r o u s I ong i t u d i n a l k e e l s , p a r t I y w i t h s i d e s s t r o n g l y r e t i cu l a t e rug o s e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . · .. .. . . . . . . .. . .. . .. . . . .. . .. .. 7 . i n c a i c u s n . sp . •

8

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no.

D i s t a l pa r t of 738 B, 741 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . P r on o t u m w i t h d i s c mo r e p r o t r u d i n g ( F i g . 7 4 1 B ) . . . . . . . . . . . . . . . . . . . . . 4 . r a di a l i s n . s p . P r on o t u m w i t h d i s c l e s s p r o t r u d i n g ( F i g . 7 4 1 A ) . 8 . bra c hycerus ( K i e f f e r ) S eg m e n t 5 o f f r on t t a r s u s w i t h t w o r o w s o f l a m e l . . 9 . trin i t a tis R i c h a rd s r ow o f l a me l l a e S e g m e n t 5 o f f r on t t a r su s w i t h .

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MALES

O n l y t h e m a l e of N.

trin i t a tis R i c h a r d s

i s k n ow n .

1 . Neo dryin u s a c u ticollis ( K i ef fe r )

Dryi n u s a c u ticollis K i effer 1 90 5 b : Neodryin u s

e t Marsha I I

9.

(Psilodryi n u s ) a c u ticollis

( K i effe r ) :

K i effer i n K i ef f e r

1 906 : 497 .

Neodryin u s a c u ticollis

( K i effe r ) : R i ch a r d s 1 953 : 6 2 . ( K i effe r ) : O l m i a n d C u r rado 1 976a :

Psilodryi n u s a c u ticollis

1 85 .

F E i' sp . or

n o t a u l i ces s h o r t e r t h a n b re a d t h of t h e oce l l i ( 2 : 3 ) ; s c u t e l l u m a n d met a n o t u m s h i n y , h a i r y , p u n c t a t e , smoo t h ; p ropodeum re t i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t h o u t d a r k t ra n s ver s a l b a n d s ; dors a l p rocess of gonoforceps l o n g , b r o a d , w i t ; , a p e x rou n d e d ( m a l e A : f i g . 840 ) or po i n ted ( m a l e B : f i g . 841 ) ; m a x i I l a ;' y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 .

3 . Pseu dogon a top u s n u du s R . C . L . P e r k i n s

Ps e u dogona topus n u dus R . C . L . P e r k i n s 1 9 1 2 :

12.

F E MAL E : a p terou s ; l en g t h 2 , 74 m m ; red d i s h - t e s t aceo u s , w i t h pet i o l e b l ac k a n d a b domen b ro w n - re d d i s h ; a n t e n n a e b r ow n , w i t h seg men t s 1 -2 y e l l ow ; b o d y h a i r l es s , s h i n y , g r a n u l a t ed ; h e a d e x c a v a ted ; p ro n o t u m c rossed b y a s t ro n g t r a n s v e rs a l i m p re s s i o n ; m e t a n o t u m w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s rou n d e d ; meso-me t a p l e u r a l s u t u re o b so l e t e ; met a t ho r a x + p ropod e u m w i t h o u t med i a n f u rrow ; mesop l e u r a , met a p l e u r a a n d pos t e r i o r s u rface o f p ropodeum t r a n s verse l y s t r i a t e ; se g m e n t 4 of f ro n t t a r s u s a s l o n g a s segmen t 1 ; e n l a r g e d c l a w ( F i g . 842 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l a e ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 842 ) w i t h 2 rows of 1 4 l a m e l l ae ; apex w i t h a g r o u p o f 6 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 246


I n st . ,

no.

37,

1 984

F i g . 842 - C he l a of Pseudogo n a topus nudus R . C . L . Perk i n s ( t y p e )

M AL E : u n k n o w n L O C U S T Y P I C U S : P ek a l o n g a n ( J a v a , I n d o n es i a ) T Y P I CA L M A T E R I A L : l ec t o t y p e F ! ( d es i g n a ted b y O l m i 1 982 ) i n B . HOST S : i n I n dones i a Dicran otropis s p . ( R . C . L . Perk i n s 1 980 ) ; i n t h e P h i l i p p i nes Soga tella furcifera ( Ho rv a t h ) ( C h a n d r a 1 980b ; I . R . R . I . E n to mo l o g y D e p a r t m e n t 1 980 ) , Nil aparvata l u gens ( St a l ) , Stenocra n u s s p .

1 247

Gon a topod i n ae

F i g . 843

-

C he l a o f Pseu dogon a top u s ma lesiae n .

sp .

( type)

1 248

Mem . A m e r . E n ! .

I ns ! . ,

n o . 37 ,

1 984

( C h a n d r a 1 980b ) . Accord i n g to C h a n d r a ( 1 980b ) i n t he P h i l i p p i nes occ a s i o n a l l y Ps . n u dus c a n p a r a s i t i ze a l so C i ca de l l i d s : Nephotettix n igropictus ( S t a l ) , Nephotettix virescens ( D i s t a n t ) and Recilia dors a lis ( Mo t s c h u l s k y ) . T h i s l a s t i n forma t i o n s m u s t b e c h eck e d . D I ST R I BU T I O N : O r i e n t a l reg i on : I N D O N E S I A : P ek a l on g a n ( J a v a ) , B ! P H I L I P P I N E S ( C h a n d r a 1 980b ; I . R . R . I . E n tomo l o g y D e p a r t m e n t 1 980 ) .

4 . Pseu dogon a top u s m a lesiae n . sp .

F E M A L E : a p t e ro u s ; l e n g t h 3 , 0 m m ; t e s t a c eo u s , w i t h p e t i o l e b l ac k ; a n t e n n a e t e s t aceou s , w i t h segmen t 1 0 w h i t i s h ; head v e r y e xc a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re , h a i r l ess ; p rono t u m crossed b y a s t ro n g t ra n s v ersa l i m p ress i o n , smoo t h , h a i r l es s , s h i n y ; s c u t u m a n d s c u t e l l u m s h i n y , smoo t h a n d w i t h o u t sc u l p t u re ; m e t a n o t u m smoo t h , s h i n y , w i t ho u t sc u l p t u re , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s po i n ted ; me t a t ho r a x + p ropode u m s h i n y , smoo t h , w i t h o u t sc u l p t u re , except for n u me ro u s t r a n s v e rs a l s t r i a e o n pos t e r i o r s u rface ; p l eu r a w i t ho u t t ra n sv e rs a l k ee l s ; meso- me t a p l eu r a l s u t u re d i s t i nc t a n d comp l e t e ; segmen t 1 of fron t t a rs u s s ho r t e r t h a n segmen t 4 ( 1 3 : 1 6 ) ; en l a r g e d c l a w ( F i g . 843 ) w i t h s U b a p i c a l tooth a n d 7 l a me l l ae ; seg m e n t 5 of fron t t a rs u s ( F i g . 843 ) w i t h 2 row s of 1 9 l a me l l a e ; a p e x w i t h a g ro u p of 1 5 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n ( see notes o n p a g e 1 24 1 ) . L O C U S T Y P I C U S : G u no n g M u l u ( Sa r a w a k , M a l a y s i a ) T Y P I CAL M A T E R I AL : ho l o t y p e F ! i n BM . HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i on : M A LAYS I A : G u n o n g M u l u ( Sa r a w a k ) , BM !

5 . Pseu dogon a top u s validus n . sp .

F E MA L E : a p terou s ; l e n g t h 2 , 5 m m ; head t e s t aceo u s-redd i s h or b ro w n redd i s h , w i t h m a n d i b l e s , c l y p e u s a n d fron t p a r t of v e r t e x y e l l o w ; a n te n n a e b row n , w i t h s e g m e n t s 1 -2 y e l l o w ; t horax a n d p ropod e u m red d i s h or b ro w n-re d d i s h ; p e t i o l e b l ac k ; a b domen b ro w n-red d i s h ; l eg s y e l l ow ; head ( F i g . 844 C ) exca v a t e d , smoo t h , s h i n y , w i t ho u t sc u l p t u re ; p ron o t u m ( F i g . 8 4 4 B ) crossed b y a s t ro n g t r a n svers a l i m p ress i o n , smoo t h , s h i n y , w i t ho u t scu l p t u re ; sc u t u m a n d sc u te l l u m smoo t h , s h i n y , w i t h o u t scu l p t u re ; m e t a n o t u m n o t t r a n s verse I y s t r i a t e , w i t h s i des p r o t r u d i n g ; p ro t r u s i on s p o i n ted ; meso-me t a p l eu r a l s u t u re o n l y d i s t i nc t on pos t e r i o r h a l f ; m e t a t ho r a x + p ropodeum ( F i g . 844 A ) smoo t h , s h i n y , w i t ho u t sc u l p t u re ; mesop l e u r a , met a p l eu r a a n d pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; segmen t 1 of fro n t t a r s u s a s l o n g a s segme n t 4 ; en l arged c l a w ( F i g . 845 ) w i t h s u b a p i c a l t oo t h a n d 5 l a m e l l ae ; segmen t 5 of f ro n t t a rs u s

1 249

Gon a topod i n a e

c

Fig.

844 - F em a l e of Pse u dogon a top us validus n . s p . t h o r a x a n d p ropode u m ; C : he a d .

( type) : A , B :

( F i g . 845 ) w i t h t w o rows of 1 3 l a me l l a e ; apex w i t h a group of 9 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I C U S : Spooner R o a d ( M a l a y a , M a l a y s i a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n BM ; 1 p a r a t y p e F ! i n OL . HOST S : u n k n o w n D I ST R I B U T I O N : O r i en t a l reg i o n : SR I L A N KA : Y a k k a l a , O L ! MALAYS I A : Spooner R o a d ( K u a l a L u m p u r , M a l a y a ) , BM !

6 . Pseu dogon a top us l a n k a e P o n o m a r e n k o

Pseu dogon a top us lankae P o n o m a r e n k o 1 98 1 b : 8 8 1 .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 32 m m ; r e d d i s h - b r o w n , w i t h a n t e n n a e tes t aceou s ; l eg s w i t h t roc h a n ters a n d fore c o x a e p a r t l y w h i t i s h ; h i n d s i d e of h i n d femora w h i t i s h ; a p e x of p ropodeum w h i t i s h ; h e a d d u l l , a l u t aceou s ; o c c i p i t a l c a r i n a i n v i s i b l e ; f ro n s w i t h a com p l e t e med i a n k ee l ; POL = 2 ; OL = 3 ; OOL = 1 0 ; head e x c a v a ted ; temp l es p rom i n e n t ; a n ten n a e d i s t a l l y t h i c k e n e d ; a n t e n n a I segmen t s i n fo l l o w i n g p ro po r t i o n s : 1 4 : 8 : 1 4 : 9 : 9 : 6 : 6 : 6 : 6 : 1 1 ; p r o n o t u m c rossed b y a s t ro n g t ra n s v ersa l i m p ress i o n , s h i n y , smoo t h , w i t ho u t scu l p t u r e ; scu t u m smoo t h , s h i n y , w i t ho u t scu l p t u r e , w i t h som" t r a n s v e r s a l kee l s o n s i des ; s c u te l l u m s h i n y , smoo t h , w i t ho u t scu l p t u re ; t ho r a x h a i r l es s ; met a n o t u m f i a t , wea k l y ho l l o w


1 250

F i g . 845

-

I ns! . ,

no.

37 ,

1 984

C h e l a of Pseudogon a top u s v a l idus n . sp .

( t y pe )

1 25 1

Gona topod i n ae

F i g . 846 - T h o r a x a n d p ropodeum o f fem a l es o f Pseudogona topus mUlrl n . s p . ( t y pe ) ( A ) a n d l a n k a e Ponomarenko ( t y p e ) ( B )

b e h i n d t h e s c u te l l u m , smoot h , s h i n y , w i t h o u t scu l p t u re , w i t h s i de s protru d i n g ; p ro t r u s i o n s rou n ded ; meso-me t a p ! e u r a l s u t u re d i s t i nc t a n d comp l e te ; m e t a t horax + propode u m ( F i g . 8 4 6 B ) smoot h , s h i n y , w i t h o u t sc u l p t u re , excep t for n u merous t r a n s vers a l s t r i a e on poster i or s u rf ace of p ropode u m ; p l e u r a w i t h o u t t r a n s versa l s t r i a e ; d i sc of m e t a t horax + propodeum w i t h a s t ro n g med i a n f u r row ; fore t a r s a l s e g me n t s i n fo l l ow i n g proport i on s : 1 5 : 4 : 6 : 20 : 3 1 ; en l arged c l a w ( F i g . 847 ) w i t h s U b a p i c a l too t h a n d 8 l ame l l a e ; segment 5 of fron t t a rs u s ( F i g . 847 ) w i t h t w o row s of 1 1 + 1 0 l ame l l ae ; a p e x w i t h a g ro u p of n e a r l y 20 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS TYP I CUS : G i l i m a l e ( S r i Lan k a ) TYP I CAL M A T E R I A L : h o l o t y p e F ! i n W A . HOST S : u n k no w n D I ST R I B U T I O N : O r i en t a l reg i on : SR I LAN K A : G i l i m a l e ( I n d u r u w a J u n g l e , R a t . D i s t . ) , WA ! W a t u r a w a Forest ( 5 i n h a r a j a J u n g l e , R a t . D i s t . ) , OL ! N OT E S : Ps . l a n k a e i s very s i m i l a r to Ps . ma lesia e : t h e on l y d i fference i s t h e a b sence o f med i a n f u rrow o n m e t a t horax + propo d e u m .

7 . Pseu dogon a top u s m u iri n .

sp .

F E M AL E : a p t e ro u s ; l e n g t h 2 , 5 m m ; red d i s h - t e s t a ceou s , w i t h pe t i o l e b l ac k a n d a b domen b ro w n -red d i s h ; a n te n n a e b row n , w i t h segmen t s 1 -2 y e l l ow ; head du I I , e x c a v a ted , weak I y g r a n u I a ted ; p ro n o t u m crossed b y a s t ro n g t r a n s v e r s a l i m p ress i o n , s h i n y , w i t h o u t sc u l p t u r e ; m e t a n o t u m s h i n y , w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s rou n ded ( F i g . 846 A ) ; meso-met a p l eu r a l s u t u re w e a k l y d i s t i nc t ; p o s t e r i o r s u rface of p ropod e u m t r a n s verse l y

Mem . Amer.

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F i g . 847 - C h e l a of Pseu dogon a top u s l a n k a e P o n o m a r e n k o ( t y pe )

striate;

segme n t 1 of fron t t a r s u s as l o n g as s e g m e n t 4 ; en l a r ged c l a w ( F i g . 848 ) w i t h s u b a p i c a l too t h a n d 6-7 l a m e l l ae ; seg m e n t 5 of fro n t

1 2 53

Go n a t o pod i n a e

F i g . 848

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C h e l a of Pse u dogoll a top u s m u iri n . s p .

( type )

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n o . 37 ,

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F i g . 849 - C h e l a of Pseudogo n a topus soga t a e R o h w e r ( t y p e )

t a r s u s ( F i g . 848 ) w i t h 2 rows o f 1 5- 1 7 l a m e l l ae ; a p e x w i t h a g ro u p of 1 1 - 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I CU S : Pek a l on g a n ( J av a , I n do n es i a ) TYP I CA L M A T E R I A L : ho l ot y pe F ! i n OL . HOST S : u n k n o w n D I S T R I B UT I O N : O r i e n t a l reg i on : I N D O N E S I A : Pek a l on g a n ( Ja v a ) , OL ! N O T E : t h e spec i es i s n a m e d i n honor of M r . F . M u i r .

8 . Pseu dogona topus hospes R . C . L .

Perk i n s

See A u s t ra l i a n spec i es

9 . Pseudogo n a topus soga t a e R o h w e r n . eme n d .

Pseu dogon a top u s soga tea Rohwer 1 920 : 1 60 ( soga tea i s p r i n t i n g -error : t h e t y p e i s l a b e l l ed soga tae ) .

1 25 5

Gona topod i n a e

F E MA L E : a p terou s ; l e n g t h 3 , 0 m m ; head b row n , w i t h c l y pe u s a n d m a n d i b l e s y e l l ow ; a n t e n n a e b row n , w i t h segmen t 1 y e l l ow a n d 2 - 3 l i g h t ; t horax a n d p ropodeum b ro w n , w i t h sc u t u m redd i s h a n d a p e x of p ropodeum y e l l o w ; p e t i o l e b l ac k ; a b domen b ro w n - d a r k ; l eg s y e l l ow - b row n ; b o d y ha i r l es s ; h e a d e xc a v a ted , s h i n y , g r a n u l a ted ; p ro n o t u m crossed b y a s t r o n g t r a n sversa l i m p ress i on , weak l y g r a n u l a ted ; s c u t u m d u l l , g r a n u l a ted ; m e t a n o t u m f l a t , t r a n sverse l y s t r i a t e , w i t h s i des ro u n ded ; m e t a t h o r a x + p ropodeum d u l l , as scu l p t u red as sc u t u m ; mesop l e u r a , m e t a p l e u r a a n d poste r i or s u rface of p ropodeum t r a n sverse l y s t r i a te ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 of fro n t t a r s u s s l i g h t l y l on ge r t h a n s e g me n t 4 ( 1 0 : 8 ) ; en l arged c l a w ( F i g . 849 ) w i t h s u b a p i ca l too t h a n d 4 l a me l l a e ; seg m e n t 5 of fro n t t a r s u s ( F i g . 849 ) w i t h 2 ro w s of 23 l a me l l ae exten d i n g con t i n uo u s l y to the a p e x ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k n o w n ( t h e m a l e a l l ot y p e desc r i b e d b y Rohwer 1 920 i s rea l l y a m a l e of Haplogon a top u s orien t a l i s Rohwer : i t h a s m a x i I l a r y p a l p i w i t h 2 seg m e n t s a n d l a b i a l p a l p i w i t h 1 segme n t ) . LOCUS TYP I CU S : Pusa ( I n d i a ) T Y P I CAL MAT E R I AL : ho l o t y p e F ! i n WA ( to g e t h e r w i t h t h e a l l o t y p e M ! n o t b e l o n g i n g to t h i s s p ec i es ) . HOST S : i n I nd i a Soga ta s p . ( Ro h w e r 1 920 ) . D I ST R I BU T I O N : O r i en t a l reg i on : I N D I A : P u s a ( B i h a r ) , WA ! C u t t ack ( O r i ss,a ) ( K u r i a n 1 954b ) .

1 0 . Pseu dogon a top u s p u s a n u s n .

F E MA L E :

a p terou s ;

vertex b row n ;

l en g t h 3 , 0-3 , 5 m m ;

abdomen b r ow n ;

sp .

test aceous-red d i s h ;

head w i t h

a n t e n n a I segmen ts 3- 1 0 b row n ;

s i des of

sc u t u m a n d m e t a n o t u m d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p roport i on s : 8 : 7 : 1 5 : 8 : 7 : 7 : 7 : 7 : 6 : 9 ; head exc a v a t e d , d u l l , g r a n u l a t e d ; frons w i t h a com p l et e med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 3 ; O L � 4 ; OOL � 9 ; p ron o t u m crossed b y a s t ro n g t r a n sv e r s a l i m p ress i on , d u l l , g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted ; met a n o t u m f l a t , t r a n s verse l y s t r i a t e , w i t h s i des r o u n d e d ; m e t a t horax + p ro p o d e u m d u l l , g r a n u l a ted , w i t h poster i or s u rface a n d p l e u ra t r a n sverse l y s t r i a t e ; meso-meta p l e u r a l s u t u re obso l e te ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropor t i on s : 1 9 : 2 : 6 : 1 4 : 23 ; e n l a rged c l a w ( F i g . 850 ) w i t h s U b a p i c a l tooth a n d 4 l a me l l a e ; s e g m e n t 5 of fron t t a rs u s ( F i g . 850 ) w i t h 3 rows of 1 2 l a me l l ae ; a pex w i t h a g ro u p of 6 l a me l l a e ; m a x i I l a r y p a l p i w i t h 3 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I CU S : Pusa ( I n d i a ) T Y P I C A L M A T e R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n N O . HOS T S :

u n k no w n

M e m . Amer . E n t .

1 256

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F i g . 850

-

I nst. ,

n o . 37 ,

1 984

I ND I A : Pusa ( B i h a r , NO !

C h e l a of Pseudogona top u s p u s a n us n . s p .

( p ar a t y p e )

Go n a topod i n a e

1 257

NEARCT I C SPEC I E S O F PSEUDOGONA TOPUS

E i g h t s p ec i es are k n ow n .

KEY T O T H E SPEC I ES

F EMALES

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B o d y r ed d i s h ; a n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m mo r e i n c l i n ed ( F i g . 8 5 5 B ) ; s e gmen t 4 o f f r o n t t a r s u s a s l on g a s o r s l i g h t I y s h o r t e r t h a n s eg m e n t 1 ( mo r e t h a n 8 5 % ) . . . . . . . . . . . . . . • • • . . . • • . . . . • • • • • . . • • • • . . • • • • • 3 . sjoes tedti ( K i e f f e r ) B od y y e l l o w - r e d d i s h ; a n t e r i o r su r f a c e o f me t a t h o r a x + p r op o d e u m l e s s t a r s u s sh o r t e r

5

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. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . freytagi n e w n a me Me s o - me t a p l eu ra l su t u r e ob s o l e t e . . . . . . • • . • • . • . . . . • • . • • • . • • • . . 2 T h o r a x a n d p r op od e u m f u l l y b l a c k ( a t m o s t a p ex o f p r o p od eu m I i g h t ) • • • • • • • • . • • • • • • • • • • • • • • • 2 . c a n a densis n . s p . T h o r a x a n d p r op od e u m f u I I Y o r p a r t I y r edd i s h o r t e s t a c e ou s . • . 3 B o d y y e l l ow - t e s t a c e ou s o r r e d d i s h , w i t h p e t i o l e b l a c k ; o c c a s i on a l l y a b d omen b r own o r p a r t l y b l a c k ; o c c a s i on a l l y h e a d w i t h v e r t ex b r own ; o c c a s i on a l l y a n t e n n a e p a r t l y b r own o r b l a c k

8 5 8 8 ) ; s eg m e n t 4 o f f r on t ( l e s s t han 8 0 % ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . arizonicus R . C . L . P e r k i n s Me t a t h o r a x + p r op od e u m r e d d i s h , w i t h b r own i r r eg u l a r s p o t s . . . . . . . . . . . . . . . . . . . . . . . . . 5 . ontarianus n . sp . Me t a t h o r a x + p r op od e u m b l a ck , a t mo s t w i t h a p ex o f p r op od e u m I i g h t a n d w i t h me t a n o t u m p a r t l y r ed d i s h • . • . . . . . • • • • . • • . . . • . . 6 S e g me n t 4 o f f r o n t t a r su s l on g e r t h a n 1 • • • • . . • • • • • • . • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . s tenocra n i R . C . L . P e rk i n s S e g m e n t 4 o f f r o n t t a r s u s s h o r t e r t h a n s e gmen t 1 • • • • • . . . ' " • • 7 Scu t um b l a ck ; 0 , 7 t i me s a s l . . . . . . . . . . S c u t u m y e l l ow ; 0 , 7 5 t i me s a s . . . . . . . . . . . . . .

i n c l i n ed ( F i g . t h a n s eg m e n t 1

s eg m e n t 4 o f f r o n t t a r s u s s h o r t e r ( a p p r ox i ma t e l y on g a s s eg m e n t 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 . a u toxen o b i u s R . C . L . P e rk i n s s eg m e n t 4 o f f r o n t t a r s u s l o n g e r ( mo r e t h a n l on g a s s e gmen t 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 . m a gn u s B r own

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MALES

2

3

D o r s a l p r o c e s s o f g on o f o r c e p s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g s 860 , 863 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 D o r s a l p r o c e s s o f g on o f o r c ep s n ea r l y a s l o n g a s g on o f o r c ep s ( F i g s 854 , 857 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 T h o r a x a n d p r op od eu m b l a c k • • • • 6 . sten ocra n i R . C . L . P e rk i n s T h o rax a n d p r op od e u m b r own - r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . arizonicus R . C . L . P e rk i n s D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex p o i n t ed ( F i g . 8 5 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . sjoestedti ( K i e f f e r ) D o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p ex r o u n d ed ( F i g . •

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T h e ma l es of t he o t h e r spec i es are u n k no w n .

1 . Pseu dogon a top us freytagi n e w n a me

Pseu dogona top u s similis F r e y t a g 1 978 : Pseu dogon a top u s similis ( B r u e s 1 906 ) .

1 39 ;

p reocc u p i ed b y

F E MA L E : a p terou s ; l e n g t h 1 , 0-4 , 0 m m ; b ro w n or b ro w n - y e l l ow , w i t h p e t i o l e b l ac k ; a n t e n n ae b ro w n - b l ac k , w i t h seg m e n t 1 -2 a n d 1 0 y e l l ow o r w h i te ; m a n d i b l es a n d l eg s y e l l o w ; h e a d s h i n y , exc a v a t ed , smoo t h , g r a n u I a t ed ; prono t u m crossed b y a s t ro n g t r a n sversa I i m p ress i o n , smoo t h , s h i n y , w i t ho u t sc u l p t u re ; s c u t u m s h i n y , smoo t h , w i t ho u t scu l p t u re , w i t h some t ra c k s of k ee l s ; sc u t e l l u m s h i n y , smoo t h , w i t ho u t sc u l p t u r e ; met a no t u m w i t ho u t t ra n s v e r s a l s t r i a e , w i t h s i des p r o t r u d i n g ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; p l eu r a and p o s te r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; seg m e n t 1 of fron t t a rs u s a s l o n g as segmen t 4 ; en l arged c l a w ( F i g . 8 5 1 ) w i t h s u b a p i c a l t oo t h a n d 7-9 l a me l l ae ; segme n t 5 o f f ro n t t a r s u s ( F i g . 85 1 ) w i t h t w o rows o f 8 + 1 0 l a me l l a e ; apex w i t h a g ro u p of 1 3 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Lex i n g to n ( K y . , U . S . A . ) TYP I C A L M A T E R I AL : h o l o t y p e F a n d 1 0 p a r a t y p e s F F i n WA ; 7 p a r a t y pes FF i n K E . HOST S : i n U . S . A . Delp h a codes l u tulenta ( V a n D u zee ) ( F re y t a g 1 978 ) D I ST R I BU T I O N : Nearct i c reg i o n : U . S . A . : 29 . 1 m i . N of M i n era l W e l l s ( J a c k C o . , Tex . ) , BY ! 2 . 8 m i . 5 o f Somerv i l i e ( W a s h i n g ton Co . , Tex . ) , O L ! Lex i n g to n ( K y . ) , O L ! C M ! WA ! KE !

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851

1 259

- C he l a of Pseu dogon a top u s freytagi n . n a me f rom L e x i n g t o n ( Ken t u c k y )

N O T E S : Ps . freytagi w a s reare d b y F re y t a g ( 1 978 ) con t i n u o u s l y i n l a bo r a t o r y for o n e y e a r . T he l y to k u s rep ro d uc t i on occ u r s i n t h i s s p ec i e s .

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F i g . 852 - T horax and p ropodeum of fem a l es of Pseudogo n a top u s s te n ocra n i R . C . L . Perk i n s from P r i nce A l bert ( Sa s k a t c h e w a n ) ( A ) a n d c a n a densis n . s p . ( t y p e ) ( B ) .

A p p ro x i m a te l y 30 d a y s a re n eeded to com p l e t e a I i f e c y c l e from e g g to a d u l t . R e a r i n g con d i t i o n s w e re u s u a l l y m a i n t a i n ed at 3 2 ° C a n d a 1 6-hour day. T h e spec i es i s n a m e d i n honor o f M r . P a u l H . Frey t a g .

2 . Pseudogona top u s c a n a densis n . s p .

F E M A L E : a p tero u s ; l en g t h 2 , 8 1 -3 , 00 m m ; b l ac k ; head w i t h m a n d i b l es , c l y pe u s a n d a n te r i or reg i o n of frons ( more a l o n g orb i t s ) tes t aceou s ; a n t en n ae brown , w i t h s e g m e n t s 1 -'7. testaceo u s ; l eg s test aceou s , w i t h fore c l u b s of femora a n d fore t i b i ae brow n ; a n te n n ae d i s t a l l y t h i ck e n e d ; a n ten n a l segmen t s i n fo l l ow i n g proport i o n s : 7 : 6 : 1 1 : 8 : 7 : 7 : 6 , 5 : 6 : 5 , 5 : 9 ; h e a d d u l l , g r a n u l a t e d , e x c a v a ted ; fro n s w i t h med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L ; 2 ; OL ; 3 ; OOL ; 9 ; p ro n o t u m crossed by a s t ron g t r a n s vers a l i mp ress i on , d u l l , w e a k l y g ra n u l a ted ; sc u t u m d u l l , g r a n u l a t ed ; m e t a n o t u m t r a n s verse l y s t r i a t e , w i t h s i des p rotr u d i n g ; p ro t r u s i on s rou n ded ( F i g . 852 B ) ; me t a t horax + propodeum d u l l , g r a n u l a t e d , w i t h n u merous s t ron g t r a n s versa l s t r i a e on poster i or s u rface ; p l e ura w i t h weak t r a n sversa l s t r i ae ; d i sc a n d a n ter i or s u rface of m e t a t horax + p ropodeum w i t h a s t ron g track of med i a n f u r row ; meso-

Gon a topod i n a e

F i g . 8 5 3 - C h e l a of Pseu dogon a top u s c a n a densis n .

1 26 1

sp .

( type)

m e t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l seg m e n t s i n fo l l ow i n g p roport i o n s : 1 5 : 3 : 5 : 1 1 : 1 7 ; e n l a r ged c l a w ( F i g . 853 ) w i t h s u b a p i c a l t oo t h a n d 4-5 l a m e l l ae ; segme n t 5 of fron t t a rs u s ( F i g . 853 ) w i t h t wo row s of 1 5- 1 7 l a m e l l ae ; a p e x w i t h a g ro u p o f 6-1 0 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4


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854 - M a l e g en i t a l ia of Pseu dogon a top u s c a n a densis n. C a l dor ( C a l i for n i a )

sp . from

segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 4 m m ; b l ac k ; m a n d i b l es y e l l o w , w i t h t ee t h b ro w n ; a n t e n n a e a n d l e g s b row n ; a n t e n n a e n o t d i s t a l l y t h i ck e n e d ; a n te n n a l s e g me n t s i n fo l l o w i n g p ropor t i on s : 7 : 5 : 1 1 : 1 2 : 1 2 : 1 2 : 1 0 : 1 1 : 1 1 : 1 2 ; a n t en n a l segmen t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 1 2 : 3 ) ; h e a d s h i n y , g r a n u l a t ed ; f r o n s w i t ho u t m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 9 ; O l = 5 ; OOl = 4 ; temp l es d i s t i nc t ; occ i p u t conca v e ; s c u t u m d u l l , g ra n u l a ted ; n o t a u l i ce s comp l e t e , p o s t er i or l y sep a r a t e d a n d very n ea r ; scu t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , w i t ho u t scu l p t u re or v e r y w e a k l y p u n c t a t e ; p ropodeum d u l l , ret i c u l a t e r u g o s e ; fore-w i n g h y a l i n e , w i t ho u t d a r k t ra n s v e rs a l b a n d s ; d i s t a l p a r t of ra d i a l v e i n m u c h l on g e r t h a n prox i m a l p a r t ; r a d i a l c e l l open ; dorsa l p rocess of g o n oforceps ( F i g . 854 ) n ea r l y a s l o n g as gonoforc ep s , very b roa d , w i t h d i s t a l reg i on s h o w i n g p a p i I l a e ; d i s t a l m a rg i n serra t e ; m a x i I l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . l O C U S T Y P I C U S : Red Deer ( A l be rt a , C a n a d a ) T Y P I CAL MAT E R I A L :

h o l ot y p e F !

and 1

p a r a t y pe F !

i n OT ;

3 p a r a t ypes

1 26 3

Gon a topod i n ae

F i g . 855 - F e m a l e of Pseu dogo n a topus sjoestedti ( K i effer ) ( t y p e ) : A , t ho r a x a n d p ropod eu m ; C : hea d .

B:

F F ! i n T E ; 1 parat y p e F ! i n AM ; 2 parat ypes ( 1 F , 1 M ) ! i n O L . HOST S : u n k n o w n D I ST R I BU T I O N : Nearc t i c reg i on : CANADA : N o r m a n W e l l s ( N . W . T . ) , a T ! R e d Deer ( A l t a . ) , O T ! U . S . A . : C a l dor ( E I Dora do Co . , C a l i f . ) , OL l T E ! S a n S i meon ( Sa n L u i s Ob i spo C o . , C a l i f . ) , A M !

3 . Pseudogo n a topus sjoestedti ( K i effer )

n . com b .

Gon a top u s sjoestedti K i effer 1 904b : 525 . Pseu dogon a top u s american us R . C . L . Perk i n s 1 905 : 37 ( n . s y n . ) ; locus typicus C o l u m b u s ( O h i o , U . S . A . ) .

F E M A L E : a p t e rou s ; l e n g t h 3 , 47-3 , 56 m m ; red d i s h , w i t h pe t i o l e b l ac k ; a b domen p a r t l y b ro w n i s h ; head ( F i g . 855 C ) exca v a ted , s h i n y , g ra n u l a t e d ; p rono t u m ( F i g . 855 A , B ) s h i n y , crossed b y a s t ro n g t r a n svers a l i m p ress i o n , g r a n u l a ted ; sc u t u m a n d s c u te l l u m s h i n y , smoot h ; m e t a n o t u m t r a n sverse l y s t r i a te , w i t h s i des rou n d e d ; meso-me t a p l e u r a l s u t u re obso l et e ; m e t a t ho r a x + propode u m ( F i g . 855 A , B ) g r an u l a ted , w i t h


1 264

F i g . 856

-

I n st . ,

n o . 37 ,

1 984

C h e l a of Pseudogona topus sjoestedti ( K i effer ) ( t y p e )

Gon a topod i n a e

1 265

F i g . 857 - M a l e gen i ta I i a of Pseu dogon a top u s sjoestedti ( K i effer ) from L e x i n g ton ( K e n t u c k y ) ( r i g h t h a l f removed )

t r a c k of med i a n f u r row ; a n t e r i or s u rf ace v e r y i n c l i n ed ; p l eu r a not t r a n s v erse l y s t r i a t e ; pos t e r i o r s u rf a c e of p ropodeum t ra n s v erse l y s t r i a t e ; segmen t 4 o f fron t t a r s u s a s l o n g a s o r s l i g h t I y shorter t h a n segmen t 1 ( 1 4 : 1 6 i n h o l o t y p e of Ps . sjoes tedti ; 29 : 34 i n h o l o t y p e of Ps . america n u s ) ; e n l arged c l a w ( F i g . 856 ) w i t h s u b a p i c a l t oo t h a n d 4-7 l a me l l ae ; segmen t 5 of fron t t a rs u s ( F i g . 856 ) w i t h t wo rows of 1 1 -24 l a me l l ae ; a p e x w i t h a g ro u p of 6-1 1 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l ab i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 1 , 8 m m ; b l ac k ; c l y p e u s t e s t a ceou s ; m a n d i b l es y e l l ow ; a n ten n a e , a b domen a n d l eg s b row n ; t a r s i y e l l o w ; h e a d s h i n y , w e a k l y g ra n u l a ted ; frons w i t hou t med i a n k ee l ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e ; tem p l es d i s t i n c t ; POL � 7 ; O L = 3 ; OOL � 3 ; sc u t u m d u l l , g ra n u l a ted ; not a u l i ces com p l e t e , pos t e r i or l y s e p a r a t e d ; sc u t e l l u m a n d met a n o t u m s h i n y , smoo t h , w i t h o u t sc u l p t u r e ; p ropodeum w e a k l y ret i c u l a t e ru gose , w i t ho u t t r a n s v e rsa l or l o n g i t u d i n a l k ee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n sv e rs a l b a n d s ; r a d i a l c e l l open ; d i s t a l p a r t of rad i a l v e i n l on g e r t h a n p rox i m a l p a r t ; dors a l p rocess of gonoforceps ( F i g . 857 )

w i t h a p e x p o i n ted ; m a x i I l a ry p a l p i w i t h 4 segmen t s ;

l ab i a l

1 266

Mem . A m e r . E n ! .

A

I n s! . ,

no.

37 ,

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c

Q;J}

F i g . 858 - F e m a l e of Ps e u do go n a t op u s arizonicus R . C . L . P e r k i n s from Mon t gomery ( Georg i a ) : A , B : t horax a n d propode u m ; C : h ea d .

p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S TYP I C U S : T e x a s ( U . S . A . ) TYP I CA L MAT E R I AL : h o l o t y p e F ! of G . sjoestedti i n ST ; l ec t o t y pe F ! ( d es i g n a t e d b y O l m i 1 982 ) of Ps . cunerica n u s i n 3 . HOST S : i n U . S . A . Liburnia sp . ( R . C . L . P e r k i n s 1 906c ) D I ST R I B U T I ON : N e a rc t i c reg ion : U . S . A . : Tex a s , ST ! Waco ( Mc L e n n a n C o . , T e x . ) , O L ! BY ! Co l u m b u s ( O h i o ) , B ! L e x : n g ton ( K y . ) , K E ! H i g h l a n d s ( N . C . ) , O T ! Sea C l i ff ( Lo n g I s l a n d , N . Y . ) , C M !

4 . Pseu dogon a top u s arizon icus R . C . L . P e r k i n s

Pse u dogon a top u s arizonicus R . C . L . P e r k i n s 1 907 : 3 6 . F E MA L E : a p t e ro u s ; l en g t h 2 , 1 9-2 , 40 m m ; y e l l o w - t e s t aceo u s , w i t h p e t i o l e b l ac k ; occa s i on a l l y a b domen b row n ; h e a d ( F i g . 858 C ) exc a v a t e d , s h i n y , g r a n u l a ted ; p ron o t u m ( F i g . 858 A , B ) crossed b y a s t ro n g t ra n sversa l i m p re ss i on , s h i n y , g r a n u l a t e d ; m e t a n o t u m w i t h s i d es rou n d e d ; m e t a t h o r a x + propo d e u m ( F i g . 858 A , B ) s h i n y , g r a n u l a ted ; a n t er i o r s u rface of m e t a t ho r a x + p ropod e u m v e r y i n c l i ne d ; p l e u r a not t r a n svers e l y s t r i a t e ; pos t e r i o r s u rface o f p ropod e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l s u t ure o b so l e t e ; seg m e n t 4 of fron t t a rs u s shorter t h a n segmen t 1 ( 20 : 25 i n h o l o t y p e ; 9 : 1 2 i n a s p e c i men from Mon t g omery ) ; e n l a rg ed c l a w ( F i g . 859 ) w i t h s u b a p i c a l too t h a n d 4-5 l a m e l l ae ; segmen t 5 of f r o n t t a r s u s ( F i g . 859 ) w i t h t wo row s of 1 0- 1 7 l a me l l ae ; a p e x w i t h a g ro u p o f 5- 1 0 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ;

Gon a topod i n ae

F i g . 859

-

C he l a of Pseudogona topus arizonicus R . C . L . Mon t gomery ( Georg i a )

1 267

Perk i n s from

1 268


I n st . ,

no .

37 ,

1 984

F i g . 860 - M a l e g en i t a l i a o f Pseu dogona topus arizon icus R . C . L . from N og a l es ( A r i zo n a )

Perk i n s

t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 3 m m ; b ro w n-red d i s h , w i t h l eg s ye l l ow ; a n t e n n a e n o t d i s t a l l y t h i ck e n ed ; a n t e n n a I segme n t s i n f o l l o w i n g propor t i o n s : 5 : 6 : 1 0 : 1 0 : 1 0 : 1 0 : 9 : 8 : 8 : 1 0 ; a n ten n a l s e g m e n t 3 n e a r l y f i ve t i mes a s l o n g a s b roa d ; head s h i n y , weak l y g r a n u l a ted ; t em p l es s hor t , b u t d i s t i nc t ; occ i p i t a l ca r i n a i n v i s i b l e ; fron t a l l i n e weak , b u t d i s t i nc t ; P O L = 7 ; OL = 2 ; O O L = 3 , 5 ; s c u t u m weak l y g r a n u l a ted ; not a u l i ces com p l et e , poster i or l y s e p a r a ted ; m i n i m u m d i s t a n c e b e t ween t h e n o t a u l i ces s h o r t e r t h a n b rea d t h of t h e oce l l i ( 2 : 3 ) ; s c u te l l u m a n d m e t a no t u m s h i n y , smoo t h , w i t h o u t scu l p t u r e ; p ropode u m ret i c u l a t e ru gose ; a n t e r i or s u rface w i t h m ed i a n f u rrow ; fore- w i n g h y a l i n e , w i t ho u t d a r k t r a n svers a l b a n d s ; d i s t a l p a r t of r a d i a l ve i n l o n g e r t h a n p rox i ma l p a r t ( 20 : 1 1 ) ; dors a l process of gonoforceps ( F i g . 860 ) very shor t , m u c h shorter t h a n g o noforce p s ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : Nog a l es ( A r i z . , U . S . A . )

1 26 9


F i g . 861

-

C he l a of Pseu dogon a top u s ontari a n u s n .

sp.

( type)

T Y P I CAL M A T E R I A L : l ec t o t y p e F ! a n d 2 p a ra l ec t o t y pes ( 1 F , 1 M ) ! ( d es i g n a te d b y O l m i 1 98 2 ) i n B . HOST S : i n U . S . A . Stoba era s p . ? ( R . C . L . P e r k i n s 1 90 7 ) a n d Sa cch arosydne s a ccharivora . D I S T R I BU T I ON : N e a rc t i c reg i o n : U . S . A . : Nog a l es ( A r i z . ) , B ! F e l l sm e re ( F l a . ) , O L ! WA ! S a p e l o I s . ( G a . ) , O L !. WA ! M o n t g o m e r y ( G a . ) , I T !

5 . Pseu dogon a top u s on taria n u s n .

sp .

F E MAL E : a p t e rou s ; l e n g t h 3 , 1 8-3 , 62 m m ; h e a d b ro w n , w i t h occ i p u t , c l y p e u s , m a n d i b l es a n d fro n t p a r t of v e r t e x ( more a l o n g o rb i t s ) t es t aceous or redd i sh ; a n ten n ae b row n , w i t h seg m e n t s 1 -2 t e s t a c eo u s ; p rono t u m red d i s h - d a r k ; sc u t u m w i t h pos t e r i o r h a l f redd i s h ; a n te r i or h a l f brow n ; s c u t e l l u m b l ac k ; m e t a t ho r a x + p ro p o d e u m red d i s h-d a rk o r red d i s h - b ro w n , w i t h a p e x t e s t aceo u s ; p e t i o l e b l ac k ; a b domen red d i s h d a rk ; l e g s t e s t aceo u s-red d i sh , w i t h c o x a e and c l u b s of femora p a r t l y b ro w n ; a n te n n a e d i s t a l l y t h i ckened ; a n t en n a l seg men t s i n f o l l ow i n g p ro p o r t i on s : 9 : 5 : 1 4 : 8 : 7 : 7 : 6 : 6 : 5 : 9 ; h e a d e x c a v a ted , g r a n u l a ted , d u l l ; f r o n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; OL = 3; O O L = 8; p ro no t u m c rossed b y a s t ro n g t r a n s v e r s a I i m p res s i on , d u I I , g r a n u l a te d ; sc u t u m d u l l , g r a n u l a t ed ; scu t e l l u m s h i n y , smoo t h ; m e t a n o t u m r u g ose , w i t h s i des p r o t ru d i n g ; p ro t r u s i on s rou n ded ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a t ed , w i t h n u merou s t r a n s v e rs a l s t r i ae on p l eu r a a n d p o s t e r i o r s u rf a c e ; meso-me t a p l eu r a l s u t u r e obso l e t e ; m e t a t ho r a x +

1 270

Fig.

Mem. Amer. E n ! .

I ns! . ,

n o . 37 ,

1 984

862 - C h e l a of Pseudogona topus stenocra n i R . C . L .

Perk i n s from O h i o

propo d e u m v e r y h u mped , a s i n P a l aearc t i c Gon a top u s l u n a tus K l u g ; fore t a rs a l seg men t s in fo l l ow i n g p ropor t i o n s : 1 5 : 3 : 5 : 1 5 : 2 1 ; en l arged c l a w w i t h s u b a p i c a l t oo t h a n d 6 l a me l l a e ; segme n t 5 o f fron t t a rs u s w i t h t w o r o w s of n ea r l y 30 l a me l l ae e x t en d i n g con t i n uo u s l y t o t h e a p e x ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n


L O C U S TYP I C U S : G o l den L . ( O n t a r i o , C a n a d a ) T Y P I CAL MAT E R I AL : ho l ot y p e F ! a n d 1 p a r a t y p e F ! i n AM ;

1

paratype F !

1 27 1

i n OT ;

paratype F !

i n OL .

HOST S : u n k n o w n D I ST R I BU T I O N : N e a rc t i c reg i on : CANAD A : G o l d e n L . ( O n t . ) , OT ! O L ! U . S . A . : K a l b f l e i sc h F i e l d S t a t i on ( H u n t i n g ton , L o n g I s l a n d , N . Y . ) , A M !

6 . Pseudogona topus stenocra n i R . C . L .

Perk i n s

Pseu dogon a top u s sten ocra n i R . C . L . P e r k i n s 1 905 : 38 . Pseu dogona topus stenocra n i v a r . d u b io s u s R . C . L . P e r k i n s 1 905 : 39 ( s y n . p roposed b y F re y t a g 1 978 ) ; locus typ i c u s C o l u m b u s ( Oh i o , U . S . A . ) . Pseudogon a top u s iowensis F e n to n 1 924 : 1 90 ( s y n . p roposed b y F re y t a g 1 978 ) ; locus typ icus Ames ( I o w a , U . S . A . ) .

F E MAL E : a p t e rou s ; l e n g t h 2 , 62-3 , 29 m m ; h e a d t e s t a ceous or y e l l o w-b row n ; occ a s i o n a l l y r e d d i s h - b r ow n ; u s u a l l y w i t h v e r t ex b row n i s h ; f r o n t p a r t of v e r t e x , c l y p e u s and m a n d i b l es y e l l o w ; a n t e n n a e u s u a l l y b ro w n , w i t h s e g " l e n t s 1 -3 a n d 1 0 y e l l ow ; occas i o n a l l y s e g m e n t 1 0 w h i t e ; occ a s i o n a l l y a n t e n n a e fu l l y t e s t a c eo u s ; p ron o t u m redd i s h , occas i o n a l l y w i t h b r o w n spots o n i d es a n d b e t ween c o l l a r a n d d i sc ; s c u t u m redd i s h , o cc a s i o n a l l y w i t h s i de s b l a ck ; occ a s i on a l l y sc u t u m f u l l y b l ack ; sc u t e l l u m a n d m e t a t ho r a x + p ro p o d e u m b l a c k ; p e t i o l e b l ac k ; a b domen red d i s h o r b ro w n ; l e g s y e l l o w ; h e a d excav a te d , s h i n y or d u l l , g r a n u l a t ed ; p ro n o t u m c rossed by a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , w e a k l y g ra n u l a t ed ; sc u t u m d u l l , g r a n u l a t e d ; m e t a n o t u m w i t h s i des rou n d e d ( F i g . 852 A ) ; m e t a t ho r a x + p ro p o d e u m d u l l , g r a n u l a te d ; p l e u r a a n d pos t e r i o r s u rface of p ropod e u m t ra n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; seg men t 1 of f ro n t t a r s u s s h o r t e r t h a n seg m e n t 4 ( 25 : 26 i n l ec t o t y p e o f Ps . s tenocra n i ; 1 1 : 1 3 i n ho l o t y pe of Ps . iO\J D : M a t l e y Bog ( New Forest , H a n t s Sou t h ) , BM ! SPA I N : E I P a r d o ( M a d r i d ) , M D ! I T ALY : C a n t a l u p a ( To r i no ) , O L l Mon d a g no l a ( F rabosa Sop ra n a , C u neo ) , O l ! Bosco d e l l a P a r t ec i p a n z a ( T r i no Verce l l ese , Verc e l l i ) , O l ! A L G E R I A : Bon a ( A b d u l - N o u r 1 974 ) .

O R I E N T A L S P E C I E S O F DONISTHOi2PINA

O ne s p ec i es i s k n ow n .

G o n a t o po d i n ae

Fig.

1 309

894 - Fem a l e of UonisthoriJ i n a a s i a ti c a n . s p .

Don i s thorp i n a a s i a t i c a n .

( t y pe )

sp .

F E MA L E ( F i g s 894 , 895 ) : a p t e rou s ; l en g t h 3 m m ; t e s t aceou s , w i t h p e t i o l e b l ac k j head exc a v a ted , s h i n y , a l u t aceo u s ; p ronot u m sh i n y , h a i r l ess , a l u t aceo u s , c rossed by a s t ron g t r a n s v e r s a l i m p res s ion ; scu t u m very s l e n d e r , s h i n y , w i t ho u t sc u l p t u re ; m e t a n o t u m s h i n y , w i t h o u t sc u l p t u re , not st r i a t e ; m e t a t ho r a x + p ropod e u m s h i n y , w i t ho u t scu l p t u re ; p l e u r a a n d poste r i o r s u rf a c e o f propo d e u m t r a n s verse l y s t r i a t e ; meso-me t ap l e u r a l s u t u re o b so l e t e ; fore t a r s a l se g m e n t s i n fo l l o w i n g p roport i o n s : 1 3 : 2 : 4 : 1 3 : 1 9 ; e n l arged c l a w ( F i g . 896 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l a e ;

1310

Mem . Amer.

Ent.

I ns t . ,

no . 37 ,

1 984

F i g . 895 - F em a l e of Don isthorpina a s ia tica n . sp .

( type)

segme n t 5 o f f ron t t a rs u s ( F i g . 896 ) w i t h 1 row o f 1 2 l a m e l l ae ; a p e x w i t h a g ro u p of 8 l a me l l a e ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : Gom b a k ( M a l a y a , M a l a y s i a ) TYP I C A L M A T E R I A L : h o l o t y p e F ! i n B M . HOST S : u n k n o w n D I STR I B U T I O N : O r i e n t a l reg i o n : MALAYS I A : Gom b a k

7 . A gona to,rJoides R . C . L .

( Ma l ay a ) ,

B fv\ !

Perk i n s

A gona topoides R . C . L . P e r k i n s 1 907 : 33 .

T Y P E S P E C I E S : Agona topoides synchroIilus R . C . L . P e r k i n s 1 907 , mono t y p i c . F E M A L E ( F i g s 906 , 907 ) : a p terou s ; m a x i l l a ry p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen ts ; p ro n o t u m ( F i g . 907 ) c rossed b y a s t ron g t r a n s v e rsa l i mp ress i on ; e n l a r g e d c l a w ( F i g . 897 ) w i t h s u b a p i c a l too t h a n d l a m e l l a e ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 899 , 900 ) : f u l l y w i n ge d ; m a x i l l a r y p a l p i w i t h 5 seg m en t s ; l a b i a l p a l p i w i t h 3 s e g m e n ts ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e o r v i s i b l e on l y b e h i n d t h e oce l l i o r o n t h e s i de s o f t h e pos te r i or oce l l i , b u t n e v e r r e a c h i n g temp l e s ; temp l es d i s t i nc t ; t i b i a l s p u r s 1 , 1 , 2 . D I ST R I B U T I O N : P a l a e a rc t i c , E t h i o p i a n , O r i en t a l , N e a r c t i c , Neo t rop i c HOST S : Delp h a cidae


F i g . 896

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C he l a of Don isthorpina a s i a tica n . sp .

1 31 1

( type)

1 31 2

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SPEC I ES :

I ns! . ,

n o . 37 ,

1 984

15

PALAEA R C T I C S P E C I E S OF A GONA TOPOIDES

T h ree spec i es are k n ow n .

K E Y TO T H E SPEC I E S

F E MA L E S

2

M e t a n o t u m h o l l ow b e h i n d t h e s c u t e l l u m ; m e s o -m e t a p d i s t i n c t , s t r on g . . . . . . . . . . . 1 . felix n . sp . M e t a n o t u m n o t h o l l ow b eh i n d t h e s cu t e l l u m , a l mo s t -me t ap l eu r a l s u t u r e ob s o l e t e o r i n d i s t i n c t . . . . . . . B od y a l mo s t f u l l y b l a ck 2 . solidus ( H a u p t ) B od y a l mo s t f u l l y r ed d i s h - t e s t a c e ou s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . can a riensis n . sp . •

•

.

l eu ra l

su t u r e

f l a t ; meso . . . . . . . . . . . .2

.

. . . . . . . . . . . . .

·

MALES

D o r sa l p r o c e s s of g on o f o r c ep s ( F i g . · . . . . . . . . . . . . . . . . D o r sa l p r o c e s s o f g on o f o r c ep s ( F i g . . . . . . . . . . . . . . . . . ·

T h e m a l e of A .

g on o f o r c e p s 9 0 1 ) ; n o t au . . . . . . . . . .2. g on o f o r c e p s 90 3 ) ; n o t au . . . . . . . . . . 3.

v e ry s h o r t , mu ch s h o r t e r t h a n l i c e s j o i n t ; ra d i a l c e l l s ma l I solidus ( H a u p t ) v e r y l on g , n ea r l y a s l on g a s l i c e s s ep a ra t ed ; ra d i a l c e l l w i d e can ariensis n . sp .

felix i s u n k n ow n .

1 . A gona topoides felix n .

sp.

F E M A L E : a p te rou s ; l e n g t h 2 , 1 8- 2 , 68 m m ; red d i s h - t e s t a ceou s , w i t h p et i o l e b l ac k a n d a b domen b row n ; a n te n n a e w e ak l y , d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n ts i n fo l l o w i n g p ropor t i o n s : 8 : 5 : 1 2 : 6 : 6 : 6 : 5 : 4 : 4 : 6 ; head d u l l , f u l l y g r a n u l a t e d , w e a k l y e x c a v a t e d ; occ i p i t a l c a r i n a i n v i s i b l e ; fron t a l l i ne a b se n t ; POL = 2 ; O L = 3 ; OOL = 8 ; p ro n o t u m s h i n y , crossed by a s t ron g t r a n s v e rs a l i m p ress i o n , w i t h i n d i s t i nc t l o n g i t u d i n a l s t r i ae ;

1 31 3

Go n a topod i n ae

F i g . 897

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C h e l a of Agona to, )oicies fel i x

n .

sp .

( t y pe )

1 31 4

Fig.

M e m . Amer . E n t .

I nst . ,

no . 3 7 ,

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898 - C he l a of Agon a topoides solidus ( H a u p t ) from S a n i co d i A l f i a n o N a t t a ( I ta l y )

Gon a topod i n a e

F i g . 899

-

1315

M a l e o f ilgon D. to , ) oides sol i d u s ( H a u p t ) from M a sserano ( I t a l y )

s c u t u m g r a n u l a ted , d u l l ; s c u t e l l u m d i s t i n c t ; m e t a n o t u m t r a n s verse l y s t r i a t e , h o l l o w be h i n d t h e sc u te l l u m , w i t h s i des p r o t r u d i n g ; protru s i on s pO i n t ed ; meso-me t a p l eu r a l s u t u r e d i s t i nc t , com p l e t e ; m e t a t horax + p ropode u m s h i n y ; a n t e r i o r s u rf a c e ',. i t ll l on g i t u d i n a l , i n d i s t i nc t s t r i ae ; pos ter i o r s u rface tra n s v e rse l y s t r i a t e ; fore t a rsa l seg men ts i n fo l l ow i n g proport i o n s : 1 2 : 2 : 5 : 1 3 : 20 ; en l arged c l a w ( F i g . 897 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l ae ( i n a d d i t i o n t o 1 b r i s t l e ) ; seg men t 5 of fron t t a rs u s ( F i g . 897 ) w i t h 2 row s o f 1 0 l a m e l l a e ; a p e x w i t h a g ro u p o f 1 2 l am e l l a e ; m a x i l l a r y p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : I� i o C a b r a ( Ca n ary I s l a n d s ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 2 p a r a t y p es F F ! i n M D ; 2 p a r a t y pes F F ! in O L . HOST S : u n k no w n D I ST R I B U T I O N : Pa l ae a rc t i c reg i o n : C A I"ARY I SL A N D S : R i o C a b r a ( F u e r te v e n t u r a ) , M D ! O L !

1316

Fig.

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no . 37 ,

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900 - M a l e of A gona to,'Joides solidus ( H a u p t ) from M a sserano ( I t a l y )

2 . Ilgon a to[Joides solidus ( H a u p t ) n . com b .

A llogon a topus Agona topoides locus typicus Agona topoides

solidus H a u p t 1 938 : 2 7 . joh a n n a e C u r r a do a n d O l m i 1 975 : 28 ( n . sy n . ) ; San i co d i A l f i a no N a t t a ( I t a l y ) . joh a n n a e C u rr a d o a n d O l m i : O l m i 1 98 1 : 1 1 6 .

F E MALE : a p t e rou s ; l en g t h 3 , 4 m m ; h e a d b ro w n , w i t h m a n d i b l es , c l y p e u s a n d f ron t p a r t of v ertex y e l l ow ; a n t e n n a e b row n-b l ac k , w i t h s e g m e n t s 1 -3 y e l l ow ; t ho r a x a n d propode u m b l ac k ; a b domen b ro w n -b l ac k ; l e g s y e l l o w , w i t h p a r t of coxae a n d p a r t of c l u b s of femora b l ac k ; h e a d e x c a v a ted , s h i n y , gra n u l a ted , h a i r l ess ; prono t u m h a i r l es s , g r a n u l ated , crossed b y a s t ron g t r a n s v e rsa l i m press i o n ; s c u t u m s h i n y , g r a n u l a ted , w i t h some l o n g i t u d i n a l k ee l s ; sc u t e l l u m s h i n y , g ra n u l a te d ; m e t a n o t u m tra n s verse l y s t r i a t e ; m e t a t horax + propodeum s h i n y , g r a n u l a ted ; a n te r i or s u rface of m e t a t ho r a x + p ropodeum w i t h track of med i a n f u rrow ; p l e u ra a n d pos t e r i o r s u rf ace o f p ropod e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l e u ra l s u t u re o b so l e t e ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropor t i on s : 1 4 : 2 : 3 : 9 : 1 4 ; en l a r g e d c l a w ( F i g . 898 ) w i t h s u b a p i c a l too t h a n d 6-7 l a me l l a e ; seg m e n t 5 o f f ro n t t a r s u s ( F i g . 898 ) w i t h t wo rows o f 1 9 l a me l l ae ; a p e x w i t h a g ro u p of 4-5 l am e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . r,1ALE ( F i g s 899 , 900 ) : f u l l y w i n g e d ; l en g t h 3 m m ; head b l ac k , w i t h

Gonatopod i n ae

1 31 7

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( Ha u p t )

from

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m a n d i b l es test aceo u s ; a n t e n n a e b l a c k - b row n ; t horax a n d propod e u m b l ack ; a b do m e n b row n ; l e g s y e l l ow - b row n , w i t h t a r s i l i g h t ; a n t en n a l seg men t 3 more t h a n four t i mes as l o n g as b ro a d ( 5 , 6 ) ; h e a d s h i n y , h a i r y , g ra n u l a ted ; f ro n t a l l i n e a b s en t ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m s h i n y , g r a n u l a ted ; not a u l i ce s comp l e te , poste r i o r l y j o i n t ; s c u t e l l u m a n d m e t a n o t u m s h i n y , wea k l y p u n c t a te ; p ropod e u m w i t h pos t e r i or s u rface ret i c u l a te ru gose ; a n te r i o r s u rface w i t h areo l ae sma l l a n d w i t h med i a n f u rrow ; poster i or s u rface w i t h areo l ae w i de ; dorsa l p rocess of gonoforceps ( F i g . 90 1 ) short , d i s t a l l y s l e n d e r ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a � i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS TYP I CU S : Bel l i nchen ( German y ) TYP I CA L M A T E R I AL : h o l o t y p e F ! of A . solidus i n HA ; h o l o t y p e F ! o f A . joh a n n a e i n E N . HOST S : i n I t a I y Mega delp h a x sp . a n d L a odelp h a x s tria tell u s ( F a l i e n ) Welp h a cida e ) ; i n Germ a n y Muirodelph a x a u bei ( P e r r i s ) ( = L ib urnia a u b e i ) ( H a u p t 1 938 ) . D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : G E R M AN Y : B e l l i nc h e n ( O d e r ) , HA ! I TALY : M asserano ( Verce l l i ) , O L ! C a s te l l a l fero ( As t i ) , E N ! Sa n i co d i A l f i a n o N a t t a ( A l es s a n d r i a ) , E N !

3 . Agona tofJoides c a n ariensis n .

sp .

F E MA L E : a p t erou s ; l e n g t h 3 , 56 m m ; red d i s h - t e s t aceou s , w i t h pos ter i or s u rface of p ropod e u m d a r k ; pe t i o l e b l ack ; a b domen b row n ; a n t e n n ae b ro w n , w i t h s e g m e n t s 1 -2 test aceou s ; a n t e n n a e w e a k l y d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n t s i n fo l l o w i n g p ropor t i on s : 7 : 5 : 1 3 : 7 : 7 : 6 : 5 : 5 : 5 : 6 , 5 ; h e a d sh i n y , exca v a ted , wea k l y g r a n u l a ted ; fron t a l l i n e presen t ; occ i p i t a l ca r i n a i n v i s i b l e ; POL = 1 ; O L = 2 ; O O L = 7 ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i mp ress i o n , d u l l , g r a n u l a ted ; s c u t u m a n d scu te l l u m d u l l , g r a n u l a ted ; m e t a n o t u m n o t h o l l o w b e h i n d t h e sc u te l l u m , a l most f l a t , t r a n s v erse l y s t r i a t e , w i t h s i des rou n d e d ; m e t a t horax + p ropo d e u m d u l l , g r a n u l a t e d , w i t h pos ter i or s u rface a n d p l e u r a t ra n s v e rse l y s t r i a te ; meso-me t a p l eu r a l s u t u r e i n d i s t i n ct ; fore t a r s a l seg men t s i n fo l l ow i n g proport i on s : 1 4 : 2 : 4 : 1 0 : 1 6 ; en l a rged c l a w ( F i g . 902 ) w i t h s u b a p i c a l toot h a n d 7 l am e l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 902 ) w i t h t w o row s of 20 l a m e l l ae ; a p e x w i t h a g ro u p of 9 l a m e l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 5 m m ; b l ac k ; m a n d i b l es b l ac k , w i t h tee t h test aceou s ; a n t e n n ae b row n ; l eg s b ro w n , w i t h fore t i b i ae test aceo u s ; a n te n n ae n o t d i s t a l l y t h i c k ened ; a n t e n n a I se g m e n t s i n fo l l ow i n g proport i on s : 6 : 4 , 5 : 9 : 9 : 9 : 8 : 8 : 8 : 7 : 9 ; a n t e n n a I se g m e n t 3 four t i mes a s long as b road ; h e a d d u l l , g r a n u l a t e d ; occ i p i t a l c a r i n a i n v i s i b l e ; f ro n s w i t ho u t k ee l s ; POL = 7 ; O L = 3 ; OOL = 4 ; scu t u m d u l l , g r a n u l a ted ; n o t a u l i ces comp l e te , pos ter i or l y s e p a r a t e d ; sc u t e l l u m w e a k l y g r a n u l a ted ; m e t a n o t u m s h i n y , w i t ho u t scu l p t u re ; p ropodeum ret i c u l a t e

Go n a topod i n ae

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902 - C h e l a of ilgon a topoides c a n arien sis n . s p .

( type )

r u gose ; dorsa l s u r face w i t h med i a n f u rrow a n d w i t h v e r y s m a l l a reo l ae ; poster i o r s u rface w i t h w i de areo l ae ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n s v e rs a l b a n d s ; gonoforceps w i t h dors a l p rocess ( F i g . 903 ) l on g , n a rrow , n e a r l y a s l on g a s gonoforce p s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : B a rra nco d e l os J u ncos ( C a n a r y I s l a n d s ) TYP I CA L M AT E R I AL : h o l o t y p e F ! i n M D ; 1 p a r a t y p e M ! i n O L .

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F i g . 903 - M a l e g en i t a l i a of A gona to,'Joides c a n a riensis n . s p . f ro m C . no de l a s M e rcedes ( C a n a r y I s l a n d s ) ( r i g h t h a l f remov e d )

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HOST S : u n k n o w n 0 1 S T R I B U T I O N : P a I a e a r c t i c r e g i on : C A N A R Y I SLANOS : B a rr a n co de l o s J u ncos ( T e n e r i fe ) , M O ! C . n o d e l as Mercedes ( T e n e r i fe ) , O L !

E T H I O P I A N SP E C I E S O F AGONA TOPOIDES

F i ve spec i e s are k n ow n .

K E Y T O T H E SPEC I E S

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E n l a rg ed c l aw w i t h a p ex v e ry b r oad

2

(F i g . 905 ) 1 fu s c u s n . sp . E n l a rg ed c l aw w i t h a p e x t a p e r i n g , n o t b r o a d ( F i g s 9 0 8 , 9 0 9 , 91 0 , 91 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2 Me t a t h o ra x + p r o p o d e u m w i t h a s t r o n g med i a n f u r r ow ( F i g . 9 0 6 ) .

3

2 . a c u m in a tu s n . sp . Me t a t h o ra x + p r o p o d e u m w i t h o u t s t r o n g med i a n f u r r ow Me t a n o t u m w i t h s i d e s r o u n d ed ; me t a t h o ra x + p r o p o d e u m b l a ck

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MALES U n k nown 1 . A go n a topoides fu s c u s n . sp .

F E M A L E ( F i g . 904 A ) : a p t e r ou s ; l en g t h 3 , 1 2-4 , 00 m m ; head , a n t e n n a e , t hora x , p ropod e u m , a b domen a n d l eg s tes t aceous-re d d i s h or redd i s h - d a r k ; p et i o l e b l ac k ; occa s i o n a l l y a b do m e n b rown i s h ; h e a d f l a t , d u l l , h a i r y , g r a n u l a t e d ; p ron o t u m d u l l , h a i ry , w i t h n u merous w e a k l o n g i t u d i n a l k ee l s , crossed b y a s t ro n g t r a n s v ersa I i m pres s i on ; s c u t u m d u I I , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m d u l l , t r a n s v erse l y s t r i a t e , ho l l o w b e h i n d t h e sc u te l l um , w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s po i n ted ; met a t ho r a x + p ropodeum d u l l ; a n te r i o r s u rface w i t h n u merous l o n g i t u d i n a l k ee l s ; p l e u r a

1 322

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F i g . 904 - F e m a l e of Agona topoides fu scus n . s p . from Van R e e n e n ( So u t h A f r i c a ) ( A ) ; B : he a d ; C : t h o r a x a n d p ropodeum l a tera l l y v i ewed .

a n d pos t e r i o r s u rf a ce n o t or w e a k l y t ra n s v e rse l y s t r i a t e ; meso- m e t a p l e u r a l s u t u re comp l e t e , d i s t i nc t ; fore t a rsa l segmen t 1 s l i g h t l y s h o r t e r t h a n seg m e n t 4 ( 1 4 : 1 5 ) ; e n l a r g e d c l a w ( F i g . 905 ) w i t h a pe x v e r y b ro a d , w i t h s u b a p i c a l too t h a n d 5-7 l am e l l a e ; segmen t 5 of fro n t t a r s u s ( F i g . 905 ) w i t h t w o ro w s of 1 2- 1 7 l a me l l ae ( s ho r t e s t row exte n d i n g o r not e x t en d i n g b e y o n d 0 , 5 l en g t h of segmen t ) ; a p e x w i t h a g ro u p of 1 0-20 l a m e l l a e ; m a x i I l a ry p a l p i w i t h 5 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : Mosse l B a y ( So u t h Afr i c a ) TYP I CA L MAT E R I AL : ho l o t y p e F ! a n d 3 p a r a t y pe s F F ! i n B M ; 1 p a r a t y p e F! in OL .

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1 323

( t y pe )

1 324

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1 984

F em a l e of Agona topoides a c um i n a t u s n .

s ;=> .

( type)

HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i on : SOU T H A F R I CA : Mosse l B a y ( C a p e P rov i n ce ) , B M ! K a t b e r g ( E C a p e P ro v i n ce ) , OL l BM ! V a n Reenen ( D r a k e n s Berg , N a t a l ) , B M !

2 . A gona topoides a c umin a tu s n . s p .

F E MALE

( F i g s 906 , 907 ) :

a p tero u s ;

l e n g t h 3 , 1 2 mm ;

red d i s h- b ro w n ,

with

1 325

Gon a topod i n ae

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Fe m a l e of Agona top o i cies a c um in a t u s n . s p .

( type )

m a n d i b l e s , c l y p e u s a n d front p a r t of vertex y e l l ow ; a n ten n ae y e l l ow , w i t h segmen ts 8- 1 0 brow n ; p e t i o l e b l ac k ; t a r s i a n d a p e x of p ropodeum y e l l o w ; h e a d d u l l , exc a v a t ed , g r a n u l a ted ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , w i t h s h o r t , i rreg u l a r kee l s ; sc u t u m s c u l p t u red b y l o n g i t u d i n a l k ee l s ; sc u te l l u m smoo t h a n d s h i n y ; m e t a n o t u m w i t h s i des p ro t r u d i n g ; p r o t r u s i o n s p o i n t ed ; m e t a t ho r a x + p ropodeum s h i n y , as scu I p t u re d as p ro n o t u m , wi th a s t ro n g med i a n f u rrow ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; poste r i or s u rf ace of p ropode u m t r a n s verse l y s t r i a t e ; p l e u r a not t r a n sv e rse l y s t r i a t e ; fore t a rsa l seg men ts i n fo l l ow i n g p ropor t i on s : 9 : 2 : 4 : 1 0 : 1 6 ; en l arged c l a w ( F i g . 908 ) w i t h s u b a p i c a l t oo t h a n d 7 l a m e l l ae ; segme n t 5 o f fron t t a r s u s ( F i g . 908 ) w i t h t w o rows of 1 7 l a me l l ae ; apex w i t h a g ro u p of 1 6 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 5 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : U p e m b a N a t i on a l P a r k ( Z a i re ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n T V . HOST S : u n k n o w n D I ST R I BU T I ON : E t h i op i a n reg i on : Z A I R E : U p e m b a N a t i o n a l P a r k , T V !

3 . A gona topoides cafer n . s p .

1 32 6

Mem . A m e r .

Fig.

908

-

En! .

I ns ! . ,

no .

37 ,

1 984

C he l a of Agon a topoides acumin a t u s n . sp .

( t ype )


F i g . 909 - C he l a of A go n a to;)oides c a fer n . s p .

1 327

( type)

F E MAL E : a p terou s ; l e n g t h 3 , 62 m m ; h e a d a n d a n te n n ae redd i s h - d a r k ; a n t e n n a I s e g m e n t 1 0 w h i t e ; p ron o t u m a n d scu t u m t e s t a c eo u s ; sc u te l l u m a n d m e t a t ho r a x + propodeum b l ac k , e x c e p t f o r a p e x of p ro p o d e u m r e d d i s h ; p e t i o l e b l ac k ; a b d o m e n red d i s h - d a r k ; l e g s t e s t aceo u s ; h e a d exc a v ated , d u I I , g r a n u I a t e d ; p ro no t u m crossed b y a s t ro n g t r a n sversa I i m p re s s i o n , s h i n y , smoot h , w i t h some l o n g i t u d i n a l kee l s ; m e t a n o t u m s h o r t , h o l l o w b e h i n d t h e s c u te l l u m , w i t h s i des rou n d e d ; m e t a t h o r a x + p ropodeurn s h i n y , w i t h o u t scu l p t u re ; pos t e r i or s u rf ace of propode u m a n d p l eu r a t r a nsverse l y s t r i a te ; meso- m e t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; fore t a r s a l segmen t 1 shorter t h a n seg m e n t 4 ( 1 5 : 1 7 ) ; e n l arged c l a w ( F i g . 909 ) w i t h a p e x t a p er i n g , n o t b ro a d , w i t h s u b a p i c a l t oo t h a n d 8 l a m e l l ae ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 909 ) w i t h t w o rows of 30 l a me l l ae ; a p e x w i t h a g ro u p of 25 l a me l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : M a tjesfon t e i n ( So u t h Afr i c a ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n B M . HOSTS : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : SOU T H A F R I C A : M a t jesfon t e i n P rov i n ce ) , B M !

( Cape

4 . A gona topoides ruber n . s p .

F E MA L E : a p terou s ; l e n g t h 3 , 43 m m ; red d i s h - b ro w n , w i t h p et i o l e d a r k ; a n te n n a e a n d t a r s i l i g h t ; h e a d e x c a v a t e d , s h i n y , smoo t h , h a i r l ess , w i t h o u t sc u l p t u r e ; occ i p u t h a i r y ; p ron o t u m crossed by a s t ro n g

1 328

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9 1 0 - C h e l a of A gona topoiJes I'u J)eI' n . s p .

( type)

t r a n s v e rs a l i m p ress i o n , s h i n y , smoot h , h a i r l es s , w i t h o u t scu l p t u re ; s c u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m smoo t h , n o t t r a n sve rse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h o u t s c u l p t u re ; p l e u r a a n d pos t e r i o r s u r'face of p ropode u m t r a n sverse I y s t r i a t e ; meso-me tap l e u ra I s u t u re obso l e te ; fore t a r s a l segmen ts i n fo l l o w i n g p ropo r t i o n s : 1 4 : 2 : 4 : 1 1 : 1 7 ; e n l a r g e d c l a w ( F i g . 9 1 0 ) w i t h s U b a p i c a l too t h a n d 5 l a me l l ae ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 9 1 0 ) w i t h two rows of 1 3 l a me l l ae ; a p e x w i t h a g ro u p of 1 3 l a me l l a e ; m a x i I l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i " i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n L O C U S TYP I C U S : M i l nerton ( Sou t h A f r i c a ) T Y P I CAL MA T E R I AL : h o l o t y p e F ! i n B lv\ . HOSTS : u n k n o w n , D I STR I I3 U T I O N : E t h i o p i a n reg i o n : S O U T H A F R I C A : ''' d l n erton , 13 M !

5 . A gon a toi)oides ceres n . s p .

F E M A L E : a p terou s ; l e n g t h 3 , 75 mrn ; red d i s h - te s t aceo u s , w i t h p e t i o l e b l ac k ; head exca v a t e d , d u " , g r a n u l a te d ; p ro no t u m d u " , g ra n u l a t e d , crossed b y a s t r o n g t r a n s v e rs a l i m p re ss i o n ; scu t u m d u l l , g ra n u l a t e d ; m e t a n o t u m t r a n s v e rse l y s t r i a t e , h o l l o w b e h i n d t h e sc u t e l l u m , w i t h s i d e s p r o t r u d i n g ; p ro t r u s i on s ro u n d e d ; rne t a t ho r a x + p ropode u m d u l l , g ra n u l a t e d ; p o s t e r i o r s u rface of p ropodeurn a n d p l eu r a t r a n s v erse l y s t r i a t e ; a n te r i o r s u rface of propo d e u m w i t h n u me ro u s weak

l o n g i t u d i n a l k ee l s ;

meso-me t a p l e u r a l s u t u r e

Gon a topod i n a e

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1 330


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d i s t i n c t , com p l e t e ; fore t a r s a l s e g m e n t 1 shorter t h a n segmen t 4 ( 1 3 : 1 6 ) ; e n l a r g e d c l a w ( F i g . 9 1 1 ) w i t h a pex t a p er i n g , not b road , w i t h s u b a p i c a l too t h a n d 6- 1 2 l a m e l l ae ; segment 5 o f front t a r s u s ( F i g . 9 1 1 ) w i t h t w o r o w s of 1 3-23 l ame l l ae ; a p e x w i t h a g r o u p o f 1 2- 1 8 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Ceres ( So u t h A fr i c a ) TYP I CA L MAT E R I AL : ho l o t y p e F ! a n d 1 p a ra t y pe F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I O N : E t h i op i an reg i o n : SO U T H A F R I CA : Ceres ( C a p e P rov i nce ) , BM ! Port S t . Johns ( Po n do l a n d ) , BM !

O R I E NT A L S P E C I E S OF AGONA TOPOIDES

F o u r s pec i es a re k n ow n .


F E MA L E S

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1 . benga lensis n . s p . D i s c o f m e t a t h o ra x + p r op od eu m w i t h o u t t ra n s v e r s a l k e e l s ; p o s t e r i o r k e e l s p r e s en t on l y on p o s t e r i o r s u r f a c e o f p r op od eu m ; o c c a s i o n a l l y s h o r t k e e l s a r e v i s i b l e o n t h e g r o u n d o f t h e m e d i a n f u r r ow 2 V e r t ex o f h ea d , me t a n o t u m a n d me t a t h o ra x + p r o p o d e u m v e ry s h i n y , smo o t h , w i t h o u t s c u l p t u r e . . . . . . . . . . . . . 2 . born e a n u s n . s p . V e r t ex o f h e a d , me t a n o t u m a n d me t a t h o ra x + p r o p o d e u m d u l I , g r a n u I a t ed 3 A n t en n a e s h o r t , mo r e d i s t a l l y t h i c k en ed ; a n t en n a l s e g men t 9 •

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s l i g h t l y l on g e r t h a n b r o a d . . 3 . insularis n . s p . A n t en n a e l on g , l e s s d i s t a l l y t h i c k e n ed ; a n t en n a l a p p r o x i ma t e l y t h r e e t i me s a s l on g a s b r o a d 4 . pyrill a e ( Ma n i ) •

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1 33 1

l on g a s b r o a d ; d o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p ex s h a rp ( F i g . 9 1 4 ) . . . . . . . . . . . . . . . . . . 2 . born e a n u s n . s p . H e a d b r own ; a n t en n a l s eg m e n t 3 l e s s t h a n t h r e e t i me s a s l on g a s b r o a d ; d o r s a l p r o c e s s o f g o n o f o r c e p s w i t h a p ex r o u n d ed ( F i g . 9 1 7 ) . . . . . . . . . . . . . . . . . . 4 . pyri l l a e ( Ma n i )

T he m a l es of A .

F i g . 91 2

-

insularis a n d bengalen sis a re u n k n ow n .

C he l a of Agona topoides benga lensis n .

sp.

( type)

1 332

Mem . Amer .

Ent.

I ns! . ,

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1 984

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C he i a of !l gona topoides b orn e a n u s n . s p .

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1 33 3

1 . Agona topoides benga len sis n . sp .

F E MA L E :

a p tero u s ;

l e n g t h 3 , 50-5 , 3 1

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y e l l o w - t e s t a ceou s ,

w i t h t horax

a n d p ropode u m red d i s h- b row n ; p e t i o l e b l ac k ; h e a d s h i n y , exc a v a ted , h a i r l es s , w i t h o u t scu l p t u re ; prono t u m crossed b y a s t rong t r a n svers a l i mp res s i on , s h i n y , smoot h , h a i r l es s , w i t h o u t scu l p t u re , w i t h some l a tera l k ee l s ; s c u t u m s h i n y , smoo t h , w i t h some l on g i t u d i n a l kee l s ; sc u t e l l u m s h i n y , smoo t h , rec t a n g u l a r ; m e t a n o t u m t r a n s verse l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i on s ro u n ded ; m e t a t h o r a x + propodeum s h i n y , w i t h a s t ro n g m e d i a n f u rrow ; poster i o r s u rface a n d d i sc of m e t a t h o r a x + propodeum t r a n s verse l y s t r i a t e ; p l e u r a not t r a nsverse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; fore t a r s a l segme n t s i n fo l l ow i n g p ropor t i on s : 1 6 : 4 : 5 : 20 : 3 3 ; e n l a rged c l a w ( F i g . 9 1 2 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 9 1 2 ) w i t h 1 row of 1 3 l a m e l l a e ; apex w i t h a g ro u p o f 2 0 l a m e l l ae ; m a x i l l ary p a l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : M i s h m i H i l l s ( A ss a m , I n d i a ) T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n BM . HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i on : I N D I A : M i s h m i H i l l s ( D e l a i V a l l ey , Assam ) , B M !

2 . Agona topoides born e a n u s n .

sp .

F E MA L E : a p tero u s ; l e n g t h 3 , 52 m m ; t e s t a ceou s - b row n ; a n te n n ae d i s t a l l y weak l y t h i c kened ; a n ten n a l segme n t s i n fo l l ow i n g p ropor t i on s : 1 2 : 7 : 2 2 : 1 2 : 1 1 : 9 : 7 : 7 : 7 : 1 1 ; h e a d very exca v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re ; occ i p i t a l c a r i n a i n v i s i b l e ; fron t a l l i n e com p l e te ; POL = 1 ; O L 3 ; OOL = 1 0 ; p ro n o t u m crossed b y a s t ro n g t ra n svers a l i mp ress i o n , w i t h o u t scu l p t u re , h a i r l es s ; scu t u m a n d s c u te l l u m s h i n y , smoot h , w i t ho u t scu l p t u re ; s i d es of scu t u m w i t h t wo l a t e r a l poi n t s ; m e t a n o t u m smoot h , s h i n y , w i t h o u t sc u l p t u r e , ho l l ow b e h i n d t h e s c u t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; m e t a t ho r a x + p ropodeum s h i n y , smoot h , w i t h o u t scu l p t u re , w i t h a s t r o n g med i a n f u r row ; pos t e r i o r s u rface o f p ropode u m a n d p l e u ra t r a n sverse l y s t r i a t e ; fore t a r s a l segmen t s i n fo l l ow i n g proport i o n s : 1 9 : 5 : 8 : 22 : 3 3 ; en l a rged c l a w ( F i g . 9 1 3 ) w i t h s u b a p i c a l too t h a n d 8 l a me l l ae ; segment 5 o f f ron t t a r s u s ( F i g . 9 1 3 ) w i t h t w o rows of 1 1 + 2 ( p ro x i m a l ) l am e l l a e ; a pex w i t h a g ro u p of n e a r l y 1 8 l ame l l a e ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 87 m m ; head , a n t e n n a e a n d l eg s test aceou s ; t h o r a x , propode u m a n d a b domen b ro w n ; a n ten n a l segme n t s i n fo l l ow i n g proport i o n s : 6 : 6 : 1 2 : 8 : 8 : 8 : 8 : 8 : 7 : 1 0 ; a n t e n n a I segme n t 3 more

1 334


I n s! . ,

no . 3 7 ,

1 984

F i g . 9 1 4 - M a l e gen i t a l i a of Agon a topoides born e a n u s n . s p .

( paratype)

t h a n f o u r t i mes a s l on g a s b ro a d ( 4 , 8 ) ; h e a d h a i ry , s h i n y , a l u t aceo u s ; fron t a l l i ne a b sen t ; frons w e a k l y g r a n u l ated ; occ i p i t a l ca r i n a a b s en t ; temp les very s h o r t , red uced ; POL = 6 ; O L = 2 , 5 ; OOL = 2 ; b re a d t h o f t h e oce l l i : 3 ; s c u t u m h a i r y , s h i n y , g r a n u l a t ed ; n o t a u l i ces com p l e t e , poster i or l y sep a r a ted ; m i n i m u m d i s t a n ce b e t w ee n t h e n o t a u l i ces s l i g h t l y s horter t h a n b rea d t h of t h e oce l l i ( 2 , 5 : 3 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoot h , w i t h o u t scu l p t u re ; p ropode u m ret i c u l a te r u gose , w i t ho u t l o n g i t u d i n a l kee l s on pos ter i or s u rface ; fore-w i n g h y a l i n e , w i t ho u t d a r k t ra n s v e r s a l b a n d s ; dors a l p rocess of gonoforceps ( F i g . 9 1 4 ) l o n g , b roa d , p o i n te d ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l spurs 1 , 1 , 2 . LOCUS TYP I C U S : G u no n g M u l u ( Sa r a w a k , M a l a y s i a ) T Y P I C A L M A T E R I A L : h o l o t y p e F ! a n d 1 p a ra t y pe M ! i n SM . HOSTS : u n k n o w n D I STR I BU T I O N : O r i e n t a l reg i o n : MALAYS I A : G u no n g M u l u ( Sa r a w a k ) , BM !

3 . Agona topoides in s u l a ris n . s p .

1 335


Fig.

9 1 5 - C he l a of Agon a topoides insul aris n . sp .

( t y pe )

F E MA L E : a p terou s ; l e n g t h 3 , 2 m m ; t e s t a ceou 5 , w i t h p e t i o l e b l ack ; a n te n n a e b row n , w i t h seg m e n t s 1 -3 a n d 1 0 t e s t aceo u s ; a n t en n ae d i s t a I I y t h i ck e n e d ; a n te n n a l s e g me n t s i n fo l l ow i n g p roport i on s : 1 0 : 7 : 1 8 : 1 0 : 9 :

1 336


Fig. 916

-

I n st . ,

no. 37,

1 984

C he l a of Agon a topoides pyrillae ( M a n i ) ( t y p e )

8 : 8 : 6 : 5 , 5 : 1 0 ; a n te n n a l segmen t 9 s l i g h t l y l on ge r t h a n b ro a d ( 5 , 5 : 4 ) ; head d u l l , very e x c a v a t e d , g r a n u l a ted ; fron t a l l i n e com p l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 1 ; O L � 3 ; OOL � 1 2 ; p ron o t u m c rossed b y a s t ro n g t r a n svers a l i m p ress i o n , s h i n y , weak l y g r a n u l a ted ; sc u t u m s h i n y , w i t hou t sc u l p t u r e ; m e t a n o t u m h o l l ow b e h i n d t h e sc u t e l l u m , w e a k l y t r a n s verse l y s t r i a t e ; m e t a n o t u m w i t h s i des p ro t ru d i n g ; p ro t r u s i o n s p o i n ted ; fore t a rs a l segmen t s i n fo l l ow i n g p ropo r t i on s : 1 6 : 4 : 6 : 1 9 : 30 ; e n l a r g e d c l a w ( F i g . 9 1 5 ) w i t h s u b a p i c a l too t h a n d 8 l a me l l a e ; segmen t 5 of fron t t a r s u s ( F i g . 9 1 5 ) w i t h t w o rows of 3 ( p rox i m a l ) + 1 2 l a m e l l ae ; a p e x w i t h a g ro u p of 1 2 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I C U S : U l h i t i y a O y a ( S r i L a n k a ) T Y P I CA L MAT E R I A L : h o l ot y pe F ! i n WA . HOST S : u n k n o w n D I ST R I B U T I ON : O r i en t a l reg i o n : S R I LANKA : U l h i t i y a O y a ( 1 5 m i . N N E of M a h i y a n g a n a , B a d . D i s t . ) , WA !

4 . Agon a topoides pyril l a e ( M a n i )

n .

com b .

Gon a topod i n a e

1 337

F i g . 9 1 7 - M a l e g en i t a l i a of Agon a topoides pyrill a e ( M a n i ) ( a l l o t y p e )

Ps e u dogon a top u s pyril l a e M a n i 1 942 : 1 6 1 . Pseu dogon a top u s pyrillae M a n i : K u r i a n 1 954a : 2 5 8 .

F E MA L E : a p tero u s ; l e n g t h 4 , 2 m m ; t e s t aceo u s-red d i s h , w i t h a b domen b l ac k ; a n te n n a e d i s t a l l y t h i ck e n ed ; a n t e n n a I segme n t s i n fo l l o w i n g p ro p or t i on s : 1 1 : 8 : 30 : 20 : 1 5 : 1 3 : 1 0 : 1 0 : 9 : 1 1 ; a n te n n a l segmen t 3 a p p ro x i m a te l y t h ree t i mes as l on g as b ro a d ( 9 : 3 ) ; h e a d e x c a v a ted , d u l l , g r a n u l a te d ; f ro n t a l l i n e comp l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; O L = 3 ; O O L = 1 4 ; p ro n o t u m crossed b y a s t ro n g t r a n sversa l i mp res s i on , s h i n y , w e a k l y g r a n u l a t e d ; sc u t u m s h i n y , w e a k l y g ra n u l a t e d ; s c u te l l u m s h i n y , i n d i s t i nc t ; m e t a n o t u m ho l l o w b eh i n d t h e sc u te l l u m , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s po i n ted ; m e t a t horax + p ropodeum d u l l , g r a n u l a ted , w i t h a s t ro n g med i a n f u r ro w , w i t h s t ro n g t r a n sversa l s t r i ae on pos t e r i o r s u rface ; d i sc ru gose , n o t t r a n sv erse I y s t r i a t e ; meso-

1 338


I n st . ,

no.

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1 984

m e t a p l e u r a l s u t u re d i s t i n c t , comp l e t e ; mesop l eu r a not t r a n s verse l y s t r i a t e ; me t a p l e u r a t r a n s v e rs e l y s t r i a t e ; fore t a r s a l segme n t s i n fo l l ow i n g p roport i on s : 2 2 : 5 : 7 : 25 : 40 ; en l a rged c l a w ( F i g . 9 1 6 ) w i t h s u b a p i c a l tooth a n d 1 1 l a me l l ae ; seg men t 5 of fron t t a rs u s ( F i g . 9 1 6 ) w i t h 1 row of 1 5 l a m e l l a e ; a p e x w i t h a g ro u p of a p prox i m a te l y 9 l a m e l l ae ; seg men t 3 of front t a r s u s p ro d u c e d i n to a hook ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : f u l l y w i n ged ; l e n g t h 2 , 4 m m ; b row n , w i t h a n te n n a e , m a n d i b l es a n d l e g s t e s t aceou s ; a n t e n n ae n o t d i s t a l l y t h i c k e n ed ; a n te n n a l s e g m e n t s i n fo l l ow i n g p roport i on s : 7 : 6 : 8 : 7 : 7 : 6 : 6 : 6 : 6 : 9 ; a n t en n a I s e g m e n t 3 l es s t h a n t h re e t i me s as l o n g as b ro a d ( 8 : 3 ) ; h e a d d u l l , gra n u l ated , w i t hou t fron t a l l i n e ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e ; temp l e s d i s t i nc t ; P O L = 8 ; O L = 4 ; O O L = 4 ; sc u t u m s h i n y , g ra n u l a ted ; not a u l i ces com p l e t e , poste r i o r l y sep a r a te d ; m i n i m u m d i s t a nce b e t w een t h e notau l i ces s horter t h a n b re a d t h of the ocel l i ( 1 , 5 : 3 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoo t h , p u n c t a te , w i t h o u t s c u l p t u re a mong p u n c t u res ; propode u m d u l l , ret i c u l a t e r u g o s e , w i t h o u t t r a n s versa l o r l on g i t u d i n a l k ee l s ; dors a l s u rface w i t h a m ed i a n n a rrow f u r ro w ; fore-w i n g h y a l i ne , w i t h o u t d a r k t r a n s v e rs a l b a n d s ; r a d i a l c e l l open ; d i s t a l p a r t of r ad i a l v e i n l on g e r t h a n p rox i m a l p a r t ( 2 3 : 1 5 ) ; d o r s a I p rocess of gonoforceps ( F i g . 9 1 7 ) l o n g , d i s tB l l y b roadened ; m a x i l l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 3 s e g me n ts ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS TYP I CU S : K a r n a l ( I n d i a ) T Y P I CA L M A T E R I AL : h o l o t y pe F ! a n d a l l o t y p e M ! i n N O . HOST S : i n I n d i a Pyril l a s p . ( M a n i 1 942 ) . D I ST R I BU T I O N : O r i e n t a l reg i on : I N D I A : K a rn a l ( P u n j a b ) , N O ! D a u ra l a F a rm ( Meeru t ) , N O !

N E A RC T I C S P E C I E S O F AGONA TOPOIDES

T w o s p ec i es a re k n ow n .


F E MA L E S

S e g m e n t 4 o f f r o n t t a r s u s s h o r t e r t h a n s eg me n t 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . synchromus R . C . L . P e r k i n s S e gm e n t 4 o f f r o n t t a r s u s l on g e r t h a n s eg m e n t 1 2 . c a lidus n . s p . •

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Go n a topod i n ae

1 339

MALES

U n k no w n

1 . Agona topoides synchromus R . C . L .

Perk i n s

A gona topoides synchromus R . C . L . Perk i n s 1 907 :

33 .

F E MAL E : a p terou s ; l e n g t h 4 , 0 m m ; redd i s h - t e s t aceou s , w i t h pet i o l e b l ac k ; body n e a r l y h a i r l es s ; h e a d exc a v a ted ; p ron o t u m crossed b y a s t ron g t r a n s v e rs a l i m p ress i on ; p o s t e r i or s u rface of p ropodeum t ra n sverse I y s t r i a t e ; segme n t 5 of f ro n t t a r s u s l on g e r t h a n s e g m e n t 4 ( 76 : 50 ) ; se g m e n t 4 of fron t t a r s u s shorter t h a n segmen t 1 ( 1 04 : 1 40 ) ; en l a rged c l a w w i t h s u b a p i c a l too t h a n d 5 l ame l l a e ; segmen t 5 of fron t t a r s u s w i t h two row s o f 8 + 9 l a m e l l ae ; apex w i t h a g ro u p o f 2 l a me l l a e ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) T Y P I CAL MAT E R I A L : l ec t o t y pe F ! a n d 2 p a r a l ec t o t y pes F F ! ( d e s i g n a ted b y O l m i 1 98 2 ) i n B . HOSTS : i n U . S . A . Bosta era ( � Cochise ) a p a c h e a n u s K i rk . ( Delph a cida e ) ( R . C . L . Perk i n s 1 907 ) ; Bosta era n a s u ta B a l l ( T ho m p son 1 95 4 ) D I ST R I BU T I O N : N e a rc t i c reg i on : U . S . A . : Nog a l e s ( A r i z . ) , 8 ! •

2 . Agona topoides ca lidus

n .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 06 m m ; tes t aceo u s , w i t h pe t i o l e b l ac k ; a n te n n ae b ro w n , w i t h seg m e n t s 1 -3 a n d p a r t of 4 t e s t aceo u s ; segmen t 1 0 a n d p a r t of 9 w h i te ; a n t e n n a e d i s t a l l y t h i c kened ; a n t en n a l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 8 : 5 : 1 5 : 8 : 8 : 6 : 6 : 6 : 6 : 9 ; head exc a v a ted , s h i n y , g ra n u l a ted ; fron t a l l i n e com p l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 1 ; OL � 2; OOL � 9; p ro n o t u m crossed by a s t ro n g t r a n svers a l i m p res s i on , s h i n y , smoo t h , w i t ho u t scu l p t u re ; scu t u m d u l l , g ra n u l a t e d , w i t h o u t l a tera l po i n ted p ro t r u s i on s ; sc u te l l u m i n v i s i b l e ; m e t a n o t u m n o t f l a t , t r a n s verse l y w e a k l y s t r i a te , w i t h s i des p ro t ru d i n g ; p rot r u s i on s po i n te d ; meso-met a p l e u r a l s u t u r e obso l e t e ; m e t a t ho r a x + p ropodeum s h i n y , w i t h o u t sc u l p t u re among s t r i ae a n d on a n te r i o r s u r f ace o f p ropode u m ; p o s te r i or s u rface o f p ropodeum a n d p l e u r a t r a n sv erse I y s t r i a t e ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 1 0 : 3 : 6 : 1 5 : 24 ; e n l a r ged c l aw ( F i g . 9 1 8 ) w i t h s u b a p i c a l too t h a n d 1 5 l a me l l a e ; seg men t 5 of f ron t

1 340


Fig.

I ns! . ,

no.

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1 984

9 1 8 - C he l a of Agona topoides c a lidus n . s p .

( type )

t a rs u s ( F i g . 9 1 8 ) w i t h t wo row s of n e a r l y 50 l ame l l ae e x t en d i n g co n t i n uo u s l y t o t h e apex ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 .

Gona topod i n a e

F i g . 91 9

-

1 34 1

C he l a of Agon a topoides longichel a t u s ( F e n ton ) { t y p e )

MALE : u n k n o w n L O C U S T Y P I C U S : W a s h i n g ton ( D . C . , U . S . A . ) TYP I CA L MAT E R I AL : ho l o t y pe F ! i n WA . HOST S : u n k n o w n D I ST R I BU T I O N : N e a rc t i c reg i o n :

U.

S . A . : W a s h i n g ton

(D.

C. ) ,

WA!

1 342


I ns ! . ,

n o . 37 ,

1 984

N E O T R O P I C SP E C I E S OF A GONA TOPOIDES

O n e s p ec i es i s k n ow n .

A gon a top o id e s l ongich e l a t u s ( Fe n ton ) n . com b .

Dicondylus l o ngich e l a tu s F e n ton 1 927 : 4 .

F E MAL E : a p terou s ; l e n g t h 3 , 3 1 m m ; h e a d red d i sh - b row n , w i t h m a n d i b l es a n d c l y p e u s y e l l ow ; fron t p a r t of vertex l i g h t ; a n te n n ae ye l l ow ; p ro n o t u m redd i s h ; co l l a r w i t h b ro w n spo t s ; s c u t u m , s c u t e l l u m , m et a t horax + p ropodeum a n d pet i o l e b l ac k ; a b domen redd i s h- b row n ; l eg s y e l l o w redd i s h ; h e a d e x c a v a t e d , d u l l , g r a n u l a t e d , w i t h s h o r t t r a n s v e rs a l kee l s n e a r med i a n fron t a l l i ne ; p rono t u m crossed b y a s t ro n g t ra n s vers a l i m p re s s i on , s h i n y , smoo t h , w i t ho u t scu l p t u r e , w i t h some l a tera l kee l s ; scu t u m d u l l , w i t h l on g i t u d i n a l k e e l s ; s c u te l l u m s h i n y , smoot h ; met a t horax + propodeum d u l l , w i t h o u t scu l p t u re ; m e t a no t u m , p l e u r a and pos ter i or s u rface of p ropodeum t r a n s v erse l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t ; fore t a rs a l seg m e n t s i n fo l l ow i n g p roport i on s : 1 4 : 2 : 5 : 1 5 : 2 5 ; e n l a r g e d c l a w ( F i g . 9 1 9 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l a e ; segmen t 5 of f ro n t t a rs u s ( F i g . 9 1 9 ) w i t h 2 row s of 1 0 + 1 3 l a me l l ae ; a p e x w i t h a g ro u p of a t l e a s t 1 5 l ame l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : P e r n a m b uco ( B r a z i I ) TYP I CAL MAT E R I AL : ho l o t y p e F ! i n WA . HOSTS : u n k no w n D I S T R I B U T I O N : Neot rop i c reg i o n : BRAZ I L : Pern a m b uco ( Bo n i t o ) , WA !

8 . Ap terodryin u s R . C . L .

Perk i n s

Ap terodryin u s R . C . L . Perk i n s 1 907 : 1 4 . Dolichochelys O l m i a n d C u rrado 1 976a : 1 77 ( n . s y n . ) ; t y p e s p ec i es Dolichochelys longicornis ( K i effer 1 904 ) , o r i g . d es i g . Noga top u s Ponomarenko 1 979a : 357 ( n . s y n . ) ; t y p e s p ec i es Noga top u s gob iensis Ponom a renko 1 97 9 , o r i g . d e s i g .

T Y P E S P E C I E S : Apterodryin u s torvus R . C . L . Perk i n s 1 907 , monot y p i c . F E MA L E ( F i g s 926 , 927 ) : a p terou s ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; labial

p a l p i w i t h 3 segme n t s ; e n l a rged c l a w ( F i g . 920 )

w i th subap ical

G o n a to pod i n a e

F i g . 920

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1 343

C he l a of Ap terodryinus gob iensis ( Ponom a r e n ko ) ( t y p e )

too t h a n d l a m e l l ae ; p ro n o t u m ( F i g . 927 ) crossed b y a s t ro n g t r a n sv e r s a l i m p ress i o n ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; m a x i I l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3

1 344

Mem . Amer .

En! .

I ns! . ,

no . 37 ,

1 984

segme n t s ; tem p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e or on l y v i s i b l e beh i n d t he oce l l i o r o n t h e s i des o f t h e pos t e r i or oce l l i , b u t never reac h i n g tem p l es ; occ i p u t concave ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N : wor l d w i de HOST S : A c a n a loniida e , Fl a tidae SPEC I ES : 1 7

PALAEA R C T I C S P E C I E S O F APTERODR YINUS

O n e spec i es i s k n o w n .

Ap terodryin u s gobiensis ( Ponom a ren k o ) n . com b .

Noga top u s gobiensis Ponom arenko 1 979a : 357 .

F E MAL E : a p t e ro u s ; l e n g t h 3 , 5 m m ; tes t aceou s , w i t h pe t i o l e b l ac k ; a n t e n n ae d i s t a l l y t h i c kened ; a n t e n n a l segme n t s i n fo l l ow i n g p ropor t i o n s : 7 : 4 : 1 1 : 7 : 6 : 6 : 5 : 5 : 6 : 7 ; head d u l l , exca v a ted , g ra n u l a t ed ; occ i p i t a l ca r i n a i n v i s i b l e ; fron t a l l i n e com p l e t e ; P O L � 1 ; OL � 2 ; OOL � 7 ; p ro n o t u m crossed by a s t ron g t r a n s versa l i m p ress i on , s h i n y , w i t h o u t scu l p t u re ; m e t a no t u m h o l l o w beh i n d t he scu t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; m e t a t horax + p ropodeum s h i n y , w i t hou t scu l p t u re , excep t for pos t e r i o r s u rface of p ropodeum a n d p l e u r a t r a n s v erse I y s t r i a t e ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 1 4 : 3 : 4 : 1 6 : 2 3 ; en l arged c l a w ( F i g . 920 ) w i t h s u b a p i c a l t oo t h and 1 0 l a m e l l ae ; segme n t 5 of f ron t t a rs u s ( F i g . 920 ) w i t h 2 row s of 26 l ame l l ae ; a pe x w i t h a g ro u p of n e a r l y 1 1 l a me l l ae ; m ax i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : h o l o t ype F ! i n L N . H OS T S : u n k no w n D I S T R I B U T I O N : P a l aearc t i c reg i o n : M O N GO L I A : S h a ra-K h u l s n y - B u l a g ( B a j a n - K ho n g o r a y m a g , C e n t r a l Mongo l i a ) , L N ! NO T E S : A . go b iensis i s v e r y s i m i l a r t o Agon a topoides felix n . s p . from C a n a r y I s l a n d s : the on l y d i fferences concern t he n u m ber of m a x i l l a r y p a l p a l segmen t s , t h e h e a d ( more e x c a v a t e d i n A . gobien sis ) a n d t he a n t e r i o r s u rf ace of met a t ho r a x + p ropodeum ( w i t ho u t scu l p t ure i n A . go b iens is , w i t h w e a k l o n g i t u d i n a l s t r i a e i n Ag. felix ) .

E T H I O P I AN S P E C I E S O F APTERODR YINUS

Gon a topod i n ae

1 345

T h ree s p ec i es a re k n o w n .


F E MA L E S

M e s o - me t a p l e u ra l s u t u r e ob s o l e t e 1 . somerseti n . s p . Me s o -me t a p l eu ra l s u t u r e d i s t i n c t , c omp l e t e 2 A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m l e s s i n c l i n ed ( F i g . 9 2 2 A ) ; me t a t h o ra x + p r op o d e u m a l mo s t f u l l y b l a c k ; e n l a rg ed . . . . . . . . . . . . c l a w w i t h a p ex v e r y b en t ( F i g . 9 2 3 ) . . . . . . . . . . • . . . 2 . richardsi n . s p . A n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m m o r e i n c l i n ed ( F i g . 9 2 2 8 ) ; me t a t h o ra x + p r op o d e u m f u l l y f e r ru g i n ou s ; e n l a r g ed c l a w w i t h a p ex I e s s b en t ( F i g . 9 2 4 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . rufu l u s n . s p . •

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MALES

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1 . A p terodryin u s somerseti n .

sp .

F E M AL E : a p tero u s ; l e n g t h 4 , 37 m m ; head b row n , w i t h front p a r t o f ver te x , c l y p e u s a n d m a n d i b l e s y e l l o w ; a n t e n n ae test aceo u s ; p r o n o t u m w i t h co l l a r b l ack ; d i sc b ro w n - re d d i s h ; scu t u m y e l l o w , w i t h a n te r i o r m a rg i n b l ac k ; sc u te l l u m a n d met a t ho r a x + p ropodeum b l ac k ; a b domen b ro w n b l ac k ; l eg s tes taceous , w i t h p a r t o f c o x a e a n d p a r t o f c l u b s o f femora b row n ; head exca v a ted , s h i n y , w i t ho u t scu l p t u re , except for occ i p u t a n d f r o n t p a rt o f v e r t e x gra n u l a t e d ; p ro n o t u m c rossed b y a s t ron g t ra n s v e rs a l i m p ress i o n , d u l l , weak l y g r a n u l a t e d ; sc u t u m d u l l , g r a n u l a t ed , l a tera l l y w i t h t w o po i n t ed p ro t r u s i o n s ; m e t a n o t u m t r a n s v erse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re obso l e te ; m e t a t h o r a x + p ropodeum s h i n y , w i t h a n t e r i o r s u rface w e a k l y a l u t aceou s ; pos t er i or s u rface of p ropodeum t r a n s verse l y s t r i a t e ; p l e u r a g r a n u l a t e d , w i t h some t r a n s v e rsa l k ee l s ; fore t a r s a l s e g m e n t 1 as l o n g as seg m e n t 4 ; en l arged c l a w ( F i g . 9 2 1 ) w i t h s u b a p i c a l t oo t h a n d 9 l ame l l a e ; segmen t 5 o f f ro n t t a r s u s ( F i g . 92 1 ) w i t h t w o r o w s o f 3 ( p ro x i m a l ) + 1 6 l am e l l ae ; a p e x w i t h a g ro u p o f 1 4 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 346


Fig.

921

I nst . ,

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1 984

- C he l a of Apterodryin u s somerseti n . s p .

( type)

MALE : u n k n o w n LOCUS T Y P I C U S : Somerset E a s t ( So u t h A f r i ca ) T Y P I CA L M AT E R I A L : ho l o t y pe F ! i n B M . HOSTS : u n k no w n D I ST R I BU T I O N : E t h i o p i a n reg i o n : SOU T H A F R I CA : Somerset E a s t P rov i n ce ) , B M !

( Cape

2 . Ap terodryinus rich ardsi n . s p .

F E MAL E : a p te ro u s : l e n g t h 4 , 8 1 m m ; head , p rono t u m , scu t u m a n d s c u te l l u m redd i s h- d a r k ; a n t e n n ae t e s t a ceou s , w i t h segmen t 1 0 w h i te ; m e t a t horax + p ropodeu m b l ac k - b row n , excep t for m e t a n o t u m a n d a p e x of p ropodeum redd i s h ; p e t i o l e b l ac k ; a b domen and l eg s t e s t aceo u s ; head excav a ted , d u I I , g r a n u I a ted ; p ro n o t u m crossed b y a s t ro n g t r a n sversa I i m p ress i on , d u l l , g ra n u l a ted ; s c u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m h o l l ow b eh i n d t h e s c u te l l u m , t r a n s v erse l y s t r i ate ; m e t a t horax + p ropod e u m ( F i g . 922 A ) d u l l , g r an u l a t e d ; a n t e r i o r s u rface w i t h s o m e w e a k l o n g i t u d i n a l kee l s ; p l e u r a a n d pos ter i or s u rf a ce of p ropodeum t r a n s verse l y s t r i a te ; meso-me t a p l e u ra l s u t u re d i s t i n c t , com p l e t e ; fore t a rsa l segmen t 1 shorter t h a n segmen t 4 ( 9 : 1 3 ) ; en l a rged c l aw ( F i g . 9 2 3 ) s t rong l y c u r ve d , w i t h s u b a p i ca l too t h , 4 l a me l l ae a n d , i n a d d i t i on , a tooth l oc a te d on t h e p rox i m a l t h i rd of t h e i n tern a l s i d e ; segme n t 5 of front t a r s u s ( F i g . 923) w i t h t w o rows of 32 l a m e l l ae ; a p e x w i t h a g r o u p of n e a r l y 40 l ame l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : M o s s e l B a y ( So u t h A f r i c a ) T Y P I CA L M A T E R I A L : h o l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I ON : E t h i o p i a n reg i on : SOUTH A F R I CA : Mos s e l B a y ( C a p e P rov i nce ) , B M !

Go n a t opod i n ae

1 347

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F i g . 922 - Meso t hora x , m e t a t horax a n d p ropodeum ( l a t e ra l l y v i ewed ) o f fema l e s of Ap terodryinus richa rdsi n . sp . ( t y p e ) ( A ) a n d rufu l u s n . s p . ( type) ( 8 )

NOTE :

t he s p ec i es i s n amed i n honor of M r . O w a i n W .

3 . Ap terodryinus rufu l u s n .

R i c h a rds .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 7 5-4 , 8 1 m m ; ferr u g i nou s , w i t h p e t i o l e b l a ck ; head e x c a v a t e d , d u l l , g r a n u l a t ed ; p ron o t u m crossed by a s t ron g t r a n s ve rs a l i m p res s i o n ; co l l a r s h i n y , weak l y g r a n u l a ted ; d i sc d u l l , s t ro n g l y g r a n u l a ted ; scu t u m d u l l , g r a n u l a t e d ; m e t a n o t u m d u l l , t r a n s v erse l y s t r i a t e , h o l l o w b e h i n d t h e s c u t e l l u m ; m e t a t horax + p ropodeu m ( F i g . 9 2 2 B ) d u l l , gra n u l a ted ; a n t e r i o r s u rface of m e t a t horax + p ropode u m v e r y i nc I i ned ; pos t er i or s u rf ace of p ropodeum t ra n s versel y s t r i a t e ; p l eu r a n o t t r a n sverse l y s t r i a te ; meso-me t a p l e u r a l s u t u re d i s t i n c t , comp l e t e ; fore t a r s a l segme n t 1 s horter t h a n segmen t 4 ( 1 4 : 1 6 ) ; en l a rged c l a w ( F i g . 924 ) w i t h s u b a p i c a l too t h and 8 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 924 ) w i t h t w o row s of 2 4 l a m e l l a e ; a p e x w i t h a g ro u p of 2 0 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : P o r t SI . Jo h n s ( Sou t h Afr i c a ) TYP I CAL M A T E R I A L : HOSTS : u n k n o w n

holotype F !

and 1

p a ra t y pe F !

i n BM .

1 348

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I nst . ,

no.

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1 984

F i g . 923 - C he l a o f A p terodryin u s richd rdsi n . s p .

D I S T R I BU T I O N : E t h i o p i a n reg i o n : ( Po n do l a n d ) , B M !

SOU T H A F R I CA :

( type)

Port St . Johns

O F, I E N T A L S P E C I E S O F A PTERODR YINUS

T wo s p ec i es are k n ow n .


F E MA L E S

A n t e n n a e v e ry as

l on g ,

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Gon a topod i n ae

Fig.

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1 349

( t y pe )

1 350


I ns! . ,

no.

37 ,

1 984

1 . a s i a ticus n . s p . A n t e n n a e s h o r t , w i t h s eg men t 4 s l i g h t I y l on g e r t h a n s e g m e n t 2 . . . . . . . . . . . . . . . . . . . 2 . insul aris n . s p . ( s e e A u s t ra l i a n s p e c i e s ) .

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MALES

U n k no w n

1 . A p terodryin u s a siaticus n .

sp.

F E MAL E : a p t e ro u s ; l e n g t h 6 m m ; y e l l ow-ferr u g i nou s , w i t h pe t i o l e b l ac k ; a n t en n a l segmen t s i n fol l ow i n g p ropor t i o n s : 9 : 8 : 40 : 32 : 2 5 : 1 7 : 1 1 : 8 : 7 : 1 0 ; head e x c a v a te d , s h i n y , w i t ho u t sc u l p t u re , e x c e p t for occ i p u t a n d fron t p a r t o f v e r t e x g r a n u I a t e d ; p ro n o t u m crossed b y a s t ro n g t r a n sversa I i mp ress i o n , s h i n y , h a i r l es s , g ra n u l a ted ; scu t u m g ra n u l a ted , w i t h t w o l a tera l p o i n te d p ro t ru s i on s ; sc u te l l u m s h i n y , d i s t i nc t ; met a n o t u m t ra n s v e r se l y s t r i a t e , n o t h o l l ow beh i n d t h e s c u te l l u m ; m e t a t horax + p ropode u m s h i n y , w i t ho u t sc u l p t u re ; p l e u r a , poster i or a n d a n t e r ior s u rface of m e t a t ho r a x + p ropod e u m t r a n sverse I y s t r i a te ; meso-me t a p I e u r a I s u t u re d i s t i nc t , corc p l e te ; fore t a r s a l segmen t 1 shorter t h a n segmen t 4 ( 1 5 : 20 ) ; en l a rged c l a w ( F i g . 925 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; segme n t 5 o f fron t t a r s u s ( F i g . 925 ) w i t h 2 rows of 2 2 + 1 9 l a me l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 3 0 l am e l l a e ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E ; u n k n ow n L O C U S T Y P I C U S : T o n d a n o ( S u l a w e s i , I n dones i a ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! i n B M . H O ST S ; u n k n o w n D I ST R I BU T I ON : O r i e n t a l reg ion : I N DO N ES I A : T o n d a no ( Su l awes i ) , BM !

2 . Ap terodryinus ins ularis n . ( see A u s t r a l i a n reg i on )

sp .

N E A R C T I C S P E C I E S OF APTERODR YINUS

T h ree s p ec i es are k n ow n .


Gon a topod i n ae

F i g . 925

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C he l a o f Apterodryin u s a s i a ticus n .

1 35 1

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( type)

1 35 2

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Ent.

I nst . ,

n o . 37 ,

1 984

F E MA L E S

D i s c o f p r on o t u m p o s t e r i o r l y s t r o n g l y c omp r e s s ed i n t o a med i a n , a cu t e , ra i s ed ca r i n a . 1 . torvus R . C . L . P e rk i n s D i s c o f p r on o t u m r ou n d ed , n o t c omp r e s s ed i n t o a med i a n , a c u t e , . . . . . . .2 ra i s ed c a r i n a Me t a n o t u m n o t h o l l ow b e h i n d t h e s c u t e l l u m . . . . . . . . . 2 . secundus ( F en t on ) Me t a n o t u m h o l l ow b e h i n d t h e s c u t e l l u m . . . . . . . . . . . . . . . . • • 3 . c a liforn icus ( A s h m e a d ) .

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MALES

F i g . 926

-

F e m a l e of Apterodryin u s torvus R . C . L .

( A r i zo n a )

Perk i n s f r o m Nog a l es

Gon a topod i n ae

F i g . 927

-

1 353

F e m a l e of A p terodryin u s torvus R . C . L . Perk i n s from Nog a l es ( A r i zo n a )

U nknown

1 . A p terodryinus torvus R . C . L . Perk i n s

A p terodryin u s torvus R . C . L . Perk i n s 1 907 :

14.

F E MA L E ( F i g s 926 , 9 27 ) : a p te ro u s : l e n g t h 3 , 43-4 , 68 m m ; head b l ac k , w i t h m a n d i b l e s , c l y pe u s a n d m a l a r s p ace y e l l ow ; occ a s i on a l l y head f u l l y f e r r u g i n o u s ; a n ten n ae f u I I y redd i s h ; occ a s i o n a I I y a n te n n ae b ro w n , w i t h segmen t s 9- 1 0 o r 8- 1 0 w h i t i s h ; t ho r a x a n d p ropodeum b l a c k ; p ro n o t u m w i t h w e a k red d i s h n u a nces on t h e s i des ; occ a s i o n a l l y a l so on d i sc a n d co l l a r ; a b domen red d i s h o r b l ac k ; l eg s ferru g i nous o r y e l l ow , w i t h coxae , t roc h a n ters , c l u b s of femora a n d t i b i ae b ro w n i s h ; head e x c a v a t e d , d u l l , h a i ry , g r a n u l a t e d ; occ i p i t a l ca r i n a on l y v i s i b l e b e h i n d t he oce l l i ; p ro n o t u m g r a n u l a te d , crossed b y a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , h a i r l es s , w i t h d i sc poster i or l y s t ron g l y com p ressed i n to a med i a n , a c u t e , r a i sed c a r i n a ; sc u t u m s h i n y , h a i r l es s , g r a n u l a ted ; m e t a n o t u m t r a n s v e r se l y s t r i a t e ; meta t h o r a x + p ropode u m s h i n y , w i t h a n te r i o r s u rf ace smoot h , w i t ho u t scu l p t u re ; p l eu ra a n d pos t e r i o r s u r f ace of p ropodeum

1 354

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Ent.

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no . 37 ,

1 984

F i g . 928 - C he l a of Apterodryin u5 torvu5 R . C . L . Perk i n s from Nog a l es ( A r i zon a )

Gon a topod i n ae

F i g . 929

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1 355

F ore t i b i a of fem a l e of Ap terodryinus secundus

( F e n ton ) ( t y p e )

t ra n s v e r se l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t ; mesop l e u ra a n d m e t a p l e u r a l oc a te d o n d i fferen t p l a nes ; fore t i b i a e n o t c u rved ; fore t a r s a l segmen t s i n fo l l o w i n g proport i o n s : 1 6 : 2 : 5 : 1 9 : 2 9 ; en l a rg e d c l a w ( F i g . 928 ) w i t h s u b a p i c a l too t h a n d 1 2 l am e l l ae ; segmen t 5 of fron t t a r s u s ( F i g . 928 ) w i t h t w o row s o f 27 l a m e l l a e ; a p e x w i t h a g ro u p of a t l ea s t 2 5 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) TYP I C A L M A T E R I AL : l ec t o t y pe F ! a n d 2 p a r a l e c t o t y pes F F ! ( des i g n a t e d b y O l m i 1 982 ) i n B ; 1 p a r a l e c t o t y p e F ! i n BM . HOST S : i n U . S . A . Mistharnop h a n tia son ora n a (Fl a tidae) a n d Amphiscep a imm a c u l a ta (A c a n a lon iida e ) ( R . C . L . Perk i n s 1 90 7 ) . D I ST R I B U T I O N : Nearc t i c re g i o n : U . S . A . : No g a l e s ( A r i z . ) , B ! BM ! OL ! Rock C reek ( Gordo n v i l l e , G r a y so n Co . , Tex . ) , T E ! F a r m i n g d a l e ( N . Y . ) , TW !

2 . Ap terodryi n u s secundus ( F e n to n )

n . com b .

E u c a mptonyx secundus F e n t o n 1 927 : 5 .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 7 5-5 , 40 m m ; fer r u g i n o u s - y e l l ow , w i t h pe t i o l e b l ac k ; occa s i o n a l l y a b do m e n b l ac k ; a n te n n a e y e l l ow , w i t h seg men t s 3-6 d a r k e n e d ; h e a d more or l ess e x c a v a t e d , s h i n y , g r a n u l a t e d ; occ i p i t a l c a r i n a i n comp l e t e ; p ron o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n , s h i n y , g ra n u l a t e d , w i t h d i sc rou n de d ; scu t u m d u l l , g r a n u l a t ed ; scu t e l l u m invisible;

meta thorax

+

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g ra n u l a te d ;

metanotu m ,

p l eura ,

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930 - C h e l a o f Apterodryin u s secundus ( F en ton ) ( t y p e )


1 357

pos ter i o r and a n t e r i o r s u r f ace of m e t a t ho r a x + propodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t ; meso p l e u r a a n d met a p l e u r a l o c a t e d o n t h e s a m e p l a n e ; fore t i b i ae ( F i g . 929 ) s t ron g l y c u r v e d ; fore t a rs a l segme n t s i n fo l l ow i n g proport i o n s : 20 : 4 : 1 0 : 3 3 : 50 ; e n l a rged c l a w ( F i g . 930 ) w i t h s u b a p i c a l too t h a n d 7- 1 3 l a m e l l a e ; segment 5 of f ro n t t a r s u s ( F i g . 930 ) w i t h t w o rows o f 36-47 l a m e l l ae ; a pex w i t h a g ro u p o f a t l ea s t 20-30 l ame l l ae ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : Western O h i o ( U . S . A . ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n WA . HOST S : i n U . S . A . A c a n a lonia con ica ( Sa y ) ( A c a n a loniida e ) . D I S T R I B U T I O N : Nea rc t i c reg i on : U . S . A . : P resco t t ( A r i z . ) , WA ! P i nery C a n . ( C h i r . M ts . , A r i z . ) , OT ! P or t a l ( Coc h i se C o . , A r i z . ) , A M ! S a n t a A n a R e f u g e ( A l a mo , Tex . ) , WA ! L a w re n ce ( Do u g l a s C o . , K a n s . ) , W A ! Western O h i o , Wi, ! A rc h bo l d B i o i . S t a . ( L a k e P l ac i d , H i g h l a n d s C o . , F l a . ) , O L ! P l u m mers I s . ( M d . ) , WA ! P o u g h k eeps i e ( N . Y . ) , T W !

3 . Ap terodryin u s c a l iforn ic u s ( A shmea d )

n . com b .

Gona top u s c a liforn i c u s A s h m e a d 1 893 : 85 .

F E MALE : a p t e ro u s ; l e n g t h 3 , 00-3 , 75 mm ; head f u l l y test aceo u s ; occ a s i o n a l l y b row n , w i t h f ro n t p ar t of vertex a l o n g orb i t s , c l y p e u s a n d m a n d i b l e s tes t a ceou s ; a n t en n ae f u l l y testaceo u s ; occas i o n a l l y b row n , w i t h v e n t r a l face of segmen t 1 y e l l o w ; t h o r a x a n d propodeum f u l l y b ro w n or re d d i s h-test aceo u s ; occ a s i o n a l l y red d i sh-test aceou s , w i t h p ropodeum b row n ; a b domen b row n ; l eg s test aceo u s or b row n ; a n te n n a e d i s t a l l y t h i c k e ne d ; a n t en n a l segme n t s i n f o l l ow i n g p ropo r t i on s : 9 : 6 : 1 3 : 8 : 7 : 5 : 5 : 5 : 5 : 7 ; head sh i n y , exca v a ted , h a i r y , g r a n u l a ted ; p ron o t u m crossed b y a s t r o n g t r a n svers a l i m press i o n , ro u n ded , w i t h d i sc a l u t aceou s ; co l l a r smoo t h , w i t h o u t scu l p t u r e ; sc u te l l u m d i s t i n c t ; m e t a n o t u m ho l l o w b eh i n d t h e sc u te l l u m ; m e t a t h o r a x + p ropodeum g r a n u l a t e d ; p l e u ra a n d p os t e r i o r s u rface of propodeum t r a n s v e rse l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t , com p l e t e ; segme n t 1 of fron t t a rs u s shorter t h a n seg men t 4 ( 2 1 : 27 ) ; en l a rged c l a w ( F i g . 9 3 1 ) w i t h s u b a p i c a l tooth a n d 7 - 1 1 l a m e l l ae ; segme n t 5 o f f ro n t t a r s u s ( F i g . 9 3 1 ) w i t h t w o rows o f 1 6- 1 9 l a me l l ae ; a pex w i t h a g ro u p of a t l e a s t 1 4- 1 8 l a me l l a e ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segmen ts ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u nknown L O C U S TYP I C U S : Los A n g e l e s ( C a l i f . , U . S . A . ) T Y P I CAL MAT E R I A L : ho l o t y p e F ! i n WA . HOST S : u n k n o w n D I S T R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : L o s A n g e l es ( C a l i f . ) , WA ! M a r s h

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S eg m e n t 5 o f f r o n t t a r s u s w i t h d i s t a l p a r t w e a k I y b e n t ( F i g s 932 , 933 ) • • • • • • • • • • . • . . . . • • • • • • . • • . . • 2 S e g m e n t 5 o f f r o n t t a r s u s w i t h d i s t a l p a r t s t r on g l y b en t ( F i g s 934 , 9 3 5 , 936 , 940 , 9 4 1 ) . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . . . . 3 Me s o -me t a p l eu r a l s u t u r e o b s o l e t e ; b od y mo s t l y y e l l o w - f e r ru g i n o u s . . . . . . . . . . . . . . . . . 1 . citri n u s n . s p . Me s o -me t a p l e u ra l s u t u r e d i s t i n c t , c omp l e t e ; b o d y mo s t l y b l a ck • • . . • • . . . • • • • • • • • . . . • • . • 2 . arn a udi n . s p . Me s o -me t a p l eu r a l s u t u r e o b s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . longicornis ( K i e f f e r ) Me s o -me t a p l eu r a l s u t u r e d i s t i n c t , c omp l e t e • . . • • . . • • • • • • . • • • . • 4 Me t a t h o r a x + p r op od e u m w i t h a n t e r i o r s u r f a c e f u l l y s m o o t h , w i t h o u t s c u l p t u r e ; s eg m e n t 4 o f f r o n t t a r s u s s l i g h t I y s h o r t e r t h a n s eg m e n t 1 . . . . . . . . . . . . . . . 7 . bellottii n . s p . Me t a t h o ra x + p r o p o d e u m w i t h a n t e r i o r s u r f a c e a t I e a s t p a r t I y g ra n u l a t ed o r t ra n s v e r s e l y s t r i a t e ; s egmen t 4 o f f r o n t t a r s u s l on g e r t h a n s e g men t 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 B o d y mo s t l y b l a ck . . . . . . . . . . . . 5 . t ij u c a n u s ( A r l e ) B od y f u l l y o r m o s t l y t e s t a c e o u s o r r ed d i s h ; o c c a s i on a l l y f e r rug i n ou s • • • • . • • • • • • • • • • • • • • • • • • • . • • . . • . • • . • • . . • • . • • • • • • • • • 6 S e g m e n t 5 o f f r on t t a r s u s w i t h t w o r ow s o f l a me l l a e ( F i g . 9 3 4 ) ; b od y d u l l . . . . . . . . . . . . . . 3 . tes taceus ( C a m e r o n ) S eg m e n t 5 o f f r o n t t a r s u s w i t h 1 r ow o f l a me l l a e ( F i g . 9 3 5 ) b od y sh i n y • • • . • • • . • . . • • . • • . . • 4 . menkei n . s p .

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F E MALE : a p t e r o u s ; l e n g t h 3 , 37-4 , 37 m m ; y e l l ow-fer r u g i no u s , w i t h p e t i o l e b l ac k ; a n t e n n ae l o n g , s l e n d e r ; h e a d exc a v a te d , s h i n y , g r a n u l a t e d ; p ro n o t u m d u l l , crossed b y a s t ro n g t r a n s v e r s a l i m p re s s i on , g r a n u l a t ed ;

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F i g . 933 - C he l a of A p terodryin u s a rn a udi n .

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s u rf a ce a n d p a r t o f a n t e r i o r s u rface o f me t a t ho r a x + p ropodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u ra l s u t u re o b so l e te ; fore t a rs a l se g m e n t 1 shorter than s u b a p i ca l too t h w i t h 2 rows of l a me l l ae ; m a x i I

segmen t 4 ( 1 5 : 25 ) ; en l a rged c l a w ( F i g . 932 ) w i t h a n d 6-7 l am e l l ae ; segmen t 5 o f f ro n t t a r s u s ( F i g . 932 ) 1 2- 1 9 l am e l l ae ; a p e x w i t h a g ro u p of a t l ea s t 20-30 l a ry p a l p i w i t h 6 s e g men t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ;

1 362

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t i b i a l sp urs 1 , 0 , 1 . M A L E : u n k no w n LOCUS T Y P I C U S : S. Jose ( Co s t a R i c a ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n O L ; I p a r a t y p e F ! i n W A . HOST S : u n k no w n D I ST R I B U T I ON : Neo t rop i c reg i o n : COSTA R I CA : S . Jos e , O L ! C U B A : S . Rangel , WA!

2 . Ap terodryin u s arn a udi n . s p .

F E MA L E : a p te ro u s ; l e n g t h 3 , 75-4 , 37 m m ; head b l ack , w i t h c l y p e u s , ma l a r s p ace a n d m a n d i b l es y e l l ow ; a n t e n n a e b ro w n , w i t h s e g m e n t s 1 -2 a n d 6- 1 0 t e s taceo u s ; t ho r a x a n d p ropodeum b l ack ; a b d omen b l ack-b row n ; l eg s test aceo u s , w i t h coxae b l ack i s h ; h e a d exc a v a t e d , d u l l , g r a n u l a t e d ; p rono t u m d u l l , g r a n u l a ted , crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n ; s c u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; scu te l l u m i n v i s i b l e ; met a t ho r a x + p ropodeum d u l l , g ra n u l a t e d , w i t h n u merous l on g i t u d i n a l k ee l s o n pos t e r i o r s u rface a n d p l e u r a ; meso-me t a p l e u r a l s u t u r e d i s t i nc t , comp l e t e ; seg m e n t 1 o f f ro n t t a r s u s s horter t h a n s e g me n t 4 ( 1 0 : 1 5 ) ; e n l a r g e d c l a w ( F i g . 9 3 3 ) w i t h s u b a p i c a l too t h a n d 8 l ame l l a e ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 933 ) w i t h 2 row s of 1 4 + 1 4 l a m e l l ae ; apex w i t h a g ro u p of at l ea s t 40 l ame l l ae ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l ab i a l p a l p i w i t h 3 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Academy B a y ( Sa n t a C r u z I s l a n d , Ga l a p agos I s l a n d s ) TYP I CAL MAT E R I AL : H OST S : u n k n o w n

holotype F!

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3 . A p terodryinus tes ta ce u s ( C ameron )

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Gona top u s testaceus C a meron 1 888 : 440 . A go n a topoides jujuyen sis O g l ob l i n 1 95 1 : 488 ( n o s d y n . ) ; locus typ ic u s Cerro Pera l e s ( Argen t i n a ) .

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934 - C he l a of Ap terodryin u s tes taceus ( A rge n t i n a )

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scu t e l l u m ; m e t a t horax + propodeu m s h i n y , g r a n u l a t e d ; p l e u r a , pos ter i or s u rface of p ropode u m a n d p a r t of a n t e r i or s u rface t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i nc t , comp l e t e ; fore t a rs a l seg men t 1 shorter t h a n segment 4 ( 24 : 3 1 ) ; en l a rged c l a w ( F i g . 934 ) w i t h s u b a p i c a l t oo t h a n d 6-1 1 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 934 ) w i t h 2 rows of 1 9-37 l a m e l l ae ; apex w i t h a group of 20-40 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C US : Vo l c a n de C h i r i q u i ( P a n a m a )

1 364

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TYP I CAL M A T E R I AL : ho l o t y p e F ! of G . te s t a c e u s i n B M ; h o l o t y p e F ! o f A . juju ye n sis i n T U . HOST S : u n k no w n D I ST R I B U T I O N : Neo t rop i c reg i o n : PANAMA : Vo l c a n de C h i r i q u i , B M ! G a m boa ( C a n a l Zone ) , L E ! Taboga I s l a n d , WA ! B u g a b a , B M ! Per a l e s ( J u j u y ) , T U ! B u enos A i res , O L l

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coxae , c l u b s o f femora a n d fore t i b i ae b row n ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n ten n a l segme n t s i n fo l l ow i n g p roport i o n s : 9 : 6 : 2 3 :

15: 13:

1 0 : 8 : 5 : 5 : 9 ; h e a d s h i n y , exca v a ted , g ra n u l a ted ; fron t a l l i n e com p l e te ; occ i p i t a l ca r i n a i n comp l e t e , on l y v i s i b l e be h i n d t h e oce l l i a n d on t h e s i des of t he poste r i or oce l l i , b u t n e v e r reac h i n g t e m p l es ; POL � 2 ; O L 1 , 5 ; OOL � 1 0 ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rsa l i m p ress i on , s h i n y , w e a k l y g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted ; s c u te l l u m s hort , t r a n sverse ; m e t a n o t u m t r a n s v erse l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s p o i n te d ; m e t a t horax + p ropode u m s h i n y , w i t h p l e u r a a n d post e r i o r s u rf ace t r a n sverse l y s t r i a t e ; a n te r i or s u rface a l most w i t h o u t sc u l p t u r e ; meso-met a p l eu r a l s u t u re d i s t i nc t , com p l e t e ; fore t a r s a l segments in fo l l ow i n g p ropor t i o n s : 1 7 : 3 : 6: 20 : 30 ; e n l a rged c l a w ( F i g . 935 ) w i t h s U b a p i c a l too t h a n d 6 l am e l l ae ; segme n t 5 of front t a rs u s ( F i g . 935 ) w i t h 1 row of 1 3 l a m e l l ae ; a p e x w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : L a Lo l a ( C o s t a R i c a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n WA . H O ST S : u n k n o w n D I ST R I BU T I O N : Neo t rop i c reg i on : C O S T A R I CA : L a Lo l a , W A ! N OT E : t h e s p ec i e s i s n a med i n honor o f M r . A r no l d M e n k e .

5 . A p terodryi n u s tij u c a n u s

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Pseu dogon a top u s tij u c a n u s A r l e 1 935 : 5 2 .

F E MA L E : a p terou s ; l e n g t h 6 m m ; h e a d b l ac k , w i t h m a n d i b l es tes t aceou s ; c l y p e u s l a ter a l l y test aceou s , med i a l l y b row n ; a n te n n ae t e s t aceo u s ; t ho r a x a n d p ropodeum b l ac k , w i t h sc u t u m y e l l o w ; a b domen b ro w n - t e s t aceou s ; l eg s testaceou s , w i t h cox ae , troc h a n ters a n d femora p a r t I y b row n ; a n t e n n a e

1 366

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d i s t a l l y t h i c kened ; a n t e n n a I segmen t s i n fo l l ow i n g p ropor t i o n s : 1 0 : 8 : 32 : 20 : 1 7 : 1 2 : 8 : 7 : 5 . 5 : 9 ; head e x c a v a t e d , d u l l , g r a n u l a t e d , l o n g i t u d i n a l l y weak l y s t r i a t e ; f ro n t a l l i n e com p l e te ; occ i p i t a l c a r i n a i n com p l e te , on l y v i s i b l e b e h i n d t h e ocel l i a n d o n t h e s i des of t h e pos ter i or oce l l i , b u t never reac h i n g tem p l es ; pron o t u m s h i n y , crossed b y a s t ron g t r a n sversa I i m p ress i on , a s scu I p t u red a s head ; s c u t u m d u I I , g r a n u l a ted ; sc u te l l u m very sho r t , t r a n s v erse ; m e t a n o t u m n o t f l a t , n o t t r a n sverse l y s t r i a t e , w i t h t wo l a tera l po i n t s ; met a t ho r a x + p t�opodeum d u l l , g ra n u l a t e d , w i t h pos t e r i or s u rface s t ron g l y t r a n sverse l y s t r i a t e ; p l eu r a n o t t r a n s v erse l y s tr i a te ; meso - m e t a p l e u r a l s u t u re d i s t i n c t , com p l e te ; fore t a r s a l segmen t s i n fo l l o w i n g p ropor t i on s : 2 2 : 3 : 6 : 27 : 37 ; e n l arged c l a w ( F i g . 936 ) w i t h s u b a p i c a l too t h a n d 9-1 2 l a m e l l ae ; segme n t 5 of fron t t a r s u s ( F i g . 936 ) w i t h t w o row s o f nea r l y 3 6 l a me l l a e ; a p e x w i t h a g ro u p of n e a r l y 1 2 l a me l l ae ; m a x i l l a l'y p a l p i w i t h 6 segme n ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( F i g s 937 , 938 ) : f u l l y w i n ged ; l e n g t h 2 , 68 m m ; b l ac k ; m a n d i b l es t e s t aceo u s ; l eg s b row n , w i t h t a r s i l i g h t ; a n t e n n ae n o t d i s t a l l y t h i ck e n e d ; a n ten n a l segmen t s i n fol l ow i n g p roport i on s : 6 : 5 : 1 1 : 1 1 : 1 0 : 1 0 : 9 : 9 : 8 :

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8 ; a n t e n n a I s e g m e n t 3 more t h a n t h ree t i mes as l o n g as b ro a d ( 1 6 : 3 , 5 ) ; h e a d d u l l , g .' a n u l a t ed ; f l'o n t a l l i n e a b s e n t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 6 , 5 ; O L = 2 , 5 ; OOL = 4 ; t em p l e s w e a k l y d i s t i n c t ; sc u t u m d u l l , g r a n u l a te d ; n o t a u l i ces com p l e te , p o s t e r i o r l y separ�a t e cJ ; m i n i m u m d i s t a n c e b e t w e e n t h e n o t a u l i c es as l o n g as b r e a d t h of t h e oc e l l i ; sc u t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , weak l y p u n c t a t e , w i t ho u t scu l p t u re a m o n g p u n c t u re s ; p ro p o d e u m d u l l , f u l l y ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t ho u t d a rk t ra n s v e rsa l b a n d s ; r a d i a l c e l l c l osed : d i s t a l p a r t of r a d i a l v e i n t w i ce a s l o n g a s p ro x i m a l p a r t ( 24 : 1 2 ) ; gonoforceps w i t h a s h o r t dors a l p .'ocess ( F i g . 939 ) , w i t h a p e x b ro a d ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : P i co d a T i j u c a ( 3 r az i l ) ( ho l o t y p e ) ; F l o res t a d a T i j u c a ( B r a z i I ) ( n eo t y p e ) TYP I CA L MAT E R I AL : ho l o t y p e F l o s t ( p erson a l com m u n i c a t i on o f M r . A r l e ) ; n eo t y ;:> e F ' ( h ereby d es i g n a te d ) i n T W . HOST S : u n k n o w n D I STR I B U T I O N : Neo t rop i c reg i o n : BliAZ I L : F l ores t a d a T i j u c a ( G u a n a b a .'a ) , T W ' P i co d a T i j uc a ( G u a n a b a r a ) ( A r l e 1 935 ) .

6 . A p terodryin u s longicornis

( K i effer ) n . comb .

Gon a topus longicornis K i effer 1 904a : 357 . Dicondylus longicol'n is ( K i effer ) : K i ef f e r 1 9 1 4 b : 78 . Dolichochelys longicornis ( K i effer ) : O l m i a n d C u r-rado 1 976a :

1 78 .

1 368

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F E MALE : ap terou s ; l e n g t h 5 m m ; y e l i ow - t e s t a c eo u s , w i t h p e t i o l e b l ac k ; a n te n n a l segme n t s i n fo l l o w i n g p ro p o r t i o n s : 8 : 8 : 3 1 : 23 : 1 8 : 1 4 : 9 : 8 : 7 : 1 0 ; h e a d s h i n y or d u l l , smoo t h , h a i r l es s , w i t ho u t scu l p t u r e , excep t for occ i p u t and p a r t o f v e r t e x g ra n u l a te d ; POL � 1 , 5 ; OL � 2 , 5 ; OOL � 1 3 ; p rono t u m crossed b y a s t ro n g t r a n s vers a l i m p ress i o n , s h i n y , h a i r l e ss , g r a n u l a t e d ; sc u t u m d u l l , g r a n u l a te d , w i t h t w o l a t e ra l , p o i n te d protrus i o n s ; scute l l um weak l y v i s i b l e ; met anotum s h i n y or d u l l , not h o l l o w b e h i n d t h e sc u t e l l u m , t r a n sverse l y s t r i a t e ; m e t a t h or a x + p ro p o d e u m s h i n y o r d u l l , w i t h o u t s c u l p t u re ; p l e u ra , pos t e r i o r s u rface of m e t a t h o r a x + propo d e u m a n d p a r' t of a n t e r i o r s u rface t r a n sverse l y s t r i a t e ; meso m e t a p l e u ra l s u t u re o b so l e t e ; fore t a r s a l segmen t s i n fo l l o w i n g p roport i o n s : 20 : 3 : 5 : 25 : 37 ; e n l a r g e d c l a w ( F i g . 940 ) w i t h s u b a p i c a l t oo t h a n d 7 l a me l l a e ; se g m e n t 5 of fro n t t a r' s u s ( F i g . 940 ) w i t h t w o rows of 29 l a me l l ae ; a pex w i t h a g r o u p of 25 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u nk nown L O C U S T Y P I C U S : Coro i co a n d C h i l u m a n i ( Bo l i v i a )

Gon a topod i n ae

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H O ST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : BRAZ I L : Nov a T e u t o n i a ( S . t a C a t a r i n a ) , BM ' B OL I V I A : Coro i co a n d C h i l u m a n i , G E !

7 . A p terodryin u s bellottii n . s p .

1 370

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F E MAL E : a p terou s ; l e n g t h 4 , 8 m m ; head b row n , w i t h m a n d i b l e s , c l y p e u s a n d a n te r i or re g i o n of f ro n s t e s t a c eo u s ; i n a d d i t i o n l a te r a l reg i o n s o f f r o n s ( more a l o n g orb i t s ) testaceo u s ; a n te n n a e b row n , w i t h seg m e n t s 1 a n d 8- 1 0 y e l l ow ; propec t u s redd i s h - t e s t aceo u s ; pro n o t u m redd i s h testaceo u s , w i t h t r a n sv e rs a l i m p ress i o n d a r k ; scu t u m red d i s h - t e s t aceo u s ; sc u te l l u m , m e t a t ho r a x + propo de u m , mesop l e u r a a n d m e t a p l eu r a b l ac k ; a b domen t e s t a c eo u s ; l eg s t e s t aceo u s , w i t h p a r t of coxae a n d p a r t of c l u b s o f femora b l ac k ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t en n a l segmen t s i n f o l l ow i n g propor t i o n s : 1 4 : 8 : 24 : 1 5 : 1 1 : 1 1 : 9 : 9 : 9 : 1 3 ; h e a d v ery exc a v a t ed , s h i n y , smoo t h , w i t ho u t scu l p t u re , w i t h t r a c k s o f t r a n sversa l , short k ee l s o n a n te r i or h a l f of f r o n s a n d o n t h e s i des o f fron t a l l i n e ; fron t a l l i n e comp l e te ; occ i p i t a l c a r i n a i ncomp l e t e , o n l y v i s i b l e b eh i n d t h e oce l l i ; P O L = 2 ; O l = 4 ; O O l = 1 2 ; p os t e r i o r oce l l i touched b y t h e t r a c k o f occ i p i t a l c a r i n a ; p ron o t u m w i t h a s t ron g t r a n s v e rs a l i m pres s i o n , s h i n y , smoo t h , w i t ho u t sc u l p t u re ; sc u t u m a n d sc u te l l u m s h i n y , smoo t h , w i t h o u t scu l p t u re ; s c u t u m w i t h a l a t e r a l s t r o n g p o i n t ; meso-m e t a p l eu r a l s u t u re d i s t i n c t , comp l e t e ; m e t a n o t u m reduced to a short t r a n s v e rs a l k ee l , wea k l y v i s i b l e b e h i n d t he sc u te l l u m ; m e t a t h o r a x + p ropode u m s h i n y , smoot h , w i t h o u t sc u l p t u re , except for t r a n svers a l st r i ae o n p l e u ra a n d pos t e r i o r s u r f a c e ; fore t a r s a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 25 : 5 : 8 : 24 : 38 ; e n l a rged c l a w ( F i g . 9/+ 1 ) w i t h s u b a p i c a l too t h a n d 7 l a m e l l ae ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 941 ) w i t h t w o rows o f n e a r l y 1 8 l a me l l ae ; apex w i t h a g r o u p o f n e a r' l y 9 l am e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n l O C U S T Y P I C U S : R o a d F ac a t a t i v a -Ano l a i m a ( C o l omb i a ) TYP I CAL M A T E R I AL : h o l o t y p e F ! i n A M . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : ( C u n d i n a m a rc a ) , AM ! NOTES:

COLOMB I A :

R o a d F ac a t a t i v a-Ano l a i m a

t h e spec i es i s n a med i n honor of M r .

Tony Be l l o t t i .

AUSTRAL I A N S P E C I ES OF A PTEPODl2Y1NUS

T w o spec i es are k no w n .


F E MA L E S

Me s o -me t a p l eu r a l t h e s c u t e l l um •

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I ns t . ,

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Me s o -me t a p l e u r a l s u t u r e v e r y s t r o n g and d i s t i n c t h o l l ow b e h i n d t h e s c u t e l l u m . 1 . n o togeicus n . sp .

me t a n o t u m

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1 . A p terodryin u s notogeicus n .

sp.

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I ns t . ,

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37 ,

1 984

re g i on of f ro n s g r a n u l a t e d ; fron t a l l i n e com p l e t e ; POL = 2 ; OL = 3 ; OOL 9; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m sh i n y , crossed b y a s t ro n g t ra n s ve rs a l i m p ress i o n , w e a k l y t r a n sverse l y s t r i a t e ; scu t u m s h i n y , w i t h w e a k l on g i t u d i n a l k ee l s ; sc u te l l u m smoo t h , s h i n y , w i t ho u t sc u l p t u r e ; m e t a no t u m t r a n s v erse l y s t r i a t e , ho l l o w b e h i n d t he s c u te l l u m ; meso m e t a p l e u r a l s u t u re d i s t i n c t , com p l e te ; m e t a t ho r a x + p ropodeum s h i n y , w i t ho u t sc u l p t u re , w i t h p os t e r i o r s u rface t r a n sverse l y s t r i a t e ; a n te r i or s u rface w i t h a w e a k med i a n f u r ro w ; mesop l e u r a n o t t r a n s verse l y s t r i a te ; m e t a p l e u r a t r a n sverse l y s t r i a te ; fore t a rs a l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 1 8 : 3 : 6 : 1 6 : 2 7 ; e n l a r ged c l a w ( F i g . 942 ) w i t h s U b a p i c a l too t h a n d 9 l a m e l l ae ; segmen t 5 o f fro n t t a rs u s ( F i g . 942 ) w i t h 1 row of 1 3 l a m e l l ae ; apex w i t h a g ro u p of 1 7 l a m e l l a e ; m a x i I l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I C U S : Perth ( A u s t ra l i a ) T Y P I CA L MAT E R I AL : ho l o t y pe F ! i n B M ; 1 p a r a t y p e F ! i n O L . HOSTS : u n k n o w n D I ST R I BU T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : P e r t h ( Western A u s t r a l i a ) , BM !

2 . A p terodryin u s insul aris n . s p .

F E MAL E : a p terou s ; l e n g t h 2 , 5 m m ; y e l l o w - t e s t aceou s , w i t h p e t i o l e b l ac k a n d a b domen b row n i s h ; a n te n n a l segme n t s i n fo l l ow i n g p roport i o n s : 8 : 5 :

1 1 : 6 , 5 : 5 : 4 : 5 ( l a s t t h ree segmen t s m i s s i n g i n t he o n l y k n o w n spec i me n ) ; h e a d s h i n y , exca v a ted , h a i r l ess , w i t h o u t scu l p t u re , exce p t for o cc i p u t a n d f ro n t p a r t of v e r t e x weak I y g r a n u I a ted ; pron o t u m c rossed b y a s t r o n g t r a n s v e rs a l i mp ress i o n , s h i n y , w i t ho u t s c u l p t u re ; scu t u m s h i n y , w i t ho u t sc u l p t u re , l a tera l l y w i t ho u t po i n t s ; scu t e l l u m w e a k l y v i s i b l e ; m e t a n o t u m n o t ho l l o w beh i n d t he sc u te l l u m ; m e t a t ho r a x + propodeum s h i n y , w i t ho u t sc u l p t u re ; p l eu r a and p os t e r i o r s u rf ace of propodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 of f ro n t t a rs u s a s l o n g a s segmen t 4 ; e n l a rged c l a w ( F i g . 943 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; seg m e n t 5 o f f ro n t t a rs u s ( F i g . 943 ) w i t h t wo ro w s of 2 + 1 3 l a m e l l ae ; a pex w i t h a g ro u p o f a t l ea s t 1 6 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C U S : L a r a t I s l a n d ( I n dones i a ) TYP I C A L M A T E R I A L : ho l o t y p e F ! i n B . HOST S : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n reg i o n : I N DONES I A : L a r a t I s l a n d ( T a n i m b a r I s l an d s ) , B !

9 . Dicondylus H a l i d a y

Gon a topod i n a e

1 3 75

nec L a beo C u v i e r 1 8 1 7 : 1 94 . Dicondyl u s H a l i d a y i n C u r t i s 1 8 29-30 : 1 1 0 . Dicondyl u s H a l i d a y 1 833 : 2 73 . L a beo H a l i d a y 1 83 3 : 273 ( s y n . p roposed by R i c h a r d s 1 937 ) ; t y p e s p ec i e s L a beo v i tripennis H a l i d a y 1 83 3 , monot y p i c . n ec L a bea B l a n c h a r d 1 840 : 430 . L a beola H a l d e m a n 1 84 2 : 1 92 ( s y n . p roposed b y K ro m b e i n 1 979 ) , new name. La bea B l a n c h a r d : D a l l a Torre 1 898 : 542 . L a beo H a l i d a y : K i effer i n K i ef f e r et M a r' s h a l l 1 905 : 220 . Trigo n a topus K i effer 1 9 1 3c : 325 ( n . sy n . ) ; t y p e spec i es Trigon a top u s b ifarius ( K i ef f e r 1 906 ) , o r i ,) . d es i ,] . L a � erius I( i e ffer 1 9 1 4 b : 59 ( s y n . p roposed b y R i c h a rd s 1 9 37 ) ; new name. Trigon a topus !< i effer : K i ef f e r 1 9 1 4b : 7 2 . Dicondyl u s H a l i d a y : K i ef f e r 1 9 1 4b : 7 2 . L a berinus O g l ob l i n 1 932 : 266 ( m i s p e l l i n g o r l a p s u s ) . Dicondylus H a l i d a y : R i c h a r d s 1 935 : 1 47 . L a berius K i effer : R i c h a r ds 1 9 36a : 1 69 . Dicon dyl u s H a l i d a y : R i c h a r d s 1 937 : 1 1 8 . Idologon a topus O g l ob l i n 1 95 3 : 1 1 1 ( n . s y n . ) ; t y p e spec i es Idologon a top u s nigrithorax O g l o b l i n 1 95 3 , o r i g . des i g .

TYPE S P E C I E S : Dicondylus bicolor ( H a l i d a y 1 828 ) , d e s i g . b y K i effer ( 1 9 1 4b ) • F E MA L E ( F i g s 947 , 948 ) : a p terou s ; m a x i l l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; p r o n o t u m ( F i g . 948 ) n o t crossed b y a s t r o n g t r a n s v e rs a l i m p ress i o n or w ea k l y i m p ressed ; e n l a rged c l a w ( F i g . 944 ) w i t h s u b a p i c a l too t h a n d l a m e l l ae ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i g s 952 , 953 ) : f u l l y w i n g ed ; m a x i l l a r y p a l p i w i t h 2-4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen ts ; tem p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d the oce l l i o r o n t h e s i des o f t h e poste r i o r oce l l i , b u t n e v e r reac h i n g temp l e s ; occ i p u t con c a ve ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N : wor l d w i de HOST S : Delp h a cidae SPEC I ES : 1 3

PALAEARCT I C S P E C I E S O F D ICOND YLUS

T w o s p ec i e s are k n ow n .


1 37 6

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F E MA L E S

Me t a t h o ra x + p r o p od e u m , s c u t e l l u m a n d p a r t o f t h e f i r s t a b d om i n a l t e rg i t e r e d d i s h o r y e l l ow - r ed d i s h . . . . . . . . . . . . . . . . . . . dich rom u s ( K i e f f e r ) · • • • • . • • . • . . . . . • • . . . . • . . • • • • • 1 . Me t a t h o r a x + p r o p od e u m , s c u t e l l u m a n d f i r s t a b d om i n a l t e rg i t e b l a c k ( e x c e p t f o r t h e r ed d i s h a p ex o f p r o p o d eu m ) • • . . • . • • • . • . • • ·

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b icolor ( H a I i d a y )

MALES

M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s n e a r l y a s l o n g a s b r ead t h o f t h e o c e l l i ( F i g . 9 4 5 B ) . . • • • • • . . • • • • • . . • • • • • • • • • . · • . . . . . • • . . . • . . , . . . . . . . • . . . . . 2 . b icolor ( Ha I i d a y ) M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s s h o r t e r t h a n b ,' ea d t h o f t h e o c e l l i ( F i g . 9 4 5 A ) • • • l . dichrom u s ( K i e f f e r )

1 . Dicondyl u s dichrom u s

( K i ef f e r )

Gon a to p u s dichrom u s K i effer i n K i e f f e r et M a rs h a l l 1 906 : 505 . Dicondylus dichrom u s ( K i ef f er ) : Ber l a n d 1 928 : 1 46 . Gon a to p u s rufescens H e l l e n 1 93 5 : 8 ( s y n . p roposed b y He l l en

1 95 3 ) ; nomen n u d u m . D i c ondyl u s helleni R a a t i k a i n e n

1 96 1 : 1 26 ( n . s y n . ) ; locus ( F i n l and ) . Dicondy l u s dichrom u s ( K i effe r ) : R a a t i k a i n en 1 96 1 : 1 3 1 . typ i c u s L a i h i a

D i c on dy l u s s i b iric u s Ponomarenko 1 965 : typ i c u s R o m a n o v k a ( S i b i r i a , U . S . S . R . ) .

624

( n . sy n . ) ; locus

( K i ef f er ) : Moc z a r 1 965 : 393 . R a a t i k a i ne n 1 97 0 : 3 2 1 . ffaplogon a top u s p a c e i C u rr a d o a n d O l m i 1 97 7 : 3 1 7 ( n . s y n . ) ; l o c u s typ i c u s G r i so l i a ( I t a l y ) . Pseu dogon a topus rufu s Ponomaren k o 1 97 9 a : 355 ( n . s y n . ) ; l o c u s typ i c u s 2 5 K m N of B u l g a n ( Mo n g o l i a ) . Dicon dyl u s dichrom u s

Dicondylus helleni R a a t i k a i n en :

( K i ef f er ) : Mocza r 1 983b : 206 . ( R a a t i k a i n e n ) : Moc z a r 1 983b : 206 .

Dicondyl u s dichrom u s Dicondy l u s helleni

F E MAL E : a p t e rou s ; l e n g t h 2 , 50-3 , 25 m m ; red d i s h or y e l l ow - t e s t a ceou s , w i t h a n te n n a I se g m e n t s 3-9 b row n ; occ a s i on a l l y ver tex o f head b row n ; p e t i o l e b l ac k ; a b domen red d i s h - y e l l ow o r b ro w n - b l ac k , occ a s i o n a l l y w i t h f i rs t t e r g i t e red d i s h ;

h e a d d u l l , exc a v a ted ,

g r a n u l a ted ;

p ron o t u m n o t

.

•

1 37 7


P/

/ //// /

F i g . 944 - C he l a o f Diconclyl u s dichrom u s

( K i e f fe r ) f r o m L a i h i a ( F i n l a n d )

c r'osseu

b y a s t ro n g t r a n s v e rs a l i m p ress i o n o r we a k l y i m p ressed , d u l l , g r a n u l a t e d ; s c u te l l u m d u l l , w i t ho u t scu l p t u re ; met a n o t u m w i t h s i de s p r o t r u d i n g ; p ro t r u s i on s po i n ted ; met a t h o r a x + p ropodeum d u l l , g ra n u l a t ed , w i t h pos t er i or s u rface a n d p l e u ra t r a n s verse l y s t r i a t e ; meso me t a p l e u ra l s u t u re obso l e t e ; seg m e n t 1 o f f ro n t t a rs u s a p p rox i m a te l y t w i ce as l o n g as segmen t 4 ( 1 2 : 6 ) ; en l a r g e d c l a w ( F i g . 944) w i t h s u b a p i c a l too t h a n d 4-5 l ame l l a e ; segmen t 5 of f ron t t a rs u s ( F i g . 944 ) w i t h 2 rows o f 6-9 l a me l l ae ; a p e x w i t h a g ro u p o f 3-6 l a me l l ae ; m a x i l l a ry p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : f u l l y w i n ged ; l e n g t h 2 , 37-2 , 62 mm ; b l ac k ; m a n d i b l e s y e l l ow ; a n te n n a e a n d t e g u l ae b ro w n ; l e g s b ro w n - y e l l ow ; a n t e n n a I s e g me n t 3 f o u r t i mes or m o r e t h a n f o u r t i mes a s l on g a s b ro a d ( 5 , 25 ) ; h e a d s h i n y , h a i r y , g r a n u l a te d ; f ro n t a l l i n e a b s e n t ; t em p l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; occ i p u t conc a v e ; sc u t u m s h i n y , g r a n u l a ted ; n o t a u l i ces d i s t i n c t , comp l e te , pos t e r i or l y sep a r a ted ; m i n i m u m d i s t a nce b e t ween t h e n o t a u l i ces ( F i g . 945 A) m u c h s ho r t e r t h a n b r e a d t h o f the oce l l i ; s c u t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , p u n c t a t e ; p ropodeum ret i c u l a t e ru gose , w i t h o u t k ee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n s v e rs a l b a n d s ; dors a l p rocess o f gonoforceps ( F i g . 946 A ) w i t h p a d d l e- s h a p ed a p e x ; m a x i I l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S :

Corse ( F r a n c e )

1 378


I ns ! . ,

no . 37 ,

1 984

F i g . 945 - N o t a u l i ces of m a l es o f Dicondylus dich rom u s ( K i ef f e r ) f rom Csevharaszt ( H u ngary ) ( A )

a n d b icolor ( H a l i d a y ) f rom Germ a n y

(B).

T Y P I C A L M A T E R I AL : h o l o t y p e F ! o f G . dichromus i n B U ; h o l o t y p e F ! a n d 3 p a r a t y p e s ( 1 F , 2 M M ) ! of D. helleni i n H E ; o t h e r p a r a t y pes of D . hellen i i n H E a n d i n R a a t i k a i n e n ' s col l ec t i o n ; ho l o t y pe F ! a n d a l l o t y p e M ! of D . sibiric u s i n MO ; 2 p a ra t y pes ( 1 F , 1 M ) ! o f D . s i b iric u s i n B�� ; h o l o t y p e F ! of H. p a ce i i n V R ; h o l o t y p e F ! of Ps . rufus i n L N . HOST S : i n F i n I a n d Megadelp h a x sordidulus ( S t a I ) , Un k a nodes excisa ( Me l . ) ( = Delp h a x excisu s ) ( Delp h a c ida e ) ( R a a t i k a i ne n 1 960 , 1 96 1 , 1 970 ) . B I OLOGY : e g g , l a rv a , cocoon , p u p a a n d l i fe cy c l e desc r i b e d b y R a a t i k a i nen ( 1 96 1 ) . N A T U RAL E N E M I ES : i n F i n l a n d A c hol'olop h u s gracilipes ( K ramer ) ( A carin a , Erythraeida e ) ( R a a t i k a i n e n 1 96 1 ) . D I ST R I B U T I ON : P a l a e a rc t i c reg i on : M O N GOL I A : 2 5 K m N of Bu l g a n ( R i ve r U I j a s t a y n-Go l , H o v d a y m a g , W e s t Mon g o l i a ) , L N ! 1 1 K m W f r o m Somon Baj a n n u u r ( Bu l g a n a i m a k ) , B U ! 3 K m N from Somon Uenc i n T a l U e n c ( C hovd a i m a k ) , B U ! Somon Baj a n zo g t ( C en t r a l a i m a k ) , B U ! U l a n - B a a to r ( C en t r a l a i m ak ) , B U ! U . S . S . R . : Romanov k a ( Ye r a v n e s n k i y D i s t r i c t , B u r y a t A . S . S . R . , S i b i r i a ) , B M ! WA ! M O ! F I NLA N D : L a i h i a ( V a a s a n ) , H E ! T v a rm i n n e , H E ! H a n go ( U u de n ma ) , H E ! H e n r i k sb e r g ( R a a t i k a i ne n 1 96 1 ) ; K a u n i a i ne n ( R a a t i k a i n en 1 96 1 ) ; T a i p a l sa a r i ( R a a t i k a i n en 1 96 1 ) ; SWEDE N : A n ge l ho l m ( K r i s t i a n s t a d ) , H E ! H U NGAR Y : M a t ra sz t i m re , B U ! Csev h a r a sz t , B U ! V a l ko , B U ! B u k k-h g s . , B U ! Hosszu herc ( B u k k ) , B U ! G E RMANY : B er l i n , E B ! A U ST R I A : Vos l a u , P ! D E N M A R K , C O ! HOLLA N D : T e rsche l l i n g , L E ! SW I T Z E R L AN D : Vercor i n ( V a l a i s ) , LO ! Les P o n t s ( N e u c h a te l ) , LO ! F R A N C E : Corse , B U ! SPA I N : Sa l ou ( T a rr a g on a ) , BM ! O L ! I TALY : Bu sso l en o ( T or i n o ) , O L ! G r i so l i a ( Cosenza ) , V R !

2 . Dicondylus b icolor ( H a l i d a y )

Dryin u s b icolor H a l i d a y 1 828 i n C u r t i s 1 828 : 206 . Dicondylus bicolor H a l i d a y i n C u rt i s 1 829-30 : 1 1 0 . L a beo vitripennis H a l i d a y 1 833 : 273 ( s y n . p roposed b y R i c h a rds 1 939 ) ; locus typicus I r e i a n d . A n t aeon excis u s Wes t wood 1 833b : 497 ( s y n . proposed by R i c h a rd s 1 939 ) ; locus typicus C a m b r i d g e ( E n g l a n d ) .

Gon a topod i n a e

Fig.

1 379

946 - M a l e g e n i t a l i a of Dicondylus dichromus ( K i effe r ) from L a i h i a

( F i n l an d ) ( A )

a n d b icolor ( H a l i d a y ) from Born t a l removed )

( Ge r m a n y ) ( 3 ) ( r i g h t h a l f

Gon a topus conjunctus K i effer i n K i effer et M a r s h a l l 1 90 5 : 1 1 9 ( n . s y n . ) ; locus typ i c u s S t a v a n ger ( Norw a y ) . Gon a top u s b ifarius K i effer i n K i effer e t M a r s h a l l 1 906 : 502 ( n. s y n . ) ; locus typ i c u s Corse ( F r a n ce ) . Gon a top u s b icolor ( H a l i d a y ) : K i effer i n K i effer e t M a r s h a l l 1 906 : 50S . C a b eo excisus ( We s t wood ) : K i effer 1 907b : 1 5 . Pla tygo n a topus b ifa rius ( K i effer ) : K i effer 1 907b : 1 9 . Trigon a top u s b ifarius ( K i effer ) : K i effer 1 9 1 3c : 325 . Dicondylus b icolor ( H a l i d a y ) : K i effer 1 9 1 4b : 7 3 . Gon a topus decretorius H a u p t 1 9 1 6 a : 4 6 ( n . s y n . ) ; loc u s typ ic u s S w i n e m u n d e ( Ge r m a n y ) . L a berius excis u s ( We s t wood ) : B er l a n d 1 925 : 1 45 . Trigon a top u s b ifa rius ( K i e ffer ) : Ber l a n d 1 92 5 : 1 45 . L a berius vitripennis ( H a l i d a y ) : R i c h a rd s 1 935 : 1 45 . Dicondylus b icolor ( Ha l i d a y ) : R i ch a rds 1 9 35 : 1 47 . Dicondylus b icolor ( H a l i d a y ) : B e h r i sch 1 95 2 : 34. D icondylus lindbergi H e i k i n he i mo 1 957 : 78 ( n . sy n . ) ; locus typicus L a i h i a ( F i n l a nd ) . Dicondyl us lindbergi H e i k i n h e i m o : R a a t i k a i n en 1 96 1 : 1 3 1 . Dicondylus b icolor ( H a l i d a y ) : R a a t i k a i n e n 1 96 1 : 1 3 1 . Haplogon a top u s b ifa ri u s ( K i effer ) : Moc z a r 1 965 : 391 .

1 380

Mem . A m e r . E n ! .

I ns!. ,

n o . 37 ,

1 984

F i g . 947 - F e m a l e of Dicondyl u s bicolor ( H a l i d a y ) from L a i h i a

(Fin land)

Dicondyl u s lindb ergi H e i k i n h e i mo : R a a t i k a i n e n 1 967 : 88 . Gon a topus conjunctus K i effe r : R a a t i k a i n e n 1 967 : 88 . Dicondylus b icolor ( H a l i d a y ) : N a g y 1 9 67 a : 1 24 . Haplogon a top u s b ifarius ( K i effer ) : C u r r a d o a n d O l m i 1 974b : 4 1 . Dicondyl u s b icolor ( H a l i d a y ) : W a l off 1 974 : 1 04 . Dicondyl u s b icolor ( H a l i d a y ) : W a l of f 1 97 5 : 662 . Dicondyl u s b icolor ( H a l i d a y ) : J . F . P e r k i n s 1 976 : 3 1 .

F E MALE ( F i g s 947 , 948 ) : a p t e ro u s ; l e n g t h 2 , 75-3 , 00 m m ; h e a d b ro w n b l ac k , w i t h m a n d i b l e s , c l y pe u s a n d f ro n t p a r t of v e r t e x ( more a l o n g o rb i t s ) y e l l o w ; a n te n n a e b row n , w i t h segme n t s 1 -2 a n d occ a s i o n a l l y 1 0 y e l l o w o r w h i t i s h ; p rono t u m f u l l y y e l l ow-tes t a ceou s ; occ a s i o n a l l y co l l a r b l ac k a n d d i sc y e l l o w ; r a re l y pro n o t u m a l mo s t f u l l y b l ac k ; sc u t u m y e l l o w , w i t h s i des b l ac k ; sc u te l l u m a n d m e t a t ho r a x + p ropod e u m b l ac k , w i t h a p e x of p ropode u m y e l l o w ; a b domen b l ac k ; l eg s y e l l o w or y e l l ow b row n , occ a s i o n a l l y w i t h c l u b s o f femora f u l l y o r p a r t l y b row n i s h ; head e xc a v a ted , g r a n u I a t e d ; p ro no t u m n o t crossed by a s t ro n g t ra n s versa I i m p res s i o n or w e a k l y i m p ressed , s h i n y , g r a n u l a te d ; scu t u m g r a n u l a t e d ;

Gon a topod i n ae

1 38 1

F i g . 9 4 8 - F em a l e of Dicondyl u s bicolor ( H a l i d a y ) f rom L a i h i a

met a n o t u m t r a n s v e rs e l y s t r i a te ,

w i t h s i des p r o t r u d i n g ;

( F i n l an d )

p ro t r u s i o n s

p o i n t ed ; pos t e r i o r s u rface of p ropodeum a n d p l eu ra t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; occ a s i o n a l l y d i s t i n c t , b u t v e r y w e a k ; fore t a rs a l s e g m e n t 1 a p p ro x i m a te l y t w i ce as l o n g as segmen t ' 4 ; en l a r g e d c l a w ( F i g . 949 ) w i t h s u b a p i c a l too t h a n d 4 - 5 l a m e l l ae ; segme n t 5 of f ro n t t a rs u s ( F i g . 949 ) w i t h t w o row s o f 6-9 l ame l l ae ; a p e x w i t h a g ro u p o f 3-6 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 2-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . Ivt" L ,, : f u l l y w i n ged ; l e n g t h 1 , 57-2 , 00 mm ; h e a d b l ac k , w i t h m a n d i b l es y e l l o w ; a n te n n ae b l a c k - b ro w n , w i t h p a r t of seg men t s 1 -2 y e l l ow ; t ho r a x a n d p ro p o d e u m b l ack ; a b domen b l a c k - b ro w n ; l eg s y e l l o w , w i t h p a r t o f c o x a e a n d p a r t of femora b row n ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a l se g m e n t s i n fo l l o w i n g p ropor t i o n s : 5 : 4 , 5 : 7 : 8 : 7 , 5 : 7 , 5 : 6 , 5 : 6 , 5 : 6 : 8 , 5 ; a n t e n n a l s e g m e n t 3 f o u r t i mes o r more t h a n fou r t i mes a s l o n g a s b ro a d ( 4 , 0 ) ; h e a d s h i n y , g r a n u l a t ed ; fron t a l l i n e a b sen t ; t em p l es d i s t i n c t ; no s h i n y , smoo t h a re a i s v i s i b l e on t h e s i des of t he poster i o r oce l l i ; occ i p i t a l ca r i n a i n v i s i b l e ; scu t u m s h i n y ; n o t a u l i ces com p l e t e , poste r i o r l y sep a r a ted ; m i n i m u m d i s t a n c e b e t ween t he n o t a u l i ce s ( F i g . 945 B ) n e a r l y a s l o n g a s b r ea d t h of the oce l l i ; s c u te l l u m a n d met a n o t u m s h i n y , w ea k l y p u n c t a t e ; p ropodeum re t i c u l a t e r u gose ; fore-w i n g h y a l i ne , w i t h o u t d a rk t r a n s v e rs a l b a n d s ; dors a l p rocess of g o n oforceps ( F i g . 946 G ) w i t h p a d d l e- s h a p e d a p e x ; m a x i I l a r y p a l p i w i t h 3 segme n t s ; l ab i a l p a l p i w i t h 2 segmen ts ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S T Y P I C US : u n c er t a i n ( D e a l i n E n g l a n d accord i n g t o Mocz a r 1 965 ; D a ren t h Wood i n E n g l a n d accord i n g to R i c h a r d s 1 939 ) . T Y P I C A L MAT E R I AL : ho l o t y p e F ! a n d 1 p a r a t y p e F ! of D . b icolor i n D U ; ho l o t y p e F ! of G. conjunctus i n P ; h o l o t y p e F ! a n d 1 p a ra t y pe F ! of D . lindb ergi i n HE ; t he a l l ot y p e M ! o f D . lindb ergi i n H E i s rea l l y a m a l e

1 382

F i g . 949

M e m . Amer .

-

Ent.

I ns t . ,

n o . 37 ,

1 984

C he l a of Dicondylus bicolor ( Ha l i d a y ) from L a i h i a ( F i n l a n d )

Go n a topo d i n a e

1 383

o f Pseu dogon a top us dis tin c t u s ( K i effe r ) ; ho l o t y p e F ! of G . b ifarius i n BU ; h o l o t y p e M ! of L . vi tripe n n is i n D U ; h o l o t y p e M of A . excisus i n B M . HOST S : i n F i n l a n d lavesella pellucida ( F . ) ( H ek i n h e i mo 1 957 ; R a a t i k a i n en 1 967 ) , lavesella o b s c urella ( Boh . ) , Megadelp h a x sordidu l u s ( S t a l ) , Dicra n o tropis h a m a t a ( Boh . ) , Stiroma 'J icarina ta ( H . S . ) , Criom orp h u s a l b om argin a tu s C u r t . ( R a a t i k a i n e n 1 967 ) , Un k a n odes excis a ( M e l . ) ( = Delph a x excis u s ) ( L i n db e r g 1 950 ) ; i n I ta I y D icranotropis h a m a t a ( Bo h . ) ( C u rrado a n d O l m i 1 974b ) ; i n E n g l a n d lavesella pellucida ( F . ) , Criomorph u s albomargin a t u s C u r t i s , Hyledelp h a x elega n t u l u s ( Boh . ) ( W a I off 1 975 ) ; Dicra n o tropis h a m a t a ( Bo h . ) ( H a s s a n 1 93 9 ; W a l of f 1 975 ) ; D i tropis p teridis ( Sp i n . ) ( R i c h a rd s 1 933 ) ; Rib a u todelp h a x s p . ( = Delp h a codes coll inus Bo h . ) ( R i c h a rd s 1 93 9 ) ; Gra vesteiniella boldi ( Scot t ) ( R i c h a r d s 1 948 ) ; i n F r a nce Delp h a x collin a e Bo h . ( M a n ev a l 1 93 5 ) a n d Stiroma sp . ; i n Germ a n y Criomorp h u s b ic arin a tu s ( H . S . ) ( S t r u b i n g 1 95 6 ) , Un k a n odes excisa ( Me l . ) ( = Delp h a x excis u s ) ( H a u p t 1 9 1 6a ) ; i n Be l g i u m I2ib a u todelph a x p u n gens ( R i b a u t ) ( C . F . M . d e n B i e m a n l e g . ) ; i n Ho l l a n d Dicra n o tropis h a m a ta ( Bo h . ) , Rib a u todelp h a x collin u s ( Boh . ) , Delp h a cinus mesomelas ( Bo h . ) ( C . F . M . den B i e m a n l e g . ) . B I OLOGY : e g g , l a r v a , cocoo n , p u p a a n d l i fe c y c l e desc r i bed b y R a a t i k a i n e n ( 1 96 7 ) . U n i v o l t i n e i n F i n l a n d ( R a a t i k a i n en 1 967 ) . D I S T R I B U T I O N : P a l a e a rc t i c reg i o n : F I N L A N D : L a i h i a ( V a a s a n ) , H E ! WA ! K o k k o l a ( V a a sa n ) , H E ! Y l i s t a ro ( He i k i n h e i mo 1 957 ) ; U u s i k i rk k o ( He i k i n he i mo 1 957 ) ; Ec kero ( A l a n d ) , H E ! NORWAY : S t a v a n g e r , P ! H U NGARY : B u k k s z e n t l l e k , B U ! H u n g a r y ( N a g y 1 967a ) ; fI I I

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1 1 54 - C h e l a of Gon a top u s m a urus K i effer ( t y pe )

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Gona topus ma urus K i effer 1 906b : 336 . Pla tygon a top u s m a u r u s ( K i effer ) : K i effer 1 9 1 4b : 94 . C h a l cogon a top u s m a urus ( K i effer ) : Mocz a r 1 96 5 : 389 .

F E M AL E : a p tero u s ; l e n g t h 3 , 1 2 m m ; h e a d b l a c k - b row n , w i t h m a n d i b l e s y e l l ow ; a n t e n n a e b row n , w i t h segmen t s 1 -2 y e l l ow ; t h o r a x a n d p ro p o d e u m b l a c k - b ro w n ; a b domen b row n ; l eg s b row n , w i t h t a r s i y e l l ow ; h e a d sh i n y , w i t h occ i p u t a n d f ro n s h a i r y , w i t h o u t scu l p t u re , excep t for occ i p u t w h i ch i s g r a n u l a ted ; v e r t e x weak l y exc a v a t e d , a l mo s t f l a t ; frons more e x c a v a ted ; p ron o t u m crossed b y a t r a n s v e r s a l

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A p e x of 1 1 68 ) A p e x of 1 1 73 ) 6 S e g m en t 5 of f r on t t a r s u s w i t h 2 - 3 r o w s of l a me l l a e ex t e n d i n g b e y o n d 0 , 5 l e n g t h o f s eg men t ( F i g . 1 1 7 3 ) . . . . . . . . . . . . . . . . . . . . . . 7 S e g m e n t 5 of f r on t t a r s u s w i t h 1 - 2 r o w s of l a me l l a e ; i f t w o •

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r o w s , t h e s h o r t e s t r o w d o e s n o t ex t e n d b ey o n d 0 , 5 l e n g t h o f s e g men t ( F i g . 1 1 7 7 ) . . . . . . . . . . . . . . . 22 7 L a b i a l p a l p i w i t h 2 s e g men t s . . . . . . 8 L a b i a l p a l p i w i t h 3 s eg m e n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 8 T h o r a x a n d p r o p od e u m mo s t l y b l a ck . . . . . . . . . . . . . . . . . . . . . . . . .9 Th o r ax and p r op od e u m f u l l y f e r ru g i n ou s o r t e s t a c e ou s . . . . .10 9 H e ad ex c a v a t ed 9 . c on t ortulus P a t t o n H ea d f l a t 1 0 . p l a n ip e s n . sp . 1 0 Me t a t h o r a x + p r o p od eu m u s u a l l y w i t h o u t s c u l p t u r e , ex c ep t f o r s t r i a e on p l e u r a a n d p o s t e r i o r s u r f a c e ; s e gmen t 5 o f f r on t t a r s u s w i t h l a me l l a e a t d i s t a l h a l f ( F i g . 1 1 7 7 ) . . 1 2 . agropyrus F e n t on Me t a t h o r ax + p r op od e u m u s u a l l y g ra n u l a t ed , ex c e p t f o r s t r i a e on p l e u r a a n d p o s t e r i o r su r f a c e ; s e g m e n t 5 of f r' on t t a r s u s w i t h l a me l l a e b eg i n n i n g a t p r ox i ma l h a l f ( F i g . 1 1 7 6 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 . mimoides ( R . C . L . P e r k i n s ) 1 1 P o s t e r i o r s u r f a c e o f p r op od e u m n o t t r a n s v e r s e l y s t r i a t e o r s t r i a t e on l y o n p o s t e r i o r h a l f . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 P o s t e r i o r su r f a c e o f p r op o d e u m f u l l y t r a n sv e r s e l y s t r i a t e . . . . . . 14 1 2 B od y p a r t I y b l a ck . . . 6 . echo ( R . C . L . P e r k i n s ) .

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B o d y f u l l y t e s t a c e ou s o r r ed d i s h -d a rk , w i t h p e t i o l e b l a c k a n d a b d omen i n s ome sp e c i me n s b r own . . . . . . . . . . . . . 1 3 M e t a n o t u m f l a t ( F i g . 1 1 7 0 A ) . . 7 . elongatus n . sp . Me t a n o t u m n o t f l a t ( F i g . 1 1 7 0 B ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 . fre yta gi n . sp . H e a d , t h o r a x a n d p r op od e u m f u l l y t e s t a c e o u s o r y e l l ow t e s t a c e ou s o r r e d d i s h - t e s t a c e ou s o r f e r r u g i n ou s o r r ed d i s h 1 5 H ea d , t h o r a x a n d p r o p od e u m a t l ea s t p a r t l y b l a ck o r b r own . . . . 1 9 E n l a r g ed c l a w mo r e c u rv ed ( F i g . 1 1 8 1 ) ; s e g men t 5 o f f r on t t a r s u s w i t h t w o sh o r t e r r o w s of l a me l l a e ( F i g . 1 1 8 1 ) . . . . . 1 3 . fre quens ( R . C . L . P e r k i n s ) E n l a rg e d c l a w l e s s c u rv ed ( F i g s 1 1 8 3 , 1 1 8 4 ) ; s e g men t 5 o f f r on t t a r s u s w i t h t w o l on g e r r o w s of l a me l l a e ( F i g s 1 1 8 3 , 1 1 8 4 ) . . 16 H ea d mo r e e x c a v a t ed ( F i g . 1 1 8 2 A , 0 ) 17 .

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H ea d l e s s e x c a v a t ed , a l mo s t f l a t 1 7 S c u t u m l a t e r a l l y w i t h t wo p o i n t s

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. . . . . . . . . 18 ( on e p e r s i d e ) ( F i g . 1 1 8 2 B ) ; .

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1 67 2

I ns! . ,

no.

37,

1 984

m e t a n o t u m l a t e r a l l y w i t h o u t p o i n t s ( F i g . 1 1 8 2 6 ) ; m e t a t h o ra x p r o p o d eu m g r a n u l a t ed a n d w i t h a n t e r i o r s u r f a c e l e s s i n c l i n ed ( F i g . 1 1 8 2 C ) ; s eg m e n t 4 0 f f r o n t t a r s u s l o n g e r t h a n s eg m e n t 1 4 . areol a tu s ( F e n t o n ) S c u t u m a n d m e t a n o t u m l a t e ra l l y w i t h t wo p o i n t s ( F i g . 1 1 8 2 E ) ; m e t a t h o rax + p r op o d e u m w i t h o u t s c u l p t u r e , ex c ep t f o r t r a n s v e r s a l s t r i a e , a n d a n t e r i o r s u r f a c e mo r e i n c l i n ed ( F i g . .

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1 1 8 2 F ) ; s eg m e n t 4 o f f r o n t s egmen t 1 . . . . . . . . . . . . . . . . . . 1 8 M e t a t h o ra x + p r op o d e u m w i t h (F i g . 1 1 85 A ) . . . . . . . . . . . . . . Me t a t h o rax + p r o p o d e u m w i t h ( F i g . 1 1 86 H ) 1 9 S c u t u m y e l l ow-b r own o r r ed d S c u t u m b l a ck .

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t a r s u s a p p r ox i ma t e l y a s l on g a s . . 1 5 . h arpax ( K r omb e i n ) a n t e r i o r s u r f a c e m o r e i n c l i n ed . . 1 6 . solitarius ( R . C . L . P e rk i n s ) a n t e r i o r s u r f a c e l e s s i n c l i n ed 1 7 . herbarum ( R . C . L . P e rk i n s ) i sh .

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20 21

2 0 M e t a t h o r a x + p r op o d e u m w i t h a n t e r i o r s u r f a c e mo r e i n c l i n ed ( F i g . 1 2 7 1 A ) . . . . . . . . . . . . . . . . 1 8 . p ara leptias ( R . C . L . P e rk i n s )

21

M e t a t h o ra x + p r o p o d e u m w i t h a n t e r i o r s u r f a c e l e s s i n c l i n ed ( F i g . 1 2 7 1 6 ) . . . . . . . . . . . . . . . . 1 9 . pseu dochrom u s ( R . C . L . P e rk i n s ) P r o p o d e u m f u l l y y e l l ow - r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

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1 9 . pseudochrom u s ( R . C . L . P e r k i n s ) P r o n o t u m a t l ea s t p a r t l y b l a ck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 0 . flavi&ons A s h me a d 2 2 P o s t e r i o r s u r f a c e o f p r op o d e u m w i t h ou t t ra n s v e r s a l s t r i a e o r wi t h s t r i ae o n I y o n p ost er i or ha I f 23 P o s t e r i o r s u r f a c e o f p r op o d e u m f u l l y a n d s t r on g l y t ra n s v e r s e l y st r i a t e 27 2 3 H e a d l e s s e x c a v a t ed , a l mo s t f l a t ( F i g . 1 1 8 6 A ) ; m e s o m e t a p l eu r a l s u t u r e d i s t i n c t a n d c o mp l e t e . . . . . . . . . . . . . . . . . . . . . 2 4 H e a d mo r e e x c a v a t ed ( F i g . 1 1 8 6 B , 0 ) ; m e s o -m e t a p l eu r a l s u t u r e o b s o l e t e o r on l y w i t h p r ox i ma l t h i rd d i s t i n c t . . . . . . . . . . . . . . . . 2 5 .

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2 4 Me t an o t um f l a t , p r o p o d e u m r ed d i Me t a n o t u m n o t f i n c l i n ed d own t .

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2 5 L ab i a l Lab i a l

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pa l p i pa l p i 2 6 S eg men t 5 o f l en g t h ( F i g . S eg me n t 5 o f 1 1 96 ) . . . . . . . 2 7 L ab i a l p a l p i Lab i a l pa l p i 2 8 Me t a t h o rax +

.

m o r e t h a n 0 , 5 a s l on g a s s c u t e l l u m ; t h o r a x a n d s h - t e s t a c e ou s . 2 7 . innitidus ( R . C . L . P e rk i n s ) l a t ; a n t e r i o r s u r f a c e o f m e t a t h o r a x + p r op od e u m o s c u t e l l u m ; t h o ra x a n d p r op od e u m b l a ck . . . . . . . . 2 8 . brooks; n . s p . w i t h 3 s eg m en t s . 2 1 . rapax ( R . C . L . P e rk i n s ) w i t h 2 s eg m e n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 .

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f r on t t a r s u s w i t h l a me l l a e o f n ea r l y t h e s a m e 1 1 9 4 ) . . . . . . . . . . . 2 2 . a sh m e a di K i e f f e r f r o n t t a r s u s w i t h l a m e l l a e l on g e r n ea r b a s e ( F . . . . . . . . . . . . . . . . . 2 3 . ombrodes ( R . C . L . P e rk i n s ) w i t h 3 s eg men t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . w i t h 2 s eg me n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . p r op o d e u m b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

ig . . . 28 . . 31 . . 29

M e t a t h o ra x + p r op o d e u m r e d d i s h -y e l l o w . . . . . . . . . . . . . . . . . . . . . . . . 3 0 2 9 M e s o -m e t a p l e u r a I s u t u r e o b s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

1 673

Gon a topod i n ae

p araleptias ( R . C . L . P e rk i n s ) a n d c o mp l e t e monilis n . s p . 30 S e g men t 4 of I y s h o r t e r t h a n s e g me n t 1 argyrias ( R . C . L . P e rk i n s ) S eg men t 4 o f t h a n s eg m e n t 1 . . . . . . . . 2 6 . curriei K r omb e i n 3 1 H ea d l e s s e x c a v a t ed ( F i g . 1 1 8 6 E ) ; b od y y e l l o w- t e s t a c e ou s , w i t h p e t i 0 I e b l a ck , a b d omen a n d a n t e n n a e p a r t I y b r own i s h 1 2 . agropyrus F en t o n H ea d mo r e ex c a v a t ed ; a t l ea s t s c u t u m , s c u t e l l u m a n d m e t a t h o ra x + p r o p od e u m p a r t l y b l a ck 2 9 . p a l lidiceps ( R . C . L . P e rk i n s ) •

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18. su t u r e d i s t i n c t 24. f r on t t a r s u s s l i g h t 25 . f r o n t t a r s u s l on g e r •

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Me s o -m e t a p l eu r a l .

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MALES

A n t en n a l

2

s eg m e n t 3 l e s s . . . . . . . . . . . . . . . . . . . . . . . A n t en n a l s eg m e n t 3 mo r e P r o p o d e u m f u l l y smo o t h ,

l o n g a s b r oad 1 7 . herb arum ( R . C . L . P e rk i n s ) t h r e e t i me s a s l on g a s b r oad 2 , n o t r e t i cu l a t e r u g o s e 2 . cyphonotus B ra d I e y P r o p od eu m d u I I , r e t i c u I a t e r u g o s e 3 D o r s a l p r o c e s s o f g on o f o r c e p s mu c h sh o r t e r t h a n g on o f o r c ep s ( F i g . 1 1 9 5 ) ; b a s i v o l s e l l a w i t h ou t n o t ch ( F i g . 1 1 9 5 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 2 . a shmeadi K i e f f e r D o r s a l p r o c e s s o f g o n o f o r c ep s n ea r l y a s l on g a s g o n o f o r c e p s ( F i g s 1 1 8 0 , 1 1 9 7 ) ; b a s i v o l s e l l a w i t h n o t ch ( F i g s 1 1 8 0 , 1 1 9 7 ) 4 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h v e ry b r oad d i s t a l r e g i on a n d w i t h d i s t a l ma r g i n s l i g h t l y s e r r a t e ( F i g . 1 1 9 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 . ombrodes ( R . C . L . P e rk i n s ) D o r s a l p r o c e s s o f g o n o f o r c ep s f u l l y s l e n d e r o r a t mo s t w i t h s l i g h t l y b r oa d e n ed d i s t a l r e g i o n ( F ' g . 1 1 8 0 ) . . . . . . . . . . . . . . . . . . 1 2 . agropyrus F en t o n •

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t h an . . . . . . t han sh i n y

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T h e m a l es of t h e o t h e r s p ec i es of Gon a top u s a re u n k now n . F re y t a g ( 1 977 ) desc r i bed' t w o new s p ec i es of Neogon a topu s ( fentoni and grandis ) . On I y the ma l es a r e k n ow n . They a re t r e a t e d as nomina incerta e sedis because I fee l the m o r p h o l og i c c h a r a c t e r i s t i c s d o not s u ff i c i en t l y i n d i c a t e t h a t the two s p ec i es a r e of the gen u s Gon a top u s .

1 . Gon a top u s cla vicornis ( F e n ton )

Cyrtogo n a topus cla vicornis F e n ton 1 9 27 : 3 .

n.

comb .

Mem . A mer .

1 674

Fig.

F E MA L E :

En\.

I n s\ . ,

no.

37,

1 984

1 1 62 - C h e l a of Gon a top u s c 1 a vicornis ( F e n t on ) ( t y pe )

a p t e rou s ;

l e n g t h 3 , 30 m m ;

h e a d b row n - b l ac k ,

w i t h f ron t p a r t of

v e r t e x , c l y p e u s and m a n d i b l es y e l l ow ; a n t en n ae b ro w n , w i t h se g m e n t s 1 -2 y e l l ow ; t ho r a x a n d p ro p o d e u m b l ac k ; a b d omen b row n - b l ac k ; l e g s y e l l o w red d i s h ; head ( F i g . 1 079 B, C ) f l a t , s h i n y , g r a n u l a t ed ; t em p l es v e r y s h or t , l ess t h a n 0 , 5 a s l o n g as eyes ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u r e ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m n o t v i s i b l e ; a n t e r i o r s u rf a ce of m e t a t ho r a x + p ropodeum very i n c l i n e d ; m e t a t h o r a x + p ro p o d e u m sh i n y , w i t h ou t scu l p t u r e ; pos t e r i o r su rface o f p ro p o d e u m t r a n sverse l y s t r i a t e ; p l eu r a n o t t r a n sverse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; m e t a t h o r a x + p ropod e u m w i t h a s t ro n g med i a n f u rrow ;

fore t a r s a l s e g me n t s

Gon a topod i n a e

Fig.

1 1 63

-

C h e l a of Gon a top u s cyphonotus Br a d l ey from C h i I I i w a c k Co l u m b i a )

1 675

( Br i t i sh

Mem . A me r .

1 676

Fig.

1 1 64

-

Ent.

I nst . ,

n o . 37,

1 984

R a d i a l v e i n of m a l e of Gon a top u s cyphonotus B r a d l ey f rom Ye l l o w k n i fe ( N . W. T . , C a n a d a )

i n f o l l ow i n g p ro p or t i on s : 1 3 : 2 : 3 : 5 : 8 ; fore t roch a n t e r s n ea r l y as l o n g a s fore c o x a e , a b o u t t w i c e a s l o n g a s b ro a d ( 1 2 : 5 ) ; e n l a rged c l a w ( F i g . 1 1 62 ) w i t ho u t s u b a p i c a l t o o t h a n d l a me l l ae , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4 b r i s t l es ; seg m e n t 5 o f f ro n t t a r s u s ( F i g . 1 1 62 ) w i t h 1 row of v e r y l o n g 7 l a m e l l ae e x t en d i n g con t i n u ou s l y t o apex w h i c h has 5 l a m e l l ae ; a p e x n o t b ro a dened ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : B ro w n sv i l l e ( Tex . , U . S . A . ) TYP I CAL M A T E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! HOST S : u n k no w n D I ST R I B U T I ON : N e a rc t i c reg i o n :

U.

S. A . :

i n WA .

B ro w n s v i l l e ( T ex . ) ,

WA !

2 . Gona top u s cyphonotus B r a d l e y

Gon a top u s cyphonotus B r a d l e y 1 906 : 380 . Gon a topu5 fou tsi W h i t t a k e r 1 9 28 : 388 ( n . s y n . ) ; locus typ ic u s C h i l l i w ack ( B . C . , C a n ada ) .

F E MAL E : a p terou s ; l e n g t h 1 , 68-3 , 25 m m ; h e a d b ro w n o r t e s t aceo u s - b ro w n , w i t h f ro n t p a r t of v e r t e x , c l y pe u s a n d m a n d i b l es y e l l o w ; a n t en n ae b ro w n , w i t h s e g me n t s 1 -2 o r 1 -5 y e l l ow ; t ho r a x , p ro p o d e u m a n d a b domen b l ac k ; r a re l y t ho r a x a n d p ropodeum or on I y p ro t h o r a x t e s t aceo u s-d a r k ; l eg s y e l l o w , w i t h cox a e a n d , i n some spec i me n s , c l u b s o f femora b row n ; h e a d f l a t , g r a n u l a t ed , s h i n y ; p ro no t u m crossed b y a t ra n s v e rs a l i m p re s s i o n , s h i n y , h a i r l es s , g r a n u l a ted ; sc u t u m g r an u l a te d ; met a t ho r a x + p ropodeum d u l l , g r a n u l a t ed ; m e t a n o t u m t r a n s v erse l y s t r i a t e ; p o s t e r i o r s u rf ac e of p ropodeu m and p l e u r a more o r l es s w e a k l y t ra n sverse l y s t r i a t e ; meso m e t a p l eu ra l s u t u re o b so l e t e ; segmen t 1 of f r o n t t a r s u s t w i ce a s l o n g as segmen t 4 ( 1 2 : 6 ) ; e n l a r ge d c l aw ( F i g . 1 1 63 ) w i t h o u t su b a p i c a l t oo t h a n d l a m e l l ae , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d 6 peg - l i k e h g a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 63 ) w i t h 2 rows of 1 5- 1 9 l a m e l l ae ; a p ex b ro a d e n ed , w i t h a grou p o f 8- 1 0 l a m e l l ae ; m a x i l l a r y p a l p i

Gon a topod i n ae

Fig.

1 67 7

1 1 65 - M a l e g e n i t a l i a of Gon a top u s cyp hon o t u s B r a d l ey f rom Y e l l ow k n i fe ( N . W . T . , C a n a d a ) ( r i g h t h a l f removed )

w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M AL E : f u l l y w i n ged ; l en g t h 2 , 3 1 -2 , 8 1 mm ; b l ac k ; a b domen , p ropodeum a n d l eg s b a n d e d ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a I segmen t s i n fo l l ow i n g p ropor t i o n s : 7 : 5 : 1 1 : 1 0 : 1 0 : 9 : 9 : 9 : 8 : 1 0 ; a n t e n n a I se g m e n t 3 more t h a n t h ree t i me s as l o n g as b road ( 5 , 6 ) ; h e a d g ra n u l a t e d , d u l l ; f ro n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 8 ; OL = 3 ;

1 678

Mem .

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Ame r .

Ent.

I nst . ,

no.

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1 1 66 - C h e l a of Gon a topus mayori n .

1 984

sp .

( type)

Gon a topod i n ae

1 679

OOl = 2; b re a d t h of the oce l l i : 4; scu t u m d u l l , g r a n u l a te d ; n o t a u l i ces i n com p l e t e , reach i n g a p p rox i m a t e l y 0 , 5 l en g t h of scu t u m ; sc u t e l l u m a n d m e t a n o t u m sh i n y , w i t h o u t scu l p t u re o r v e r y w e a k l y g r a n u l a t e d ; p ro p o d e u m sh i n y , smoot h , w i t ho u t scu l p t u re ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n sv e r s a l b a n d s ; r a d i a l v e i n : f i g . 1 1 64 ; gonoforceps ( F i g . 1 1 65 ) w i t h a d o rs a l p rocess s h o r t a n d n a rrow ; b a s i v o l se l l a w i t h o u t n o t c h ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 . l O C U S TYP I C U S : Dow n i e C reek ( B . C . , C a n a d a ) TYP I CAL MAT E R I AL : h o l o t y p e F ! of G. cyphonotus i n I T ; h o l o t y p e F ! a n d 2 p a r a t y pes F F ! of G. fo u t s i i n BM ; 3 p a r a t y p es F F ! of G . fou tsi i n LA . HOST S : i n C a n a d a j\:'a crosteles fascifron s ( S t a l ) . D I ST R I B U T I O N : N e a rc t i c reg i on : CANA D A : Ye l l o w k n i fe ( N . W . T . ) , O T ' O l ' C h i l l i w a c k ( B . C . ) , B ty', ! lA � Dow n i e C reek ( Se l k i r k Mo u n t a i n s , B . C . ) , I T ! M i ss i on C i t y ( B . C . ) , O l ! O T ' S l a v e l a k e ( A l t a . ) , W A ! C a s t o r ( A l t a . ) , O T ! W a i n w r i g h t ( A l t a . ) , O l ' O T ' l a n c a s t e r P a rk ( A l t a . ) , O T ! A l d e r F l a t s ( A l t a . ) , OT � F i s h e r B r a n c h ( M a n . ) , O T ! W i n n i p eg ( M a n . ) , O T ! F l o F r a n c e s ( On l o ) , OT ! U . S. A . : Homestead ( F l a . ) , O T !

3 . Gon a top u s mayori n .

sp .

F E M AL E : a p t e rou s ; l en g t h 3 , 00 m m ; f u l l y t e s t aceo u s , w i t h p e t i o l e b l a c k ; h e a d f l a t , sh i n y , g r a n u l a t e d ; p ro n o t u m crossed by a t r a n s v e r s a l i m p ress i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re ; scu t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m sh i n y and smoo t h , w i t h o u t scu l p t u re ; m e t a n o t u m ( F i g . 1 1 86 C ) s h i n y a n d smoo t h , w i t h o u t K �e l s ; m e t a t ho r a x + p ro p o d e u m smooth a n d sh i n y , w i t ho u t sc u l p t u re , w i t h n u merous t r a n s v e rs a l k ee l s o n p os t e r i o r s u rf ace a n d p l e u r a ; viSible;

meso-met a p l eu r a l s u t u re w e a k l y

seg m e n t 1 of f ron t t a rsu s a p p rox i m a t e l y

t w i ce as

l o n g a s seg men t

4 ( 1 2 : 6 ) ; e n l a r g e d c l a w ( F i g . 1 1 66 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4 b r i s t l es ; segmen t 5 of front t a r s u s ( F i g . 1 1 66 ) w i t h i n n e r s i d e p ro x i m a l l y not se r r a t e , w i t h 2 rows of 1 0 l a me l l ae ; a p e x w i t h a g r o u p of 9 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S TYP I C U S : E I M a y o r ( C a l i f . U . S . A . ) T Y P I CAL M A T E R I AL : h o l o t y p e F � i n C A . HOS T S : u n k no w n D I ST R I B U T I ON : N e a rc t i c reg i o n : U . S . A . : E I M a y o r ( C a l i f . ) , C A '

4 . Gon a to o u s portalen sis n .

sp .

F E MA L E : a p t e ro u s ; l e n g t h. 1 , 60 - 2 , 65 m m ; h e a d b l ac k or b row n , w i t h a n t er i or h a l f o f f r o n s , c l y p e u s a n d m a n d i b l es t e s t aceou s ; a n t e n n a e b row n ,

Mem .

1 680

Fig.

Am e r .

Ent .

I nst . ,

no.

37,

1 984

1 1 67 - C h e l a of Gonatop u s porta lensis n . sp .

( type)


Fig.

1 1 68 - C h e l a o f Gon a top u s tran slucidus n .

1 68 1

n a me ( t y p e )

w i t h se g me n t s 1 -3 tes t aceo u s ; t h o r a x , p ropod e u m , a b domen , l e g s u su a l l y t e s t aceou s ; i n some spec i me n s mesosoma d a r k ened ; p e t i o l e b l ac k ; a n t e n n ae d i s t a l l y t h i ck e n e d ; a n t en n a l segmen t s i n f o l l ow i n g p ropor t i o n s : 6 : 4 : 9 : 5 : 4 , 5 : 5 : 4 : 4 : 4 : 6 ; h e a d f l a t , d u l l , g r a n u l a t e d ; frons w i t h a s h o r t i n comp l e t e me d i a n k ee l ; occ i p i t a l ca r i n a i n v i s i b l e ; p r o n o t u m crossed b y a t r an s v e rs a l i m p re s s i on , d u l l , w e a k l y g r a n u l a ted ; sc u t u m a n d m e t a t h o r a x +

1 682

Me m . Amer .

Ent.

I ns t . ,

no.

37,

1 984

p ropode u m d u I I , w e a k I y g r a n u I a ted ; poste r i o r s u r f ace of propod e u m a n d p l e u r a n o t t r a n s v e r se l y s t r i a te ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a r s a l se gmen t s in fo l l o w i n g propor t i o n s : 1 1 : 2: 3: 5 , 5 : 1 0 ; P O L = 3 ; OL 2 ; OOL = 6 ; e n l a r g e d c l a w ( F i g . 1 1 67 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d w i t h 5-6 peg - l i k e h a i r s ; s e g me n t 5 of fron t t a r s u s ( F i g . 1 1 67 ) w i t h i n n e r s i d e p rox i m a l l y n o t ser r a t e , w i t h 2 rows of 1 1 - 1 2 l a me l l a e ; apex w i t h a g r o u p of 6-7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0, 1 . MAL E : u n k no w n LOCUS T Y P I C U S : Por t a l ( A r i z . U . S . A . ) TYP I C AL MAT E R I AL : ho l o t y p e F ! a n d 2 p a r a t y pes F F ! i n O T ; 1 p a r a t y p e F ! i n T E ; 1 p a r a t y p e F ! i n A M ; 3 p a r a t y p es F F ! i n OL . HOST S : u n k n o w n D I S T R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : S a g e h e n C reek ( N e v a d a C o . , C a l i f . ) , OL ! Om i r a ( L a s se n C o . , C a l i f . ) , T E ! Port a l ( C a v e C reek C y n . , Ar i z . ) , OL l O T ! W a r d ( C o l o . ) , AM ! K e r rv i l l e ( Tex . ) , O T ! OL !

5 . Gon a top u s translueidus n .

n a me

n e e Gon a top u s testaeeus C a meron 1 888 : 440 . E u e a mptonyx testaeeus R . C . L . Perk i n s 1 907 : 28 .

F E M AL E : a p t e rou s ; l e n g t h 2 , 87-4 , 02 m m ; y e l l ow - t e s t aceo u s or red d i s h - d a r k , w i t h pe t i o l e b l ack , head , i n some spec i me n s , b r ow n i sh ; a n t e n n ae f u l l y t e s t aceous or w i t h segme n t s 1 -4 y e l l ow - t e s t ac eo u s a n d 5- 1 0 b row n i sh ; h e a d exc a v a t ed or a l mo s t f l a t , d u l l or s h i n y , s t r o n g l y or w e a k l y g r a n u l a ted ; p ro n o t u m crossed by a s t r o n g t r a n sversa I i m p re s s i o n , g r a n u I a ted ; m e t a t h o r a x + p ropodeum g r a n u l a t e d ; p os t e r i o r su rf ace a n d p l e u r a t r a n s v e r se l y s t r i a t e ; a n t e r i or su rf ace w i t h l o n g i t u d i n a l k ee l s ; meso - met a p l e u r a l s u t u re obso l e t e ; segmen t 4 of fron t t a rs u s as l o n g as seg m e n t 1 ; en l a r g e d c l a w ( F i g . 1 1 68 ) w i t h 4 - 9 l a me l l a e , w i t h o u t s u b a p i c a l too t h a n d a sm a l l t o o t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow ; a p e x n o t po i n t ed and bent a t r i g h t a n g l e ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 1 68 ) w i t h t w o rows of 1 5 - 1 6 l a me l l a e ; a p e x w i t h a group of a t l e a s t 20-40 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen ts ; t i b i a l spurs 1 , 0, 1 . MALE : u n k no w n LOC U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) T Y P I C A L MAT E R I AL : ho l o t y pe F ! i n B . H O ST S : u n k no w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : D a n v i l l e ( C a l i f . ) , O L ! Nog a l e s ( A r i z . ) , B ! Sab i n a C a n y on ( Ar. i z . ) , WA ! Bowmer R a n c h ( Be l l C o . , Tex . ) , O T ! F a rm i n g d a l e ( N . Y . ) , T W !

1 68 3

Gon a topod i n a e

Fig.

1 1 69

-

C h e l a o f Gon a top u s e c h o ( R . C . L . Perk i n s ) ( t y p e )

6 . Gona top u s echo ( R . C . L .

Perk i n s )

Chalcogon a top u s echo R . C . L . Perk i n s 1 90 7 : 23 .

n . com b .

Mem . Ame r .

1 684

Fig.

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no.

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1 984

1 1 70 - Thorax sp .

and p ropodeum of f e m a l es of Gona top u s elonga tus n . ( t y pe ) ( A ) a n d freyt a gi n . sp . ( t y p e ) ( B ) .

F E M A L E : a p t e ro u s ; l e n g t h 4 , 2 1 m m ; h e a d a n d p rono t u m y e l l ow - redd i sh ; a n te n n a e y e l l ow-red d i s h , w i t h seg men t s 5 - 1 0 d a r k e n e d ; scu t u m , scu t e l l u m , me t a t h o r a x + p ropod e u m , p e t i o l e a n d a b domen b l ac k ; l e g s y e l l ow - redd i sh ; h e a d e x c a v a t e d , sh i n y , g r a n u l a ted j p r o no t u m crossed by a t r a n s v e r s a l i m p ress i o n , g r a n u l a t ed ; m e t a t h o r a x + p ropodeum g r a n u l a t ed ; pos ter i o r s u r f a c e of p ropodeum t r a n sverse l y s t r i a t e o n l y o n poste r i o r h a l f ; segmen t 4 of f r o n t t a r s u s l on g e r t h a n segmen t 1 ( 40 : 30 ) ; e n l a rged c l a w ( F i g . 1 1 69 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 1 1 peg - l i k e h a i rs ; segmen t 5 of f r o n t t a r s u s ( F i g . 1 1 69 ) w i t h 2 rows of 7 + 1 6 l a me l l a e ex t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; a p ex w i t h a g ro u p o f a t l ea s t 1 0 l a me l l ae ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Nog a l es ( A r i z . , U . S . A . ) TYP I CAL MAT E R I AL : l ec to t y p e F ! ( des i g n a t ed b y O l m i 1 982 ) i n B . HOSTS : u n k no w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B !

7 . Gon a top u s elonga t u s n . sp .

F E MAL E :

a p terou s ;

l en g t h 4 , 37-4 , 75 m m ;

red d i s h -d a rk or f e rr u g i n o u s ,

w i th

1 68 5

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p e t i o l e b l ac k , a n te n n ae b l ac k , w i t h seg men t s 1 -4 b r ow n ; a n t en n ae d i s t a l l y t h i ck ened ; a n t e n n a I segme n t s i n fo l l o w i n g p ropor t i on : 1 1 : 7 : 26 : 1 6 : 1 0 : 1 0 : 8 : 8 : 7 : 1 0 ; h e a d s h i n y , w e a k l y g ra n u l a t ed , exc a v a t ed ; f ron s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O ' � 2 ; OL = 2 ; OOL = 1 0 ; p ro n o t u m c rossed b y a t ra n s v e r s a l i m p ress i o n , d u l l , w e a k l y g r a n u l a t ed ; m e t a n o t u m ( F i g . 1 1 70 A ) f l a t , g r a n u l a t ed , t r a n s v e r se l y s t r i a te ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a ted ; p l eu ra n o t s t r i a te t r a n s v e r se l y ; p o s t e r i o r s u rf ac e o f p ro p o d e u m not s t r i a t e t r a n s v erse I y , on I y w i t h a f e w s t r i a e n e a r a pex ; meso-met a p l e u r a l s u t u re o b so l e t e ; fore t a r s a l seg men t s i n fo l l ow i n g p roport i o n s : 1 5 : 3 : 5 : 20 : 32 ; en l a r g e d c l a w ( F i g . 1 1 7 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d w i t h 1 0- 1 7 s m a l l l a me l l a e , p l u s 1 b r i s t l e ; s e g m e n t 5 of f r o n t t a r s u s ( F i g . 1 1 7 1 ) w i t h 2 rows of 29-4 l am e l l ae ; a pex w i t h a g ro u p of a t l e a s t 1 8-30 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u nknown L O C U S T Y P I C U S : T a l l a h a ssee ( F l o r i d a , U . S . A . ) T YP I CAL M A T E R I AL : h o l o t y p e F ! i n WA ; 1 p a r a t y p e F ! i n O L ; 1 p a r a t y p e F ! i n O T ; 1 p a r a t y p e F ! i n AM . HOSTS: u nknown D I ST R I B U T I O N : N e a rc t i c reg i o n :

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F E MAL E : a p t ero u s ; l en g t h 2 , 50-3 , 37 m m ; tes t a ceou s , w i t h p e t i o l e b l ack ; some t i mes a b domen b row n ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l seg men t s i n fo l l ow i n g p ropor t i o n s : 7 : 4 : 1 0 : 8 : 6 : 6 : 5 : 5 : 4 , 5 : 6 ; a n t en n ae tes t ac eo u s , w i t h s e g me n t s 4- 1 0 or 6- 1 0 b row n ; h e a d a l mo s t f l a t , d u l l , g r a n u l a ted ; f r o n s w i t h a med i a n k ee l , occ i p i t a l c a r i n a i n v i s i b l e ; P O L 2; O L = 1 , 5 ; O O L = 8 ; p r o n o t u m cros sed b y a t r a n s v e r s a I i m p ress i o n , d u I I , g r a n u l a ted , h a i r l es s ; sc u t u m , s c u t e l l u m , m e t a n o t u m ( F i g . 1 1 70 B ) , m e t a t h o r a x + p ropodeum a n d p l eu r a d u l l , f u l l y g r a n u l a ted , w i t h o u t k ee l s o r o t h e r sc u l p t u r e , e x c e p t for p o s t e r i o r s u rface o f p ropodeum w h e re there are some t ra n sv e rs a l k ee l s on s i des of pos t e r i or h a l f ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ropo r t i on s : 1 1 : 2 : 3 , 5 : 9 : 1 5 ; en l a r g e d c l a w ( F i g . 1 1 72 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 5 peg - l i k e h a i r s ; seg m e n t 5 o f fron t t a r s u s ( F i g . 1 1 72 ) w i t h 2 rows o f 2+6 o r 1 +3 o r 2+5 l a m e l l ae ( u s u a l l y l oc a ted on a d i s t i n c t p rom i n e n ce ) e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ; a p ex w i t h a g r o u p of 9- 1 5 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S T Y P I C U S : C h a n n i n g ( T ex a s , U . S . A . ) T Y P I C A L M A T E R I AL :

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in O T ; 1 p a r a t y p e F ! in C M . HOSTS : u n k nown D I S T R I B U T I ON : N e a r c t i c reg i on : U . S . A . : M i l n es a n d ( Roosev e l t C o . , N . M ex . ) , WA ! C h a n n i n g ( O l d h a m C o . , Tex . ) , \VA ! L i m p i a C n . ( F t . D a v i s , T ex . ) , O T ! G r a n i t e M t . ( Tex . ) , X C M ! N O T E S : t h e spec i e s i s n a med i n h o n o r of M r . P a u l H . F rey t a g .

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Gona top u s con tort u l u s P a t t o n 1 879 : 65 . Neogon a top u s contor t u l u s ( P a t ton ) : F re y t a g 1 97 7 : 5 7 2 .

F E M AL E : a p terou s ; l en g t h 2 , 37-3 , 62 mm ; h e a d b l ac k or b ro w n - r e d d i s h , u s u a l l y w i t h occ i p u t , f r o n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n te n n ae b row n , w i t h seg men t s 1 -2 y e l l ow ; p ro n o t u m b ro w n o r b l ac k or t e s t aceo u s , u s u a l l y w i t h d i sc b row n i sh o r b l ac k ; scu t u m b r o w n o r b l ac k o r y e l l o w ; s c u t e l l u m a n d m e t a t h or a x + p ro p o d e u m b l ac k o r b l a c k - b row n , w i t h a p ex of p ro p o d e u m r e d d i s h ; a b domen b row n - b I ack or b r o w n -redd i s h ; l e g s t e s t aceou s ; h e a d exc a v a ted , s h i n y , w e a k l y g r a n u l a t ed ; p ro n o t u m c rossed by a t r a n s v e rs a l i m p re s s i o n , s h i n y , w e a k l y g r a n u l a t ed ; scu t u m s h i n y , l a ter a l l y w i t h 2 p o i n t s ( F i g . 1 1 86 F ) , w i t h some l o n g i t u d i n a l k ee l s ; met a n o t u m t r a n s v e r se l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , w i t h o u t scu l p t u r e ; p l e u r a a n d p os t e r i or s u rf a c e of p ro p o d e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; seg men t 1 o f f ron t t a r s u s a s l o n g a s seg men t 4 ; en l a r g e d c l a w ( F i g s 1 1 73 , 1 1 74 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d 5-6 p eg - l i k e h a i r s ; seg men t 5 o f f r o n t t a r s u s ( F i g . 1 1 73 ) w i t h 2-3 row s o f 2 1 -26 l a me l l ae e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ; a p ex w i t h a g ro u p of 7 - 1 6 l a me l l a e ; m a x i I l a ry p a l p i w i t h 4-5 segme n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l spurs 1 , 0, 1 . MAL E : u n k n o w n L O C U S TYP I C U S : W a t e r b u ry ( U . S . A . ) TYP I CA L M A T E R I A L : h o l o t y p e F ! i n PH . HOSTS : i n U . S . A . � a ta ll1s sayi ( Fe n ton 1 9 1 8 a , 1 9 1 8 b ) a n d E u tettix ten ella Baker ( Se v er i n 1 924 ) ; in C a n a d a Endria inimica S a y . D I ST R I B U T I O N : N e a rc t i c reg i on : C ANADA : B l a n c h a r d R . ( m i . 93 H a y n e s H w a y , B . C . ) , O T ! C r i s t o p h e r L a k e ( Sa s k . ) , O T ! W i n n i p eg ( M a n . ) , OT ! O L ! P i l o t M o u n d ( M a n . ) , O T ! U . S . A . : M i l V a l l ey ( M a r i n Co . , C a l i f . ) , C A ! P i n e V a l l ey ( Sa n D i ego C o . , C a l i f . ) , C A ! S a g e h e n C reek ( Ne v a d a Co . , C a l i f . ) , T E ! D r y de n ( T e r re l l Co . , T ex . ) , W A ! 1 1 , 2 m i . S of Leve l l a n d ( H ock l e y C o . , Tex . ) , BY ! A l f a l fa C o . ( Ok l a . ) , WA ! A n n A r b o r ( M i c h . ) , T W ! S i ou x C i t y ( I ow a ) , WA ! C o l u m b u s ( Oh i o ) , OL ! S w a n n a n o s ( N . C . ) , A M ! W a t e r b u ry ( Co n n . ) , P H ! S a l i s b u ry C e n t er ( N . Y . ) , OL ! TW ! M E X I C O : C a r r ( C u e r n a v ac a , More l o s ) , O T ! O L !

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u s u a l l y w i t h o u t sc u l p t u re ; p l e u r a n o t t ra n s verse l y s t r i a t e ; pos t e r i o r su rface of propode u m t r a n s v erse I y s t r i a t e ; meso-me t a p I e u r a I s u t u re obso l e t e ; segmen t 1 of fro n t t a rs u s s l i g h t l y l on ge r t h a n s e g m e n t 4 ( 1 0 : 9 ) ; e n l a r g e d c l a w ( F i g . 1 1 77 ) w i t ho u t s U b a p i c a l too t h , w i t h a s m a l l too t h a t t he e n d o f a l on g i t u d i n a l f u r ro w a n d 4-6 peg - l i k e h a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 77 ) w i t h 1 row of 9- 1 2 l a m e l l a e or t w o rows of 4 ( p rox i m a l or d i s t a l ) + 2 1 l a m e l l ae ; a p e x w i t h a g ro u p of 7 l a m e l l ae ( 2 of


1 697

t hese very l o n g ) ; m a x i l l a r y p a l p i w i t h 3-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i g s 1 1 78 , 1 1 79 ) : fu l l y w i n g e d ; l e n g t h 2 , 0-2 , 5 m m ; b l a c k ; m a n d i b l es y e l l ow ; a n te n n ae b row n ; teg u l ae y e l l o w ; l e g s b row n , n o t b a n ded , w i t h t i b i ae a n d t a r s i l i g h t ; a n te n n a l segme n t s i n fo l l o w i n g p ropor t i o n s : 6 : 6 : 1 4 : 1 4 : 1 4 : 1 4 : 1 2 : 1 1 : 1 0 : 1 3 ; a n ten n a l s e g me n t 3 more t h a n f i ve t i mes a s l o n g a s b r o a d ( 6 , 5 ) ; h e a d s h i n y , we a k l y p u n c t a t e , h a i r y , w i t ho u t sc u l p t u r e , except for p u n c t u res ; fro n t a l I i n e a b se n t ; POL � 1 0 ; OL � 3 ; OOL � 3 ; b rea d t h of t h e oce l l i : 4 ; temp l e s d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m d u l l , g r a n u l a ted , h a i r y ; n o t a u l i ces comp l e t e , poster i o r l y s e p a r a t e d ; m i n i m u m d i s t a nce b e t w een t h e n o t a u l i ces shorter t h a n b rea d t h of oce l l i ( 2 : 4 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoot h , w e a k l y p u n c t a t e ; p ropodeum re t i c u l a te r u g ose , w i t ho u t k e e l s ; dors a l p rocess o f gonoforceps ( F i g . 1 1 80 ) n ea r l y a s l o n g a s gonoforceps , not b roadened a p i c a l l y , s l ender ; b a s i vo l se l l a w i t h n o tc h ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Ames ( I ow a , U . S . A . ) T Y P I CAL M A T E R I A L : h o l o t y p e F ! , a l l o t y p e M ' a n d 2 p a r a t y p es F F ! of G . a gropyru s i n W A . HOST S : i n U . S . A . Ps ammotettix affin is ( G . a n d B . ) ( F en t o n 1 9 2 1 ) ; Ps ammotettix stria t u s , Gra minella n igrifron s ( F orbes ) , L a ta l u s sayi ( F re y i a g 1 97 7 ) ; i n C a n a d a Endria inimica . O I ST R I BUT I O N : N e a rc t i c reg i on : CANADA : S t . D a v i d s ( O n t . ) , OT ' O t t a w a ( O n t . ) , O T ! U . S . A . : S u n n y s i de ( H u a c h u c a M t s . , A r i z . ) , OT ! 25 , 1 m i . �J of A n g l e t o n ( B razor i a C o . , T e x . ) , OL ! 5 , 3 m i . N of La G r a n g e ( F a y e t t e Co . , T e x . ) , BY ! O L ! B i g B e n d N . P . ( T ex . ) , OT ! 2 , 8 m i . S of Somer v i l l e ( W a s h i n g to n Co . , Te x . ) , BY ! W a co ( Me Len n a n Co . , Te x . ) , B Y ! 25 , 4 m i . W of Refu g i o ( Bee C o . , Te x . ) , Te x . ) , BY ! W i scon s i n ( F rey t a g 1 977 ) ; Ames ( I o w a ) , WA ! Le x i n g ton ( K y . ) , OL ! P t . V a l l e y ( G a . ) , WA ! I t h a c a ( N . Y . ) , TW !

1 3 . Gon a topus fre quens

(R.C.L.

Perk i n s ) n . co m b .

Chalcogon a to o u s fre q uens R . C . L . Perk i n s 1 907 : 2 1 . A gon a top u s p a llidicornis R . C . L . Perk i n s 1 90 7 : 29 ( n . s y n . ) ; locus typ i c u s N og a l es ( A r i z . , U. S . A . ) . A gona top u s p a ll idicornis R . C . L . P e r k i n s v a r . d u 6 iosus R . C . L . P e r k i n s 1 90 7 : 30 ( n . s y n . ) ; loc u s typ icus Nog a l es ( A r i z . , U . S . A . ) .

F E MALE : a p terou s ; l en g t h 2 , 70-4 , 02 m m ; y e l l o w - t e s t aceo u s , w i t h pe t i o l e b l ac k a n d a b domen b row n i s h ; a n t en n ae t e s t a ceou s , w i t h segmen t s 4- 1 0 d a rk e n e d ; h e a d e xc a v a ted , d u l l , g r a n u l a t e d ; pron o t u m crossed b y a t r a n svers a l i m p ress i o n , s h i n y , w i t ho u t sc u l p t u re or w e a k l y g r a n u l a ted ; s c u t u m g ra n u l a te d ;

m e t a n o t u m we a k l y t r a n sve rse l y s t r i a t e ;

me t a t horax +

Mem .

1 698

Fig.

1 1 81

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Ame r . E n t .

I ns t . ,

C he l a of Gon a top us frequens ( A r i zo n a )

no. 37,

(R.C.L.

1 984

Pe � k i n s ) from N og a l es

Gona topod i n ae

1 699

c

F i g . 1 1 82 - F e m a l e of Gon a top u s areol a t u s ( F e n ton ) ( t y p e ) : A : h e a d ; B : m e so t h o r a x a n d m e t a t h o ra x ; C : t ho r a x a n d p ropode u m ; fema l e of Gon a top us harpax ( K romb e i n ) ( t y p e ) : ::l : h e a d ; E : meso t h o r a x a n d met a t hor a x ; F : t h o r a x a n d p ropodeurn .

1 700


I nst . ,

no.

37 ,

1 984

p ropodeum s h i n y , w i t h o u t sc u l p t u re ; p l e u r a a n d poster i or s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t ure o b so l e t e ; seg m e n t of fro n t t a rs u s a s l on g a s seg m e n t 4 ; en l a rged c l a w ( F i g . 1 1 8 1 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rro w a n d 2-5 p e g - I i k e h a i rs ; seg men t 5 of fro n t t a r s u s ( F i g . 1 1 8 1 ) w i t h 2 r o w s of 7 - 1 3 l a me l l ae ( e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ) ; a p e x w i t h a g r o u p of 7- 1 3 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n knonw L O C U S T Y P I CU S : No g a l es ( A r i z . U . S . A . ) TYP I CAL M A T E R I A L : l ec t o t y pe F ! a n d 3 p a r a l ec t o t y pes F F ! ( d es i g n a ted b y O l m i 1 98 2 ) o f CU. fre quens i n B ; l ec t o t y p e F ! a n d 1 p a r a l ec t o t y pe F ! ( des i g n a t e d b y O l m i 1 982 ) o f A . pa llidicornis i n B ; t y p i c a l m a te r i a l o f A .

pall idicornis v a r . d u b io s u s l os t . HOST S : i n U . S . A . n o t i d en t i f i e d C i c a d e l l i d s . D I S T R I B U T I O N : N e a rc t i c roeg i o n : U . S . A . : T a ss a j a r a H o t ( C a l i f . ) , CA l N og a l es ( A r i z . ) , O L ! B ! A r i zo n a , B M !

1 4 . Gon a top u s areol a t u s ( F en ton )

n . com b .

Chalcogon a top u s areol a tu s F e n ton 1 927 : 7 .

F E M AL E : a p terou s ; l en g t h 3 , 4 3-4 , 1 0 m m ; ferr u g i n ou s , w i t h p e t i o l e b l a ck ; h e a d ( F i g . 1 1 82 A ) exc a v a t e d , d u l l , h a i r l e s s , gra n u l a t e d ; p ron o t u m crossed b y a t r a n s vers a l i m p res s i on , d u l l , g r a n u l a ted ; s c u t u m d u l l , g r a n u l a ted , l a t e ra l l y w i t h 2 po i n t s ( F i g . 1 1 82 B ) ; s c u te l l u m d u l l a n d g r a n u l a ted ; m e t a n o t u m t r a n s verse l y s t r i a t e ; m e t a t ho r a x + propodeum ( F i g . 1 1 82 C ) d u l l , gra n u l a t e d ; p l e u r a a n d pos t e r i or s u rface of p ropodeum t r a n s v e rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 1 4 : 3 : 4 : 20 : 30 ; e n l arged c l a w ( F i g . 1 1 83 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 1 1 - 1 2 peg - l i k e h a i rs ; segme n t 5 o f fro n t t a rs u s ( F i g . 1 1 83 ) w i t h 2 rows of 26-36 l a me l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; apex w i t h a g r o u p o f 1 3-20 l a me l l a e ( some of t h ese very l o n g ) ; m a x i l l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : F a l l s C h u rc h ( V a . , U . S . A . ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n IVA . HOST S : u n k no w n D I ST R I B UT I O N : N e a rc t i c reg i o n : U . S . A . : U n i ve r s i t y of I l l i n o i s ( Woods , I I I . ) , O L ' C o l u m b i a ( S . C . ) , T W ! F a l l s C h u rc h ( V a . ) , ( F e n ton 1 927 ) ; G r a s s y me a d ( V a . ) , WA ! W a s h i n g to n ( D . C . ) , WA ! C h e v y ( C h a s e L a k e , M d . ) , WA '

Gon a topod i n a e

Fig.

1 1 83

-

C he l a o f Gon a top u s areol a tu s ( F en ton ) ( t y p e )

1 70 1

Mem . A m e r . E n t . " I n s t . ,

1 702

Fig.

1 1 84

-

no.

37,

1984

C h e l a of Gon a top u s h a rp ax ( K rombe i n ) ( t y p e )

1 703

Gon a topod i n ae

1 5 . Gon a top u s h a rp a x ( K rom b e i n ) n . com b .

Chalcogon a top u s h a rpax Krombe i n 1 9 56 :

1 58 .

F E MALE : a p t e ro u s ; l en g t h 3 , 25-3 , 70 m m ; tes t aceous o r redd i s h - d a r k , w i t h p e t i o l e b l a ck a n d a b domen b ro w n - red d i s h ; h e a d ( F i g . 1 1 82 D ) exca v a ted , s h i n y , w i t h o u t sc u l p t ure , e xc e p t for occ i p u t a n d fron t p a r t of v e r t e x w h i c h a re g ra n u l a te d ; pron o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , v e r y s h i n y , s moo t h , h a i r l ess , w i t h o u t scu l p t u re ; scu t u m s h i n y , w i t ho u t scu l p t u re ; m e t a n o t u m s h i n y , w i t h some t r a n s v e r a I s t r i a e ; m e t a t h o r a x + p ro p o d e u m ( F i g . 1 1 82 F ) s h i n y , w i t h o u t sc u l p t u re ; s c u t u m l a ter a l l y w i t h 2 po i n t s ( F i g . 1 1 82 E ) ; m e t a n o t u m a l so l a te ra l l y w i t h 2 poi n t s ( F i g . 1 1 8 2 E ) ; meso-met a p l eu r a l s u t u re obso l e t e ; p l e u r a a n d p o s t e r i or s u r face of p ropode u m t r a n s v ers e l y s t r i a te ; seg m e n t 4 of fron t t a rs u s n e a r l y a s l o n g a s s e g m e n t 1 ; e n l a r ged c l a w ( F i g . 1 1 84 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 7 peg - l i k e h a i rs ; seg m e n t 5 of fron t t a rs u s ( F i g . 1 1 84 ) w i t h 2 rows of 5-6 + 24 l a m e l l a e e x t e n d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g ro u p of 7-20 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown L O C U S TYP I C U S : L o s t R i ver S t a te P a rk ( W . V a . , U . S . A . ) TYP I C AL MAT E R I AL : h o l o t y p e F ! i n WA ! HOSTS : u n k no w n D I ST R I BU T I O N : N e a rc t i c reg i o n : C A NA D A : P r o E d w ard Co . ( On ! . ) , U . S . A . : Lost R i v e r S t a t e P a rk ( IV . V a . ) , WA !

1 6 . Gon a top u s solitarius

(R.C.L.

Perk i n s )

OT !

n . com b .

Ch alcogon a topus solitarius R . C . L . Perk i n s 1 90 7 : 2 2 . Ch alcogon a topus xestocep h a l u s R . C . L . Perk i n s 1 907 : 24 ; loc u s typ icus N og a l es ( A r i z . , U. S. A . ) Agona top u s ferrugin e u s R . C . L . P e rk i n s 1 907 : 30 ( n . s y n . ) ; loc u s typ i c u s Nog a l e s ( A r i z . , U . S . A . ) A gona top u s picescens R . C . L . Perk i n s 1 907 : 30 ( n . s y n . ) ; locus typicus Nog a l es ( A r i z . , U. S. A . ) Chalcogon a top u s xestocep h a l u s R . C . L . P e rk i n s : K i effer 1 9 1 4 b : 1 0 1 . A gona top u s ferrugin e u s R . C . L . P e rk i n s : K i effer 1 9 1 4 b : 1 32 . A gon a top u s p icescen s R . C . L . Perk i n s : K i effer 1 9 1 4 b : 1 3 3 .

F E MA L E : a p t e rou s ; l e n g t h 3 , 6-5 , 1 m m ; redd i s h-ferru g i n ou s , w i t h p e t i o l e b l ac k ; a n t e n n ae red d i sh-ferru g i no u s , w i t h s e g m e n t s 4 - 1 0 o r 6- 1 0 o r 7- 1 0

1 704

Fig.

Mem . A m e r .

Ent.

I ns! . ,

no.

37 ,

1 984

1 1 8 5 - F em a l e of Gon a top us solitarius ( R . C . L . P e rk i n s ) from N og a l es ( A r i zo n a ) ; A : t h o r a x a n d p ropode u m ; B : c h e l a .

b r ow n ; a b domen occ a s i on a l l y b r o w n i s h ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l s e g m e n t s i n fo l l o w i n g p roport i o n s : 1 4 : 6 : 1 7 : 1 0 : 8 : 9 : 7 : 6 : 6 : 9 ; h e a d f l a t , d u l l , g r a n u l a te d ; fron t a l l i n e comp l et e ; tem p l es d i s t i n ct ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; OL = 2 ; O O L = 1 0 ; p ron o t u m s h i n y , smoot h , w i t h o u t scu l p t u re , crossed b y a s t ro n g t r a n s ver s a l i m p res s i o n ; scu t u m w i t h l on g i t u d i n a l k ee l s ; m e t a n o t u m t r a n sverse l y s t r i a t e ; m e t a t ho r a x + p ro p o d e u m ( F i g . 1 1 85 A ) d u l l , g r a n u l a ted , w i t h p l e u ra a n d p o s te r i or s u rface t r a n sve rse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l ete ; seg m e n t 4 of fro n t t a r s u s as l on g as or s l i g h t l y l on ge r t h a n seg m e n t 1 ( 37 : 3 5 ) ; en l a r g e d c l a w ( F i g . 1 1 85 B ) w i t h o u t s U b a p i c a l toot h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5-8 peg - l i k e h a i rs ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 1 1 85 B ) w i t h 2 rows o f 20-28 l a me l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g r o u p of a t l ea s t 1 2- 1 6 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 segmen t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k nown L O C U S TYP I C U S : No g a l e s ( A r i z . , U . S . A . ) TYP I CAL MA T E R I A L : h o l o t y p e F ! of Ch . solitarius i n B ; ho l o t y pe F ! of Ch . xestocep h a l u s i n B ; l ec t o t y p e F ! a n d 3 p a r a l ec t o t y pes F F ! ( de s i g n a ted b y O l m i 1 9 82 ) of A . £errugin e u s i n B ; l ec t o t y p e F ! ( des i g n a ted b y O l m i 1 98 2 ) o f A . p icescens i n B . HOST S : u n k n o w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B !

1 7 . Gon a top u s herb arum ( R . C . L .

P er k i n s )

n . com b .

1 705

Gon a topod i n a e

c

�F F i g . 1 1 86 - H e a d s of fem a l es of Gon a topus innitidus ( R . C . L . P e rk i n s ) ( A ) , ombrodes ( R . C . L . P e rk i n s ) ( B ) , a shmeadi K i effer ( D ) , a gropyru s F e n t o n ( E ) a n d herb arum ( R . C . L . P e rk i n s ) ( G ) ; meso t ho r a x a n d m e t a t ho r a x of fema l es of Gon a top u s mayori n . s p . ( C ) a n d contortulus P a \ ton ( F ) ; t h o r a x a n d p ro p o d e u m of fem a l e of Gon a top u s herb arum ( R . C . L . P e r k i n s ) ( H ) .

Cha lcogon a top u s herb arum R . C . L . P e r k i n s 1 907 : 23 .

F E MAL E : a p terou s ; l e n g t h 3 , 4 7-5 , 4 9 m m ; ferr u g i no u s , w i t h pet i o l e b l ac k a n d a n t en n a l s e g m e n t s 6- 1 0 b row n ; h e a d ( F i g . 1 1 86 G ) v e r y s l i g h t l y e x c a v a ted ,

a l most f l a t ,

g r a n u l a ted ; p rono t u m crossed b y a t ra n s vers a l

i m p ress i o n , s h i n y , w i t h o u t sc u l p t u re ; sc u t u m g r a n u l a te d , l a te ra l l y w i t h 2 p o i n t s ; m e t a n o t u m t r a n s v e rse l y s t r i a t e ; m e t a t h o r a x + p ropodeum ( F i g . 1 1 86 H ) w ea k l y g ra n u l a ted ; p l eu r a a n d pos t e r i o r s u rface of p ropodeum t r a n s verse l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re o b so l e t e ; fore t a rsa l seg men t s 1 -4 i n f o l l ow i n g p ropor t i o n s : 1 4 : 3 : 5 : 1 6 ; en l a r ged c l a w ( F i g . 1 1 87 ) w i t ho u t s u b a p i c a l t oo t h , w i t h o r w i t ho u t a s m a l l too t h a t t h e e n d o f a

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1 706

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1 70 7

1 1 88 - M a l e g e n i t a l i a of Gon a top u s h e r ba r u m ( R . C . L . Perk i n s ) from Somerset ( C a l i for n i a ) ( r i g h t v o l se l l a removed )

l o n g i t u d i n a l f u rrow a n d w i t h 5-8 peg- l i k e h a i r s ; seg men t S of f ro n t t a rs u s ( F i g . 1 1 87 ) w i t h i n ne r s i de p ro x i m a l l y n o t serra t e , w i t h t wo rows o f 1 7-34 l a m e l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g ro u p of 1 0- 1 7 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 seg men t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 8-2 , 0 m m ; b l ac k ; m a n d i b l e s p a r t l y y e l l o w ; l eg s n o t b a n ded ; a n t e n n ae n o t d i s t a l l y t h i c k e n e d ; a n t en n a I segmen t s i n fo l l o w i n g p roport i on s : 4 : 4 : 7 : 6 : 6 : 6 : 6 : 5 : 5 : 8 ; a n t e n n a I s e g m e n t 3 l e ss t h a n t h ree t i mes as l o n g as b ro a d ( 7 : 3 ) ; h e a d d u l l , g r a n u l a ted ; f r o n s w i t h a com p l e t e med i a n k ee l ; occ i p i t a l c a r i n a a b sen t ; occ i p u t conca v e ; P O L 5 ; O L = 1 , 5 ; OOL = 5 ; t e m p l es p ro m i ne n t ; s c u t u m d u l l , g r a n u l a ted ; n o t a u l i ces comp l e t e , poster i o r l y sep a r a ted ; s m a l l es t d i s t a n c e b e t w e e n n o t a u l i ce s n ea r l y a s l o n g a s b re a d t h of oce l l i ( 2 : 2 ) ; sc u t e l l u m a n d m e t a no t u m smoo t h , s h i n y , w i t h o u t sc u l p t u re , f i n e l y p u n c t a t e ; propode u m ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t ho u t d a r k t r a n s vers a l b a n d s ; r a d i a l v e i n w i t h d i s t a l p a r t m u c h l on ge r t h a n prox i m a l p a r t ; r a d i a l ce l l

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1 1 89 - C he l a of Gon a topus p a ralep tias ( R . C . L . ( W yom i n g )

1 984

Perk i n s ) f rom G l en d a

o p e n ; dors a l p rocess of gonoforceps ( F i g . 1 1 88 ) v e r y l o n g a n d n a rrow ; b a s i vo l se l l a w i t h n o t c h ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 1 , 2 .

Gon a topod i n ae

1 709

LOCUS TYP I C U S : No g a l es ( A r i z . , U . S. A . ) T Y P I C AL MAT E R I A L : l ec t o t y p e F ' a n d 2 p a r a l ec t o t y pes F F ! O l m i 1 982 ) i n B .

( d es i g n a ted b y

HOSTS : u n k no w n O I ST R I BUT I O N : N e a rc t i c reg i o n : U . S . A . : S . Berd i n o C o . ( C a l i f . ) , IVA ! A p p l e V a l l e y ( C a l i f . ) , O L ! OT ! Somerset ( E I Dorado C o . , C a l i f . ) , T E ' OL ! N og a l es ( A r i z . ) , B ! B M ! O L ! C a s a G r a n d e ( A r i z . ) , BM ' W a co ( M c Len n a n C o . , Te x . ) , BY ! 5 , 3 m i . N o f L a G r a n g e ( F a y e t te Co . , T e x . ) , OL !

18.

Gon a top us paralep ti a s U ' . C . L .

Perk i n s )

n.

com b .

Cha lcogon a top u s paralep tias R . C . L . Perk i n s 1 907 : 1 8 . Cha lcogon a top u s euprepes R . C . L . Perk i n s 1 907 : 1 8 ( n . sy n . ) ; locus typ i c u s Nog a l es ( A r i z . , U . S. A . ) . Cha lcogon a top u s euprepoides R . C . L . Perk i n s 1 907 : 1 9 (n . s y n . ) ; locus typicus Nog a l e s ( A r i z . , U . S . A . ) . Cha lcogo n a top u s cuprepes ( e rro r ) R . C . L . Perk i n s : K i effer 1 9 1 4 b : 99 . Cha lcogon a top u s cuprepoides ( e rror ) R . C . L . Per k i n s : K i effer 1 9 1 4b : 99.

F E MA L E : a p tero u s ; l en g t h 3 , 66- 5 , 1 2 m m ; head b r o w n or re d d i s h , w i t h m a n d i b l es , fron t p a r t of vertex a n d s i des of c l y pe u s y e l l o w ; a n t e n n ae b r o w n w i t h seg m e n t s 5-7 y e l l ow , or y e l l o w w i t h seg m e n t s 8- 1 0 or on l y 1 0 b row n ; p ro n o t u m t e s t aceo u s or b ro w n w i t h s i des l i g h t or redd i s h ; scu t u m b ro w n -y e l l o w or b l a ck o r redd i s h ; occ a s i on a l l y re d d i s h w i t h s i des b l ac k ; sc u te l l u m , m e t a t ho r a x + p ropodeu m , p e t i o l e a n d a b domen b l a c k ; l e g s fu l l y t e s t aceo u s or y e l l o w , w i t h coxae , femora a n d t i b i a e p a r t l y b row n ; head e xc a v a t e d , s h i n y , g r a n u l a ted ; p ron o t u m crossed b y a s t ro n g t r a n s versa l i m p ress i o n , g ra n u l a t ed ; m e t a t ho r a x + p ropodeum s h i n y , w i t h o u t sc u l p t u re ; r a re l y g r a n u l a t ed a n d w i t h a t r a c k of m ed i a n f u rrow ; p o s t e r i o r s u r face of p ropodeum a n d p l e u r a t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re ob so l e t e ; seg m e n t 4 of fro n t t a r s u s l o n ge r t h a n s e g m e n t 1 ( 4 5 : 40 or 46 : 3 5 or 44 : 39 ) ; e n l a rged c l aw ( F i g s 1 1 89 , 1 1 90 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t he e n d of a l on g i t u d i n a l f u r ro w a n d 7- 1 2 peg - l i k e h a i rs ; s e g me n t 5 of fro n t t a r s u s w i t h 1 ( F i g . 1 1 90 ) o r 2 row s ( F i g . 1 1 89 ) of 1 6-36 l a m e l l ae ; apex w i t h a g rou p of at l e a s t 1 2- 1 6 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C US : Nog a l e s ( A r i z . , U . S . A . ) TYP I CAL MA T E R I A L : l ec t o t y p e F ! a n d 1 p a ra l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 982 ) of Ch . p ara leptias i n B ; ho l o t y p e F I o f Ch . e u pre p es i n B ; ho l o t y p e F ! of Ch . euprepoides i n B . HOSTS : u n k no w n D I S TR I BU T I O N : N e a rc t i c re g i o n : U . S . A . : S a n J ac i n to M t s . ( I d y l l w i l d ,


1 71 0

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1 9 84

C h e l a of Gon a topus paralep tias ( R . C . l . J ac i n t o M t s . ( C a l i forn i a )

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C a l i f . ) , Ol ' T a n b a r k F l a t ( los A n g e l es Co . , C a l i f . ) , C A ! lee S t . ( M i l l V a l l ey , C a l i f . ) , C A ! M a r i n C o . ( M i l l Va l l e y , C a l i f . ) , CA ' S t a n fo r d ( C a l i f . ) , Ol ! Nog a l es ( A r i z . ) , B ! W i l l i a m s ( A r i z . ) , W A ' G l e n d a ( W y o . ) , WA !

171 1

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1 984

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Cha lcogon a topus (Eugo n a top u s ) p s e u dochromus R . C . L . P e r k i ns 1 905 : 48

F E MA L E : ap terou s ; l e n g t h 3 , 9 1 -4 , 1 8 m m ; h e a d r e d d i s h or y e l l ow-redd i s h , w i t h b l ac k s p o t o n v e r t e x ; a n te n n ae y e l l o w-red d i s h o r b r o w n , w i t h segmen t s 1 -2 t e s t aceo u s ; p r o n o t u m y e l l o w-redd i s h ; sc u t u m , scu te l l u m , m e t a t h o r a x + p ropodeum a n d pe t i o l e b l a c k ; ab domen b l ac k o r t e s t aceo u s ; l e g s y e l l o w-redd i s h ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I seg men t s i n fo l l ow i n g p ro p or t i o n s : 1 3 : 9 : 35 : 20 : 1 5 : 1 2 : 1 1 : 1 0 : 9 : 1 2 ; h e a d exca v a ted , s h i n y , g r a n u l a t ed ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 , 5 ; O L 2 ; O O L = 1 1 ; f ro n t a l I i n e com p l e t e ; p r o no t u m c rossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , g r a n u l a ted ; scu t u m d u l l , g r a n u l a t ed ; meso m e t a p l e u r a l s u t u re obso l e t e ; m e t a t h o r a x + p ro p o d e u m g r a n u l a ted ; p o s t e r i o r s u rface t r a n s v e rse l y s t r i a t e ; seg m e n t 4 o f f ron t t a rs u s l on g e r t h a n segmen t 1 ( 37 : 3 1 ) ; en l a r ged c l a w ( F i g . 1 1 9 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 1 1 - 1 4 p e g - I i k e h a i rs ; segme n t 5 o f f r o n t t a r s u s ( F i g . 1 1 9 1 ) w i t h 2-3 rows o f 32-56 l a m e l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; a p e x w i t h a g r o u p of a t l ea s t 1 1 -30 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; , l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : Co l u m b u s ( O h i o , U . S . A . ) TYP I C A L MAT E R I A L : l ec t o t y p e F ' ( d es i g n a t ed b y O l m i 1 982 ) i n B . HOSTS : i n U . S . A . Ph lepsius sp . ( R . C . L . Per k i n s 1 906c ; O l m i 1 982 ) . D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : 4 m i . S of W a r ren ( T y l e r C o . , Tex . ) , O L ! 1 0 , 5 m i . N of H a l l et s v i l l e ( L a v ac a C o . , Tex . ) , B Y ! R e i s e l ( F a l l s Co . , T e x . ) , WA ! 3 , 8 m i . S o f Sea l y ( W a l l er C o . , Te x . ) , B Y ! 2 5 , 1 m i . N of A n g l e t o n ( B razo r i a Co . , Te x . ) , B Y ! C o l u m b u s ( O h i o ) , B ! B a r r e n I s . ( Lo n g I s l a n d , N . Y . ) , AM !

20 . Gon a top u s fl a vifrons A s h m e a d

Gon a topus fl a vifrons A s h m e a d 1 89 3 : 84 . Cha lcogon a top u s lep t i a s R . C . L . P e r k i n s 1 907 : 1 7 ( n . sy n . ) ; locus typicus Nog a l es ( A r i z . , U . S . A . ) . Cha lcogon a top u s fl a vifrons ( A s h m e a d ) : K ro m b e i n 1 9 56 : 1 59 .

F E MA L E : a p t e rou s ; l en g t h 3 , 37-4 , 50 m m ; h e a d b l ac k , w i t h c l y p e u s , m a n d i b l es a n d f ro n t p a r t of v e r t e x y e l l ow ; occ i p u t b l ac k or y e l l o w - b ro w n ; a n t e n n a e b r o w n w i t h s e g m e n t 1 y e l l ow , or y e l l o w w i t h segme n t s 8- 1 0 d a rk ; t horax a n d p ropodeum b l a c k ; i n some s p ec i m e n s p rono t u m p a r t l y red d i s h ;

Gon a topod i n a e

Fig.

1 71 3

1 1 92 - C he l a o f Gon a topus flavifrons A s h m e a d ( t y p e )

abdomen b rown-b l ac k ; l eg s y e l l ow - b ro w n , w i t h coxae , femora a n d t i b i a e brown i s h ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 8 : 7 : 1 6 : 1 4 : 1 0 : 8 : 6 , 5 : 6 : 5 : 7 , 5 ; h e a d s h i n y , e xc a v a ted , h a i r l es s , g ra n u l a ted ; occ i p i t a l c a r i n a i n v i s i b l e ; fro n t a l I i n e comp l e t e ; P O L = 2 ; OL = 2 ; OOL = 8 ; p rono t u m crossed b y a s t ro n g t r a n s vers a l i m press i on , w ea k l y g r a n u l a ted ; m e t a n o t u m t r a n s verse l y s t r i a t e ; m e t a t h o r a x + p ropod e u m d u l l , g r an u l a ted ; pos t er i or s u rface f u l l y t r a n sverse l y s t r i a t e ;


1714

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I n s ! . , n o . 37 ,

1 984

1 1 93 - C he l a of Con a to p u s r a p a x ( R . C . L .

Perk i ns ) ( type)

p l e u ra p a r t l y t r a n s v erse l y s t r i a t e ; meso-met a p l eu r a l s u t u r e obso l e t e ; se g m e n t 4 of fro n t t a r s u s l on g e r t h a n s e g m e n t 1 ( 30 : 26 ) ; en l a r g e d c l a w ( F i g . 1 1 92 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t he e n d of a l o n g i t u d i n a l f u r row a n d 8- 1 0 peg- l i k e h a i rs ; seg m e n t 5 of f ro n t t a r s u s

1 71 5

Gon a topod i n a e

( F i g . 1 1 9 2 ) w i t h 2 rows of 26-3 l a m e l l ae e i t h e r i n t e r r u p t ed or e x t en d i n g co n t i n uo u s l y t o a p e x ; a p e x w i t h a g rou p o f a t l e a s t 1 3-30 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k no w n LOCUS TYP I CU S : A l bany ( N . Y . , T Y P I CA L M A T E R I A L : ho l o t y p e F ! p a r a l ec t o t y pes F F ! ( des i g n a t ed HOST S : u n k n o w n D I ST R I B U T I O N : N e a rc t i c reg i o n : ( A r i z . ) , O L ! F l a g s t a f f ( Ar i z . ) , ( A s h m e a d 1 893 ) .

U. S. A . ) of G . fl a vifron s i n WA ; b y O l m i 1 982 ) i n B .

l ec t o t y p e F !

and 2

U . S . A . : L a s Vegas ( N e v . ) , WA ! W i l l i a m s O L ! Nog a l es ( Ar i z . ) , B ' A l b a n y ( N . Y . )

2 1 . Gon a top u s rapax ( R . C . L .

Perk i n s )

n . comb .

Ch a lcogon a top u 5 rap a x R . C . L . Perk i n s 1 907 : 20 .

F E MA L E : a p terou s ; l en g t h 5 , 30 m m ; h e a d re d d i s h - y e l l ow , w i t h s m a l l b r o w n o c e l l a r s po t ; a n te n n a e red d i sh-ye l l o w , w i t h s e g me n t s 3- 1 0 w e a k l y d a r k e n e d ; p ron o t u m re d d i s h - y e l l o w ; scu t u m re d d i s h - y e l l ow , w i t h s i des b row n ; scu t e l l u m b ro w n ; m e t a t ho r a x + p ro p o d e u m b l a c k , w i t h a n t e r i o r s u rf a ce red d i s h ; d i sc w i t h a b ro w n s p o t ; a p e x of p ro p o d e u m redd i s h ; p e t i o l e a n d a b domen b l ac k ; l eg s redd i s h - y e l l o w ; h e a d exc a v a ted , s h i n y , g r a n u l a t ed ; p r o n o t u m crossed by a t r a n s vers a l i m p ress i o n , g r a n u l a t ed ; m e t a t h o r a x + p ro p o d e u m g r a n u I a ted ; p o s t e r i o r s u rface of p ropodeum t r a n s v e rs e l y s t r i a te on l y on p o s te r i or h a l f ; seg m e n t 4 of fore t a r s i s l i g h t l y l on g e r t h a n seg men t 1 ( 50 : 4 8 ) ; e n l a rged c l a w ( F i g . 1 1 9 3 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 1 2 p e g l i k e h a i rs ; seg men t 5 of f r o n t t a rs u s ( F i g . 1 1 93 ) w i t h 2 rows of 3 + 21 l am e l l ae ( s h o r t e s t r o w n o t e x t e n d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ) ; a p e x w i t h a g r o u p of a t l e a s t 20 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : N og a l es ( Ar i z . , U . S . A . ) TYP I CAL M A T E R I A L : l ec t o t y p e F ! ( des i g n a ted by O l m i 1 982 ) i n B . HOST S : i n U . S . A . n o t i de n t i f i e d C i c a d e l l i d s . D I S T R I BU T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l e s ( A r i z . ) , B !

2 2 . Gon a topu5 a shme a di K i effer

nee Dryin u 5 � ieolor H a l i d a y 1 828 i n C u r t i s 1 8 28 : 206 . Gon a top u s ,"ieolor A s h m e a d 1 893 : 8 5 .

Mem . A m e r . E n ! .

1 71 6

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1 1 94

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I n s ! . , n o . 37 ,

1 984

C he l a of Gon a top u s ashme a di K i effer from Sp r i n g C reek ( Georg i a )

1 71 7

Gon a topod i n a e

Gon a top u s ashmea di K i e ffer i n K i e ffer e t M a r s h a l l 1 905 : 1 08 ( ne w n am e ) . 1 Neogon a top u s erythrodes R . C . L . P e r k i n s 1 905 : 4 3 ( s y n . p roposed b y F re y t a g a n d B ac k 1 977 ) ; locus typicus C o l u m b u s ( U . S . A . ) . Neogon a top u s obscurissimus R . C . L . P e r k i n s 1 905 : 43 ( s y n . p roposed by F re y t a g a n d Back 1 977 ) ; locus typicus C o l u m b u s ( U . S. A . ) . Neogon a top u s brunnescens R . C . L . P e r k i n s 1 90 5 : 44 ( s y n . p roposed b y F re y t a g a n d Back 1 977 ) ; locus typicus O h i o ( U . S . A . ) . Gon a top u s ashmeadi K i effer : K i effer 1 907b : 2 1 . Gon a top u s b icolor A s h m e a d : K i effer 1 9 1 4 b : 1 22 . Gon a topus affinis F e n ton 1 9 1 8b : 266 ( s y n . p roposed b y F re y t a g a n d B a c k 1 977 ) ; l o c u s typ icus B a y V i ew ( O h i o , U . S . A . ) . Gona topus inimicis F e n ton 1 9 1 8b : 270 ( s y n . p roposed b y F rey t a g a n d B a c k 1 977 ) ; l o c u s typicus B a y V i ew ( O h i o , U . S . A . ) . Gon a top u s similis F e n ton 1 9 2 1 : 7 1 ( s y n . p roposed b y F rey t a g a n d B a c k 1 977 ) ; l o c u s typ icus A m es ( I ow a , U . S . A . ) . Epigon a top u s ten uis F e n ton 1 927 : 8 ( n . s y n . ) ; locus typicus L a f a y e t t e ( I n d . , U . S. A . ) . Gon a top u s b icolor A s h m ea d : F re y t a g a n d Back 1 977 : 222 .

F EMALE : a p t e rou s ; t e s t aceou s ,

l en g t h 1 -4 m m ; co l o u r v e r y v a r i a b l e ;

f u l l y red d i s h

w i t h p e t i o l e b l a c k a n d a n t e n n a I seg men t s 1 -2 y e l l ow ,

3- 1 0

b row n ; i n some s pec i me n s w i t h b ro w n oce l l a r spot ; somet i mes co l o u r redd i s h- b r ow n , w i t h head b ro w n ( w i t h y e l l o w m a n d i b l es a n d c l y pe u s ) , scu t u m red d i s h - b ro w n , scu t e l l u m a n d m e t a t h o r a x + p ropodeu m b l a c k ; head ( F i g . 1 1 86 D) exc a v a ted , s h i n y o r d u l l , w e a k l y g r a n u l a t e d ; p ron o t u m c rossed b y a t r a n s v e rs a l i m p ress i o n , w i t hou t sc u l p t u re or w e a k l y g r a n u I a te d ; s c u t u m d u I I , g r a n u I a t e d ; m e t a n o t u m a n d m e t a t h o r a x + p ropod e u m w i t h o u t scu l p t u r e or w e a k l y g r a n u l a t ed ( sC U l p t u re more v i s i b l e on p o s te r i or s u rface ) ; p l eu r a a n d pos t er i or s u r f ace of p ropode u m not t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; segme n t 1 of fron t t a r s u s s l i g h t l y s h o r t e r t h a n segm e n t 4 ( 1 1 : 1 3 ) ; en l a r g e d c l a w ( F i g . 1 1 94 ) w i t hou t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t t he e n d of a l o n g i t u d i n a l

1 K i effer ( 1 90 5 ) con s i de red Gon a top u s b icolor A s h m e a d 1 893 p reoc c u p i e d b y Gon a top u s b icolor ( H a I i d a y 1 828 ) a n d p roposed t he n e w n a me Gona topus a shmeadi . T h i s i s , howev e r , a Case of seco n d a r y homonomy , b e c a u s e H a l i d a y desc r i bed h i s b icolor as Dryin u s a n d a c t u a l l y h i s /:; icolor i s b e l o n g i n g t o t h e g e n u s D icondyl u s . T h e A s h m e a d s p ec i es i s , o n t h e con t r a r y , i n t h e g e n u s Gon a top u s . Bicolor A s h m e a d s h o u l d h a v e s o p r i or i t y o v e r a s hmeadi K i effer . T h i s p rob l em shou l d h a v e , however , a n o t he r s o l u t i on on t he b as i s o f t h e r u l es o f zoo l og i c a l nomenc l a t u re . A s hmeadi K i effer and b icolor Ashmead a re i n fac t s y n on y m s . T h e s a m e K i effer ( 1 907b ) was t he f i rs t rev i se r and p l aced b icolor A s h m e a d as sy non y m of a sh me a di . G . a shmeadi so a c t u a I I y has p r i or i t y O V e r G . b icolor .

1 71 8

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f u rrow a n d 4-7 p e g - l i k e h a i rs ; s e g m e n t 5 of fron t t a rs u s ( F i g . 1 1 94 ) w i t h 1 row o f 8-20 l a m e l l ae ( n ea r l y eq u a l i n l en g t h ) ; a p e x w i t h a g r o u p o f 6-8 l a m e l l a e ( 2-3 of t h ese v e r y l o n g ) ; m a x i l l a r y p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 1 , 0-4 , 0 m m ; b l ac k or b r o w n i s h-b l ac k , w i t h t e g u l ae y e l l ow , m a n d i b l es y e l l ow , l e g s b a n ded ; a n t e n n a I s e g me n t s i n fo l l o w i n g p ropor t i o n s : 5 : 5 : 9 : 9 : 8 : 8 : 8 : 8 : 8 : 1 1 ; a n t e n n a I seg men t 3 more t h a n t h ree t i mes CIS l o n g as b road ( 4 ) ; h e a d h a i r y , s h i n y , g r a n u l a ted ; D O L � 8 ; D L � 3 ; DOL � 3 ; b r e a d t h o f t he oce l l i : 3 ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m d u l l , g r a n u l a ted , h a i r y ; n o t a u l i ces comp l e t e , p o s t er i o r l y sep a r a t ed ; m i n i m u m d i s t a n c e b e t ween n o t a u l i ces shorter t h a n b re a d t h o f oce l l i ( 2 : 3 ) ; sc u t e l l u m and m e t a n o t u m s h i n y , smoot h , w e a k l y p u n c t a t e ; p ro p o d e u m ret i c u l a t e r u g ose , w i t h o u t k ee l s ; dorsa l p rocess of gonoforceps ( F i g . 1 1 9 5 ) m u c h s h o r t e r t h a n gonoforce p s , d i s t a l l y b ro a d en e d ; b a s i vo l se l l a w i t h o u t n o t c h ; ' m a x i l l a r y p a l p i w i t h 4 - 5 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S T Y P I C US : Se l m a ( A l a b a m a , U . S . A . ) TYP I C A L MAT E R I AL : h o l o t y p e F ! of G . b icolor A s h m e a d i n \V A ; l ec t o t y pe F ' ( de s i g n a t ed b y O l m i 1 982 ) o f N . erythrodes i n B ; l ec t o t y p e F ' ( de s i g n a t ed b y D l m i 1 98 2 ) o f N . oh sc urissimus in B ; l ec t o t y p e F ! ( de s i g n a ted by D l m i 1 982 ) o f N. brunnescens i n B ; h o l o t y p e F ! a n d a l l o t y p e M ! o f G . a ffin is i n WA ; h o l o t y p e F ' o f G . inimicis i n \VA ; h o l o t y p e F ! o f G . similis i n WA ;

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h o l o t y pe F ! of E. ten u is i n WA . HOST S : i n U . S . A . Amblysell u s curtisii ( F i tc h ) ( F rey t a g a n d B a c k 1 977 ) A thys a n u s curtisii ( R . C . L . P er k i n s 1 906c ) ; Deltocep h a l u s sp . ? ( R . C . L . P e r k i n s 1 906c ) ; Deltocep h a l u s aifinis G . -B . ( F e n ton 1 9 1 8b , 1 92 1 ) ;

Deltocep h a l u s i n imicus ( S a y ) ( F en t o n 1 9 1 8b ) ; Graminella n igrifron s ( F orbes ) , Endria i n imica ( S ay ) , Ps ammotettix stri a t u s ( L . ) , Macrosteles fasc ifrons ( S t a I ) , L a ta l u s s ayi ( F i t c h ) , Deltocep h a l u s fl a vicosta ( S t a I ) ( F rey t a g a n d B a c k 1 9 77 ) . D I S T R I B U T I O N : N e a r c t i c reg i on : U . S . A . : Ames ( I o w a ) , WA ! L a f a y e t t e ( I n d . ) , WA ! C o l u m b u s ( O h i o ) , B ! B a y V i ew ( O h i o ) , WA ! Lex i n g ton ( K y . ) , O L ! Se l m a ( A l a . ) ( As h me a d 1 893 ) ; Sp r i n g C reek ( G a . ) , I T ! R a l e i g h ( N . C . ) , TW !

23 . Gon a top u s ombrodes ( R . C . L .

Perk i n s )

Neogon a top u s ombrodes R . C . L . P e r k i n s 1 905 : 43 . Gon a topus ombrodes ( R . C . L . Per k i n s ) : K i effer 1 9 1 4 b : 1 24 . Gon a top u s p u n c t a tu s Fen ton 1 9 1 8b : 268 ( sy n . p roposed b y F rey t a g 1 97 7 ) ; locus typ icus C o l u m b u s ( O h i o , U . S . A ) . nee Neogon a top u s ombrodes R . C . L . Perk i n s : S t r u b i n g 1 9 56 : 1 48 .

F E M AL E : a p terou s ; l e n g t h 2 , 45- 3 , 29 m m ; h e a d t es t aceou s , w i t h v e r t e x b l ack ; a n t e n n a e b ro w n , w i t h segmen t s 1 - 2 ye l l ow ; p r o n o t u m b l ac k , w i t h re d d i s h s p o t s on a n t e r i or m a r g i n , pos t e r i or m a r g i n , s i des a n d t r a n s v e r s a l i m p ress i o n ; scu t u m , sc u te l l u m a n d m e t a t h o r a x + p ropodeum b l ack ; a b domen b r o w n - b l ac k ; l e g s t e s t aceo u s ; head ( F i g . 1 1 86 B) e x c a v a ted , s h i n y , w e a k l y g r a n u l a t ed ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , sh i n y , g r a n u l a te d ; scu t u m g r a n u l a t e d , w i t h some l o n g i t u d i n a l k ee l s a n d l a ter a l l y w i t h 2 p o i n t s ; met a n o t u m d u l l , t r a n sverse l y s t r i a t e , g r a n u l a ted ; m e t a t h o r a x + p ropodeum du I I , scu I p t u red as m e t a n o t u m ; p l e u r a a n d pos ter i o r s u r f a c e of p ropodeum n o t t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; segmen t 1 o f f ro n t t a r s u s a s l o n g a s segm e n t 4 ; en l a r ged c l a w ( F i g . 1 1 96 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h at the end of a l on g i t u d i n a l f u rrow and 6 p e g - l i k e h a i r s ; segmen t 5 of fron t t a r s u s ( F i g . 1 1 96 ) w i t h 1 r o w of 1 4 l a me l l a e ( l a me l l ae l on g e s t n e a r p rox i m a l e n d of segmen t ) ; a p e x w i t h a g r o u p of 8 l a me l l a e ( 2 of t h ese v e r y l o n g ) ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MA L E : f u l l y w i n ged ; l e n g t h 2 , 0 -2 , 5 m m ; b l a ck o r b l ac k i s h - b r o w n o r b r ow n , w i t h a n t e n n a e b r ow n , m a n d i b l es a n d t e g u l ae y e l l o w ; l e g s n o t b a n d e d , y e l l ow , w i t h coxae b r o w n i s h ; a n t e n n a I segme n t s i n fo l l ow i n g p ropor t i o n s : 5 : 6 : 1 5 : 1 5 : 1 3 : 1 3 : 1 2 : 1 1 : 1 1 : 1 4 . a n ten n a I segmen t 3 more t h a n s i x t i mes a s l on g a s b r o a d ( 7 , 5 ) ; h e a d sh i n y , smoo t h , w i t h o u t scu l p t u re , w e a k l y p u n c t a t e , h a i r y ; fron t a l l i n e a b se n t ; P O L = 9 ; OL = 3 ; O O L = 3 ;

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b re a d t h of t h e oce l l i : 3 ; t e m p l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m h a i ry , g r a n u l a t e d , d u l l ; n o t a u l i ces com p l e t e , poster i o r l y sep a r a ted ; m i n i m u m d i s t a nce b e tween t h e n o t a u l i ces a p p rox i m a t e l y as l o n g as bre a d t h o f oce l l i ; sc u t e l l u m a n d m e t a n o t u m sh i n y , smoo t h , w i t h o u t sc u l p t u re , w e a k l y p u nc t a t e ;

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h y a l i n e ; dorsa l process of gonoforceps ( F i g . 1 1 97 ) n ea r l y a s l o n g a s gonoforcep s , a p i c a l l y b ro a de n ed , p a d d l e- s h a p e d ; b a s i vo l se l l a w i t h n o t c h ; m a x i l l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I CU S : Col u m b u s ( O h i o , U . S . A . ) TYP I C A L M A T E R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 98 2 ) of N . om 'Jrodes i n B ; ho l o t y p e F ! , a l l o t y p e M ! a n d 1 p a r a t y ;o e F ! o f G . p u n c t a t u s i n IVA . HOST S : i n U . S . A . Cic a dula sexn o t a t a F a l l . ( e rror ) ( T ho m p son 1 9 54 ; A i n s l i e 1 920 ) ; L a t a l u s sayi ( F i t c h ) ( F rey t a g 1 97 7 ) = Deltocep h a l u s sayi ( R . C . L . Perk i n s 1 906c ) ; Ps ammotettix stri a t u s ( L . ) ( F re y t a g 1 977 ) . B I OLOG Y : i n form a t i on s i n A i n s l i e ( 1 9 20 ) . D I ST R I B U T I ON : N e a rc t i c reg i on : U . S . A . : W i scon s i n ( F rey t a g 1 977 ) ; I o w a ( F rey t a g 1 977 ) ; Co l u m b u s ( O h i o ) , B ! IVA ! K e n t u c k y ( F rey t a g 1 97 7 ) . P a l a e a rc t i c re g i on : t h e record from Germ a n y b y S t r u b i n g ( 1 9 56 ) i s erroneo u s .

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F E M.A l E : a p tero us ; l e n g t h 3 , 37 m m ; h e a d b l a c k , w i t h m a n d i b l es , c l y p e u s a n d m a I a r s p a ce t e s t aceou s ; a n te n n ae b row n ; t h o r a x a n d p ropodeum b l ac k , excep t for s i des a n d po s t e r i o r s u r face of p r o n o t u m w h i c h are red d i s h ; a b domen b l ac k ; l e g s t e s t aceou s , w i t h coxae p a r t I y b row n ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t en n a I seg men t s i n fo l l o w i n g p ropor t i o n s : 9 : 6 : 1 6 : 9 : 8 : 7 : 6 : 5 : 5 : 6 ; h e a d e x c a v a ted , d u l l , g r a n u l a t ed ; fro n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; Ol = 3 , 5 ; O O l = 9 ; p ro no t u m crossed b y a s t ro n g t r a n s vers a l i m p ress i on , s h i n y , w i t h co l l a r g r a n u l a t ed a n d d i sc w i t ho u t sc u l p t u re ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a no t u m h o l l o w b e h i n d sc u t e l l u m ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted , w i t h pos t e r i o r s u rface t r a n s verse l y s t r i a t e ; p l eu r a n o t t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d c o m p l e t e ; fore t a rs a l seg m e n t s i n fo l l o w i n g p roport i o n s : 1 3 : 3 : 5 : 1 9 : 26 ; en l a rged c l a w ( F i g . 1 1 98 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h 4 b r i s t l es ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 1 1 98 ) w i t h 2 rows of 4 + 29 l a me l l a e ( s h o r t e s t row n o t

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e x t e n d i n g b e y o n d 0 , 5 l en g t h of segme n t ) ; a p e x w i t h a g ro u p of 1 4 l a m e l l a e ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown L O C U S T Y P I C U S : E I Sa i t o ( Me x i c o ) T Y P I CAL M A T E R I AL : ho l ot y pe F ' i n O T . HOST S : u n k no w n D I S T R I BU T I O N : Nearc t i c reg i on : M E X I C O :

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Ch alcogon a top u s a rgyrias R . C . L . Perk i n s 1 907 : 2 0 .

F E MAL E : a p t erou s ; l e n g t h 3 , 00-3 , 66 m m ; red d i sh- y e l l ow , w i t h p e t i o l e a n d a b domen b l a c k ; a n t e n n a I s e g me n t s 8- 1 0 b ro w n i s h ; h e a d e x c a v a t e d , s h i n y , g r a n u I a t e d ; p r o n o t u m crossed b y a t r a n sversa I i m pres s i on , g r a n u I a t e d ; s c u t u m l o n g e r t h a n b roa d ; m e t a t h o r a x + p ropodeum g r a n u l a ted ; pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; s e g me n t 4 o f fron t t a r s u s s l i g h t l y shorter t h a n s e g me n t 1 ( 30 : 32 ) ; e n l arged c l a w ( F i g . 1 1 99 ) w i t ho u t s u b a p i c a l too t h , w i t h a sma l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 0-5 peg - l i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 1 99 ) w i t h 1 row of 1 1 or 3 ( p rox i m a l ) + 1 9 l a me l l a e ; a p e x w i t h a g ro u p of 1 3- 1 4 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : Nog a l es ( A r i z . , U . S . A . ) T Y P I CAL M A T E R I A L : l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 982 ) i n B . H O ST S : u n k no w n D I ST R I BU T I O N : N e a rc t i c reg i o n : U . S . A . : T a n b a r k F l a t ( Los A n g e l es Co . , C a l i f . ) , O L ! Nog a l es ( A r i z . ) , B !

2 6 . Gon a top u s c u rriei K rombe i n

Gon a top u s curriei K ro m b e i n 1 962 : 4 .

F E MAL E : a p tero u s ; l en g t h 3 , 00-3 , 43 m m ; red d i s h - y e l l o w ; i n some spec i me n s w i t h a s m a l l b ro w n oce l l a r spot , p e t i o l e b l ac k a n d ab domen b ro w n -red d i s h ; a n t en n ae redd i s h - y e l l ow ; i n some spec i me n s w i t h segmen t 1 0 b row n ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I s e g m e n t s i n fo l l ow i n g propor t i on s : 9 : 5 : 1 7 : 1 0 : 7 : 7 : 5 : 5 : 5 : 8 ; h e a d e x c a v a t e d , s h i n y , w ea k l y g r a n u l a t e d , w i t h frons u n s c u l p t u red ; occ i p i t a l c a r i n a i n v i s i b l e ; fro n t a l l i n e comp l e t e ;

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w i t ho u t scu l p t u re ; seg m e n t 4 of f ro n t t a r sus l o n g e r t h a n segmen t 1 ( 27 : 22 ) ; e n l arged c l a w ( F i g . 1 200 ) w i t ho u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d w i t h 8- 1 1 peg - l i k e h a i rs ; segmen t 5 of fro n t t a rs u s ( F i g . 1 200 ) w i t h 1 row of 1 7 l a m e l l ae or 2 rows of 3 ( p ro x i m a l ) + 2 1 l a me l l a e ; a p e x w i t h a g ro u p of a t l e a s t 1 4 - 1 5 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : P l u m m ers I s l a n d ( M d . , U . S . A . ) TYP I C A L MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y pe F ! i n WA . H O ST S : u n k n o w n D I ST R I B U T I O N : Nearc t i c reg i o n : U . S . A . : Co l u m b i a ( Mo . ) , O L ! Lo r i n g ( V a . ) , WA ! P l u m mers I s l a n d ( M d . ) , IVA !

WA !

Dunn

1 72 7

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Fig.

1 20 2 - C h e l a of Gon a to p u s brooksi n .

27 . Gon a top u s in n i t idu s

A gon a top u s inn itidus R . C . L . ,� gon a to p u s s u tura l i s F e n t o n

( R . C . L . P e rk i n s )

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P e r k i n s 1 907 : 3 1 . 1 9 27 : 9 ( n . sy n . ) ; l o c u s typ i c u s T u cson

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F E M AL E : a p t e ro u s ; l en g t h 2 , 56-3 , 4 3 m m ; red d i s h - t e s t aceo u s , w i t h p e t i o l e b l ac k a n d a b d omen b row n , h e a d ( F i g . 1 1 86 A ) a l most f l a t , d u l l , g r a n u l a t ed ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , g r a n u l a t ed , d u l l ; sc u t u m s l e n de r , g r a n u l a ted ; m e t a n o t u m g r a n U l a t ed , n o t t r a n sverse l y s t r i a t e ; m e t a t h o r a x + p ropode u m d u l l , g r a n u l a t ed ; p l e u ra a n d pos t e r i o r s u rface of p ropod e u m n o t t r a n s v e r se l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; fore t a rsa l segmen t s i n fo l l ow i n g p ropor t i o n s : 1 5 : 2 : 5 : 1 5 : 23 ; en l a r g ed c l a w ( F i g . 1 20 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4-8 peg - l i k e h a i rs ; segmen t 5 of f ro n t t a r s u s ( F i g . 1 20 1 ) w i t h 1 row of 1 0 l a m e l l ae ; a p e x w i t h a g r o u p of 7-1 3 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n L O C U S T Y P I C U S : No g a l es ( A r i z . , U . S . A . ) TYP I C A L M A T E R I A L : l ec t o t y p e F ' ( des i g n a t ed b y O l m i 1 98 2 ) of A . i n n i tidu s i n B ; h o l o t y p e F ! of A . s u t uralis i n 'NA . HOS T S : i n U . S . A . n o t i d e n t i f i e d C i c a d e l l i d s ( R . C . L . Perk i n s 1 907 ) . D I ST R I BU T l m� : Nearc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B ' T u cson ( A r i z . ) , WA !

1 7 28

Fig.

Mem . Amer .

Ent.

I nst . ,

1 203 - C he l a of Gon a top u s p a l l idiceps ( K a n sas )

no.

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(R.C.L.

28 . Gon a top u s brooksi n .

1 984

P e rk i n s )

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sp .

F E MAL E : a p t e rou s ; l e n g t h 3 m m ; h e a d t e s t aceo u s , w i t h a b ro a d b ro w n t ra n s v e r s a I b a n d o n v e r t e x ; a n t e n n a e b ro w n , w i t h segme n t s 1 -2 t e s t aceou s ; t h o ra x , p ropode u m , pet i o l e a n d a b domen b l ac k ; l eg s t e s t a c eo u s , w i t h c l u b s o f femora d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l segmen t s i n fo l l ow i n g proport i o n s : 6 : 5 : 1 0 : 7 : 6 : 6 : 5 : 5 : 5 : 6 ; h e a d s l i g h t l y exca v a t ed , d u l l , g ra n u l a t e d ; frons w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; O L = 2 ; O O L = 7 p ro n o t u m crossed b y a t r a n s v e rsa l i m p ress i on , d u l l , g ra n u l a t e d ; sc u t u m d u l l , g r a n u l a t e d ; m e t a t ho r a x + p ropode u m d u l l , g ra n u l a t ed ; pos t e r i o r s u rface of p ro p o d e u m t r a n s v erse l y s t r i a t e o n l y on pos t e r i o r h a l f ; p l e u r a n o t t r a n sverse l y s t r i a te ; meso me t a p l e u r a l s u t u re d i s t i nc t a n d comp l e t e ; fore t a r s a l seg m e n t s i n fo l l o w i n g p ropor t i o n s : 1 1 : 2 , 5 : 4 : 1 2 : 1 9 ; e n l arged c l a w ( F i g . 1 202 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow and w i t h 2 b r i s t l es ; seg m e n t 5 of f ron t t a r s u s ( F i g . 1 20 2 ) w i t h i n ner s i de p ro x i m a l l y n o t serra t e , w i t h 1 row of 20 l a me l l ae ; a p e x w i t h a g ro u p of 1 3 l am e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2

1 7 29

Gon a topod i n ae

Fig.

1 204 - Segme n t 5 of f ro n t t a rs u s of fem a l e o f Gon a top us p a l l idiceps ( R . C . L . P e r k i n s ) from F a rgo ( No r t h D a k o t a )

segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : S a s k a toon ( Sa s k a t c h ew a n , TYP I C AL M A T E R I A L : h o l o t y p e F ! i n OT .

Canada)

HOST S : u n k no w n D I ST R I B U T I O N : N e a rc t i c reg i o n : CANADA : S a s k a toon ( Sas k . ) , O T ! NOT E S : t he s p ec i es i s n a me d i n honor of M r . A . R . Brook s .

29 . Gon a top u s p a l l i diceps

( R . C . L . P e rk i n s )

Neogon a top u s p a llidiceps R . C . L . Perk i n s 1 905 : 45 . Gon a top u s p a l lidiceps ( R . C . L . P e rk i n s ) : K i effer 1 9 1 4 b :

F E MAL E :

a p t erou s ;

l en g t h 2 , 1 9-3 , 4 3 m m ;

1 26 .

h e a d y e l l o w o r fer r u g i n o u s or

b row n , w i t h occ i p u t and f ro n t p a r t of v e r t e x t e s t aceou s ; a n t e n n ae y e l l o w , w i t h s e g m e n t s 4 - 1 0 b ro w n i s h ; p rono t u m b l ac k or y e l l o w or ferr u g i n o u s , u s u a l l y w i t h d i sc a n d m o s t p a r t of s i des b row n ; scu t u m , sc u te l l u m , m e t a t h o r a x + p ropod e u m , p e t i o l e a n d a b domen u s u a l l y b l a ck ; a p e x of p ropodeum r e d d i s h ; somet i mes scu t u m , s c u t e l l u m and m e t a to r a x + p ropodeum t e s t ac eo u s , w i t h b ro w n spo t s ; l eg s y e l l ow o r f e r ru g i n o u s ; h e a d exca v a ted , s h i n y , g r a n u l a t ed ; p r o n o t u m crossed b y a t r a n s ver s a l i m p ress i o n , s h i n y , g r a n u l a t e d ; me t a t ho r a x + p ropodeum d u l l , w e a k l y o r s t ro n g l y g r a n u l a ted ; p os t e r i or s u r f a ce of p ropodeum a n d p l eu ra t ra n sverse I y s t r i a t e ; meso m e t a p l e u ra l s u t u re obso l e t e o r w e a k l y d i s t i n c t ; fore t a rsa l s e g m e n t s i n fo l l ow i n g p ro p or t i o n s : 1 0 : 2 : 4 : 1 1 : 1 7 ; en l a r g e d c l a w ( F i g . 1 20 3 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 2-7 p e g- l i k e h a i rs ; seg m e n t 5 of f ro n t t a rs u s ( F i g s 1 20 3 , 1 204 ) w i t h o n e row of 8-2 1 l a m e l l ae ; some t i mes t h i s row of l a me l l ae i s on d i s t a l h a l f o f segmen t ; a p e x w i t h a g r o u p o f 7 - 1 3 l a me l l a e ; m a x i l l a ry p a l p i w i t h 4-5 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 730

Mem . A m e r .

Ent.

I n st . ,

no.

37 ,

1 984

MALE : u n k n o w n L O C U S TYP I C U S : A l a m e d a ( C a l i f . , U . S . A . ) TYP I C AL MAT E R I AL : l ec t o t y pe F ! ( de s i g n a t e d b y O l m i 1 982 ) i n B . HOST S : i n U . S . A . D e l t o c ep h a l u s fuscinervosus V . D . ( F rey t a g 1 97 7 )

A thys a n u s s p . ( R . C . L . Perk i n s 1 906c ) ; Endria in im ic a . D I S T R I B U T I O N : N e a rc t i c reg i o n : CANADA : C h e r r y v i l l e ( B . C . ) , T E ' She l b u r n e ( N . S . ) , OT ! u . S . A . : A l a m e d a ( C a l i f . ) , 3 ! D a n v i l l e ( C a l i f . ) , CA l F a rgo ( N . D . ) , OL! IV A ! L a w rence ( D o u g l a s C o . , K a n s . ) , OL ' J o h n son C i t y ( T e n n . ) , CA l Pa l k C a . ( F l a . ) , IVA !

N E O T R O P I C S P E C I ES OF

GONA TOPUS

T we n t y four spec i e s a re k n ow n .


F E MALES

S e g men t o f f r on t t a r s u s a p p r o x i ma t e l y t w i c e a s l o n g a s Seg men t o f f r o n t t a r s u s s h o r t e r t h a n , a s l on g a s , o r s l l on g e r t h a n s eg m e n t 4 . . . . . . . . . . . . . . . . H e a d , t h o r a x a n d p r o p o d e u m f e r r u g i n ou s . 1 . fern a n din a e n . sp . H e a d mo s t I y b l a ck ; t h o ra x a n d p r o p o d e u m b l a c k . 2 . ore vifoI'ceps K i e f f e r Me s o -m e t a p l e u r a l su t u r e d i s t i n c t . . . . . . . . . . . . . . . . . . . . . . . . . .

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. 1 2 Me s o -me t a p l eu r a l s u t u r e ob s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . S c u t u m l a t e r a l l y w i t h t wo p a i n t s ( F i g . 1 2 3 4 B ) . . . . . . . . 5 S c u t u m l a t e r a l l y w i t h o u t t wo p o i n t s ( F i g . 1 2 3 4 A ) . . . . . . . . . . . . . 9 S t a l k b e t w e e n p r on a t u m a n d m e t a t h a r a x + p r o p ad e u m v e r y s l en d e r , mo r e t h a n t w i c e a s l o n g a s p r an a t u m y e l l ow . . . . . . . . . . . S t a l k b e t w e en p r on o t u m a n d l e s s t h a n t w i c e a s l on g a s

b r oad ( F i g . 1 2 3 4 B ) ; s t a l k a n d . 3 . a gilis n . s p . m e t a t h o r a x + p r o p o d e u m l e s s s l en d e r , b r o a d ; s t a l k a n d p r o n o t u m mo s t l y

6

b l a ck An t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m w i t h

7

f u r r ow a n d w i t h n u me r o u s t r a n s v e r sa l k e e l s ; m e t a n o t u m h o l l ow b eh i n d s c u t e l l u rn . . . . . . 5 . spinol a i O l m i A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u rn srno o t h , w i t h o u t s c u l p t u r e o r t r a ck s o f med i a n f u r r ow ; m e t a n o t u rn n o t h o l l ow b eh i n d s cu t e l l u m 7 Me s op l e u r a a n d me t a p l eu ra On d i f f e r e n t p l a n e s , b e c a u s e t h e k e e l

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1 73 1

Gon a topod i n ae

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su t u r e i s v e ry b r o a d a n d 4 . p eruvianus

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9

2 3 . b oli via n u s n . sp . S c u t u m a l mo s t f u l l y y e l l ow ; v e r t e x o f h ea d s h i n y , w i t h on l y o c e l l a r t r i a n g l e g r a n u l a t e d ; s i d e s o f v e r t e x s mo o t h a n d w i t h o u t s c u l p t u r e . . . . . . . . . . . . . . . . . 2 0 . arlei n . sp . Me t a t h o ra x + p r op od e u m w i t h a s t r o n g med i a n f u r r ow . •

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2 2 . contortus n . sp . p r op od e u m w i t h ou t med i a n f u r r o w . 1 0 T h o ra x a n d p r op od e u m r ed d i s h - t e s t a c e o u s . . . . . . . . . . . . . . . 6 . willinki n . sp . T h o r a x a n d p r op od eu m f u l l y o r mo s t I y b l a ck . 1 1 Me s op l e u r a a n d m e t a p l e u ra on t h e s a m e p l a n e ·

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1 218) 15 1 3 E n l a rg ed c l a w w i t h l a m e l l a e ( F i g . 1 2 1 5 ) ; t h o ra x a n d p r op od eu m du I I , g ranu I a t ed 9 . op a cithorax n . sp . E n l a rg ed c l aw w i t h o u t l a m e l l a e , w i t h b r i s t l e s ( F i g s 1 2 1 6 , 1 2 1 7 ) ; t h o r a x a n d p r op od e u m s h i n y , s m o o t h , w i t h ou t s c u l p t u r e . 1 4 1 4 H e a d w i t h OL n e a r l y a s l o n g a s P O L ; t h o r a x a n d p r op o d e u m m o s t l y r e d d i sh - t e s t a c e ou s d a rk . . 1 0 . p urp urascens n . sp . H ea d w i t h OL n ea r l y t w i c e a s l o n g a s POL ; t h o ra x a n d p r op od e u m mo s t l y b l a c k 1 1 . townesi n . sp . 1 5 S t a l k b e t we e n p r o n o t u m a n d m e t a t h o r a x + p r op od e u m v e r y l on g , l on g e r t h a n d i s c o f p r on o t u m ( F i g . 1 2 2 1 B ) . . . . . . . . . . . . . . . . . . . . •

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. 1 2 . apica lis C a m e r on S t a l k b e t w e en p r o n o t u m a n d me t a t h o ra x + p r op od e u m s h o r t , l on g a s , o r sh o r t e r t h a n d i s c o f p r o n o t u m ( F i g . 1 2 2 1 A ) . 1 6 T h o r a x a n d p r op od e u m mo s t l y y e l l ow - t e s t a c e o u s o r r e d d i s h T h o ra x a n d p r op od e u m f u l l y o r a l mo s t f u l l y b l a c k . . . . . 1 7 Me t a t h o r a x + p r op od e u m w i t h a med i a n f u r r ow . . . . . 2 1 . cilipes K i e f f e r Me t a t h o ra x + p r o p o d e u m w i t h o u t med i a n f u r r ow . •

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1 8 H ea d f I a t 1 3. H ea d e x c a v a t e d . 1 9 H e a d , t h o r a x a n d p r op od e u m f u I w i t h ou t s c u l p t u r e ( e x c e p t f o r H ea d , t h o r a x a n d p r o p od e u m d u I 2 0 S i d e s o f m e t a n o t u m r o u n d ed ( F i .

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1 4 . fl a vipes n . s p . S i d e s o f me t a n o t u m p r o t r u d i n g 1 5 . camp b elli n . s p . P o s t e r i o r s u r f a c e o f p r op o d eu m f u l l y t ra n s v e r s e l y •

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st r i a t e. . . . . .

1 6 . tristis n . s p . P o s t e r i o r s u r f a c e o f p r op o d eu m w i t h o n l y p o s t e r i o r h a l f t ra n s v e r s e l y s t r i a t e . . . . . . 1 7 . doellojura doi ( O g l ob l i n ) 2 2 P o s t e r i o r su r f a c e o f p r op o d eu m s t r o n g l y t ra n s v e r s e l y s t r i a t e ; me t a t h o ra x + p r op od e u m w i t h o u t s c u l p t u r e a m o n g s t r i a e 1 9 . l a c u a li s n . s p . P o s t e r i o r su r f a c e o f p r op o d eu m w i t h o u t t ra n sv e r s a l s t r i a e , •

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w i t h on l y a f e w s t r i a e n ea r a p ex ; me t a t h o r a x + p r op o d e u m g ra n u l a t ed , d u l l 23 2 3 Me t a t h o ra x + p r op od e u m s l e n d e r , w i t h a n t e r i o r s u r f a c e s l i g h t l y i n c l i n ed ( F i g . 1 2 2 6 A ) 1 8 . s ilvestrii K i e f f e r Me t a t h o ra x + p r op o d e u m sq u a t , w i t h a n t e r i o r s u r f a c e m o r e i n c l i n ed ( F i g . 1 2 2 6 8 ) . . . . 2 4 . a u tumna lis n . s p . •

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MALES

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N o t au N o t au D o r sa 1 206 ) D o r sa 1 208 )

l i c e s c o mp l e t e . . 13. l i c e s i n c omp l e t e l p r o c e s s of g o n o f o r c e p s 1 . l p r o c e s s of g o n o f o r c ep s 2. .

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2 mu c h s h o r t e r t h a n p en i s ( F i g . fern a n di n a e n . s p . s l i g h t l y sh o r t e r t h a n p en i s ( F i g . breviforceps K i e f f e r •

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T h e m a l e s o f t h e o t h e r s p ec i es a r e u n k n ow n .

1 . Gon a top u s fern a n di n a e n . s p .

F E MAL E : a p terou s ; l e n g t h 1 , 87-2 , 00 m m ; t e s t a c eo u s , w i t h p e t i o l e b l ac k a n d a b domen p a r t l y b row n i s h ; h e a d f l a t , s h i n y , w i t ho u t scu l p t u re ; p ronot u m s h i n y , crossed b y a t r a n s ve rs a l f u rrow , w i t h o u t scu l p t u re , h a i r l es s ; s c u t u m d u l l , g r a n u l a ted ; m e t a n o t u m t r a n s v e rse l y s t r i a t e , met a t h o r o x + p ropodeum s h i n y , w e a k l y g r a n u l a ted ; p o s te r i o r s u rface of p ro p o d e u m a n d p l eu r a t r a n s v erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 o f f ro n t t a rs u s n e a r l y t w i ce a s l o n g a s segmen t 4 ( 8 : 4 ) ; en l a rged c l a w ( F i g . 1 205 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5 p e g- l i ke h a i r s ; segmen t 5 of fron t t a rs u s ( F i g . 1 205 ) w i t h i n n er s i de p ro x i m a l l y serra t e , w i t h 2 r o w s of 8 + 9 l a m e l l ae ; a p e x w i t h a g r o u p of 7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4

G o n a topo d i n ae

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1 205 - C he l a of Gon a top u s fern a n din a e n . s p .

1 73 3

( t y pe )

segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : fu l l y w i n ged ; l en g t h 1 , 68 m m ; b ro w n - l i g h t ; a n ten n a l seg m e n t s i n fo l l o w i n g p roport i o n s : 4 : 3 5 , 5 ; 5 , 5 : 4 , 5 : 4 : 4 : 5 : 4 , 5 : 6 ; an t e n n a l segmen t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 3 , 5 ) ; head s h i n y , h a i r y , a l u t aceou s ; fro n t a l l i ne abse n t ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 7 ; OL = 3 ; OOL = 2 ; brea d t h of t h e oce l l i : 4 scu t u m

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1 206 - M a l e g en i t a l i a o f Gon a top u 5 fern a n din a e n .

sp .

from G a l a p agos

I s l ands

s h i n y , s h i n y g ra n u l a ted , h a i r y ; n o t a u l i ce s i n v i s i b l e ; scu te l l u m a n d m e t a n o t u m s h i n y a n d smoot h ; p ro p o d e u m smoo t h a n d sh i n y , w i t h o u t sc u l p t u re , w i t h a m ed i a n f u r row o n dors a l s u rface ; dors a l p rocess o f gonoforceps ( F i g . 1 20 6 ) short a n d s l e n d e r , w i t h a p e x rounded ; m a x i l l ary p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Fer n a n d i n a I s l a n d ( G a l a p a g os I s l a n d s , E c u a dor ) TYP I CAL MA T E R I A L : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n CA ; 2 p a r a t y p e s ( 1 F , 1 M ) ! i n OL. HOST S : u n k n o w n D I S T R I B U T I O N : GALAPAGOS I SLA N D S : F e r n a n d i n a I s l a n d , C A ! OL ! S a n t a C r u z I s l a n d , CA ! OL !

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1 207 - C he l a o f Gon a top u s bre viforcep s K i e ffer ( t y p e )

2 . Gon a topus bre viforceps K i effer

Gon a top u s bre viforceps K i effer 1 904 a : Pa chygon a top u s brev i forceps K i effer ) :

356 . K i e ffer i n K i effer e t

M a rs a h a l l 1 906 : 50 . Cyrtogon a top u s breviforceps

( I< i effer ) : K i e ffer 1 907b : 1 9 . ( K i e ffer ) : K i effer 1 9 1 4 b : 70 . Cyrtogon a to p u s breviforceps ( K i e ffe r ) : B r u c h 1 9 1 5 : 443 . Netagon a top u s brasiliensis O g l o b l i n 1 9 32 : 264 ( n . s y n . ) ; l o c u s typ i c u s G u a y r a ( B r a z i I ) . Cyrtogo n a top u s breviforceps ( K i e ffe r ) : R i c h a r d s 1 969 : 80 . Cyrtogon a top us bre viforceps ( K i e ffe r ) : O l m i a n d C u rrado 1 976a : 1 65 . Cyrtogo n a top u s breviforceps

F E MA L E : a p terou s ; l e n g t h 2 , 4 3-3 , 00 m m ; b l ac k , w i t h fron t p a r t of v er t e x , c l y p e u s a n d m a n d i b l e s y e l l o w ; a n t e n n a e b ro w n , w i t h segmen t s 1 -3 y e l l ow ; l e g s y e l l o w , w i t h co xae , c l u b s of femora a n d b o t h a p exes of m i d a n d h i n d t i b i ae b row n ; h e a d f l a t , s h i n y , h a i r l es s , g r a n u l a te d ; p ro n o t u m crossed b y a t r a n svers a l i m p res s i o n , s h i n y , g r a n u l a ted ; scu t u m g r a n u l a ted , s h i n y ; m e t a n o t u m n o t t r a n s v erse I y s t r i a te ; m e t a t h o r a x + p ropodeum sh i n y , g r a n u l a te d ; p l e u r a a n d p o s te r i or s u rface of p ro p o d e u m n o t t r a n s v erse l y s t r i a t e ; meso-met a p l e u r a l s u t ure obso l e t e ; seg men t 1 o f front t a r s u s n e a r l y t w i ce a s l o n g a s segmen t 4 ( 1 24 : 7 ) ; en l a r ged c l a w ( F i g . 1 20 7 ) w i t ho u t s u b a p i c a l too t h a n d l a me l l ae , w i t h a s m a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d 3-6 p e g - I i ke h a i r s ; s e g m e n t 5 of fron t t a r s u s ( F i g . 1 207 ) w i t h i n n e r s i de p ro x i m a l l y not serra t e , w i t h 2 r o w s of 8 - 1 2

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1 20 8 - M a l e gen i t a l i a of Gon a topus breviforceps K i e f f e r f ro m R i o P i e d r a s ( Pe r u } { r i g h t ha l f remo v e d )

v e r y s m a l l l a m e l l a e ; a p e x b roadened , w i t h a g r o u p of 6-8 l am e l l ae ; m a x i l l a ry p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u rs 1 , 0 , L MALE : fu l l y w i n ged ; l e n g t h 1 , 3 1 - 2 , 00 m m ; b l ac k ; m a n d i b l e s t e s t aceo u s ; a n te n n a e a n d l e g s n o t b a n de d , b row n ; a b domen b row n ; a n te n n a e n o t d i s t a l l y t h i c j k e ned ; a n te n n a l se g m e n t s i n fo l l o w i n g p ro p or t i o n s : 5 : 4 : 8 : 8 : 7 : 7 : 6 : 6 : 6 : 7 ; a n t e n n a l seg men t 3 more t h a n t h ree t i m e s a s l o n g a s

Gon a topod i n a e

1 737

b ro a d ( 3 , 25 ) ; h e a d s h i n y , w e a k l y g r a n u l a t e d ; fron t a l I i ne a b se n t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 6 ; O L = 2 ; OOL = 2 ; b r e a d t h of t h e oce l l i : 4 ; sc u t u m w e a k l y g r a n u l a t e d ; not a u l i ces i n comp l e t e , reac h i n g a p p ro x i m a te l y 0 , 5 l e n g t h of sc u t u m ; sc u t e l l u m , m e t a n o t u m a n d p ropode u m smoo t h , s h i n y , w i t ho u t scu l p t u re ; dors a l s u r f ace o f p ropodeum w i t h a m e d i a n f u rrow ; fore w i n g s h y a l i n e , w i t ho u t d a rk t r a n sv e r s a l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ; gon oforceps ( F i g . 1 208 ) w i t h a dors a l p rocess l o n g a n d n a r row ; b a s i v o l se l l a w i t ho u t notc h ; m a x i I l a r y p a l p i w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C U S : S . t a C r u z ( A r g en t i n a ) T Y P I CAL M A T E R I A L : h o l o t y p e F ! of G. breviforceps i n G E ; h o l o t y pe F ! a n d 4 p a ra t y pes F F ! o f M. bra siliens is i n P T . HOST S : u n k n o w n D I ST R I BU T I ON : Neo t rop i c reg i on : BAHAMAS : New P rov i d ence , BM ! HA I T I : P t . P r i n ce , WA ! D O M I N I CA N R E P U B L I C : R a n c ho A rr i b a , O T ! E I R i o ( L a V e g a ) , O L ! O T ! P E R U : R i o P i e d r a s , O L ! WA ! BRAZ I L : E s c a d a ( P e r n a m b u co ) , B ! G u a y ra ( P a r a n a ) , P T ! A R G E N T I NA : S . t a C r u z ( P a t agon i a ) , GE !

3 . Gon a top u s agilis n .

sp .

F E MALE : a p terou s ; l e n g t h 4 , 0 m m ; h e a d t e s t aceou s ; a n te n n ae y e l l o w t e s t aceou s ; p ro n o t u m b l a c k or fu l l y tes t aceou s , w i t h red d i s h n u a nces on s i des ; sc u t u m and sc u te l l u m ye l l ow ; m e t a t ho r a x + p ropodeum b l ac k , w i t h a p e x r e d d i s h ; a bdomen b l ac k or re d d i s h - b ro w n ; l e g s test aceou s n , w i t h coxae a n d c l u b s of femora p a r t l y b rown or fu l l y tes t aceou s ; h e a d exca v a te d , s h i n y , w i t hou t scu l p t u r e ; p rono t u m crossed b y a s t r o n g t ra n s v e rs a l i m p ress i on , smoo t h , s h i n y , w i t ho u t scu l p t u re ; sc u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s , l a t e ra l l y w i t h two po i n t s ( F i g . 1 234 a ) ; s t a l k b e tween p rono t u m a n d me t a t ho r a x + p ropodeum more t h a n t w i ce a s l o n g a s b roa d ; m e t a n o t u m s h i n y , n o t t r a n sverse l y s t r i a t e ; m e t a t ho r a x + p ropodeum s h i n y , smoo t h , w i t ho u t scu l p t u r e ; p l e u ra a n d pos ter i o r s u rface of p ropodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l eu r a l s u t u re d i s t i nc t a n d comp l e t e ; fo re t a rsa l seg m e n t s i n fol l ow i n g p ropor t i o n s : 1 1 : 2 : 3 : 1 3 : 1 9 ; en l a r g e d c l a w ( F i g . 1 209 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 8 peg- l i k e h a i r s ; seg m e n t 5 o f f ron t t a r s u s ( F i g . 1 209 ) w i t h 2 rows of 4 + 20 l a me l l ae ; apex w i t h a g roup of 1 3 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : T i b u ( C o l om b i a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n BM ; 1 p a ra t y p e F ! i n LE . HOST S : u n k n o w n D I ST R I BU T I O N : Neo t rop i c reg i o n : PANAMA : G a m b o a ( C a n a l Zone ) , L E ! COLOMB I A : T i b u ( M a r ac a i bo B a s i n ) , B M !

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Gon a top u s pel'u v i a n u s n . sp .

F E MALE : a p terou s ; l en g t h 3 , 2 5 mrn ; h e a d b row n , w i t h occ i p u t , c l y p e u s , frons a l o n g orb i t s a n d m a n d i b l es t e s t a c eo u s ; a n t e n n a e t e s t a ceo u s , w i t h segmen t s 6- 1 0 b row n ; t h o r a x a n d p ropodeum b l ac k ; scu t u m l i g l> ' , p e t i o l e b l ac k ; a b domen b l a c k - b ro w n ; l e g s b row n , w i t h t a r s i l i g h t ; h e a d exca v a ted , s h i n y , smoot h , w i t ho u t scu l p t u r e ; p ron o t u m crossed b y a t r a n s v e rs a l i m p ress i on , h a i r y , smoo t h , w i t ho u t SCU l p t u re ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; s t a l k b e t ween p ron o t u m a n d m e t a t h o r a x + p ropode u m l es s t h a n t w i ce as l o n g as b roa d ; me t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h o u t scu l p t u re , h a i r y , w i t h n u merous t r a n s v e r s a l k ee l s o n pos t e r i o r s u rface ; p l e u r a w i t ho u t t r a n s v e r s a l k ee l s ; meso-me t a p l eu r a l s u t u r e d i s t i n c t a n d comp l e t e ; mesop l e u ra a n d met a p l e u r a o n d i f f e re n t p l a nes ; se g m e n t 1 o f fro n t t a r s u s s h o r t e r t h a n seg m e n t 4 ( 9 : 1 1 ) ; e n l a r ged c l a w ( F i g . 1 2 1 0 ) w i t ho u t s u b a p i c a l too t h , w i t h sm a l l too t h at the e n d of a

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1 2 1 0 ) w i t h i n n e r s i de pro x i m a l l y serr a t e , w i t h 2 rows of 5 + 1 7 l a m e l l ae ; a p e x w i t h a g r o u p of 1 5 l a me l l a e ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p ; w i t h 2 segmen t s ; l a b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C U S : T i ngo M a r i a ( Peru ) TYP I CAL MAT E R I A L : h o l o t y p e F ! i n C A . HOS T S : u n k n o w n D I ST R I B U T I O N : Neo t rop i c re g i o n : P E R U : T i n go M a r i a ( Monzon V a l l e y ) , CA '

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c l y p e u s , m a n d i b l es a n d fron t p a r t o f v e r te x ; a n t e n n a e t e s t aceo u s ; t h o r a x , p ropod e u m , p e t i o l e a n d a b domen b l ac k ; p ro n o t u m w i t h b ro w n s p o t s on s i des ; l e g s tes t aceo u s , w i t h cox a e , t i b i a e a n d c l u b s of femora b row n ; h e a d

Go n a topod i n ae

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1 2 1 2 - C h e l a of Gon a topu5 v,illinki n . s p .

1 74 1

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exca v a te d , s h i n y , g r a n u l a t ed ; p rono t u m crossed b y a t r a n s v e rs a l i mp ress i on , s h i n y , smoo t h , w i t hou t sc u l p t u r e ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m s h i n y and smoo t h ; me t a t hor'ax + p ropodeum

1 742

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1 984

s h i n y , w i t h n u m erous t r a n s v e r s a ! k ee l s on p o s t e r i o r s u r face , p l e u r a a n d a n te r i o r s u r f ace ; a n t e r i o r s u r f a c e w i t h t ra c k of med i a n f u rrow ; m e t a n o t u m h o l l o w b eh i n d sc u t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d co m p l e t e ; fore t a rs a l segm e n t s i n fo l l o w i n g p ropo r t i o n s : 8 : 2 : 3 : 8 : 1 4 ; en l a r g e d c l a w ( F i g . 1 2 1 1 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d 8 p e g - l i k e h a i rs ; segmen t 5 of f r o n t t a rs u s ( F i g . 1 2 1 1 ) w i t h 2 r o w s o f 5-1 6 l a m e l l a e ; a p e x w i t h a g r o u p of 1 1 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : L a Seren a ( C h i l e ) TYP I C A L M A T E R I A L : ho l o t y p e F ! i n CA ; 1 p a r a t y p e F ! i n SP . HOST S : u n k n o w n D I ST R I B U T I O N :

N e o t r o p i c reg i o n :

CH I L E :

L a Seren a ,

6 . Con a to p u s w i l l i n k i n .

CA !

Ch i l e ,

SP !

sp .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 7 5 m m ; red d i s h , w i t h a n t e n n a I seg men t s 7 - 1 0 b ro w n i s h , p e t i o l e b l ac k , a b do m e n b l ac k ( e x c e p t for red d i s h n u a nces o n t e r g i t e 1 ) ; l e g s re d d i s h , w i t h c l u b s o f femora a n d c o x a e b ro w n i s h ; h e a d e xc a v a te d , d u l l , g r a n u l a te d ; p r o n o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s , l a t e r a l l y w i t h o u t p o i n t s ; m e t a n o t u m f l a t , s h i n y , w i t h o u t scu l p t u re a n d t r a n s v e rs a l s t r i a e , not or s l i g h t l y ho l l o w b e h i n d sc u t e l l u m ; m e t a t h o r a x + p r o p o d e u m s h i n y , w i t h o u t s c u l p t u re ; me t a p l e u r a a n d p os t e r i o r s u r face of p ro p o d e u m t r a n sverse l y s t r i a t e ; mesop l e u r a n o t t r a n s v e rse l y s t r i a t e ; meso m e t a p l e u r a i s u t u re d i s t i n c t and comp l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ro p or t i o n s : 1 2 : 2 : 4 : 1 6 : 24 ; en l a r ge d c l a w ( F i g . 1 2 1 2 ) I-V i t hou t s u b a p i c a l too t h a n d s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow , on l y w i t h 9 peg - l i k e h a i rs ; a p e x of en l a r g e d c l a w p o i n t e d a n d n o t b e n t a t r i g h t a n g l e ; se g m e n t 5 o f f ro n t t a rs u s ( F i g . 1 2 1 2 ) w i t h 2 rows of 7 + 20 l a m e l l a e ; a p e x w i t h a g r o u p of 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k nown LOCUS TYP I C U S : T ucuman ( Argen t i n a ) TYP I CAL MAT E R I A L : ho l o t y p e F ! i n T U . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : I" RGE N T I NA : T u c u m a n , TIJ ! 1-1 0 T E S : t he spec i es i s n a med i n h o n o r of M r . A . W i l l i n k .

7 . Gon a top u s p s e u dorb i t a l i s n .

F E MA L E :

a p t e ro u s ;

l e n g t h 3 , 0-4 , 0 m m ;

h e a d b r ow n ,

sp .

w i t h f ro n t p a r t of


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1 2 1 3 - C h e l a of Gon a top u s pseudorb i t a lis n . s p .

1 743

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v e r t e x , c l y p e u s a n d ma n d i b l e s y e l l o w ; a n t e n n ae y e l l ow , w i t h segme n t s 7- 1 0 b row n ; pron o t u m b l a c k , w i t h re d d i s h s i d es ; scu t u m , scu t e l l u m a n d m e t a t h o r a x + p ropodeum b l a c k ; abdomen b r ow n , w i t h redd i s h n u a nces ; l e g s red d i s h ; h e a d exc a v a t e d , s h i n y , w i t h o u t s c u l p t u r e , excep t f o r occ i p u t a n d f r o n t p a r t of v e r t e x g r a n u I a ted ; p r on o t u m c rossed b y a t r a n s v e r s a I i mp ress i o n , smoo t h , h a i r y , sh i n y , w i t h o u t s c u l p t u re ; s c u t u m s h i n y , w i t h

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some l o n g i t u d i n a l kee l s ; m e t a n o t u m smoo t h , n o t t r a n s v e r se l y s t r i a t e ; m e t a t h o r a x + p ropodeum sh i n y , smoo t h , w i t h o u t scu l p t u re ; p l e u ra a n d p o s t e r i o r s u r face o f p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a a n d met a p l e u r a o n t h e s a m e p l a n e ; segm e n t 1 of f ro n t t a rs u s s h o r t e r t h a n segmen t 4 ( 1 3 : 1 9 ) ; e n l a r g e d c l a w ( F i g . 1 2 1 3 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d 1 2 p e g -- l i k e h a i rs ; seg m e n t 5 of f ro n t t a rs u s ( F i g . 1 2 1 3 ) w i t h 2 rows of 4 + 2 3 l a me l l ae ; a p e x w i t h a g ro u p of a t l e a s t 1 3 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : S . J u a n ( Co s t a R i c a ) TYP I CAL MA T E R I A L : ho l o t y p e F ! i n OL ; 1 p a r a t y p e F ! i n A M . HOST S : u n k n o w n


D I ST R I B UT I ON :

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COSTA R I C A :

1 745

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8 . Gon a top u s orb italis C a meron

Gona top us orb it alis C a m e ron 1 888 : 442 .

F E MALE :

a p tero u s ;

l e n g t h 2 , 56-4 , 06 m m ;

h e a d b l ac k or b row n ,

w i t h c l ypeus

a n d m a n d i b l es red d i s h - y e l l o w ; i n s o m e spec i mens a l so f rons p a r t l y testaceo u s ; a n te n n a e redd i s h , w i t h seg men t s 3 - 1 0 o r 7- 1 0 b row n ; t h or a x , p ropod e u m ,

pe t i o l e a n d a b domen b l ac k ;

l eg s b r o w n o r b l ac k o r t e s t a c eo u s ;

t a r s i l i g t h ; h e a d e x c a v a ted , d u l l , g r a n u l a ted ; i n some spec i me n s s h i n y a n d w i t ho u t sc u l p t u re ; p rono t u m crossed b y a t r a n s v ers a l i mp res s i on , s h i n y , smoo t h , w i t ho u t s c u l p t u re ; sc u t u m ( F i g . 1 2 34 A ) d u l l , w i t h SOme l o n g i t u d i n a l k e e l s ; m e t a n o t u m very short , smoo t h , not t r a n s verse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h 0 L1 t scu l p t u re ; p l e u r a a n d poste r i o r s u r f a c e o f propod e u m t r a n s v e rse I y s t r i a t e ; meso- m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a a n d m e t a p l eu r a o n d i f f e ren t p l a nes , b ec a u se t h e kee l a l o n g t h e meso-me t a p l e u r a l s u t u re is very b road and s t e p - s h a p e d ; segmen t 1 of f ron t t a r s u s shorter than segmen t 4 ( 1 4 : 1 8 ) ; e n l a r g e d c l a w ( F i g . 1 2 1 4 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r ro w a n d 8 - 1 1 l a m e l l ae , p l u s 1 b r i s t l e ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 2 1 4 ) w i t h 2 rows o f 28-32 l a me l l ae ; a p e x w i t h a g r o u p of a t l e a s t 1 5-30 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S TYP I C U S : C h i r i q u i vo l c a n o ( P a n a m a ) TYP I CAL MA T E R I A L : h o l o t y p e F ! i n BM . H O ST S : u n k no w n D I ST R I B U T I O N : N eo t rop i c re g i on : PANA M A : C h i r i q u i v o l c a n o , BM ! C h i r i q u i H a r t m a n n s ( 5 . t a C l a r a ) , OT ! V E N E Z U E LA : S a n E s te b a n , WA I

9 . Gon a topus op a cithorax n . sp .

F E MA L E : a p terou s ; l e n g t h 3 , 62 m m ; h e a d t e s t aceous-d a r k , w i t h m a n d i b l e s t e s t a c eo u s ; a n te n n ae t e s t a c eo u s ; t h o r a x a n d p ro p o d e u m b l ack , w i t h s i de s o f p rono t u m test aceo u s - d a r k a n d w i t h pos t e r i o r h a l f of sc u t u m y e l l o w ; a p e x of p ropode u m test aceo u s ; a b domen b l ac k ; l e g s t e s t aceo u s , w i t h coxae p a r t l y b row n , fore femora a n d fore t i b i a e a l most f u l l y b ro w n , m i d a n d h i n d femora p a r t l y b row n ; a n t e n n a e d i s t a l l y t h i c k e n ed ; a n t e n n a I segmen ts i n fo l l o w i n g p ropor t i o n s : 1 0 : 7 : 28 : 1 5 : 1 2 : 1 1 : 9 : 7 , 5 : 6 : 1 0 ; h e a d s h i n y , esc a v a t e d , smoo t h , w i t h o u t sc u l p t u re ; f rons w i t h an i n com p l e t e med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 2 ; Ol = 2 , 5 ; O O l = 8 , 5 ; p ron o t u m

1 746

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1 2 1 5 - Ch e l a of Gon a top u s op acithorax n . s p .

( type)

d u l l , crossed b y a s t ro n g t r a n svers a l i m p ress i on , g r a n u l a ted ; sc u t u m s h i n y ; met a n o t u m f l a t , g r a n u l a t e d ; m e t a t h o r a x + p ropodeum d u l l , g ra n u l a te d , w i t h p o s te r i o r su rface t r a n s v erse l y s t r i a te ; meso-me t a p l e u r a l s u t u re obso l e te ; sc u t e l l u m w i t h t w o l a tera l p o i n t s ; fore t a rs a l s e g me n t s i n fo l l o w i n g p ropor t i o n s : 1 6 : 4 : 8 : 1 7 : 30 ; en l a r ged c l a w ( F i g . 1 2 1 5 ) w i t ho u t s u b a p i c a l too t h , w i t h 3 b r i s t l e s a n d 6 l a me l l ae ; segmen t 5 of fro n t t a rs u s ( F i g . 1 2 1 5 ) w i t h i n n er s i de p ro x i m a l l y not serra t e , w i t h 1 row o f 6 l a m e l l a e ; a p e x w i t h a g r o u p of n e a r l y 23 l a me l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : M e t a t e ( Me x i co ) TYP I C AL MAT E R I A L : ho l o t y p e F ! i n T W . HOST S : u n k n o n w D I ST R I B U T I O N : N e o t rop i c reg i o n : M E X I C O : M e t a t e ( T u x tepec , O a x a c a ) , TW !

1 0 . Gon a top u s p u rp urascens n .

sp .

F E M AL E : a p t e rou s ; l e n g t h 2 , 7 5 m m ; re d d i s h - t e s t aceo u s , w i t h p e t i o l e b l ac k a n d a b domen d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l seg m e n t s i n fo l l o w i n g p ropor t i o n s : 7 : 6 : 1 9 : 1 0 : 8 : 7 : 6 : 6 : 5 : 7 ; head e xc a v a ted , s h i n y , smoo t h , w i t h o u t sc u l p t u re ; fro n s w i t h w e a k med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; O L = 2 ; OOL = 9 ; p ron o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m w i t h o u t l a te r a l p o i n t s ; meso t ho r a x a n d m e t a t ho r a x + p ropodeum s h i n y , smoo t h , w i t ho u t sc u l p t u re , excep t for pos t e r i o r s u r face o f p ropodeum a n d p a r t of me t a p l e u r a t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a rsa l segmen t s i n fo l l o w i n g proport i o n s : 1 3 : 3 : 6 : 1 2 : 23 ; en l a rged c l a w ( F i g . 1 2 1 6 ) w i t ho u t s u b a p i c a l too t h , w i t h o u t sm a l l tooth a t t he e n d of a l o n g i t u d i n a l f u r row , w i t h a p e x n o t po i n ted , rou n ded , w i t h 1 row of 5

1 74 7

Gon a topod i n a e

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