MEMOIRS of the AMERICAN ENTOMOLOGICAL INSTITUTE Number 37, Part 1 Pages I-XII, 1-946
A REVISION OF THE DRYINIDAE (HYMEN...
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MEMOIRS of the AMERICAN ENTOMOLOGICAL INSTITUTE Number 37, Part 1 Pages I-XII, 1-946
A REVISION OF THE DRYINIDAE (HYMENOPTERA) by Massimo Olmi Universit::i degli Studi della Tuscia Viterbo, Italy
The American Entomological Institute 5950 Warren Road, Ann Arbor, Michigan 48105, U. S. A. 1984
TABLE OF CONTENTS
Introduction . . . . . .
............ 1
'.Iaterial
.. . . . . . . . . . 1 .4
examined.
~cknowledgments
3 i
0
.•
.5 .7 .. ... 7 .. .. 10 · .31 . . . . 31
logy . . . . . . . . . .
~osts
attacked ••..•
::'conomic
importance.
',',orphology
and
terminology.
::ollection . • . . . . . • . . • • . • • . • ::; reserva t ion . . . . . . . . . . . . . . . "ami Iy
size and distribution . . . . . . . . .
<ey
the
to
subfami lies
of
Sdbfami Iy Aphelopinae.. :;enus Aphelopus . . . . . . . . Palaearctic species. 1.
me I a I eu cu s .•..
2.
atratus ..
3.
serra tus.
4.
camu s ••..
5. 6.
nigriceps. n epa I en sis.
7. querceus ... Ethiopian species . . . . 1.
mediocarinatus.
2.
wittei . . . . . . .
3.
incisus . . . .
Oriental
species.
. .. 43 , •. 44
5. 6.
leucopus ..
.78
di ffusus ..
. . . 46
7.
t r in ita tis .
· .47 · .50
8.
alvarenganus . . . . . .
.79 .80 .80
9.
jamaicanus . . . .
.52 .54 .56 .57
Australian
1.
townesi . . . .
2. 3.
guineensis. australis ..
4.
ca I ed on i cu s .
.60 . ... 60
5.
a I bop i c t us ..
1. 2.
nearctica.
.. .. 62 .64 . ... 64
borneanus .. maculiceps.
.81 .. .81 · .. 82
species.
.59
.. .. 65 .66 .66 3. malayanus •.. 4.ochreus . . . . . .67 . 67 5. 0 r i en t a lis • • • . . . . . . .68 6. penanganus .. 1. 2.
.32 .34
Dry i n i dae •.
· .82 · .. 83 . . . . . 83
· .• 84 Genus Crovettia . . . . . . . . . . . . . . . 84 Nearctic species . . . . . . . . . . . 84
.... 85 · .87
the I i ae ..••.•
Neotropic species • . . . . 1. barbara . . . . • . . 2.
· .89 .89
4.
neotropica •••••••.••• 92 plaumanniana. · .92 brasiliana . . . . . . . . . . . 93
5.
3.
colombiana.....
..94
7.
birmanus . . . . . • . . . .
.69
Subfami Iy Biaphelopinae.
..94
8.
phi I ippinus ..
.69
Genus Biaphelopus.......
. .95
\Iearctic
species ...
.69
1.
ru f i v en t r is.
2.
bicolor . . . .
.70 .71
3. 4.
varicornis.
.72
a I bop i c t us. species.
.73 .76 .76
~eotropic
1.
tropicalis • . . . • • • .
2.
diaphanus . . . .
· .77 . .. 77
3.
su r i namen 5 is.
4.
fen t on i . . . . • . • . . . • • . . 7 B
Palaearctic 1.
species...
.96
masneri.........
.96
Subfami Iy Conganteoninae. Genus Conganteon . . . . . . .
Palaearctic species. 1.
nepalense . . . . .
Ethiopian
species.
1.
townesi .•..
2.
vulcanicum.
Genus Chelanteon . . . .
.96
.98 .. . . . 98 .. . . . 98 · .99
.100 .103 . . . . . . 105
II
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
species . • • • . • • . . . 105
22.
insignis . . . . . . . . . . . . 193
richardsi . . . . . . . . . . . 105
23.
pseudobilobus ..••••• 193
Genus Fiorianteon ••.••••••••. 108 speci es . . . . . . . 1 09
24. 25.
rubrolobatus . • • • . . . • 197 nigrolobatus • . • • . . . . 199
junonium . . . . . . . . . . . . . 109
26.
vagans . . . . . . . . . . . . . . 201
Subfami Iy Anteoninae . . . . . . . . . 110
27.
pilosifrons . . . . . . . . . 204
Oriental 1.
Pa I aearct i c 1.
28.
politus . • . . . . • • • . • • . 206
species •••••..•• 114
29.
claripes . . . . . . . . . . . . 206
1.
aerias . . . . . . . . . . . . . . 115
30.
chiapasi • • . . . . . • • . . . 209
2.
fuscum . . . . . . . . . . . . . . . 116
31.
n i 9 r i can s ••••••••••. 21 1
Genus Deinodryinus . . . . . . . . . . . 118
32.
actuosus • • • . . . • • . . . • 211
Palaearctic species . . . . . . . 121 1. areolatus . . . • • • • • . . . . 121
33.
incaicus . . . . . . . . . . . . 214
34.
bol ivianus ••...••••• 217
Ethiopian
eminens . • . • . • . . . . . . • 217
Genus Metanteon . . . . . . . . . . . . . . 113 Neotropic
species . . . . . . . . . 123
35.
1. 2.
insulanus . . . . . . . . . . . 124 madecassus ••..•••.•• 125
36.
trinidadi •••...•••.• 220
37.
rapax . . . • • • • • . . . . . • . 223
3.
madagascariensis .••• 128
38.
croceus . • • • • . . . . . • . • 223
4.
ben 0 i t i . . • • • • • • • • . . . 1 29
39.
5.
mon t i co Ius . • • • • . • . . • 130
40.
asper ••••.•••••••... 226 schl ingeri .•.••••.•• 230
sp e c i e s •••••••••. 1 3 1
41.
alexandrae . . . • • • . . . . 230
as i at i cu s •.•.•••••••. 131
42.
carpens .••••••.•.••• 232
species . . . . . . . . . . 133
43.
per I u c en s . . • • . • . • . . . 233
1. 2.
at r i v en t r is ••••••••• 1 33 quercicolus . . . . . . . . . 143
44. 45.
i nermi s .••••••••••.• 233 paranus . • . . . • • • • • • . . 236
3.
paradoxus . . . . . . . . . . . 145
46.
hi r tic 0 rn is •••..•••. 237
Or i en t a I 1.
Nearctic
species . . . . . . . . . 147
47.
rivularis • • • . . . . . . . . 239
e I egan s • • • • . . . . . . • • • 154
48.
aequa lis . • • . . . . . . . • • 240
Neotropic 1.
2•
sp e c i 0 su s ••••••••••• 1 57
49.
minor ••.••••••..•••• 241
3.
diaphanus . . . . . . . . . . . 158
50.
ben i anus • • . . . . . • • . • • 243
4.
n i grorufus ••••...••. 159
51.
i t en e z i . . . • • • • • • . . . . 245
5.
iphias ••....•••••••• 160 bicolor • • • • • • . . • . • . . 163
52.
b i I ob 0 ide s .••••••••• 245
53. 54.
f I uv i at i i i s . . • • . • . . . 248 cuzcanus ••••••.••.•• 249
6. 7.
maximus ••••••••••••• 165
8. 9.
melanocephalus . . . . • . 168
10.
11.
a t I an t i cu s .•.••••••• 172 achterbergi .•.•••••• 174
12.
peruvianus . . • • . • • . . • 174
13. 14.
hymenaeus . . . . . . . . . . . 178 colombianus . . . . . . . . . 180
15. 16. 17.
cascus . . . . . . . . . . . . . . 182 albopictus . . . . . . . . . . 183 amoenus •••••••••••.. 185
18.
pegnai .•.••••••••••• 186
19.
pseudoamoenus •••••.• 188
noyesi •••••••••••••• 170
55. panamensi s . . . • • . . . . . 251 Genus Lonchodry i nus . . . . . . • . . . 253 Palaearctic
1. 2. 3.
ruficornis • • . . • • . • • . 258 nepalensis ..•••••••• 267
Ethiopian
1. 2.
species . . . . . . . 255
subapterus . . • . • • • • . . 256
species • . . • • • . . . 269
seyrigi •••••••••••.• 269 paul iani . . • • • • . • . . . . 271
Oriental
species •.••.••••. 271
1.
sin en sis ••••••••.... 273 Nearctic species . . . . . . . . . . 273
20.
townesi . . . . . . . . . . . . . 190
1.
f I a vu s • • • • . . • . . • . • • • 274
21.
b i I obu s . . . . . . . . . . . . . 191
2.
masneri . . . . . . . . . . . . . 277
Table of contents
bakeri . . . . . . . . . . . . . . 278 3. Neotropic species • . . . . . . . . 282 1. tricolor . . . . . . . . . . . . 282 Australian species . . . . . . . . 284 1. guineensis . . . . . . . . . . 284 2. notogeicus . . . . . . • . . . 286 :;enus Anteon . . . . . . . . . . . . . . . . . 286 Palaearctic species . . . . . . . 290 1. jurineanum . . . . . . . . . . 294 2. brachycerum . . . . . . . . . 300 3. reticulatum . . . . . . . . . 304 4. nepalense . . • . . . . . . . . 304 5. arcuatum . . . . . . . . . . . . 305 6. flavicorne . . . . . . . . . . 309 7. ephippiger . . . . . . . . . . 313 8. pubicorne . . . . . . . . . . . 316 9. scapulare . . . . . . . . . . . 322 10. infectum . . . . . . . . . . . . 325 11. tripartitum . . . . . . . . . 329 12. crassifrons . . . . . . . . . 332 13. gaul I ei . . . . . . . . . . . . . 334 14. fulviventre . . . . . . . . . 337 15. hilare . . . . • . . . . . . . . . 342 16. ingenuum . . . . . . . . . . . . 342 17. corax . . . . . . . . . . . . . . . 345 18. muni tum . . . . . . . . . . . . . 345 19. japonicum . . . . . . . . . . . 347 Ethiopian species . . . . . . . . . 348 1. brachypterum . . . . . . . . 353 2. agile . . . . . . . . . . . . . . . 354 3. maritimum . . . . . • . . . . . 357 4. kasaicum . . . . . . . . . . . . 362 5. fiorii • . . . . . . . . . . . . . 362 6. kivuanum . . . . . . . . . . . . 363 7. taylori . . . . . . . . . . . . . 366 8. dayi . . . . . . . . . . • . . . . • 367 9. canabense . . . . . . . . . . . 370 10. ugandanum . . . . . . . . . . . 372 11. kawandanum • . . . . . . . . . 374 12. afrum . . . . . . . . . . . . . . . 375 13. turneri . . . . . . . . . . . . . 378 14. townesi . . . . . . . . . . . . . 379 15. decellei . . . . . . . . . . . . 380 16. anthereon . . . . . . . . . . . 381 17. namibiense . • . . . . . . . . 383 18. gutturnium . . . . . . . . . . 384 19. arab i cum . . . . . . . . . . . . 387
20. 21 . 22. 23. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34 . 35. 36 . 37. Or i en 1. 2. 3. 4. 5. 6. 7. 8.
III
natalense . . . . . . . . . . . 388 hoyoi . . . . . . . . . . . . . . . 390 mingoae . . . . . . . . . . . . . 391 long i scapum . . . . . . . . . 392 madagascolum . . . . . . . . 393 medleri . . . . . . . . . . . . . 395 hova . . . . . . . . . . . , . . . . 397 compressicoxale . . . . . 398 inf latrix . . . . . . . . . . . 399 rufonigrum . . . . . . . . . . 402 oriphi lum . . . . . . . . . . . 403 fabulosum • . . . . . . . . • . 404 abruptum . . . . . . . . . . . . 404 zairense . . . . . . . . . . . . 406 mo sse Ii . . . . . . . . . . . . . 408 capei . . . . . . . . . . . . . . . 409 eme r i tum . . . . . • . . . . . . 41 a garambanum . . . . . . . . . . 411 t a I sp e c i e s . . • . . . . . . . 41 2 borneanum . . . . . . . . • . . 416 thai . . . . . . . . . . . . . . . . 416 s i I v i co I um . . . . . . . . . . 419 indicum . . . . . . . . . . • . . 419 mysorense . . . . . . . . • • . 422 yasumatsui . . . . . . . . . . 424 sarawaki . . . . . . . . . . . . 425 bengalense . • . . . . . . . . 428 9. k r omb e i n i . . . . . . . . . . . 429 10. mui ri . . . . . . . . . . . . . . . 432 11. laotianum . • . . . . . . . . . 435 12. fyanense . . . . . . . . . . . . 435 13. javanum . . . . . . . . . . . . . 437 14. mun roe i . . . . . . . . . . . . . 439 lankanum . . . . . . . . . . . . 441 15. nemora Ie . . . . . . . . . . . . 442 16. si Ivestre . . . . . . . . . . . 444 17. expol itum . . . . • . . . . . . 444 18. pet ersen i . . . . . . . . . . . 445 19. bauense . . . . . . . . . . . . . 446 20. debi Ie . . . . . . . . . . • . . . 447 21. 22. phi I ippinum • . . . . . . . . 448 Nearct ic species •.•... '" •. 450 1. popenoe i . . . . . . . • . . . . 452 2. arizonense . . . . . . . . . . 455 3. canadense . . . . . . • • . . . 457 4. nebulosum . . . . . . . • . . . 461 5. funestum • . . . . . . . • • • . 462
IV
Mem. Amer.
En!.
6. wasbaueri . . . . . . . . . . . 464 7. minimum • . . . . . . . . . . . . 467 8. masoni . . . . . . . . . . . . . . 469 9. rugosiceps . . . . . . . . . . 471 10. puncticeps . . . . . . '" .474 11. xanthothorax . . . . . . . . 479 12. osborni . . . . . . . . • . . . . 481 13. durangoi . . . . . . . . . . . . 484 14. ciudadi . . . . . . . . . . . . . 486 15. menkei . . . . . . . . . . . . . . 486 16. hirtifrons . . . . . . . . . . 487 Neotropic species . . . . . . . . . . 488 1. clavatum . . . . . . . . . . . . 491 2. mo I Ie . . . . . . . . . . . . . . . 492 3. nigrorubrum . . . . . . . . . 494 4. pan amen se . . . . . . . . . . . 497 5. jamaicanum . . . . . . . . . . 497 6. mexicanum . . . . . . . . . . . 499 7. oliveirai . . . . . . . . . . . 502 8. noyesi . . . . . . . . . . . . . . 503 9 . I ob a tum . . . . . . . . . . . . . 506 10. propodeale . . . . . . . . . . 506 11. chi riquense . . . . . . . . . 508 12. nycteum . . . . . . . . . . . . . 510 13. annul icorne . . . . . . . . . 511 14. albitarse . . . . . . . . . . . 512 15. v i vax . . . . . . . . . . . . . . . 514 16. p iii corn e . . . . . . . . . . . 51 5 17. conterminum . . . . . . . . . 516 pi aumann i . . . . . . . . . . . 517 18. 19. bol ivianum . . . . . . . . . . 518 micros . . . . . . . . . . . . . . 519 20. 21. surinamense . . . . . . . . . 521 Austral ian species . . . . . . . . . 522 1. gi luwense . . . . . . . . . . . 526 2. chelogynoides . . . . . . . 527 myrmecophi lum . . . . . . . 529 3. 4. reticulaticeps . . . . . . 531 au I i cum . . . . . . . . . . . . . 533 5. sedlaceki . . . . . . . . . . . 534 6. 7. anxium . . . . . . . . . . . • . . 536 8. australe . . . . . . . . . . . . 538 9. parvu I um . . . . . . . . . . . . 539 10. orientale . . . . . . . . . . . 541 11. aculeatum . . . . . . . . . . . 545 12. corruptum . . . . . . . . . . . 545 13. miles . . . . . . . . . . . . . . . 547
Ins!.,
no. 37,
1984
14. destructor . . . . . . . . . . 547 15. caledonianum . . . . . . . . 551 16. typicum . . . . . . . . . . . . . 553 17. bougainvi I lei . . . . . . . 555 18. curradoi . . . . . . . . . . . . 556 19. firmum . . . . . . . . . . . . . . 558 20. leiosomum . . . . . . . . . . . 560 21. ni t idum . . . . . . . . . . . . . 561 22. bouceki . . . . . . . . . . . . . 564 23. mundum . . . . . . . . . . . . . . 565 24. bel lator . . . . . . . . . . . . 568 n i gri corne . . . . . . . , .. 568 25. fij ianum . • . . . . . . . . . . 569 26. dub i um . . . . . . . . . . . . . . 570 27. cor i aceum • . . . . . . . . . . 571 28. su ck lin 9 i . . . . . . . . . . . 573 29. bi smarckense . . . . . . . . 575 30. gloriosum . . . . . . . . . . . 575 31 . vulsum . . . . . . . . . . . . . . 576 32. 33. I uctuosum . . . . . . . . . . . 577 34. hagen i . . . . . . . . . . . . . . 578 haustum . . . . . . . . . . . . . 580 35. coral I inum . . . . . . . . . . 581 36. 37. tasmanianum . . . . . . . . . 582 permirum . . . . . . . . . . . . 583 38. involutum . . . . . . . . . . . 584 39. 40. superbum . . . . . . . . . . . . 585 41. rufiscapum . . . . . . . . . . 586 42. gi raul t i . . . . . . . . . . . . 587 Genus Prioranteon . . . . . . . . . . . . 589 Palaearctic species . . . . . . . . 589 1. biroi . . . . . . . . . . . . . . . 591 Ethiopian species . . . . . . . . . . 593 1. richardsi . . . . . . . . . . . 594 2. casalei . . . . . . . . . . . . . 596 Subfami Iy Bocchinae . . . . . . . . . . 599 Genus Bocchoides . . . . . . . . . . . . . 601 Ethiopian species . . . . . . . . . . 602 1. beki Iyensis . . . . . . . . . 602 Genus Chelothel ius . . . . . . . . . . . 604 Palaearctic species . . . . . . . . 605 1. gryps . . . . . . . . . . . . . . . 607 2. berlandi . . . . . . . . . . . . 609 Genus Bocchus . . . . . . . . . . . . . . . . 611 Palaearctic species . . . . . . . . 613 1. ita lieu s . . . . . . . . . . . . 6 1 4 2. umber . . . . . . . . . . . . . . . 614
v
Table of contents
3.
szelenyi . . . . . . . . . . . . 617
1.
4.
slovacus . • • . • • . . . . • • 618
2.
mi rabi lis . . • • . . • . . • . 687 infuscatus • . . . . . . . . . 690
5. 6.
pagl ianoi • • . . . . . . • . . 620 scaramozzinoi . . . . . . . 621
3.
dentatus . . . . . . . . . . . . 690
4.
townesi . . . • . . . . . . . . . 692
7. 8.
scobiolae • . • • . . . • . • • 622 europaeus . . . . . . . . . . . 624
5.
migratorius . . . . . . • . . 694
Oriental
9.
parthicus . . . . . . . . . . . 624
1.
species . . . . . . . . . . . 695
phi I ippinus . . . . . . . . . 697
species . . . . . . . . . . 627
2.
asiaticus . . . . . . . . . . . 697
1.
richardsi . . . . . . . . . . . 628
3.
malayanus . . . . . . . . . . . 699
2.
bini . . . . . . . . . . . . . . . . 629
Nearctic
3.
s ey rig i . • . . . . . . . . . . . 629
1.
:::thiopian
Jriental
species . . . . . . . . . . . 632
species . . . . . . . . . . . 700
perkinsi . . . . . . . . . . . . 700
Neotropic 1.
species . . . . . . . . . . 702
variegatus . . • . . • • . . . 703
1. 2.
laotianus . . . . . . . . . . 635 mu I u en sis . . . . . . . . . . . 635
2.
minimus . . . . . . • . . • . . . 704
3.
pedunculatus •.•••••• 636
3.
macilentus . . . . . . . . . . 706
4.
rubricus . . . . . . . . • . . • 639
'<earctic
species . . . . . . . . . . . 641
1.
mirabilis . • . • . . . . . . . 643
2. 3. 4. 5.
testaceus . . . . . • . . . . . 643 wasbaueri . . . • . . . • . . . 644
a r i z on i cu s . . . . . . . . . . 645
4.
rufus . . . . . . • . . • . . • . . 710
5.
bruchi . . . . • . . . . . . . . . 714
6.
clarus . • . . . • . • • . . • • . 715
Australian
species . . . . . . . . . 717
1.
flavus . . . . . . . . . . . . . . 718
2.
koebelei ••.••••••••• 721
f lavipes . . • . . . . • . . . . 650
Subfami Iy Dryininae . . . . . . . . . . 723
6. 7.
flavicoll is ••••.•••. 650
Genus Dryinus . . . . . . . . . . . . . . . . 728
I at i c ep s . . . . . . . . . . . . 652
8.
dubius • • . . . • • . . . . . . . 653
Palaearctic species . . . . . . . . 729 1. sanderi . . . . . . . . . . . . . 731
species . . . . . . . . . 655
2.
canariensis . . . . . . . . . 734
austral iae . . . . . . . . . . 656
3.
collaris . . . . . . . . . . . . 735
guineensi s ••••..••.. 658
4.
nepalensis . . . . . . . . . . 740
minimus . . . . . . . . . . . • . 658
5.
tarraconensis . . . . . . . 742
6.
koreanus . . . . . . . . . . . . 747
7.
an t i quu s . . • . . . • • . . • • 751
>\ustral ian
1. 2. 3. 4.
robustus . . . . . . . . . . . . 660
5.
bouceki . . • • . . . . • . . . • 661
:=""us Mirodryinus . . . . . . . . . . . . 663 "alaearctic species . . . . . . . . 664 1. gobiensis . . . . . . . . . . . 666
Ethiopian 1.
species . . . . . . . . . . 751
bisulcatus . . . • . . . . . . 754
2.
seyrigi . . . . . . . . . . . . . 756 alticolus . . . . . . . . . . . 758 spangleri . . . . . . . . . . • 762
3.
ungulatus . . . . . . . . . . . 671
3. 4.
4.
atlanticus . . . . . . . . . . 672
5.
deceptor . . . • • . . . . • • . 762
5.
ponomarenkoae . . • . . . . 674
6.
mahensis . . . . . . . . . . . . 764
:=enus Radi imancus . . . • • • • . . . . . 675
7.
sierranus . . . . . . . . . . . 766
species . . . . . . . . 676
8.
pretorianus . . . . . . . . . 767
1. olmii . . . . . . . . . . . . . . . 676 :::enus Mystrophorus . . . . . . . . . . . 678
9.
saussurei . . . . . . . . . . . 770
10.
hova • • • . . . . . . . . • . . . • 772
species . . . . . . . 678
11.
basilewskyi . . . . . . . . . 774
formicaeformis . . . • • . 678 Iy Thaumatodryininae .. 682
12.
chamaeleo . . • • . . . • • • . 775
13. 14.
schoutedeni . . . . . . . . • 778 mayogo . . . . • . . . . • . . . . 780
15.
orophi Ius • • . . . . • . . . • 781
2.
xerophi Ius • • . . . . • . . . 668
Palaearctic
Palaearctic 1. 5'~bfami
:=enus Thaumatodryinus • • • . . . . • 683 Ethiopian
species . . . . . . . . . 686
VI
Mem. Amer. Ent.
16 . mod est us . . . . . . • • . . . . 785 17. i ncertus . . . . . . . . . . . . 785 18. burgeoni . . . . . . . . . . . . 786 19. afer . . . . . . . . . . . . . . . . 787 Oriental species . . . . . . . . . . 791 1. irregularis . . . . . . . . . 794 2. kiefferi . . . . . . . . . . . . 795 3. longipes . . . . . . . . . . . . 799 4. pulcher . . . . . . . . . . . . . 799 5. ex iii s . . . . . . . . . . . . . . 802 6. k r omb e i n i . . . . . . . . . . . 802 7. stantoni . . . . . . . . . . . . 806 8. trifasciatus . . . . . . . . 807 9. latus . . . . . . . . . . . . . . . 808 10. indi cus . . . . . . . . . . . . . 810 11. praeclarus . . . . . . . . . . 814 12. fulvus . . . . . . . . . . . . . . 815 13. lucens . . . . . . . . . . . . . . 817 14. browni . . . . . . . . . . . . . . 817 15. lankanus . . . . . . . . . . . . 821 Nearctic species . . . . . . . . . . 822 1. mexicanus . . . . . . . . . . . 823 2. crawfordi . . . . . . . . . . . 825 americanus . . . . . . . . . . 826 3. alatus . . . . . . . . . . . • . . 828 4. Neotropic species . . . . . . , .. 831 1. constans . . . . . . . • . . . . 833 citricolus . . . . . . . . . . 835 2. f I a von i 9 e r . . . . . . . . . . 838 3. well ingensi s . . . . . . . . 839 4. surinamensi 5 • • • • • • • • 839 5. striatus . . . . . . . . . . . . 845 6. argentinus . . . . . . . . . . 846 7. 8. caraibicus • . . . . . . . . . 849 9. napensi s . . . . . . . . . . . . 850 opaci frons . . . . . . . . . . 851 10. 11. grand is . . . • • . . . . . , .. 852 a I va renganu s . . . . . . . . 857 12. 13. sin op en sis . . . . . . . . . . 858 14. bel i zen sis . . . . . . . . . • 858 anti I leanus . . . . . . . . . 860 15. 16. t eu t on i a e . . • . . . . . . . . 862 17. catarinae . . . • • • . . . . . 864 18. kimseyae . . . . . . . . . . . . 864 19. palaeodominicanus ... 866 Austral ian species . . . . . . . . 868 1. pallidus . . . . . . . . . . . . 871
Inst.,
no. 37,1984
2. speciosus . . . . . . . . . . . 874 3. subtilis . . . . . . . . . . . . 875 4. pacificus . . . . . . . . . . . 876 5. montanus . . . . . . . . . . . . 878 6. p seudophanes . . . . . . . . 878 7. insularis . . . . . . . . . . . 880 8. bouceki . . . . . . . . . . . . . 880 9. k 0 eb e lei . . . . . . . . . . . . 882 10. guineensis . . . . . . . . . . 884 11. gigas . . . . . . . . . . . . . . . 885 12. bi smarcki . . . . . . . . . . . 887 13. aterrimus . . . . . . . . . . . 888 14. biaki . . . . . . . . . . . . . . . 890 15. punctatus . . . . . . . . . . . 893 16. areolatus . . . . . . . . . . . 894 17.leptias . . . . . . . . . . . . . 895 18. quatei . . . . . . . . • . . . . . 897 19. dahmsi . . . . . . . . . . . . . . 897 20. glaber . . • . . . . . . . . . . . 900 21. austral ianus . . . . . . . . 900 22. venator . . . . . . . . . . . . . 903 23. papuanus . . . . . . . . . . . . 904 Genus Richardsidryinus . . . . . . . 907 Palaearctic species . . . . . . . 908 1. albrecht i . • . . . . . . . . . 909 maroccanus . . . . . • • . . . 910 2. corsicus . . . . . . . . . . . . 913 3. Ethiopian species . . . . . . . . . 914 1. e r ra t i cu s . . . . . . . . . . . 91 6 ca r i n i c ep s . . . . . . . . . . 91 8 2. 3. undu I a t us . . . . . . . . . . . 919 Ori ental speci es . . . . . . . . . . 922 1. indianus . . . . . . . . . . . . 922 2. pyri Ilae . . . . . . . . . . . . 925 Nearctic species . . . . . . . . . . 927 1. nearcticus . . . . . . . . . . 927 2. wetmorei . . . . . . . . . . . . 929 3. canadensis . . . . . . . . . . 930 Austral ian species . . . . . . . . 931 1. t r iii n ea t us • . . . . . . . . 932 Genus Tridryinus . . . . . . . . . . . . . 932 Ethiopian species . . . . . . . . . 934 1. ugandanus . . . . . . . . . . . 935 2. ampuliciformis . . . . . . 937 Nearctic species . . . . . . . . . . 940 1. serratus . . . . . . . . . . . . 941 2. testaceus . . • . . . . . . . . 941
VII
Table of contents
oculatus • . . • . . . • . • . . 945 3. Neotropic species •••.•••.• 947 1. maxiexophthalmicus .. 948 2.
ruficauda ..•••••.•.• 951
3.
poecilopterae ••••••• 952
4.
maximus . . • • • • . . . . . • • 954
5 •
s t ria tic ep s ••••••••• 956
6.
ru f i c ep s •••.•.•.•.•. 958
7. 8.
4.
barbarus ••.••••••.• 1018
Nearctic
species .•••.•.•. 1018
1.
inconsul tus . . . . . • . • 1019
2. 3.
do r sa lis ••••.•••••• 1 021 solaris . . . . . . • . • . . • ;f)24
4.
favreauae •••••••••. 1024
Neotropic
species ••.•.•.. 1027
1.
forestalis . • . • . . . • . 1027
nigroflavus ••.••••.. 961
2.
caraibicus .•.•.•.•• 1029
exophthalmicus •••••• 962
3.
ruber • • . . . . . . . . . . • . 1031
9.
quechuanus • . . . . • . • • • 965
4.
aterrimus • . • . • . • . • . 1031
10.
pi cescen s . • . • . • . . • . . 965
11.
andinus ..•.•.•••••.• 967
gibbosus • . • • . • . . • . . • 969 12. Genus Megadryinus .•••••.•.•.• 971
1.
species .•.•.•.•• 1033
amoenus .•.•••••.•.• 1 034
Neotropic
species • . . . . . • . 1037
1.
autumnalis •.•.•.•.• 1041
magnificus ••••.••... 971
2. 3.
bol ivianus . . • . . • • . • 1041 neotropicus •••....• 1044
4.
bicolor •••••••••.•. 1045
Genus Alphadryinus .•••••.•••• 973 Palaearctic
species . . • . . . • 973
1.
b ru e s i ••••••••••..•• 975
2.
balticus • . • . • • . • . . . • 978
Ethiopian
species ••••.•... 980
1. aberrans .•.••••••••• 980 Oriental species . . . • • • . • • . 982 1. asiaticus •••.•.•.••• 982 2.
Nearctic
species ..•.•.••• 971
Neotropic 1.
Genus Perodry i nuS . . • . • . • . . . • 1033
gibbosus . . . • . • . • • . . . 985
Neotropic
Palaearctic 1.
species . • . . . . 1046
zherichini •..•.•••. 1046
Subfami ly Transdryininae . . . . 1048 Genus Transdryinus •••••.•.•• 1048 Australian species . . • . • . . 1048 1.
notogeicus • . . . . • . . . 1048
species • • . • . • . . . 985
Subfami Iy Gonatopodinae ..•.• 1050 Genus Neodryinus • . . . • • . . • . • . 1065
1.
lamellatus •.•.•.•.•• 986
2.
panamensis . . . • . • . • . • 989
3.
parvus • • . • . • . . . . . . • • 989
4. 5.
pegnai . • . . • • • . • . • • . • 991 brasi I ianus . . • • • . . . . 992
6. 7.
rufus •••.••••••.•••• 995 ferrugineus •..•.•••• 995
8.
piceus • . • . • . • • • • . . . . 996
Australian
Genus Cretodryinus • . . . . • . . . • 1046
species ••.••••• 998 1. planus . . • . • . • • . • . • • . 998 Genus Mesodryinus ••.••.•••.• 1000 Palaearctic species .••••. 1001 1. dayi • • . • . • . . • . . . • . . 1003 2. niger •••••••••••••• 1005 Ethiopian species . • . . . . • . 1007 1. aethiopicus ••..•.•• 1008 Or i en t a I sp e c i e s . • . • . • • . • 1 01 1 1. 0 r i en t a lis • . • • . . . • . 1 0 1 1 2. peterseni •••••••••. 1015 3. viet. • . . • • . • . . • . . . • 1016
Ethiopian
species ••••.•.• 1067
1.
an t i quu s . . • . • • • • . • . 1069
2.
gigas • • . • . • . • • . . . . . 1071
Oriental species •.••.•.•. 1073 1.leptopus . . . . . . • . • • . 1076 2.
reticulatus . . • . . • . • 1076
3.
ch e I at us . • . • • . • • . . . 1 079
4.
rubescens . • • • . . • . . . 1081
5.
dolosus . • . • • • • . . • . • 1081
6.
robustus . • . • • . . . . • . 1082
7.
javanus . . • • • . . • . . . . 1085
8.
pseudodiffusus .•••• 1086
9.
diffusus . . . . • • . • • . . 1089
Nearctic species ••.•..•.• 1091 1.
typhlocybae • . . . . . . . 1091
2.
ball i • . • . . • . • • . . . . • 1095
Neotropic
species .•.•..•• 1097
1.
acuticol I is . • . • . . . . 1098
2.
dominicanus •.•..•.• 1101
Mem.
VIII
Amer.
Ent.
Inst.,
no.
37,
1984
albosignatus . . . . . . . 1 101 radialis . . . . . . . . . . . 1104
E t h i op i an
5.
peruvianus •••••.•.• 1107
Oriental
6.
maculicornis . . . . . . . 1107
3. 4.
7.
incaicus . . . . . . . . . . . 1109
8.
brachycerus ••••••.. 11 11
9.
trinitatis . . . . . . . . . 1111
Australian 1.
species •.•.•.. 1115
cheesmani . . . . . . . . . . 1116
2.
nelsoni . . . . . . . . . . . . 1118
3.
koebelei . . . . . . . . . . . 1120
4.
amboni . . . . . . . . . . . . . 1123
Genus Adryinus . . . . . . . . . . . . . . 1126 Ethiopian species . . . . . . . . 1126 1. cerrutii . . . . . . . . • . . 1128
1. 1.
sp e c i e s . . . . . . • . 1 1 77
ugandanus . . . . . . . . . . 1177 species . . . . . . . . . 1180
sinensis . . . . . . . . . . . 1180
Nearctic
species . . . . . . . . . 1180
1.
americanus . . . . . . . . . 1182
2.
mimus . . . . . . . . . . . . . . 1183
Neotropic 1.
species . . . . . . . . 1186
cubensis . . . . . . . . . . . 1188
Austral ian 1.
species . . . . . . . 1189
vitiensis . . . . . . . . . . 1191
Genus Pseudogonatopus . . . . . . . 1191 Palaearctic species . . . . . . 1193
1. 2.
fulgori . . . . . . . . . . . . 1194 camelinus . . . . . . . . . . 1197 rosellae •••...••.•• 1199
2.
nigerianus • . . • • . . . . 1130
3.
3.
bellicosus . . . . . . . . . 1132 owen i . . . • • . . . . . . . . . 1 135
4.
flavifemur . . . . . . . . 1202
5.
ortholabi s . . . . . . . . . 1203
4. 5.
6.
f
species . . . . . . . . . 1138
7.
dromedarius . . . . . . . . 1209
1.
gracilis . . . . . . . . . . . 1138
8.
albosignatus . . . . . • • 1209
2.
meridionalis . . . . . . . 1139
9. 10.
augustae . • • • • . . . . . . 1212 distinctus . . . . . . . . . 1212
1 1. 12.
P r i e sn e r i •••..•.••• 1 2 1 6 septemdentatus . . . . . 1217
rugidorsal is •••••.. 1135
Nearctic
Genus Echthrodelphax • . . . . . . . 1140 Palaearctic species . . . . . . 1140
0
ca r i lei . . . . • • • . . . 1 206
1. 2.
fa i r ch i I d i i . . . . . . . . 1 1 41 ba e t i cu s • . . . . . . . . . . 1 142
3.
longicollis ....•••• 1144
4.
hortusensis . . . . . . . . 1147
1.
pilosus . . . . . . • • . . . . 1222
5.
italicus . . . . . . . . . . . 1150
2.
perkinsiellavorus •• 1224
Ethiopian species . . . . . . . . 1152 1. migratorius . . . . . . . . 1152
3.
seyrigi . . . . . . . . . . . . 1225
4.
polli . . . • • . . . • • . . . . 1226
5.
ruens . . . . . . . . . . . . . . 1228 acutus . . • . . . . . . . • . . 1230
2. afer . . . . . . . . . . . . . . . 1155 3. obscurus . . . • . . . . . . . 1155 Ori enta I speci es • • • . . . . . • 1157 1. ru f us . . . . • . • . . . . . . . 1 1 57 2. fairchildii . . • . . . . . 1158 3. laotianus •...•••••• 1158 Nearctic species . . . . . • . . . 1160 1. krombeini . . . . . . . . . . 1160 Australian species . . . • . • • 1161 1. fa i r ch i I d i i . • • • . . . . 1 1 61 2. nigricoll is . . . • • . . • 1166 bi fasciatus . . . . . . • . 1169 4. pel op s . • . . . . . • • • . . • 1 1 69 Genus Acrodontochelys . . . . . . . 1172 Palaearctic species . . . . . . 1174 1. b ou c ek i . . . . . . . . . . . . 1 1 74
13.ligusticus •••..••.. 1219 Ethiopian
6.
species . . . . . . . . 1221
7 •
sp i ra cu I a r is .•••••. 1 233
8.
similis • • . . . . • . • • . . 1234
Oriental 1.
species . . . . . . . . . 1235
sarawaki . • . • . . • • . • • 1238
2.
sarawakensis . . . . . . . 1241
3.
nudus •.•••.•.••..•• 1245
4. 5.
malesiae . . . . . . . . . . . 1248 validus . . . . • • . . • . • • 1248
6. 7.
lankae • • . . . • • . . . . . . 1249 muiri . . . . . . . . . . . . . . 1251
8.
hospes . . . . • • . . . . . . . 1254
9.
sogatae . . . . . . . . . . . . 1254
10.
pusanus . . . . . . . . . . . . 1255
Nearctic
species .•.•••••• 1257
IX
Table of contents
1. 2. 3. 4. 5.
1258 freytagi canadensis . . . • • • . . . 1260 s joe s t ed t i . . . . . . . . . 1 263 ari zon i cus . . . • . . . . . 1266 ontarianus . . • • . . . . . 1269
6.
stenocrani . . . . . . . . . 1271
7. 8.
au t ox en ob ius .••.••. 1273 magnus . • . . . . . . . . • . • 1275
Neotropic
species . . . . . • . . 1276
1.
variistriatus . • . . . . 1276
2.
flavus . . . . . . . . • . . . . 1277
3.
delphacidi s . . . • • • . . 1279
4.
muesebeck i . . . . . . . . • 1280
5.
morenoi . . . • . . . . . . . . 1284
Australian species •••••.. 1284 1. nigricans . . . . . . . . . . 1287 2.
hospes . . . . . . . . . . • . • 1 290
3.
dichromus . . . . . . . . . . 1293
Neotropic 1.
species . . . . . • • . 1342
longichelatus
1342
Genus Apterodryinus . . . . . . . . • 1342 Palaearctic 1.
species . . . . • • 1344
gobiensis . . . • . • . . . . 1344
Ethiopian
species . . . . . . . . 1344
1.
somerset i . • . . . • • • . . 1345
2.
richardsi . . . . . . . . . • 1346
3.
rufulus • . • . . . . . . . • • 1347
Oriental 1. 2.
species . . . • • • • . . 1348
as i at i cu s . . . . . . . . • • 1 350 insularis . . . • . • • • • . 1350
Nearctic species . . . . • . . . . 1350 1. torvus . . . . . . . . . . . . . 1353 2.
secundus . . . . . . . • . . . 1355
3.
cal ifornicus . . . . . . . 1357
Neotropic
species ••••.••. 1358
1.
citrinus . . . . . . • • . . • 1359 arnaudi • . . . . . . . . • . . 1362
4.
saccharetorum . . . . . . 1294
2.
5.
kurandae •.••••••••. 1296
3 .
t est a c eu s ••••••..•• 1 362
6.
juncetorum . . . . . . . . . 1298
4.
menkei . . . . . • . • . . . • . 1364
7.
plurisulcatus . . . . . . 1299
5.
tijucanus . . . . . . • . . . 1365
8. opacus . . . . . . . . . . . . . 1302 '::;enus Donisthorpina . . . . . . . . . 1304
6. 7.
longicornis . . . . . . • . 1367 bellottii . . . . . . . . . . 1369
Palaearctic 1.
Oriental 1.
species •••..• 1305
pa I I ida . . . . . . • . • . • • 1306 species . . . . . . . . . 1308
asiatica • . . • • • • . . • . 1309
::;enus Agonatopoides . . . . . . . . . 1310 Palaearctic 1.
2. 3.
species •••.•• 1312
felix . . . . . . . . . . • . . 1312 sol idus . . . . . . . . . . . . 1316
canariensis . . . . . . • . 1318 Ethiopian species . . . . . . . . 1321
Austral ian species •••.••• 1371 1. not ogei cus . . . . . . . . . 1372 2.
insularis ••.••••••• 1374
Genus Dicondylus . . . . . • . . . . . . 1374 Palaearctic
dichromus . • • . • • . . . . 1376
2.
bicolor . . . . . . . . . . . . 1378
Ethiopian 1.
Oriental
fuscus • . . . . . . . . . . 1321
2. 3.
acuminatus . . . . . . . . . 1324 cafer . . . . . . . . . . . . . . 1325
Nearctic
4.
ruber . • . • • • . . . . . • • • 1327
Neot rop i c
5.
ceres . . . . . . . . . . . . . . 1328 species . . . . . . . . . 1330
1. 1.
1.
species . • . . . . . . 1383
pallidior . . . . . . . . . . 1383
1.
Oriental
species . . . . . . 1375
1.
species . . . . . . . . . 1384
indianus . . . • . • • • . • . 1384 species . . . . . . . . • 1387
americanus • . . . . . . . . 1388 spec i es •.••••.. 1390
nigrithorax . . . . . . . . 1390
Austral ian
species . . . . . . . 1392
1.
bengalensis . . . . . . . . 1333
2.
borneanus . . . . . . . . • . 1333
2.
rufus . . . . . . . . . . • . . . 1398
3.
insularis . . . . . . . . . . 1334
3. 4.
dubius . . . . . • . • . . • • . 1399 kiefferi . . . . . . . . . . . 1399
5.
primitivus . . . • . . • . . 1401
4. pyri Ilae • . • • . . . . . . . 1336 Nearctic species . . . . . . . . • 1338
1.
perkinsi . . . . . . . . . . • 1394
1.
synchromus . . . • . . • • . 1339
6.
oceanicus . . . . . • . . . . 1403
2.
calidus . . . . . . . . . . . . 1339
7.
alpinus . . . . . . . . . . . . 1404
x
Mem.
Amer.
En!.
Genus Haplogonatopus . • . • . . • . 1408 Palaearct i c
speci es ...••• 1408
1. 2.
apicalis ••.•••••.•• 1409 oratorius . . . . . . . . . . 1409
3•
at ra t us ••..••••.. • '. 1 41 3
Ethiopian
species . . . . . . . . 1415
Ins!.,
no. 37,
1984
Austral ian species . . . . . . . 1473 1. anomalus . . . . . . . . . . . 1473 Genus Trichogonatopus . . . . . . . 1473 Nearctic species . . . . . . . . . 1475 1. simulator . . . . . . . . . . 1475 Neotropic species . . . . . . . . 1475 1.
rich a rd s i • . . . • • . . . . 1 476
species . . . . . . . . . 1416
2.
hi sp i du s •..••••.••• 1478
ap i ca lis .•••••••••. 1 41 7
3.
pall idi tarsi s . . . . . . 1481
species . . . . . . . . 1421
4.
raptor . . . . • • • . . . • • • 1485
1.
insularis . . . . . . . . . . 1423
5.
albomarginatus . . . . . 1485
2.
hernandezae . . . . . . . . 1423
6.
neotropicus . . . • . . • . 1487
Australian species . . . . . . . 1424 1. v i t i en sis . . . . . . . . . . 1 425
7.
rubriceps . . . . . . . ... 1488
1.
katangae . . . . . . . . . . . 1415
Oriental 1.
Neotropic
2.
apicalis . . . . . . . . . . . 1427
Genus Pentagonatopus . . • • . • . • 1427 Nearctic 1.
species . . . . . . . . . 1428
borealis •••...••••. 1428
Genus Epigonatopus • . . . . • . . • • 1490 Austral ian species . . . . . . . 1490 1.
fallax . . . . . . . . . . . . . 1491
2.
sol i tarius . . . . . . . . . 1493 Genus Esagonatopus . . . • • • • . . • 14 93
montanus . . . . . . . . . . . 1430
Nearctic species . . . . . . . . . 1494 1. niger . . . . . . . . . . . . . . 1495
Genus Plectrogonatopoides ... 1431 Palaearctic species . . . . . . 1431
Genus Gonatopus . . • • • . . . • • • . . 1498
1. ch 0 r tin is •...•••••• 1431 Nearctic species . . . . . . . . . 1433 1. nearcticus . . . . . . . . . 1433
Palaearctic species . . . . . . 1502 1. vistosus . . . . . . . . . . . 1508 2. horvathi . . . . . . . . . . . 1509
Austral ian 1.
species . . . . . . . 1429
Genus Tetrodontochelys . . . . . . 1434 Palaearctic
species . . . . . . 1436
2.
perdebilis . . . . . . . . . 1497
3•
dod e r
4.
bi I ineatus .••...••• 1513 distinguendus . . . . . . 1513
0
i .••..••••••• 1 51 1
1.
p ed est r is ..••••••.. 1 437
5.
2.
sakai i . . . . . • • • • • . • . 1441
6.
striatus . . . . . . . . . . . 1518
3.
pulicarius . . . . . . . . . 1442
7.
vollenhoveni
4. 5. 6.
pseudoaudax •••••••. 1446 aegyptiacus . . . . . . . . 1446 audax ..•.••••••••.• 1 447
8. 9 •
iranicus . . . . . . . . . . . 1523 pop ov i • • . . . . • • • • . • . 1 523
10.
a Ib
Ethiopian species . . . . . ... 1449 1. ochreus . . . . . . . . . . . . 1450
11. 12.
suarezi . . . . . . . . . . . . 1528 dimidiatus . . . . . . . . . 1528
2.obscurus . . . . . . . . . . . 1452
13.
atlanticus . . . . . . . . . 1530
3.
14.
uni I ineatus . . . . . . . . 1532
rubripes . . . . . . . . . . . 1454
0
1521
lin ea t us . . . . • • . 1 527
4.
zai rensi s ••.•••...• 1454
15.
aegyptiacus . . . . . . . . 1533
5.
nigrior •.••.••.•••• 1458
16.
plumbeus . . . . . . . . . . . 1535
6.
afer . . . . . • • • • . . . . . . 1458
17.
barbatel Ius .•..•••. 1538
7.
varipes . . . . . . . . . . . . 1459
18.
ater •...•••..•••••• 1538
Oriental species . . . . . . . . . 1461 1. lucens . . . . . . . . . . . . . 1461
19.
I indbergi •••.•..••• 1541 lunatus • . . . • . . . • . . . 1541
20.
species . . . . . . . . . 1463
21 •
t en e r i f e i •.•••••..• 1 548
1.
plesius . . . . . . . . . . . . 1465
canari ensi s .•••••.. 1550
2.
unicus ..••••••••••• 1468
22. 23.
wit t me r i •...••••••• 1 551
3.
pecul iaris . . . . . . . . . 1470
24.
nigrosignatus . . . . . . 1554
Nearctic
Table of contents
25. planiceps •••••••••. 26. campestris •.•....•• 27. subtilis •••..•.•.•. 28. spectrum .••.• •.•• •• 29. bernard i . • . . . • . . . • . 30. formicarius ••.•.••. 31. mongolicus • . . • . . . . . 32. longus . • . . • . • . • . . . . 33. nivosus • . . • . . . . . . • . 34. brunneicollis .•.•.. 35. chersonesius •...••. 36. sepsoides . • . . . . . • • . 37. rhaensis • . . . . . . . . • . 38. barbieri •••.••....• 39. schenklingi . . . • . . . . 40. graecus • . . . • . . . . • . . Ethiopian species . . . . . . . . 1. meridionalis .•.••.. 2. longiusculus ••••... 3. austral is •• .•• ..•.. 4. pilosipes . . . . . . . . . . 5. guigliae ••.••.••••. 6. seyrigi . • . . . . . . . . • . 7. ankaratrus . . . . . . . . . 8. capensis •.••.••.... 9. mo sse Ii . . . • . . . . . . . • 10. rubrithorax . . . . . . . . 11. hyalinus ••.•.•••••. 12. ridens . . . . . -. . . . . . . . 13 • c ommu n is ••..••.•••. 14. luteipes . . . . . . . . . . . 15. bekilyanus . . . . . . . . . 16. incogni tus . . • . . . . . . 17. I eptothorax . . . . . . . . 18. a e t hi op i cu s ..•.••.. 19. tananarivei . . . . . . . . 20 . a rma t us . . . • . • • • . . . . 21. okahandjae . . . . . . . . . 22 . j oh n s i . . . . . . . . . . . . . 23. taylori . . . . . . . . . . . . 24. upembanus . . . . . . . . . . 25. benoi t i • • • . . . . . . . • . 26. trochantericus ...•. 27. opacus . . . . . . . . . . . . . 28. patrizii .••••••••.. 29. turneri . . . . . . . . . . . . 30. ericeti . . . . . . . . . . . .
1554 1556 1558 1560 1562 1564 1567 1567 1570 1570 1572 1573 1582 1585 1587 1587 1587 1590 1590 1593 1595 1596 1598 1601 1601 1 603 1605 1607 1607 16 10 1611 1612 1613 1615 1619 1620 1 623 1623 1 626 1628 1628 1631 1631 1634 1634 1637 1637
XI
Oriental species . . . . . . . . . 1. yasumatsui •..••.•.• 2. attenuatus . . . . . . . . . 3. schenkl ingi . . . . . • . • 4. mysorensis . . . . . . . . . 5. plebeius . . . . . . . . . . . 6. perpolitus . . . . . . . . . 7. javanus . . . . . . . . . . . . 8. cristatus . . . . . . . . . . 9. abjectus . . . . . . . . . . . 10. daunus . . . . . . . . . . . . . 11. phi I ippinus •.••..•• 12. maurus . . . . . . . . . . . . . 13. medius . . . • • . . • • . . . • 14. lucidus . . . . . . . . . . . . 15. magnus . . . • . • . . . • . . . 16. sitae . . . . . . . . . . . . . . 17. indianus . . . . . . . . . . . 18. thai . • • . . • . • . . . . . . . Nearctic species . . . . . . . . . 1. clavicornis . . . . . • . . 2. cyphonotus . . . . . . . . . 3. mayori . . . . . . . . . . . . . 4. portalensis . . . . . . . . 5. translucidus . . . . . . . 6. echo . . . . . . . . . . . . . . . 7. elongatus •••.••••.. 8. freytagi . . . . • . • • • . . 9. contortulus . . . . . . . . 10. planipes . . . . . . . . . . . 1 1. m i mo ide s . . . . . • . . • . . 12. agropyrus . . . . . . . . . . 13. frequens . . . . . . . . . . . 14. areolatus . . . . . . . . . . 15. harpax ••.•••..••... 16. solitarius . . . . . . . . . 17. herbarum . . . . . . . . . . . 18. paraleptias . . . . . . . . 19. pseudochromus . . . . . . 20. flavifrons . . . . . . . . . 21. rap ax .•..••.....••• 22. ashmeadi . • . . . . • • . . . 23.0mb rodes . . . . . . . . . . . 24. moni lis ••..••..•••. 25. argyrias . . . . . . . . . . . 26. curriei . . . . . . . . . . . . 27. innitidus . . . . . . . . . .
1639 1641 1641 1645 1647 1647 1650 1652 1652 1655 1656 1656 1660 1662 1662 1665 1665 1667 1668 1670 1673 1676 1679 1679 1682 1683 1684 1685 1688 1689 1 693 1695 1697 1700 1 703 1703 1704 1709 1712 1712 1715 1715 1720 1723 1724 1724 1 727
XII
Mem.
Amer.
En!.
28.
brooksi . . . . . . . • . • • . 1728
29.
pal I idiceps •••••••. 1729
Neotropic
species .•••.•.. 1730
1.
fernandinae • . . . . . . . 1732
2.
breviforceps .•••••• 1735
Ins!.,
no. 37,
1984
cont ortus . . . • • • . . . . 1768
22. 23.
bol ivianus . • • . . . . . . 1768
24.
autumnalis .•••••••• 1770
Australian
species . . . • . . . 1771
1.
melanias . . • • . . . . . . . 1772 williamsi . . . . . • • • . • 1774
3.
a9ilis • • . . . . • • . . . . • 1737
2.
4.
peruvianus . • . . • . . . . 1738
3.
zealandicus . . . . • • • . 1776
5. 6.
sp i no I a i . . . . . • • • • • • 1 739 wi II inki . . . . . . • • . . . 1783
4. 5.
9i9as •...•••••••••• 1779 p I i ca t us . • . . . . . . . . . 1 783
7.
pseudorbitalis . . . . • 1742
6.
rossi • . • . • . . . . . . . . . 1783
8.
orbitalis .••••••... 1745
7. 8.
naumann i • . . . . . . • • • • 1785 papuanus . . . . . • . . . . . 1786
9.
opacithorax . . . . • • • • 1745
10.
purpurascens •..•••• 1746
1 1. 12.
town e 5 i . . . . . . . . • . . . 1 748 apicalis . . . . . • • • • • • 1750
13. 14.
bartletti ..•.•••••• 1751 flavipes . . . . . • . • • . . 1752
9.optabilis . • . • • • . . . . 1786 10. 11.
decoratus . . . . . • . . . • 1788 austral iae . . . • • . . . • 1790
12.
pulcherrimus ..••••• 1791
Genus Gynochelys . . . . . • . . . • . . 1792 Ethiopian
species . . . . • • . . 1793
15.
campbelli .•••.••••• 1755
16.
tristis . • • • • . . . . • . . 1757
17.
doellojuradoi ••...• 1757
Subfami Iy Apodryininae . . • . . . 1796
18.
si Ivestri i •.••••..• 1759 lacualis . • . . • • . . . . • 1762
Genus Apodryinus .•••••••••.. 1796
1.
braunsi . . . . • . . • • • • • 1793
Neotropic species . . • . • . . • 1796 19. 1. masneri •..••••••••. 1796 20. arlei ••••.•..•••••• 1764 cilipes . • . . • . . • • . . • 1764 21 • Nomina dubia and incertae sedis species • • . . • . . . • . . . . . . . . . . • • . . 1798 Nomina nuda • • • • • • • • • • • • • . • • • • • • • • • • • • . . . . . . . . . • • • • . . . . . • • . • • • • 1829 F0 ss i I
dry i n ids . . . . . . . . . . • • • • . . . . . • . • . . . • • . . . . . . • . . . • • . . • . • . . . 1 830
Parasites of Host
hopper
dryinids . . . . . . • . • • . . . . . . . . . . . • . • . • • • . . . • . • . . . . • . • 1831 records • . . • . . • . • . . • • • . . . . . . . . . . . . • • . . • • . • . . • . . • • • • 1834
Ref e r en c e s . . . • . . . • . . . . . . . . . . . . . . . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . 1 847 Abb rev i at i on s . . . . • • • • • • • • • . . . . • . . • • • • • • • • • • . . . . . . . . • • • • • • • • . . . 1869 Index to hosts . . . • . . . . . . • . • • • . . • . . . . . . . . . . • . . . . . . . . • • . . . . . . . . • 1869 Index to the parasites and predators of dryinids . . . . . . . . . • • • • . 1875 Index
to dryinids . . • • • . • • . • . . . . . . • . . . . . • . . . • • • • . • . . . . . • . . . . . • • l877
A REVISION OF THE
DRYINIDAE
(HYMENOPTERA)
by f.iassimo Olmi
Institute of Plant Protection, Via 5.
Camillo de Lellis,
University of Tuscia, 01100 Viterbo,
Italy
I iHRODUCT I ON
The Dryinidae, one of the smaller fami I ies of the acu leate ,:jymenoptera, is included in the superfamily Chrysidoidea (sensu Day 1977). The
literature on dryinids is wide.
or unrel iable because of
inc'lr rect
Jnrel iable taxonomic descriptions. :roublesome. ~evision
of
::>y Kieffer .J,
c),
a
The more useful
key
.c,ustralian,
however,
(1939),
the family,
is a
to the European
is useless
identifications or
incorrect and
Desides Richards'
monograph of the world species
treatment of the I~earctic dryinids by
a
Oriental
it,
The nomenclature is often
literature on
the British species (1914b),
Much of
or questionable
Fenton
(1918a,
Aphelopus by Jervis (1977), some studies on
and Nearctic dryinids by
R.C.L.
Perkins
(1903,
1906a,
1906b, 1907, 1912), another revision of the British species by J.F. Perkins (1976), a study on the classification of the dryinids by Richards (1953), a -<ey
to
the Russian species by
c!ryinids by rreytag useful
Freytag
(1982).
(1977,
Kieffer's work
because of
Ponomarenko
1978,
1980),
is unreliable for
its completeness.
><earctic species by Ashmead
(1978)
and studies on Nearctic
Freytag and Back
The rudimentary
(1893)
(1977),
identifications,
Giri
but
it
and is
revision of the
is not recommended.
The object of this paper is to revise the species already t.iescribed, to assemble the biological information about them and to describe the new species represented
in
the collections.
world species reasonably Little
The user wi II
find
treatment of the
complete.
is known of the males and the keys are therefore
unsatisfactory.
The classification
association of the sexes this problem.
Only
is based mostly on
the females.
is difficult and much further work
breeding experiments can
finally
The
is needed on
answer these
questions.
MATERIAL EXAMINED
The keys and the descriptions are based on numerous collections.
Those
individuals and
material
examined
institutions from which
in
Mem.
2
Amer.
Ent.
Inst.,
material was borrowed are listed below abbreviation used throughout the text: AD
South Austral ian Museum,
AM
American Museum of Natural
AS
Academia Sinica,
37,
1984
together with
Adelaide
Zoological
no.
the appropriate
(South Austral ia)
History,
New York
Institute,
Peking
(N.Y.)
(China)
B
Bernice P. Bishop Museum,
BA
Museo Argentino de Ciencias Naturales "Bernardino Buenos Aires
Honolulu
(Hawaii) Rivadavia",
(Argentina)
BE
Department of Entomology,
University of Wageningen
BI
Ferdinando Bin collection,
Perugia
BM
British Museum
BN
Zoologisches Forschungsinstitut und Museum Alexander Koenig, (German Federal
(Natural
(Holland)
(Italy)
History),
London
(England)
BO BR
Istituto di Entomologia dell'Universita, Bologna (Italy) Department of Primary Industries, Indooroopilly (Queensland)
BU
Nat iona I Museum of Natura I History,
BY
Department of Bio!ogy,
CA
Cal ifornia Academy of SCiences,
CB
Australian National Entomology,
Bonn
r,epublic)
Internacional
(Hungary)
Waco
San Francisco
Collections of
Canberra
Budapest
Baylor University,
(Texas) (Cal ifornia)
Insects (ANIC),
CSIRO,
Division of
(Austral ia)
CI
Centro
CM
Museum of Comparative Zoology, (Massachusetts)
de Agricultura Tropical,
CO
Zoologisk Museum,
DA
Department of Entomology,
KObenhavn
Cali
(Colombia)
Harvard UniverSity,
Cambridge
(Denmark)
University of Cal ifornia,
Davis
(California) DL
Jiri
DU
National
Dlabola collection,
EB
Institut fur Pflanzenschutzforschung
Museum of
Praha
Ireland,
(Czechoslovakia) Dubl in
Landwirtschaftswissenschaften,
(I reland) der Akademie der
Eberswalde (O.O.R.)
ED
South African Sugar Association (South Africa)
Experiment Station,
EN
Istituto di Entomologia agraria ed Apicoltura dell'Universita, (I taly)
FO
Alessandro Focarile collection,
GE
Museo Civico di Storia i~aturale "Giacomo Doria",
Aosta
GV
Museum d'Histoire Naturelle,
Mount Edgecombe Torino
(Italy) Genova
(Italy)
Geneve (Switzerland)
HA
Martin-Luther-Universitat,
HE
Zoological Museum of the University,
Halle-Wittenberg
(D.D.R.)
HU IF~
Zoologisches Museum an der Humboldt-Universitat, Berlin (O.D.R.) Institut Royal des Sciences Naturelles de Belgique, Bruxelles
IT I(E
Cornell University, Ithaca (I"ew York) College of Agriculture, University of Kentucky,
KY
Entomological
LA
~Jatural
Helsinki
(Finland)
(Belgium)
Laboratory,
Kyushu University
History Museum of Los Angeles County,
(Cal ifornia)
Lexington (Kentucky)
(Japan) Los Angeles
3
Introduction
LE
Rijksmuseum
LI
Linnean Society,
van Natuurlijke Historie,
LN
Zoological
LO
(U.S.S.R. ) Museum d'Histoire naturelle,
LU
Zoological
MA
Department of Entomology,
London
Institute,
U.S.S.R.
Institute,
Leiden
(Holland)
(England)
Lund
Academy of Sciences, Lausanne
Leningrad
(Switzerland)
(Sweden) University of Wisconsin,
Madison
(Wisconsin) MD
Instituto Espanol
ME
National
de Entomologia,
MI
Museo Civico di Storia Naturale,
ML
Milwaukee Public Museum,
Museum of Victoria,
Madrid
Melbourne
(Spain)
(Victoria)
Milano (Italy)
Milwaukee (Wisconsin)
MO
Zoological
MP I: MR
Moscow Palaentological Institute, Moscow (U.S.S.R.) Claud Morley collection (part kept in Ipswich h'luseum,
Museum,
Moscow
(U.S.S.R.) Ipswich,
England) MT
Ecole Nationale Superieure Agronomique,
Montpell ier
MU
M.
(Japan)
Miura collection,
Shimane University
NA
Museo di
ND
Division of Entomology, Delhi
Zoologia dell'Universita,
Napoli
(France)
(Italy)
Indian Agricultural
Research
Institute,
New
(I ndia)
NI
Entomological
NZ
Department of Scientific and
Laboratory,
Kochi
University,
I ndustrial
Nangoku-City (Japan)
Research,
Auckland
(New
Zealand) OL OT
Massimo Olmi collection, Viterbo (Italy) Biosystematics Research Institute, Ottawa
OX
Hope Museum,
Oxford
(Canada)
(england)
P
Museum National
d 'Histoire ,',aturelle,
PG
National
\I'"atural
PH
Academy of r,atural
PK
:,adezdha Ponomarenko collection,
PL PO
Guido Pagl iano collection, Torino (I taly) Istituto di entomologia agraria dell 'Universita,
PT
Facultad de Ciencias Naturales y '''1useo,
QU RY
Queensland r'l,useum, Fortitude Valley (Queensland) Biological and Chemical Research Institute, Rydalmere
Museum
History),
Sciences,
Paris
Praha
Philadelphia r"oscow
(France)
(Czechoslovakia) (Pennsylvania) (U.S.S.R.) Portici
La Plata
(I taly)
(Argentina) (,',ew South
Wales) SA
Friedrich Sander collection,
SC
Department of Food and Agriculture,
SP
Spinola collection,
c/o
Jena
r..1useo
(D.
D.
R.)
Sacramento
f,egionale di
(California)
Scienze Natural i,
Torino
(I taly) ST SU
Naturhistoriska Riksmuseet, Stockholm (Sweden) Instituto de Aclimatacion, Almeria (Spain)
SW
Museo Nacional
SZ
Pier Luigi
TE
Department of Entomology, Station
de Historia Natural,
Scaramozzino collection,
(Texas)
Texas A.
Santiago Torino
(Chi Ie)
(I taly)
and M.
University,
College
4
Mem.
Amer.
Ent.
Inst.,
TN
Museo de Ciencias Naturales,
TO
Istituto e Museo di (I taly)
TR
Museo Regionale di Scienze Naturali,
TU
Instituto de Zoologia,
no.
S.ta Cruz
37,
1984
(Canary
Islands)
Zoologia sistematica dell'Universita,
Torino
Torino (Italy)
Universidad Nacional
de Tucuman,
Tucuman
(Argentina) TV
Musee Royal
TW
American
de I 'Afrique centrale,
UQ
Department of Entomology,
Entomological
Institute,
Tervuren
(Belgium)
Ann Arbor
(Michigan)
University of Queensland,
SI.
Lucia
(Queensland) VE
Museo Civico di Storia Naturale,
VI
Naturhistorischen Museum,
Wien
VR
"Iuseo Civico di Storia Naturale,
WA
National
WS
~jatuurhistorisch
Museum of Natural Museum,
Venezia
(Italy)
(Austria) Verona
History,
(Italy)
Washington
Maastricht
(D.C.)
(Holland)
ZO
Zoolog ische Staatssamm lung,
Munchen
YA
Keizo Yasumatsu collection,
Oonojo-shi
(German Federa I Repub I i c) (Japan)
ACKNOWLEDGMENTS
I have been much indebted to the following persons who helped with the collections, either by sending dryinids on loan, or acting as host when thei r museums were vis i ted: Kees van Achterberg (Le i den), Anders Albrecht (Helsinki), Paul H. Arnaud Jr. (San Francisco, California), Donald Azuma
(Philadelphia,
S. Baldridge (Waco, Texas), (Tervuren), Claude Besuchet Ferdinando Bin (Montpellier), A.J.M.
(Perugia),
G.R.
Brown
Pennsylvania),
Marcos Baez
(Tenerife),
Robert
Tony Bellotti (Cali), P. L. G. Benoit (Geneve), C. F. M. den Bieman (Wageningen),
Zdenek Boucek (Rydalmere),
Carnegie (Mount Edgecombe),
(London),
Ernst-G.
A.
Bournier
Burmeister (Munchen),
Achille Casale
(Torino),
Lennart
Cederholm (Lund), Raul Cortes (Santiago), I talo Currado (Torino), E.C. Dahms (Fortitude Valley), Roy Danielsson (Lund), Michael Day (London), Jean Decelle (Tervuren), (Bruxelles), Dorn
(Halle),
N.
F.
(New Delhi),
(Wien),
Ian Galloway
Delhi),
I
41
,
teeth progressing
Anteon gaullei
17
Kieffer).
larger from anterior on e to
18
Mem.
I nternally and a
Amer.
Ent.
Inst.,
no.
37,
1984
two lobes, a more proximal one, the lacinia, galea. The labium consists of a submentum and The distal end of the prernenturn bears externally the labial
the stipes bears
more distal
one,
the
a prementum. palpi, which characteristically have three segments though the number may be reduced
in particular genera and species.
palp-segments may
be concisely
maxi Ilary palp-segments;
The data as to reduction of
summarised as follows
second
number =
labial
(first number
palp-segments):
Dryininae: female and male: 6/3; ThaUluatodryininae: female and male: 6/3; Anteoninae: female and male: 6/3; Bocchinae: female and male: 6/3; Conganteoninae: female and male: 6/3, 5/3; Transdryininae: female: 6/3; male unknown; Biaphelopinae: female: 6/3, 5/3; male unknown; Aphelopus (Aphelopinae): fema I e and ma Ie: 5/2; Crovettia (Aphelopinae): fema I e and ma Ie: 5/3; Gonatopodinae: Gynochelys: female: 6/3; male unknown; Epigonatopus: female: 2/2; male unknown; Esagonatopus: female and male: 6/2; Gonatopus: female and male: 3/2, 4/2, 4/3, 5/2, 5/3, 6/3; Tetrodontochelys: female and male: 2/2, 3/2, 4/2, 5/2; Trichogonatopus: female: 6/3; male unknown; Haplogonatopus: female and male: 2/1; Dicondylus: female and male: 2/2, 3/2, 4/2; Pentagonatopus: female: 5/2; male unknown; Plectrogonatopoides: female: 5/3; male unknown; Pseudogonatopus: female and male: 2/2, 3/2, 4/2; Donisthorpina: fema Ie and ma Ie: 5/2; Agonatopoides: female and male: 5/3; Apterodryinus: female and male: 6/3; Acrodontochelys: female and male: 2/2, 3/2, 4/2; Adryinus: female: 6/3; male unknown; Echthrodelphax: fema Ie: 3/2, 4/2, 5/2, 5/3, 6/3; rna Ie: 4/2; Neodryinus: female and male: 6/3; Apodryininae: female: 4/3; male unknown. The maxillary
and
labial
palpi
are usually borne on special
stipes and prementum respectively. like a
basal
segment of the palp.
segment of the palpi
really
there can be no doubt that watching a
narcosis)". palpi
In
The question
is a palpiger or a
in the present paper. According seen by
These lobes,
to R.C.L.
it is a
the
as to whether the basal true joint
Perkins
the present paper,
(1905)
true moveable joint
species under the microscope,
is always considered
the first small
a true joint.
lobes of the
palpigers, may look
as
is not discussed "in many forms
(as can easi Iy be
it recovers from
basal
The maxillary
segment of the and
labial
palpi
Introduction
have been
largely used in
generic differentiation. palpal
segments
distinguishing pal pal
variable
segments
in
however,
Moreover,
the
last
labium shows
it now
lobes,
in
the genus.
longer of prime appears that palpi
The
importance in
the number of
except
two segments of the maxillary three central
the number of
the species and
is therefore no
the genera.
that
segments agrees between males and females,
particular, The
Gonatopodinae for
the classification of the
I t now appears,
is usually
number of palpal
19
that
in males
in
may be fused.
the outer ones being
the
paraglossae and the inner one the glossa. The Dryinidae have fi I iform or distally
thickened antennae,
segment or is small
and
with
segments. when
10 segments
the scape forms a
part of the antenna, of various
with
in both sexes.
The first
scape is articulated in a socket. The second segment or pedicel flagellum.
the
The surface of types.
ball
and socket joint for
the antennae
is closely covered
These are sometimes taxonomically
large enough
Thaumatodryininae,
the distal
The flagellum consists of eight with
important,
to be visible without great magnification. the females have tufts of
long hairs on
sense-organs especially In
the antennal
Gonatopodinae the females have longitudinal groove-I ike structures, the rhinaria (Fig. 6), on the distal
segments 5-10. segments. in
In some genera of
The number of distal
the genus
(2,
3,
4 or 5),
segments showing
but constant
in
the rhinaria
the species.
is variable
Rhinaria are
also present in the females of Apodryininae (on antenna I segments 3-10), Transdryininae (on the I as t three antenna I segmen ts) and Dryininae (on the last six antenna I segments: except for fiiesodryinus, which seem to be always without rhinaria). distal
The data as to distribution of rhinaria on
antenna I segments may
be concisely summarised as follows
(within
brackets number of rhinaria per segment):
Gonatopodinae Agonatopoides: 9th (1), 10th (2); Haplogonatopus: 9th (1), 10th (2); Acrodontochelys: no rhinaria; Tetrodontochelys: no rhinaria; Gonatopus: no rhinaria; Dicondylus: 9th (1), 10th (2); Pseudogonatopus (A): 9th (1), 10th (2); (8):
8th
(1),
9th
(1),
10th
(C):
7th
(1),
8th
(1),
9th
(2); (1),
10th
(2);
Trichogonatopus: no rh i nari a; Apterodryinus: 7th (1), 8th (1), 9th (1), 10th (2); Donisthorpina: 9th (1), 10th (2); Esagonatopus: no rhinaria; Plectrogonatopoides: no rhinaria; Echthrodelphax: 9th (1), 10th (2); Adryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Neodryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Pentagonatopus: 7th (1), 8th (1), 9th (1), 10th (2);
""
•
20
Mem . Amer. En!.
I ns!.,
no. 37,
1984
Gynochel ys : 6th ( 1 ), 7th ( 1 ), 8th (1), 9 th ( 1 ) , 10th (2); Tr an sdryininae Transdryinus : 8th ( 1) , 9th ( 1 ), 10th (2); Dry ininae Dryinus : 5th ( 1 ), 6th (1), 7th ( 1 ), 8 t h ( 1 ), 9th (1), 10 th (2); Richardsidryinus: 5th ( 1 ), 6 th ( 1) , 7th ( 1 ), 8 t h (1) , 9th ( 1 ), 10th (2); Al phadryinus : 5th (1), 6th (1), 7th (1), 8th (1), 9th (1), 10 th (2); Tr i d r y i nus: 5 t h ( 1), 6 t h ( 1), 7 t h ( 1 ), 8 t h ( 1 ), 9 t h ( 1 ), l Ot h ( 2 ) ;
Fig. 6 -
Last antennal segme nt s showing rh in ar i a Haplogo natopus oratorius Westwood).
(fema le of
Perodryinus : 5th ( 1 ) , 6 th ( 1 ) , 7th ( 1 ) , 8 th ( 1 ) , 9th ( 1 ) , 10 t h ( 2) ; Me sod ryinus: n o rh i nar i a j Megadryinu s : 5th ( 1 ) , 6th ( 1 ) , 7th (1 ) , 8th ( 1 ) , 9th ( 1 ) , 10th ( 2) ; Apodryininae Apodryinus : 3th ( 1 ) , 4th ( 1 ) , 5th ( 1 ) , 6 t h ( 1 ) , 7 th ( 1 ), 8 t h ( 1 ) , 9th ( 1 ) , lath THORAX.
( 2) ;
T h e thor ax consists of t h ree segm ent s (pr',ihorax,
mesothorax ,
I ntroduct ion
21
meta thorax} ; the two hinder ones normally bear wings. Some females of Gonatopodinae are apterous. In Gonatopodinae, Anteoninae and Bocchinae, females may be brachypterous or micropterous.
The first
abdominal
segment
(propodeurn) has become firmly attached to the thorax and is connected to the rest of the abdomen by a narrow petiole. Some authors propose the term mesosoma for the thorax + propodeum. Usually the segments of the mesosoma are easily visible. fused
(only
propectus
PROTHORAX.
less
is distinct).
The dorsal
the mesonotum, the
recognizable because the sutures are more or
Apodryininae, however, the mesosoma sclerites are completely
In
sclerite
whi Ie the
lateral
(pronotum) is rather closely attached to sclerites (propleura, composed entirely of
proepisterna) are loosely attached to their notum. The propleura form the prothorax and are termed propectus. They meet
the ven tra I part of along
the median
I ies the small
ventral
I ine.
Quite often hidden between
the first coxae,
prosternum. Anteriorly the propectus supports the head,
sharing this duty with the pronotum; posteriorly it is attached to the first coxae. The pronotum is commonly divided by an anterior transverse furrow into an a
anterior part
rather higher level
(anterior collar) and a more posterior part (disc) at (in
the plane of the mesonotum).
Sometimes a
posterior collar. Gonatopodinae no transverse furrow is visible. I n the ma les of Dryinidae as well as the fema I es of Aphelopinae, Biaphelopinae and Conganteoninae, the p ronotum is very reduced dorsa II y posterior transverse furrow
is also visible,
definining a
In some females of apterous
and
is almost
invisible from above.
This condition
is primitive.
In
the
more specialized females of Dryinidae the pronotum is distinctly visible dorsa II y. It is produced backwards and swoll en into pronotal tubercles. They may
reach back as far as the tegulae or be widely
separated from
Gonatopodinae are the pronotal tubercles absent. In females of the most specialized forms (Dryininae, Gonatopodinae, Transdryininae) the pronotum is remarkably elongated, them.
Only
in
the females of
further forward extension
is caused
of other prothoracic elements is unusually mobile. the prothorax adaptive,
in
by
the dorsal
(propleura
According
to Reid
in particular). (1941)
and
position and development The pronotum
itself
the peculiar modifications of
Gonatopodinae and Dryininae are in a large measure
being concerned
in
the raptorial
habits of the females.
In
Apodryininae the pronotum is fused with the posterior sclerites of the mesosoma. MESOTHORAX.
contains
The second
thoracic segment
the principle flight-muscles.
Only
is usually in
the
largest,
since
it
the apterous females of
Gonatopodinae is it more slender than the other thoracic segments. Dorsa II y the segment is covered by the mesonotum which is divided into two principa I sc I eri tes, the scutum and the scutellum; taxonomi sts often reserve the term mesonotum for the scutum alone. scutum
two sulCi,
the
From the front margin of the
notaulices, often converge backwards, defining a V-
shaped prescutal area. The sulci may be deep and strong and meet at about the centre of the posterior margin of the scutum. In other dryinids they extend back
to the hind margin of the scutum without meeting;
in
22
Mem.
many
groups,
they
are
weak
Amer.
and
very
Gonatopodinae median scutal line
apterous fema I es of
Aphclopinae bisects less
scutum. in
each
and
are the
37,
1984
i ncomp I ete or even
absen t.
the notau I ices are absen t. is often
each
present,
si de of
about
sulcus may
half way
connected
aspect of
anterior,
which
forwards
sternal that
It
very
narrow
are
the
it
scutum.
areas
and
and
They
I a tera I
pleural
to
are absent
regions of
is convenient
to
treat
mesopleuron
is
the
pleural sulcus, lateral
reduced,
the pleural
narrow and wh i ch
articulation Since
sulcus runs
near and
the mesopleuron
the mesopleura.
are partly
visible.
marked off by
a
Just
the
the mesepisternum
an
the
is
tegula
cutting
function
of fl ight,
forms
long
entirely of
anterior
very
iJryininae. marked.
much
while
dorsum of
the
scutellum
to Reid
(1941
be seen
l,
Gonatopodinae,
among there
prothorax,
In
meta thorax
METATHORAX.
Apodryininae and
runs from of
the
subserv ience and
the
It
the bulky,
is formed
reduced. by
In
to
and special ised.
female
the
The
almost
lateral
the prepectus,
stages of thoraces
this
among
various modifications of
is one
this
non-adapta t i ve one.
is
It
region
prepectus. The Gonatopodinae.
from
that
the
be obsol ete or strong I y
This condition
scutellum.
area and
antero-dorsal
visible.
The early
the reduction
the
the
are formed
some phylogenetic significance; of
At
or
is much
suture may
median,
transverse
the mesonotum
in
a
trochantin
the pronotum
importance.
the
the
apterous
the mesothoracic waist
The meso-metap I eura I apterous
is a
modified
The
the prepectus Can
According in
in
from
is
to
and continuous
antero-ventral
rei eased
parallel
parts by
is
epicnemium
developed
hO's assumed considerable
special isation of
area
waist connecting
thorax.
walls of
the
the mesothorax,
slender the
the scutum,
ventral
off an
has become
and
posterior part of
this
the mid
sterno-pleural sutures
trochantinal sulcus. epicnemial sulcus is towards
well
lateral
spec ies the
that
down
particularly
Gonatopodinae
developed
of each mid coxa
is
of
into two
many
in front
it off
apterous
thorax
In
seems
mesepisternum,
which
divided
It
the region prepectus
is
sulcus.
the sulcus cutting corner of
It
sternal sulcus.
vis ib I e
runs
of
the mesepimeron
well
suture, which divides mesosternum is relatively
The
are
The
into an
very
the
the forewings.
together.
divided
posterior,
d i v i ded by beneath
in
the mesothorax
them
the nieso-I~etapleural
and
two more or
the paraps i da I furrows
metapleuron.
a
the
Gonatopodinae. Arising on parapsidal furrows which
upwards from
is
and
l"ing-process
to
with
In
the
scutellar sulcus.
pair of
across the
The scuta I
the
apterous
be a
the mesothorax or
mesepimeron.
coxa
part I y
In
longitudinally
the scu tell um
with
in
large mesepisternum and a
very
obliquely
coincident
sulcus are absent
Gonato;Jodinae. parapsides. The
intimately
reduced
no.
the
the scutal
forward
apterous
lateral
On
is centrally
scutal
side of
extend
so
areas,
axi Ilae and
Axillae
area.
Ins!.,
axillae, which belong morphologically to the The scutellar sulcus runs behind the axi Ilae and, on its central front of the scutellum. The scutal sulcus divides the scutum from
triangular
part, the
a
the prescuta I
En!.
that
may
possibly
the enlargement of may
the scuturn
perhaps
the mesothorax
the
have and
be regarded
is fused
as
with
propodeum.
The metathorax,
which bears
the relatively
small
hind wings,
Introduction
is smaller and segment area,
less differentiated
is covered
usually
Gonatopodinae between
the
a
suture from In
region
metasternum fused
Vv
i th
The
tergite.
the propodeum
to
The
the a
the dorsal
is a
small
muscle of the
Dryinidae
together on
have
each
front edge of
micropterous. is
the
of
the
In
wings When
"hich
articulate
the subfamilies.
'.'eins
and
Irig.
7)
I
text
I
term
in front
of
as
the
inclined
No suture between
Apodryininae
In
the meta thorax
is
to
the first
abdominal
spiracles
lie near the
segment
is articulated
and
above
The
the point
lateral by
margins
its petiole
i nterna I organs where
they
leave
the mesosoma.
a
row
of
wings,
the fore
hooks or
hamuli,
and engaging
Dryinidae, 3-5 wing
the
follow
hamuli
margin
thorax
the
by
is
the
a
system
the
hind margin
some females of
are brachypterous or margin
dorsal margin, the
by
a
of
the
and
series of
axillary sclerites. the tegula.
(1969)
for for
wing
the
large plate,
single characteristics Townes
In
held
arise from
downfolded
is supported by
hind ones,
which
the front
the
means of
hidden
and
are found.
Some females
The membrane
imoortant
with
is outstretched
hind
are normally
.\ ing-veins provide
Two
are absent.
with
sclerites
the
and
and
is
orifice of the propodeum, through which passes petiole. In Apodryininae the propodeum is fused
costal margin, the wing is the apex.
axillary
In
wh ich
separated
metepisternum
into
the metap I euron
the name given
two pairs of
the
is not
the meSQsoma.
ventrally,
the hind wing
the
divided
groove
the metanotum,
doub Ie ba II-and-socket joi n t.
side by
the fore-wing.
:Jryinidae
is visible. is
the other segments of
NII~GS.
not
is what
+ propodeum. The metasternum is a very
abdominal
the petiole somewhat
the propodeum
transverse
wetapleuron
The
is
A
area
apterous
is probab I y
the scutellum
thorax.
second
with
the
two posterior pairs of coxae.
propodeum
attached
to
of
the
and
In
the
single central
This central
defined.
the propodeum.
behind
the other segments of
the
with
with
area
between
well
Gonatopodinae
the meta thorax
of
enter
not
the propodeum.
and metapleura
PROPODEUM. segment
is
Dorsally
consists of a
depressions.
the propodeum
ap terous
fused
the mesothorax. which
postscutellum.
the
the propodeum
the flat
surface of
narrow
with
the
indistinguishably fore
lateral
the metanotum fused
metepimeron. metanotum
by
termed
scutell um and
is more often by
has
than
metanotum.
the
defined
(1939)
Richards
by
23
tip
veins The The
the classification labelling
of
the wing-
cells which they enclose. The fore-wing of the Dryinidae pterostigma, 1-3 basal cells enclosed by pigmented veins, 2 costal vein, the metacarpus joining the pterostigma to the apex, and in SOme species an enclosed radial cell. The radial cell is open when the -adial
the
shows a
vein
is
incornplete.
In I~adiimancus
Moczar
the radial
vein
is
Gonatopodinae, Dryininae, Anteoninae, Bocchinae, -::lauIllatodryininae, Transdryininae have three basal cells (costal, median 3"ld submedian) enclosed by pigmented veins (costa, subcosta, medius, s:1!Jmea'ius, basal, nervulus). COI1ganteoninae and iliaphelopinae have two :casal cells {costal and median} enclosed by pigmented veins (costa, .o:..;:'costa, medius, basal). A;;helopinae have basal cell (costal) enclosed by ::igmented veins (costa and subcosta). The first brachial cell rarely 38sent.
Vvinged
, '"
24
Mem.
appear s
En!.
Ins!.,
no.
37,
1984
Th e male of My strophorus formicaeformis Ruthe has
to be enclosed.
fore-wings reduced,
Amer.
spoon-shaped;
the hind-wings a re absent.
"
Fig.
7 -
Fore-wing of Gonatopodinae : C
submedian cell;
=
ra di al
ve in,"
basal vein;
LEGS.
There
F cs
=
costal
cell;
M
=
median cell;
5 -
= first brachial cell; R = radial cell; t = metacarpus; r = costa," b = subcosta", e = medius", h = submedius", k =
= nervulus.
0
is a pair of
legs for each
thoracic segment.
The
legs consist
of a coxa,
trochanter, femur, tibia and a segmented tarsus. The females of the Dryinidae (except for Aphelopinae and Biaphelopinae) have relatively
elongated fore-coxae.
segments.
The
l ast tarsal
the foot-pad or arolium
T he tarsus always consists of five
segment bears the claws.
(Fig.
8).
Its dorsal
Between the claws
surface contains a
plate,
li es the
orbicula. The f ron t tarsi of most fema l e dryinids (excep t for Aphelopinae and Biaphelop inae) are formed into a chela (Fig. 9) for gripping the homopterous prey.
The chela
is composed of the 5th segment,
and an
enlarged claw. A second claw, the rudimentary claw (Fig. 8), i s reduced (in Thaumatodryininae, Bocchinae, Dryininae, Gonatopodinae) or absent (in Anteoninae, Conganteoninae and Transdryinillae) The trochanter is usua II y a srnall segment. I n the females of Dryininae, Transdryininae, Thaumatodryininae and Gonatopodinae it is very elongated. The femur usua l ly consists of a prox i ma l swol len part {club of femur} and a distal slender part {stalk of femur}. The tibiae are provided with articulated spurs o n
the ventral
apex of the tibia
in
s ide.
Typically,
the following
the
three legs have spurs at the
numbers:
1,
1,
2
(males,
females of
25
Introduction
Aphelopinae, Biaphel opin ae, Conganteoninae, Anteoninae, Thaumatodryininae, some Bocchinae, most of Dryininae ) ; 1, 1, 1 (females of Apodryininae, some Dryininae, most of Bocchinae); 1, 0, 1 (females of Transdryininae, most of Gonatopodinae) or 1, 0, 2 (some females o f Gonatopodinae). The fore-tibia always has only one apical spur. This spur i s mod ifi ed, tarsus,
Fig.
together with
into an
8 -
Arolium
the ve ntral
s i de of the first segment of the
antenna-cleaner ( Fig. 10). The first segment of the fore-
(A)
and rudimentary
claw
(C)
of
Haplogonatopus oratorius
(Westwood) .
(bas itarsus ) may be short or very elo ngated. The modified fifth segmen t of the fore-tarsus con s ist s of a proximal part (basal part, tarsus
d irec tl y distal m ay
art icul ated
part
form a
with
the fourth
proximal
anterior
segmen t of the fore-tarsus ABDOMEN.
and a more or
less
long,
free
(ap ical part) . Segments 2 or 3 (rarely 1) of the fore-tarsus (inner)
fore-tarsus may be short or very bris tles,
segment)
and
hook.
e longa ted.
The fourth The
seg men t o f
the
inner side of the fifth
the enlarged claw may bear
lamellae,
ha i rs or peg-I ike ha i rs. The abdomen consists of
ten
segments,
but this number may
not
~iem.
26
Fig. N ;
9 5th
Arner.
En!.
I ns!.,
Chela o f Anteon g aullei Kieffer t a r s al
segment).
(A ;
no.
37,
1984
aro liu m;
E ;
enlarged claw;
Introduction
be visible without careful of a dorsal
27
study or dissection.
Each
is normally
made up
tergite and a ventral plate or sternite. T he
plate or
....
;'1' ,-
""
j" "
\ ,
j
•
, •
•
,
L F i g.
10 -
Antenna-cleaner of
Jiaplogonatopus or atorius (Westwood) (female):
T = tib i a l spur.
•
spiracular openings
lie at the sides of the
segmen t
into the propodeum wh ic h
is modi fied
tergites.
The first abdominal
is fu nctiona ll y
part of t he
gaster is used f or all the abdominal segments beh in d the propodeum. Some authors propose the term metasoma for t he post-propodeal part of the abdomen. Th e petiole is the narrowed, anterior, mor e or less long stalk. Accord i ng to R ich ards (197 7 ) t he Dryinidae have 8 pairs of thorax.
The
abdom in al
ter m
spiracles.
are normall y
T erg it e 10 is
to which
ov ipos itor, segment
which
is reduced the
only 8 abdom inal Th e ninth
t he seventh
tergites
(7 gastral)
tergite is entire ly
indistinguishably fused
side of the abdomen
The eighth sternite
val vifer,
the female,
developed and v i s i ble.
and depigmented. the vent ral
In
is the
with
retracted
tergite 9.
On
last vis ible sternite.
and membranous and on each s i de is a
first
first valvula is attached. They are parts of the
is adapted for sti nging.
T he sternite of the ninth
is also redu ced and on each s ide has a
second valvifer with the
,
28
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
second valvula attached to its anterior end. The valvifers described as appendage bases. the ovipositor,
forming
immovably joined first The
valvulae.
the
The first
can be
valvulae are the active part of
lancets of the sting.
to one another and form
a
The second valvulae are
sheath
(sting-sheat)
for
the
terebra.
The two pairs of valvulae together constitute the
third valvulae each arise fror,; the posterior end of the second
valvifer.
The genital
orifice
is anterior to the first
longer act as an ovipositor.
The sting and
plates have not so far proved as useful corresponding parts of the male. gastral)
In
are exposed and normally
are available.
The first
next six sternites The external
(up
sternite
in separating
the male developed.
No data on
is not visible as a
to sternite 7 = sixth
last abdominal
species as have the
the first eight
tergites
(7
tergites 9 and
definite sclerite.
gastral)
genital ia of the male are attached
ninth segment.
valvifers which no
the modified
are fully
10
The
developed.
to the hind margin of the
The shape of the ninth sternite may
have taxonomic
importance in Dryininae, Anteoninae and Aphelopinae in separa t i ng species. The male genital ia (Fig. 11) consist of a basal ring, two lateral a median penis (consisting of two penis valves) and two volsellae on the sides of the penis. Each volsella consists of a proximal basivolsella and a d i sta I distivolsella. I n the Gonatopodinae the dorsa I end of each paramere forms an appendage termed the dorsal process of gonoforceps (Fig. 12). I ts shape has taxonomic importance in separating species. In the males of Anteoninae the parameres may bear proximally more or less large dorsal membranous lobes. LENGTH OF THE BODY. The adu I ts of the Dryinidae range in size from 1,00 mm (Gonatopus ashmeadi Kieffer) to 10,93 mm (i'legadryinus magnificus
parameres,
Richards).
Measurements of total
the front of the head (excluding
the an tennae)
to
length from
the apex of the abdomen
the sting).
COLOUR.
Most males are black.
gasters,
but head,
black,
length of adults refer to the
(exc Iud i ng
thorax
Most females
have black
to reddish-black
and propodeum are highly variable
in colour -
reddish or yellowish.
EGG AND POSTEMBRYON I C STAGES. The morpho logy of the egg has been described by Fenton (1918a). The egg appears oval or kidney-shaped and varies .in color from sculpturing on
light yellow
the chorion.
of the egg were added by Westdal,
Richardson
has been described by (1965).
The
larva
third and fourth and serves each molt, usually the oral process
Fenton
(1918a)
is apterous. instars.
It
is
but soon darkens in
the second,
(1961),
process
Westdal,
is present
located near the anterior end the host.
light brown.
pink or green.
third and fourth
The
The first-instar
the
larva
Richardson in
the second,
the
larva
larva is white after
The mature larvae are A pair of fleshy oral
instars.
In
the fifth
lobes are replaced with biting-chewing mouthparts and has disappeared.
is no
Barrett,
The structure of
and Barrett,
in
There
notes on the morphology
Raatikainen
(1980c).
A ventral
to a
but occasionally
grey or dark brown.
fragmentary
(1940),
and Jervis
to anchor the parasitoid
white,
are present
Clausen
(1965)
to dark
Additional
lobes
instar the ventral
larva has a sacciform or
-
Introduct i on
29
•
•
, .
1
"
•
Fig. ring;
,
-
11
- Ma Ie gen i ta I i a of Deinodryinus atriventris (C resson):
N -
penis;
P ;
parameres;
S ;
basivolsella;
V ;
B ;
distivolsella.
basa I
30
Mem.
embryo n ic shape. or U-shaped.
12 -
En!.
third -
T he f i fth -i nstar larva
w i der posteriorly
F i g.
The second - ,
Amer.
I ns!.,
no.
is straigh t ,
and more pointed anteriorly.
N = penis;
1984
and fourth - ins t ar l arvae are curved d i stinctly
segmen t ed,
Other notes on
Male genitalia of Gonatopus lunatus Klug :
gonoforceps;
37,
0 -
the structure
dorsa l
process of
V = dist i vo l sella. •
of the ( 1905) ,
larva were published by Mik Jeanne l
( 19 13),
(1882),
Keilin and Thompson
G i ard
(1889),
( 1915),
Haupt
R.C.L. ( 19 16,
Perk i ns 1932),
Fenton ( 1918b), Maneval (1935), Esaki and Sameshima (1939, 1940), Clausen (1940), Pruthi and Mani ( 1942), L i ndberg (1950), Behrisch (1952), Heik in heimo (1957), Raa t ika i nen ( 1961), Guagliumi (1962), Chaudhary and Ramzan
( 1968),
Ponomarenko
(1975b)
and Jerv i s
(1980c).
Introduction
31
COLLECTION
Best collecting Townes
1972;
of flying
will
be from
Malaise traps
(classical
or
for
Dryinids.
near the ground, collecting
results
instance Townes ' and rVl asner l s models: 1\1alaise 1937; Masner and Goulet 1981). They are used mostly for sampling
modified models,
Pan
i.
traps may
e.
the
methods such
be used
normally
as sweeping or Malaise trapping
communication).
Apterous
Sweeping
recommended for collecting
is not
to sample Dryinids
spectrum of species not
Dryinids are best obtained
oy
active on or
obtained
by
(Masner personal pan
adult Dryinids,
traps. as
the results
are unsatisfactory. Collecting satisfactory Dryinids, obtained rearing
nor by
their hosts.
sweeping.
parasitized
parasitization Chandra's rearing
as
pantrapping
Dryinids
hoppers collected 1980c).
hoppers
Dryinids may
will by
be studied
Chandra
parasitized
or Malaise trapping
the behavior and
For these purposes,
Adult
abilities may
models:
Adult from
methods such
techniques for studying
in
resul ts
be obtained
sweeping. in
Good
glass
best
cages
may
tubes plugged
also be obtained by
wi II
from
be
the field
Biology,
rearing
results
are not
development of by
behavior and
(for
instance
also be obtained with
holding
by
cotton.
cocoons collected
plants. 'ilinged
Dryinids or parasitized
hosts may
be attracted
oy
light
traps. P,c<ESERVATION
Small
Dryinids are best mounted on
are pushed using
through
stainless steel
pinned
pins.
away
to obscure are relaxed genitalia
from
the body
the sides and it
with
is as a
rectangular cards.
and
to open needle.
identify
many males,
prepare genitalia relaxed capsule should be removed
examination
in
Faure
glycerol
20 cc,
and clearing
GOlla topodillae, identify
many
Arabic
and
the
the mouth
a
hooked
w hi ch have very
and
c I ear
dorsal
pin,
genitalia
be
wings
so as not
the
male
be glued it
to
has the
and
in
is essential.
To
The entire male genital
50 cc,
for
dorsa I the
the
specimens
pUllout
Dryinids Can
the
Maceration
Gonato:.JoJinae,
should
have
Vihi Ie the
which
if dropped.
water
recommended
the adul ts should
is recommended;
the specimen
gum 30 g).
the
parts
examination of
(distilled
pins
pinned
antennae positioned
specimen.
Alternatively,
using
are not
males of
easy examination of
legs
they
specimens are required.
liquid
solution
is used,
thorax,
This method of mounting
advantage of protecting To
the
the
underside of
well
hooked
minute stainless steel
Larger Dryinids are best
Whichever method
to one side of the centre of
pulled
added
Plastazote strips.
10
mounted
Chloral
for
microscope
hydrate 50 g,
% potassium
hydroxide
male genitalia of processes of gonoforceps.
volsellae should
process of gonoforceps.
In
be removed many
To for
species of
Dr/ininae, Anteoninae and Aphelopinae the n i n th stern i te shou I d a I so be
32
Mem.
removed from To
Amer.
the male genital
identify females,
En!.
removed
using
Maceration
a
pin
and
in
37,
1984
capsule for microscope examination. are required.
mounted
is essential.
The fore-tarsi
cleared
This method of mounting
the examination of which
To
should be
for microscope examination
10% potassium hydroxide solution
The chela should be dehydrated, examination.
no.
examination of the chela
prepare chelae relaxed specimens balsam.
Ins!.,
in
Canada
is not necessary.
and mounted for microscope
is recommended
is important for
also for the palpi,
identifying females and males of
Gonatopodinae. The palpi should be removed using a hooked pin or two fine needles. 1939)
Richards'
is too complicated
boi ling of the speci men amount of boiling, which
method for removing and in
the mouth-parts
is not recommended.
water or very
the mouth-parts can
weak
potash.
Dryinid
larvae should be preserved
acetic acid
(absolute alcohol
After
be pulled out with
must be made to pierce the gula area and
in Carnoy's fluid
then
is based on the proper
a
fine needle,
tear forwards.
in 90% alcohol
60 ml,
(Richards
This method
after being
chloroform 30 ml
fixed
and glacial
10 ml).
FAMILY SIZE AND DISTRIBUTION
This revision records 844 species grouped
probably comprises approximately distribution of subfami I ies and Krombein,
Hurd,
North of Mexico)
(1979:
Nearctic;
Neo.
=
tables 2a and 2b. names
Catalog of Hymenoptera
in America
Eth.
Ori.
Nea.
Neo.
46 37 5 2
24 22 1 1
22 16 3 3
79 55 1
-
-
2 3 3
-
2
1 7 7
-
-
8 8
-
-
-
6 4 2
-
-
1 1
-
-
-
-
-
-
I
Geographic distribution of Pal.
in
geographical
25 20 1 3
Pal.
2a -
world fauna
The geographical
have been follows.
Anteoninae Anteon Deinodryinus Lonchodryinus Metanteon Prioranteon Aphelopinae Aphelopus Crovettia Apodryininae Apodryinus Biaphelopinae Biaphelopus
per region:
is shown
and Canadian
Smith and Burks
ncmina dubia and nUda)
The actual
1,000 species.
genera
For abbreviations of U.S.
Tab.
(except for
into 46 genera and 10 subfamilies.
=
Palaearctic;
Neotropic;
Aus.
Eth. =
I
Dryinids and = Ethiopian;
Australian.
21
Aus.
44 42 2
14 9 5 1 1
-
-
-
5 5
-
number of known Ori.
=
Oriental;
species
Nea.
=
33
Introduction
Bocchinae Bocchoides Bocchus Chelothelius Mirodryinus Mystrophorus Radiimancus Conganteoninae Chelanteon Conganteon Fiorianteon Dryininae Alphadryinus Cretodryinus Dryinus Megadryinus Mesodryinus Perodryinus Richardsidryinus Tridryinus Gonatopodinae Acrodontochelys Adryinus Agonatopoides Apterodryinus Dicondylus Donisthorpina Echthrodelphax Epigonatopus Esagonatopus Gonatopus Gynochelys Haplogonatopus Neodryinus Pentagonatopus Plectrogona topoides Pseudogonatopus Tetrodontochelys Trichogonatopus Thaumatodryininae Thaumatodryinus Transdryininae Transdryinus Tab.
2b -
Eth.
Ori.
Nea.
Neo.
18
4 1 3
4
8
-
-
-
9 2
-
5
-
1 1 2
-
-
-
1 1 15 2 1 7
2
-
2
Pal.
Neo.
~
~
8
-
1 1
-
-
26 1
23 2
-
-
-
-
15
48 8
-
4
-
-
-
-
2
1
4
-
-
-
3
3 2 67 1 5 5 3 1
4 1 3 3 59 2 2 2 3 1
3 1 2 1 5
3
-
-
-
40
3 -
1 13 6
-
Aus.
Eth. ~
51 1
4 2 1 1 3
-
30 1 1 2
18
-
-
8 7
10 1
-
-
5 5
-
1 9
-
-
-
-
-
-
5 -
-
15
76 1
5
-
-
-
2
Aus.
-
19
Palaearctic;
Neotropic;
4
-
-
Geographic distribution of
per region: Nearctic;
Pal.
1
2 29
-
-
25 1
-
-
19 1 4 4
23
-
1
12 57 1
43 1
-
-
-
-
1 7 1
-
-
-
24
12
2 7 4 2
-
-
2 9
1 4 1
3 1 1
-
8
5 -
3 3
3 1 1 1
-
-
7 6 6
-
-
-
8 1
2 2 1 1
Dryinids and number of known species ~
Ethiopian;
Australian.
Ori.
~
Oriental;
Nea.
~
34
;"em.
Amer.
t<EY TO THE
En!.
Ins!.,
no.
37,
SUBFAi\', I LIE 5 Or=-
1984
DNYldI JA~'
F Er'IALE 5
Front
tarsus not
Front
tarsus
Fig.
13 -
Female of
2 Fore-wing
chelate
chelate
wi th
(Fig.
(Fig.
13).
.2 .3
16)
Aphelopus atratus (Dalman) from Caluso (Italy). only
costal
cell
enclosed by
pigmented
veins
(Fig.
13)
1. AphelopinaeR.C.L. Perkins -
Fore-wing vei ns
with
(F i g.
3 Mesosoma
costal
composed by
completely apparent
and median
cells
14)..................... fused
obliterated,
pronotum and
segments
except
a
for
lateral
pronotum and rnesopleura; -
Mesosoma not pronoturn,
because 4 Chela
or
scutum,
the
partly
composed
metanotum and
rudimentary
rudimentary
claw
claw
mesosoma
of
• 1 O. by
ocelli
15) . . . . . . . . . . . . . . . . .
Apodryininae n. fused
segments
visible
easily
(Fig.
subf. (Fig.
16);
recognizable,
16).
.4
9)
••...•.••••• 5
8)...
• .•..••..••. 6
(Fig.
(Fig.
sutures
suture between apparent
(Fig.
propodeum always
sutures are visible;
wi thout
Chela wi th
only
15);
pigmented
subf.
suture between propectus and
absent
. .•. • . . . • • ••• •. • ••• . • •• . • • . • • , ••• '"
enclosed by
13iaphelo,;;inae n.
(Fig. the
track
ocelli
clearly 2.
35
Introduction
f'i9.
g.
14 -
Female of BiaphelopliS Inasneri
n.
15 ~ Female of ArJoc--{r/inus n,d~neri n.
sp.
sp.
(type)
from
from Godavari
t,lalalcahuello
(I'epal).
(Chile).
36
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Fig.
16 -
Female of
Conganteon vuic2nicum Benoit (type) from Ruanda.
Fig.
17 -
Fema I e of
Transdryinus notogeicus n. sp.
(type)
from
Queens I and.
Introduction
Fig.
18 -
Female of
37
j'letanteon aerias (Walker) from EI Coigo (Chile).
5 Median leg without tibial spur (tibial spurs 1,O,1)(Fig. 17); occipi tal carina absent • . . . . . • • . . • . • . 8. Tra.rJsdryininae n. subf. -
Median
leg with
tibial
spur
(tibi",1
spurs
1,1,2)(Fig.
18);
occipital
carina complete • . • . . • . . • • • . . . . . . • . . . . 4. Anteoninae ."I.C.L. Perkins 6 Antennae with tufts of long hairs on segments 5-10 (Fig. 19) .•.• ..• • . . .• . . . .••. . • .. . . . . . '" -
Antennae wi thout
7 Median
tufts
leg without
of
•....•.... long hai rs
tibial
spur
Thaumatodryininae n.
6.
(Fig.
(tibial
spurs
-
Median
leg with
tibial
four
posterior
(Fig.
teeth,
at
most
tarsus
(Fig.
5);
(tibial
spurs
progressing
enlarged
with
21)
spur
teeth,
claw
one preapical
1,0,1
1,1,1
larger
(Fig.
21)
tooth;
or
1,0,2)(Fig.
20)
00natopodinae K i ef fer
. . • • • . . . . . . . • • • • . • . • • . . . . • . • • . . . . • • . • . 9. 8 Mandible with
subf.
16) . • • • . . . . • • . . . . . • . . . . . 7
or
1,1,2)(Fig.
from anterior never with a
fifth
segment
16).8
one
to
row of
of
front
always wi th more
than one preapical lamella . . . . . . . . • . • • • . . . • • • • • • . . . . . . • . . . . . • . . . . . . • . . . . 7. Dryininae K i ef f er
-
Mandible with
1-4
large
teetn and a
(Fig.
22);
teeth
(Fi;j.
23)
front
tarsus
(Fig.
wi th veins
quadridentate mandible with
tooth between
winged; costal, (Fig.
23)
has
(Fig.
rarely
to posterior,
row of
fore-wing
pigmented veins
Fully
one
always has a
9 Fully winged; -
22);
the
two posterior
occasionally quadridentate mandible with
larger from anterior
by
(Fig.
rudimentary
strong
teeth
one preapical
with
costal
submedian
in
in
and
24) . . . . . . . . . . . . . . . . . . . . . . . 5.
the
lamella
progressing
enlarged
fifth
segment
cells
winged
cl early
claw of
only . . . . . . . . . . . . . . 9 clearly
Conganteoninae n.
fully
cell s
teeth
this case
and median
16) . . . . . . . . . . 3.
micropterous;
median and
but
three teeth
forms
enclosed
subf. fore-wing
enclosed by oigrnented
lJoccninae ,ortions:
with numerous
occipital
with
antennae
scutum,
shorter than scutellum
without sculpture among
566
Mem.
Amer.
En!.
Inst.,
no.
37,
\i
/lI
I
I l /
\
\I,
I
\
1
\
\ \
1
\
\
\
\
Fig.
punctures; scutum;
366 -
notaulices incomplete,
bands;
strong
part of radial
fore-wing
sp.
(type)
approximately 0,5
transversal
except for a central
is without sculpture; distal
reaching
posterior surface without
transversely striate, which
Chela of Anteon mundum n.
propodeum with a
posterior surface;
/
i /
,
I
1984
keel
length of
between dorsal
longitudinal
keels,
and
strongly
smooth area near dorsal
hyal ine,
without dark
surface
transversal
vein much shorter than proximal
part
567
Anteoninae
(3: 12);
fore tarsal
segment 5 of front enlarged claw prominence; 55
segments
in following proportions:
tarsus with basal
(Fig.
366)
with one bristle
segment 5 of front
tarsus
lamellae extending continuously
MALE:
part
6:
3:
6:
longer than apical
Fig.
Kassam
367 -
(12:9);
located on a proximal
(Fig.
366)
to the apex;
with some rows of nearly tibial
spurs
unknown
LOCUS TYPICUS:
12: 31; part
(NE New Guinea)
Chela of Anteon bellator n.
sp.
(type)
1,1,2.
568
Mem.
TYPICAL MATERIAL: HOSTS:
Amer.
En!.
holotype F!
Ins!.,
no.
37,
1984
in B.
unknown
DISTRIBUTION: Kainantu,
Australian
region:
NE New Guinea),
NEW GUINEA:
Kassam
(48 Km E of
B!
24. Anteon bellator n. sp.
FEMALE: fully
fully
thickened; 8:
8:
7:
OL = 3;
striate,
10;
length 3 mm;
abdomen
antennal
8:
longitudinal 5;
winged;
testaceous;
mandibles,
petiole black;
antennae and
without
OOL = 6;
fully retuculate rugose;
lateral
OPL = 7;
keels;
TL = 4;
occipital
(8:17);
13: 8:
pronotal
10:
10:
10:
frons with a median
carina complete;
pronotum shiny,
except for a smooth area near scutum;
pronotum shorter than scutum
legs
antennae not distally
segments in following proportions:
head shiny,
keel,
black;
testaceous;
POL
transversely
posterior surface of tubercles reaching
tegulae;
scutum shiny, smooth, weakly punctate, without sculpture among punctures or very weakly granulated; notaulices incomplete, reaching only approximately 0,25 without sculpture; keel
between dorsal
longitudinal distal
length of scutum;
scutellum and metanotum shiny,
propodeum reticulate rugose,
keels;
and posterior surface; fore-wing hyaline,
part of radial
vein
with a strong
transversal
posterior surface without
without
dark transversal
much shorter than proximal
tarsal segments in following proportions: 10: 4: 4: 7: front tarsus with basa! part shorter than apical part
part
bands;
(5:10);
fore
17; segment 5 of (7: 10); enlarged
claw (Fig. 367) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 367) with 2 rows of nearly 42 lamellae extending continuously MALE: unknown LOCUS TYPICUS:
U.
Watut
TYPICAL MATERIAL: HOSTS:
to the apex;
tibial
spurs 1,1,2.
(NE New Guinea)
holotype F!
in B.
unknown
DISTRIBUTION: Guinea),
Australian
region:
NEW GUlrjEA:
U.
Watut SW
(NE New
B!
25. Anteon nigricorne (R.C.L. Perkins)
Neochelogynus nigricornis R.C.L. Perkins 19Q5: 63. Anteon nigricornis (R.C.L. Perkins): Kieffer 1907b: 27 Chelogynus nigricornis (R.C.L. Perkins): Kieffer 1914b: 191.
FEMALE:
fully
winged;
wi th teeth brown;
length 2,62-2,93 mm;
antennae brown;
black;
mandibles testaceous,
tegu I ae testaceous;
legs redd i sh-
569
Anteoninae
368 -
Fig.
Anteon nigricorne (i'.C.L. Perkins) from Sandhill
Chela of
(Austral ia)
testaceous, with coxae brown; occasionally brown, with tarsi light; antennal segments in following proportions: 11: 5: 5; 5: 5: 5: 5: 5: 6;
head fully
reticulate rugose;
frons with a
median
longitudinal
5:
keel
and with 2 lateral keels near orbits directed towards antenna I sockets; occipital carina complete; POL ~ 7; OL ~ 3; OOL ~ 4; OPL ~ 4; TL ~ 4; pronotum reticulate rugose,
(8:12);
scutum
pronotal
metanotum smooth,
hairy,
shiny,
weakly
punctures;
notaulices incomplete,
of scutum;
propodeum
dorsal
reticulate
and posterior surface;
fore-wing
hyaline,
with posterior surface shorter than
tubercles reaching
without
punctate,
tegulae;
reaching only rugose,
scutum,
with
a
approximately 0,25 transversal
posterior' surface without
dark
scutellum and
without sculpture among
transversal
bands;
keel
length
between
longitudinal
distal
keels;
part of radial
(3:8); fore tarsal segments in 12; enlarged claw (Fig. 368) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 368) with 1 row of 21 lamellae; apex with a group of 7 lamellae; tibial spurs 1,1,2. vein much shorter than proximal
following proportions:
MALE: unknown LOCUS TYPICUS:
Bundaberg
TYPICAL MATERIAL: Olmi
1982)
HOSTS:
(Australia)
lectotype F!
and 3 paralectotypes FF!
(designated by
in B.
in Austral ia
DISTRIBUTION: Sandhi II,
part
8: 2: 2: 5:
B!
Eutettix sp. (R.C.L. Perkins 1906c).
Australian
region:
AUSTRALIA:
Bundaberg
(Queensland),
B!
OL!
26. Anteon fljianum n. sp.
FElj,ALE:
fully
tegulae and
winged;
length 1,25-1,56 mm;
legs yellow;
antenna I segments
black;
mandibles,
antennae,
in following proportions:
7:
570
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
369 - Chela of Anteon fijianum n. sp.
37,
1984
from Lautoka
3,5: 2: 2: 2: 2: 2: 2: 2,5: 4;
antennae distally thickened;
granulated,
area
except for a
is without sculpture; carina complete; shiny,
( 3,5:7);
with
a
OL = 3;
pronotal
without sculpture,
incomplete,
reaching
metanotum shiny, wi th a
frons
POL = 5;
with numerous transversal
than scutum smooth,
small
smooth,
longitudinal OOL = 5; keels,
head dull,
median
OPL = 4;
keel;
occipital
TL = 5;
pronotum
with posterior surface shorter
tubercles reaching
with sides very
tegulae;
scutum shiny,
weakly granulated;
length of scutum;
without sculpture;
between
Islands)
in front of the anterior ocellus which
approximately 0,3
transversa I keel
(Fiji
notaulices
scutellum and
propodeum reticulate rugose,
dorsa I and posterior surface;
posteri or
surface wi thout long i tud i na I keel s; areol ae of dorsa I surface wider than those of posterior surface; fore-wing hyaline, without dark transversal bands;
distal
part of radial
vein much shorter than
proximal
part
(1,5:4); fore tarsal segments in following proportions: 4: 1: 2,5: 2,5: 8; enlarged claw (Fig. 369) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 369) with 1 row of 6 lamellae; apex with a group of 3 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Lautoka
TYPICAL MATERIAL: F!
Islands) and 3 paratypes FF!
in 8M;
1 paratype
in OL.
HOSTS:
in Fiji
DISTRIBUTION: BM!
(Fiji
holotype F!
AUSTRAL IA:
Islands
Nesosteles sanguinescens Kirkaldy.
Australian Narrabri
region:
FIJI
ISLANDS:
(New South Wales),
Lautoka
8M!
27. Anteon dubium Fouts
(Viti
Levu),
DL!
Anteoninae
571
Anteon dubius Fouts 1935: 157.
FEMALE:
fully
winged;
with teeth brown; with coxae brown; granulated; broad
length 1,70 mm;
body shiny,
scutum granulated;
approximately 0,5 without
longitudinal
bands;
distal
very short,
head
twice as long as
reaching
scutellum and metanotum smooth;
fore-wing
part of radial
segment 4 of front
more than
mainly near scutellum;
keels;
legs yellow,
mainly on head;
notaulices incomplete,
length of scutum;
propodeum strongly rugose,
mandibles yellow,
tegulae yellow;
with sparse hairs,
pronotum granulated,
(3:1);
brown-black;
antennae yellow-brown;
hyal ine,
posterior surface
without dark transversal
vein much shorter than
proximal
part;
tarsus slightly shorter than segment 1 (17:22);
tibial
spurs 1,1,2. MAL:::
unknown
LOCUS TYPICUS:
Teivipakeka
TYPICAL MATERIAL:
(Marquesas
holotype F!
HOSTS: unknown DISTRIBUTION: Australian region: (Nukuhiva),
Islands)
in B. MARQUESAS
ISLANDS: Teivipakeka
B!
28. Anteon coriaceum (R.C.L. Perkins)
Neochelogynus coriaceus R.C.L. Perkins 1905: 65. Anteon coriaceus (R. C. L. Perk i ns): Kieffer 1907b: 25. Anteon apterus Dodd 1913: 181 (n. syn.); locus typicus Nelson (Austral ia)
Chelogynus coriaceus (R.C.L. Perkins): Kieffer 1914b: 192.
FEMALE: with and
fully
winged;
teeth brown; legs yellow;
length 2,01-2,12 mm;
antennae brown or yellow, legs sometimes brown;
without median and
lateral
keels;
body
black;
mandibles yellow,
weakly darkened; dull;
pronotum granulated,
surface shorter than
scutum
incomplete,
anterior third or fourth of scutum;
reaching
(6: 11 );
tegulae
head granulated;
scutum granu I ated;
frons
with posterior notau Ii ces scutellum and
metanotum sh i ny and smooth; propodeum rugose, ma i n I y near metanotum, with a transversal keel between dorsal and posterior surface; posterior surface fully
rugose,
without
longitudinal
keels;
fore-wing
hyal ine,
without dark transversal bands; distal part of radial vrin much shorter than proximal part; fore tarsal segments in following proportions: 6: 1,5: 2:
4:
10; enlarged claw
(Fig.
prominence;
segment 5 of front
than
part
apical
(4,5:5,5),
group of 6 lamellae;
tibial
370)
with one bristle located on
tarsus
(Fig.
370)
with basal
with one row of 12 lamellae; spurs 1,1,2.
a proximal
part shorter
apex with a
572
Fig.
Mem.
370 -
Amer.
En!.
Chela of ;jnteon coriaceum Creek
MALE:
Ins!.,
unknown
no.
(FLC.L.
(Queensland)
37,
1984
Perkins)
from Marlborough
Anteoninae
LOCUS TYPICUS:
Cairns
TYPICAL MATERIAL:
(Queensland,
lectotype F!
573
Australia) 1982) of N.
(designated by Olmi
coriaceus in S; hoiotype F! of A. apterus in AD. HOSTS: in Australia Euleimonios sp. (R.C.L. Perkins 1906c). DISTRIBUTION: Marlborough Cent.
Australian region:
(Queensland),
Queensland),
OLl
SR!
SR!
AUSTRALIA:
Cairns
Marlborough Creek
Nelson
(Cairns,
(Queensland),
S!
(2 Km N Marlborough,
N Queensland),
AD!
NOTES: A. apterus Dodd was described as wingless. I bel ieve that the single female specimen is damaged and the wings have been broken off.
29. Anteon sucklingi n. sp.
FEMALE:
unknown
MALE: fully winged; length 1,68-1,87 mm; black; mandibles testaceous; antennae brown with segments 1 or 1-2 testaceous; legs testaceous with hind coxae proximally brown; with all
thickened; 7:
7:
7:
antennal
9;
2;
=
OOL
5;
posterior half, incomplete,
OPL
lateral
=
2;
371
-
TL
keels;
=
2;
10: 5:
occipital
carina complete;
scutum shiny,smooth,
8:
gonoforceps
7:
7:
7:
frons
POL = 4;
OL
notaulices
length of scutum;
scutum without
weakly granulated in specimen
Male genital ia of Anteon sucklingi n. S:
also
weakly punctate on
strongly punctate near anterior margin;
reaching approximately 0,65
removed);
Wilhelm)
antennae not distally
with weak areolae or irregular keels;
sculpture among punctures in holotype;
Fig.
(specimen from Mt.
segments in following proportions:
head shiny,
without median and
=
sometimes
coxae and clubs of femora partly brown;
sp.
(type) (A:
(laterally viewed).
right
half
574
Mem.
from Mt.
Wilhelm;
sculpture; keel
Amer.
between
much shorter than proximal
with part
TYPICAL MATERIAL:
Fig.
smooth and
without
longitudinal
bands;
distal
gonoforceps
with an
keels;
fore-
part of radial
(Fig.
371)
inner subdistal
vein
without pointed
(New Guinea)
holotype M!
in B;
1 para type M!
in OL.
unknown
DISTRIBUTION: Wi Ihelm
(3:9);
1984
posterior surface with
two distinct
membranous process,
process; tibial spurs 1,1,2. LOCUS TYPICUS: Mt. Suckling HOSTS:
37,
without a distinct transversal
and posterior surface;
without dark transversal
proximal
no.
scutellum and metanotum shiny,
dorsal
hyal ine,
dorsal
Inst.,
propodeum reticulate rugose,
irregular areolae and keels, wing
Ent.
Australian
(NE New Guinea),
372 -
region:
NEW GUINEA:
Mt.
Suckling,
B!
Mt.
OL!
Male genitalia of
Anteon bismarckense n. half removed.
sp.
(type):
right
Anteoninae
575
30. Anteon bismarckense n. sp.
FEMALE: MALE:
unknown
fully
testaceous;
winged;
length 2,80 mm;
antennae brown,
propodeum black;
head black,
with mandibles
wi th segments 1-3 testaceous;
abdomen brown;
legs testaceous;
thorax and
antennae ha i ry;
antennal segments in following proportions: 13: 7: 10: 7: 8: 9: 9: 8: 9: 12; head shiny, fully reticulate rugose; frons with a complete median keel; = 4;
occipital
carina complete;
scutum smooth,
broad,
=
POL
=
8; OL
very strongly punctate;
similar to areolae;
5; OOL
=
7; OPL
=
6; TL
punctures of scutum very
notaulices incomplete approximately reaching
0,5 length of scutum; scutellum and metanotum strongly puctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface;
median area of posterior surface rugose;
without dark proximal
transversal
part
(3:15);
bands;
distal
gonoforceps
fore-wing
part of radial
(Fig.
372)
with a
hyal ine,
vein shorter than
dorsal
process not
curved at apex; tibial spurs 1,1,2. LOCUS TYPICUS: Yalom (Bismarck Islands) TYPICAL MATERIAL:
holotype M!
HOSTS: unknown DISTRIBUTION: Australian
CO!
in CO;
region:
1 paratype M!
BISMARCK
ISLANDS:
in OL. Yalom
(New Britain),
OL!
31. An teon gioriosum n. sp.
FEMALE: MALE:
unknown
fully
winged;
length
1,25-2,00 mm;
mandibles testaceous,
with
proximal half black; antennae brown; legs brown, with tarsi light; antennae not distally thickened; antennal segments in following proportions:
6:
4:
3:
4:
4:
4:
smooth;
occipital
or with
incomplete median keel
3; OOL smooth,
4:
4:
carina complete;
4:
6;
head dull,
frons without
and
without
fully granulated,
longitudinal
lateral
keels;
median keel
POL = 5;
OL =
=
4; OPL = 2; TL = 3; scutum, scutellum and metanotum shiny, weakly punctate, without sculpture among punctures; notaulices
incomplete,
reaching
reticulate rugose, posterior surface; area shiny, hyaline,
approximately 0,3
slightly
rugose,
nearly as rugose as
without dark transversal
shorter than
length of scutum;
proximal
part
(2:6);
bands;
lateral
areas;
distal part of radial
gonoforceps
(Fig.
proximal
membranous process and without distal
process;
tibial
LOCUS TYPICUS:
propodeum
with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median
373)
fore-wing vein much
without dorsal
inner pointed or rounded
spurs 1,1,2. Mt.
TYPICAL MATERIAL:
Glorious
(Australia)
holotype ~'.!
in 8;
1 paratype M!
in 81-::;
1 paratype
576
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
B Fig.
Male genitalia of Anteon gloriosurn n.
373 -
removed);
M!
in TV/;
HOSTS:
B:
paratype M!
gonoforceps
sp.
(type)(A:
left half
(laterally viewed).
in OL.
unknown
DISTRIBUTION:
Australian region:
SE Queensland), Queensland),
8!
BR!
AUSTRALIA:
2nd Palen Ck.
Mt.
Nebo
(SE
Mt.
Glorious
(Rain Forest,
Crossing from Rathdowney(SE
Queensland),
OL!
Strahan
(Tasmania),
TW!
32. Anteon vulsum n. sp.
FEMALE: unknown t.\ALE: fully winged; with
teeth brown;
testaceous,
with coxae and fore and
antennae not proportions: rugose;
distally 11:
frons
OL
=
rugose,
4;
6:
thickened;
7:
6:
with a
orbits directed 6;
length 2,06-2,25 mm;
antennae testaceous,
6:
antenna I segments 5:
6:
6:
10;
keel
towards antenna I sockets;
OOL
=
5,5;
OPL
approximately
=
mandibles testaceous,
4,5;
visj~le
legs
hind clubs of femora partly brown;
track of median
with areolae less
incomplete,
5:
black;
with segments 4-10 darkened;
=
TL
in following
head shiny,
and with 2 occipital
3,5;
reticulate keels near
carina complete;
scutum shiny,
near posterior margin;
reaching 0,5
fully
lateral
POL
reticulate
notaulices
length of scutum;
scutellum and
metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 areolae smaller than dark
transversal
proximal
part
bands;
(2:9);
longitudinal
areolae of distal
keels;
lateral
median
areas;
part of radial
gonoforceps
(Fig.
374)
area rugose,
fore-wing
hyaline,
with without
vein much shorter than
without dorsal
proximal
577
Anteoninae
A
Fig.
Male genitalia of Anteon
374 -
B:
removed);
membranous process and
without
Mt. Suck ling
LOCUS TYP I CUS:
TYPICAL MATERIAL:
vulsum n. sp.
gonoforceps
distal
(I a tera II y
(type)(A:
right
half
viewed).
inner process;
tibial
spurs 1,1,2.
(Nevi Gu i nea)
holotype M!
and
1 paratype M!
in B;
1 paratype Iv'i!
in OL. HOSTS:
unknown
DISTRIBUTIO;":
Australian
(NE ~,ew Guinec.),
OL!
~,EW
region:
rill.
I.
unknown
DISTRIBUTIOr,: Queensland),
Australian region:
AUSTRALIA:
Kuranda
AD!
42. An teon gira u Iti Dodd
Anteon giraulti Dodd 1914d: 251.
(Cairns,
N
588
Mem.
Fig.
384 -
Amer.
En!.
Inst., no. 37,
1984
Chela of Anteon giraulti Dodd
(type)
Anteoninae
FEMALE:
fully
winged;
abdomen brown;
antenna I segments 8;
head shiny,
shiny,
I"OL
reticulate rugose, ~
4;
~
3,5;
is smooth;
(6:11);
scutum,
mandibles testaceous;
antennae distally 10:
except for a
is smooth;
OL
reticulate rugose,
wh i ch
black;
legs yellow;
in following proportions:
ocellar triangle which complete;
length 2,4 mm;
antennae and
589
frontal ~
OOL
3;
8:
9:
7:
wide area
I ine complete; ~
OPL
6:
2,5;
6:
6:
6:
in fr'ont of
occipital ~
TL
thickened; 6:
2,5;
carina
pronotum
except for a short area near posterior margin
posterior surface of pronotum shorter than
scutellum
and metanotum shiny,
smooth,
scu tum
without
sculpture; notaulices incomplete, faintly visible near anterior margin of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal longitudinal distal
posterior surface; fore-wing
part of radial
segments 384)
and
keels;
front
in following proportions:
tarsus
(Fig.
1\-1ALE:
without dark
vein shorter than
with one bristle located on 384)
group of nearly 30
posterior surface without
hyaline,
with
7:
proximal
3: 4:
11:
a proximal
21;
tibial
(2:10);
bands;
fore tarsal
enlarged claw
prominence;
two rows of nearly 33
lamellae;
transversal
part
(Fig.
segment 5 of
lamellae;
apex with a
spurs 1,1,2.
unknown
LOCUS TYP I CUS:
Austra I i a
TYPICAL MATERIAL: HOSTS:
holotype F!
in AD.
unknown
DISTRI8UTIO~I:
Australian
,I Queensland),
region:
AUSTRALIA:
Yungaburra
(Cairns distr.,
AD!
5. Prioranteon n. gen.
TYPE SPEC I ES: Prioranteon casalei n. sp. FEMALE (Figs 385, 386): micropterous; thoracic structure simi lar to that of
Gonatopodinae: pronotum remarkably elongate; scutum forming a stalk
between
the pro- and metanotum;
propodeum segments;
humped;
located further' distally 390)
MALE:
than
i nv i si b Ie;
with 6 segments;
inner side of enlarged claw
prominence not bearing bristles; 388,
metanotum
maxillary palpi
(Fig.387)
metathorax + labial
tibial
prominence;
chela
spurs 1,1,2.
unknown
Palaearctic,
with 3
1-2 short bristles or peg-like hairs
the proximal
without rudimentary claw;
DISTRI8UTIO~:
palpi
with a proximal
Ethiopian
HOSTS: unknown SPECIES: 3
PALAEARCT I C SPEC I ES OF PRIORANTEON
(Figs 387,
590
Mem. Amer. En!.
Fig.
385 -
Ins!.,
no. 37,
1984
Female of Prioranteon casalei n.
sp.
(type)
Anteoninae
Fig.
386 -
One species
Female of
591
l'riora"teon casalei n. sp.
(type)
is known.
Prioranteon biroi n. sp.
FEMALE:
micropterous;
scutellum;
fore
testaceous,
with
teeth brown;
segments 1-3 testaceous; testaceous; black, 10:
8:
10:
10;
punctures;
segments
occipital
frons
mandibles ';,ith 4 teeth progressing
posterior; weakly
vertex flat;
punctate,
posterior
transversal
narrower than tegulae,but nearly as reduced; long as scutum humped,
disc;
very
without median
and
smooth,
tarsi
13:
13:
11:
keel,
but
POL = 7;
with Tl =
anterior one to
without sculpture or very
very
posterior lobe directed
humped,
scute II um; 389 8);
shiny,
smooth,
tegulae and
long anterior disc,
towards
carinate;
reduced
tegulae;
hollow,
without fore
reaching scutum
rugose,
sculpture,
wings on
very
nearly
as
sides of
fore wi ngs not reachi ng transversa I furrow beh i nd metanotum
invisible;
separated from scutellum by
transversely
hind 18:
posterior margin of collar almost straight,
as broad;
(Fig.
OOl = 11;
larger from
long as posterior collar of pronotum, scutellum
5:
without sculpture among
Ol = 5;
humped and
10:
impression se;oarating disc from a posterior collar,
without a
long
scutellum
pronotum shiny,
with a
with
with coxae almost fully
proportions:
POL = 4;
as
mandibles
antennae brown,
last segment of mid and
weakly punctate,
furrow;
long
black;
with posterior collar whitish-
legs testaceous,
carina complete;
median
nearly as
testaceous;
in following
head shiny,
longitudinal
12;
clypeus
pronotum black,
tegulae testaceous;
antennal
reduced,
length 3,25-3,80 mm;
clubs of femora darkened,
brown;
a
wings very
hind wings invisible;
a
metathorax + propodeum
transversal
hollow
impression
anterior surface of metathorax + propodeum shiny,
592
Mem. Amer.
Ent.
Inst.,
no. 37,
1984
i
I
j
!
\\\. /' -.
~~/
Fig. 387 -
/
Chela of Prioranteon biroi n. sp.
(type)
593
Anteoninae
partly granulated and smooth; disc of metathorax + propodeum with a distinct longitudinal median furrow; posterior surface of propodeum dull, granulated and with some weak smooth,
shiny,
complete;
fore tarsal
enlarged claw
(Fig.
the proximal
prominence; lamellae;
long);
meso-metapleural
with one bristle
into a
small
unknown
hook;
LOCUS TYPICUS:
tibial
11:
3:
6:
13:
25;
tarsus
a group of 5
387)
(Fig.
lamellae
with 2 rows
(among which
segment 2 of front
1
tarsus produced
spurs 1,1,2.
SzSztMiklos
TYPICAL MATERIAL:
pleura
located further distally than
segment 5 of front
apex with
rudimentary claw absent;
MALE:
striae;
suture distinct and
segments in following proportions:
387)
of nearly 29 very
irregular longitudinal
without sculpture;
(Hungary)
holotype F!
and 1 paratype F!
1 paratyep F!
in BU;
in OL. HOSTS:
unknown
DISTRIBUTION: NOTES:
Palaearctic region:
the species
HUNGARY:
is named in honor of the
SzSztMiklos, late Mr.
L.
BU! Biro.
ETHIOPIAN SPECIES OF PRIORANTEON
Two species are known.
KEY TO THE SPECIES
FEMALES
Posterior
surface of
propodeum strongly
. . • • • • • • . • • . . . . • • . . • . . . . • • . . • . . • . .. -
Posterior
surface of
transversely striate .•• sp.
1. richardsi n.
propodeum granulated,
kee Is. • • • • • • • . • • • • • . • . . • • • • • • • • • . •.
2.
without
casalei n.
transversal
sp.
MALES
Unknown
In the key to the Ethiopian species, the Palaearctic species PI'. biroi shou I d be near PI'. casalei, because the posterior surface of the propodeum is granu I ated and not transverse I y str i ate. Casalei and biroi NOTES:
can be distinguished as follows:
594
{v,em.
Amer.
Ent.
Inst.,
no.
t,letathorax
+ propodeum with a distinct
disc;
with
head
reaching
frons
transversal
fully
black;
furrow behind
frons black, reaching
with
a
central
furrow behind
winus
scutellum
longitudinal
yellowish
scutellum
furrow on
shorter, (Fig.
biroi n;
+ propodeum without
Metathorax
1984
longitudinal
fore
. .. . .. . .. . . . . . . . . .. . . . .. . . . . .. . -
37,
not
tl).
389
sp.
furrow on disc;
area;
fore
wings
longer,
389 A}......
(Fig.
casalei n.
.. . . . . .
sp.
1. Prioranteon richardsi n. sp.
FEMALE:
rnicropterous;
scutellum;
fore
hind wings
testaceous with
teeth brown;
segments 1-2 and proximal pronotum black, legs brown, tibiae and B:
5:
17:
with
with tarsi
12:
wings
reduced,
clypeus
testaceous;
part of coxae,
testaceous; 10:
8:
7:
antenna I segments 7:
9;
TL =
mandibles
carina complete;
larger from anterior one to posterior; smooth,
weakly punctate,
only on
sides;
pronotum
posterior transversal narrower than tegulae, nearl y
but
as
reduced;
disc;
a
behind
posterior
humped,
nearly
as
and
shiny,
scutellum;
(Fig.
24;
distally
weakly
fore
proximal
tarsal (Fig.
and
hook;
MALE:
unknown
LOCUS TYPICUS:
tibial
388)
absent;
spurs
Van Reenen
TYPICAL MATERIAL:
12;
very
OPL
=
6;
granulated
long anterior disc;
towards
posterior collar, reaching
tegulae;
hollow,
scutum
rugose,
without sculpture,
very
nearly
as
tegulae and reduced fore wings
wings
not
reaching
invisible; a
transversal
metathorax +
transversal
hollow
posterior surface strongly striate;
with
in following 2 bristles
apex
with
a
11:
tarsus
(Fig.
3BB)
group of nearly 4
segment 2 of front
South Africa)
in 8M.
pleura strongly proportions:
located further
segment 5 of front
1,1,2.
(Natal,
holotype F!
punctate,
teeth progressing
disc from a
metanotum
segments
lamellae;
rudimentary claw
small
=
pronotum shiny,
strong and complete;
prominence;
with 2 rows of nearly 43 lamellae;
OOl
anterior surface of metathorax +
granulated;
suture distinct,
striate;
into a
proportions:
strongly
3;
with 4
separated from scutellum by
enlarged claw
than
fore
389 C);
transversely carinate;
transversely
smooth,
long as broad;
impression
12:
humped
lobe directed
humped,
meso-metapleural
=
OL
vertex flat;
impression separating
scutellum
propodeum dull,
testaceous;
sculpture among punctures,
very
propodeum
7:
smooth,
long as posterior coil ar of pronoturn, scutellum
sides of scutum
furrow
with
with
stalk of femora,
posterior margin of collar almost straight,
without a
long as scutum, on
without
4;
as
antennae brown,
in following
head shiny,
=
long
mandibles
tegulae testaceous;
trochanters,
POL
occipital
black;
half of 3 testaceous;
without sculpture among punctures; 11;
nearly as
length 4,68 mm;
anterior margin and posterior collar
distal
10:
very
invisible;
tarsus produced
3:
595
Anteoninae
('ig.
388 -
Chela of Prioranteon richardsi n.
sp.
(type)
596
Mem.
Fig.
389 -
Amer.
Inst.,
no.
37,1984
Prioranteon casalei n. sp. (A), biroi n. richardsi n. sp. (C).
Reduced fore wings of sp.
HOSTS:
Ent.
(B),
unknown
DISTRIBUTION:
Ethiopian
Natal),
8M!
NOTES:
the species
region:
is named
SOUTH AFRICA:
in honor of Mr.
Van Reenen
Owain W.
(Drakensberg,
Richards.
2. Prioranteon casalei n. sp.
FEMALE long
(Figs 385,
386):
as scutellum;
mandibles testaceous, longitudinal
segments shiny,
=
with
genae,
=
OPL
5;
pronotum shiny, with a
TL
=
very
5:
clypeus and a
abdomen reddish-brown;
tarsi
13:
8;
occipital
9:
8:
testaceous;
7:
carina complete;
humped and very
than disc;
posterior margin of collar almost straight, posterior
disc from
antennal
5:
9;
head
POL = 3,5;
mandibles with 4 vertex flat;
long anterior disc;
impression separating a
6:
weakly punctate only on
trasversal but without
5:
anterior one to posterior;
without sculpture,
with
with posterior collar and
without sculpture among punctures;
larger from smooth,
8:
and
nearly as
black;
antennae brown,
testaceous;
junctions
in following proportions:
teeth progressing sides,
length 3,25-4,00 mm;
pronotum black,
tegulae
trochanters,
weakly punctate,
9;
teeth brown;
testaceous;
margins testaceous;
legs brown,
OL
with
fore wings very reduced,
invisible;
median area on frons yellowish;
segments 1--2 partly lateral
micropterous;
hind wings
a
lobe directed towards
posterior
posterior collar, tegulae;
reaching
narrower tegulae,
scutum nearly as
Anteoninae
Fig.
390 -
Chela of
597
Prioranteon casalei n. sp. (type)
long as posterior coil ar of pronotum,
hollow,
rugose,
very reduced;
scutellum humped, shiny, smooth, without sculpture, nearly as long as scutum, nearly as broad; tegulae and reduced fore wings on sides of scutum and scutellum; scutellum humped,
(Fig.
389 A);
fore w;ngs reaching metanotum
transversal
invisible;
separated from scutellum by a
furrow
behind
metathorax + propodeum
transversal
hollow
impression
598
Mem.
transversely carinate;
Amer.
En!.
Ins!.,
no.
37,
1984
anterior surface of metathorax + propodeum shiny,
smooth, without sculpture; posterior surface dul', granulated, without transversal striae; meso-metapleural suture distinct, strong and complete; pleura dull,
granulated,
with weak transversal
segments in following proportions: 390)
with 2 bristles
prominence; lamellae; absent;
9:
2: 5:
keels;
12: 21;
located further distally than
segment 5 of front
tarsus
(Fig.
390)
tarsus produced
(Fig.
the proximal with 1 row of nearly 33
apex with a group of nearly 3 lamellae;
segment 2 of front
fore tarsal
enlarged claw
rudimentary claw
into a small
hook;
tibial
spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Milnerton
TYPICAL MATERIAL: FF!
(South Africa)
holotype F!
and 7 paratypes FF!
in BM;
4 paratypes
in OL.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
SOUTH AFRICA:
BM! OL! NOTES: the species is named in honor of Mr.
Fig. 391
-
Milnerton
(Cape Town),
Achi lie Casale.
Female of Bocchoides bekilyensis Benoit
(holotype)
Bocchinae
599
5. SUBFAMILY BOCCHINAE RICHARDS
Bocchini Richards 1939: 189. Bocchini Muesebeck and Walkley 1951: 1039. Bocchinae Nagy 1967b: 334.
TYPE
Bocchus Ashmead 1893
GENUS:
FEMALE:
fully
brachypterous
winged (Fig.
(Fig.
424);
391)
or micropterous (Fig.
fore-wing
(in fully
452)
or
winged species)
with costal,
median and submedian cells clearly enclosed by pigmented veins (Fig. 391); maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front with
tarsus chelate; 1,
2,
chela
3 or 4 teeth
three
large teeth and a
(Fig.
396 A);
rarely
(Fig.
(Fig.
405)
396);
with rudimentary claw;
rudimentary
tooth between
quadridentate mandibles with
larger from anterior one to posterior; of strong
teeth
(Fig.
405);
mandibles
quadridentate mandibles usually teeth progressing
enlarged claw usually with a row
segment 5 of front
tarsus
(Fig.
405)
only with a preapical lamella, without other lamellae; pronotal directed towards the tegulae present; occipital carina complete, incomplete;
petiole usually very
long,
with
the two posterior teeth
rarely
short;
usually tubercles rarely
antennae without
tufts of long hairs; pterostigma usually very reduced, sometimes wide; tibial spurs 1,1,1, sometimes 1,1,2. MALE: fully winged (Fig. 393) or micropterous (Fig. 454); fore-wing (in fully
winged species)
enclosed by
Fig.
with costal,
pigmented veins)(Fig.
392 -
Female of
median and submedian cells clearly 393);
maxillary palpi
with 6 segments;
Bocchoides bekilyensis Benoit (holotype)
600
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
/
Fig.
393 -
Male of Bocchoides bekilyensis Benoit
(allotype):
antennal
segments 7-10 missing.
labial
palpi
with 3 segments;
mandibles with 1, 2,
3 or 4 teeth;
tridentate or quadridentate mandibles with teeth not progressing from anterior one to posterior;
occipital
carina complete;
tibial
larger spurs
1,1,2. DISTRIBUTION: Palaearctic, Ethiopian, Oriental, Nearctic, Australian HOSTS: Issidae; Cicadellidae (exc Iud i ng I diocerinae, Macropsinae and
Typhlocybinae) GENERA: 6 SPECIES:
39
KEY TO THE GENERA OF BOCCHINAE
FEMALES
Occipital head;
carina
enlarged
incomplete,
claw with
only
visible on dorsal
one preapical
lamella
(Fig.
side
of
399) .•
Bocchinae
...........................
2
Occipital
carina
preapical
lamella;
preapical
lamella
Segment with tooth
5 of
other
enlarged Enlarged front
most
segment
tarsus with (Fig.
front
of
lamella,
claw at
never
least
with
one
lamellae
preapical
(Figs
395,
lamella, 442,
451,
sometimes 456);
claw without teeth (Figs 395,442,451,456) . . . . . . . . . 3 claw wi th numerous bristles (Fig. 395 A); segment 4 of
Enlarged
least
claw withoit
451,456);
segment
segment
Fore-wing
as
long as
segment
front
1
(Fig.
395 B) . . . . . . . .
Bocchoides Beno i t
1.
brist les or
4 of
other appendixes
tarsus
than
0,5
(Figs 442, times as
1
(Figs
442,
(Fig.
452) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
complete
Fore-wing with
451,
less
reduced
........................... Fore-wing 5
one
tarsus with
Bocchus Asmead
tarsus without
group
tarsus at
long as
front
enlarged
3.
• • • • • • • • . • • • • • • • • • • • • • • • • ••
4
claw never with 5 of
one preapical
405);
405) •••••••••••
5 of
Chelothelius Reinhard
2.
enlarged
(Figs 395,405) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
front
wi th a di stal 3
at
lamellae
(Fig.
Segment
complete;
601
radial
f\1ystrophorus Forster
6.
(Figs 438,
456) •••••••••••••••••••••••• 4
449) ••••••.••••••••••••••••••••• 5
vein distinct
(Fig.
438) . . . . . . . . . . . . . . . . .
••••••••• • ••••••••• • • •••• •• 4. Mirodryinus Ponomarenko Fore-wing with no radial vein (Fig. 449) . . . . . . . . . . . . . . . . . . . . . . . • ••••••••••••••••••••••••••
Nadiimancus Moczar
5.
MALES
Temp I e s not
vis i b I e
(F i g s
393,
• • • • • • • • • • • • • • • • • • • • • • • • • •• 2
Temples distinct (Figs 402,403,440,441,454,455) . . . . . . . . . . 2 Fore-wi ng reduced (F i g. 454) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ........................... Fore-wing
3
394) . . . . . . . . . . . . . . . . . . . . . . . . . . ..
Bocchoides Ben 0 i t
1.
compl ete
Mystrophorus Forster
6.
(Figs 402,
440) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
Propodeum with areolae very
small,
440) ••.••••••••••••••••••••
4.
Propodeum wi th at
I east
least
tegulae
as
large as
part
of
(Fig.
• • • . • • • • • • • • • • • • • • • • • • • • • ••
3.
less wide
than
tegulae
(Fig.
Mirodryinus Ponomarenko the areolae very
large,
at
402) . . . . . . . . . . . . . . . . . . . . . . . . . . .
Bocchus Asmead
The ma les of Chelothelius Rei nhard and Radiimancus Moczar are unknown.
1. Bocchoides Benoit
602
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Bocchoides Benoit 1953b: 424.
Bocchoides bekilyensis Benoi t 1953, monotyp ic.
TYPE SPEC I ES: FEMALE labial
(Figs 391,392): palpi
tridentate
(Fig.
396 B);
segment 5 of front front
tarsus at
not reaching MALE
least as
palpi
visible;
394):
HOSTS:
(Fig.
lamellae (Fig.
395);
1 (Fig.
pronotal
395);
395);
segment 4 of tubercles
spurs 1,1,2. winged;
with 3 segments;
mandibles tridentate,
DISTRIBUTION:
mandibles
with numerous bristles
long as segment
fully
with 6 segments;
carina complete;
with numerous
tibial
anterior one to posterior
maxillary palpi
occipital
enlarged claw
tarsus
tegulae;
(Figs 393,
labial
fully winged;
with 3 segments;
(Fig.
maxillary palpi
occipital with
with 6 segments;
carina complete;
teeth not progressing
396 D);
tibial
temples not larger from
spurs 1,1,2.
Ethiopian
unknown
SPECIES:
1
13occhoides bekilyensis Benoi t
Fig.
394 -
Male of
Bocchoides bekilyensis Benoit (allotype): antenna I segments 7-10 missing.
603
Bocchinae
Fig.
395 -
Female of Bocchoides bekilyensis Benoit fore tarsa I segmen ts.
(holotype):
A:
chela;
B:
604
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Bocchoides bekilyensis Benoi t 1953b: 424.
FEMALE
(Figs 391,392):
fully
winged;
clypeus and mandibles testaceous; darkened;
thorax,
testaceous,
propodeum, 14:
7:
33:
distally thickened;
14:
12:
head dull,
10:
8:
hairy,
7:
with
legs brown-
antennal
8:
black,
with segments 8-10
pet iole and abdomen black;
with coxae and clubs of femora dark;
following proportions: slightly
length 6,25-6,50 mm;
antennae testaceous,
10;
rugose,
segments
in
antennae
sculptured by short
irregular keels; occipital carina complete, reaching posterior margin of eyes; temples invisible; thorax and propodeum shiny, hairy; pronotum and scutum smooth,
without sculpture;
without sculpture;
propodeum reticulate rugose; median
area rugose;
five times as as broad
enlarged claw
for", tarsal
(Fig.
segment 5 of front
395)
tarsus
extending continuously MALE
(Figs 393,
mandibles yellow;
394):
(14:2,5);
(Fig.
and
hind
to the apex; fully
4:
14:
rugose;
mandibles tridentate,
396 D);
occipital
temples
invisible;
scutum;
scutellum granulated; without
(25:7);
tibial
yellow;
9;
legs yellow,
antennal
with
segments
1-6
intermediate tooth
in
(Fig.
visible only on anterior half of
metanotum reticulate rugose;
longitudinal
propodeum
keels on posterior surface;
long and ovoidal,
more than
fore-
three times as
long as
spurs 1,1,2. in P.
unknown
DISTRIBUTION: Ethiopian region: MADAGASCAR: 8ekily, P! NOTES: B. bekilyensis is very different from the other Bocchinae. general
spurs
(1,1,2)
are similar to tibial
tridentate mandibles the species
spurs
is included
arrangement of thi s genus prQbab I y
in
is between
The
Anteoninae or as in Dryininae. Female of Dryininae. Only for the Bocchinae. The exact Bocchinae and Dryininae.
structure of the chela is simi lar to that of
Dryininae. The pterostigma is long and narrow tibial
with
in allotype);
posterior margin of eyes;
LOCUS TYP ICUS: Beki Iy (Madagascar) TYPICAL MATERIAL: holotype F! and allotype M! HOSTS:
head black,
head and scutum reticulate
reaching
notaul ices weakly
wings with pterostigma broad
tegulae
with one rudimentary
carina complete,
lamellae
spurs 1,1,2.
length 3,12 mm;
10:
long
(11:14);
without other appendixes;
with 2-3 rows of 63
(segments 7-10 missing
10:
keels;
nearly
fore trochanters nearly twice as
abdomen brown-black;
tibiae darkened;
longitudinal
1 shorter than segment 4
tibial
winged;
antennae yellow
following proportions: 5:
reticulate rugose,
395)
two
long and narrow,
with numerous bristles,
thorax and propodeum black; femora
with pterostigma
segment
scutellum smooth,
with some areolae near scutellum;
posterior surface with
fore-wing
long as broad
(8:4);
notaul ices invisible;
metanotum smooth,
2. Chelothelius Reinhard
Bocchinae
605
A
Fig.
396 -
E
Mandibles of Bocchinae:
rudimentary tooth;
B:
from anterior one to posterior; mandible; E:
quadridentate mandible with one
C:
mandible with one tooth;
tridentate mandible with one rudimentary
Chelothelius Chelothelius Chelothelius Chelothelius
Reinhard
1863:
D:
larger
tridentate
intermediate tooth.
409.
Reinhard:
Kieffer
Reinhard:
Kieffer 1914b: 56.
in Kieffer et Marshall
Reinhard:
Berland
1928:
1905:
125.
143.
Chelothelius gryps Reinhard 1863, monotypic.
TYPE SPECIES: FEMALE
A:
tridentate mandible with teeth progressing
(Figs 397,
398):
fully
winged,
maxillary palpi
with 6 segments;
labial palpi with 3 segments: occipital carina incomplete, only visible on dorsal side of head; mandibles tridentate: enlarged claw with one preapical with
lamella and some bristles
lamellae
(Fig.
399);
pronotal
(Fig.
399);
segment 5 of front
tubercles not reaching
spurs 1,1,2. MALE: unknown DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES:
2
PALAEARCTIC SPECIES OF
Two species are known.
CHELOTHELIUS
tegulae;
tarsus tibial
606
Mem.
Fig.
397 -
Amer.
Ent.
Ins!.,
no.
37,
1984
Female of Chelothelius gryps Reinhard from South France
KEY TO THE SPECIES
FEMALES
Head
fully
granulated,
scutellum dull,
wi th numerous keel s,
granulated and
•••• •••• •••••••• ••• 1. gryps Rei nhard Head shiny, granulated and with keels, wide areas
on
sides
of
ocell i
except
whi ch are
sculpture;
scutum and
sculpture,
weakly granulated,
without
berlandi
Bernard
•••••••••••••••••• • 2.
dull;
scutum and
with keels . . . . . . . .
scutellum shiny,
for
two
smooth and wi thout smooth,
almost
without
keels . . . . . . . . . . . . . . . . . . .
607
Bocchinae
MALES
Unknown
1. Chelothelius gryps Reinhard
Chelothelius gryps Rei nhard 1863: 410. Dryinus gryps (Reinhard): Dalla Torre 1898: 544. Chelotelius gryps Reinhard: Kieffer in Kieffer et Marshall 1905: 127. Chelothelius gryps Reinhard: Kieffer 1914b: 57. Chelothelius gryps Rei nhard: Ber I and 1928: 143. Chelothelius gryps Rei nhard: Bernard 1935a: 41.
FEMALE
(Figs 397,
398):
fully
winged;
length 5,0-5,4 mm;
antennae brown,
with segments 1-2 testaceous; head black, with mandibles, clypeus, a narrow frontal region near clypeus, malar space and a region around orbits reddish-testaceous;
Fig.
398 -
propectus black;
pronotum black,
with margins
Female of Chelothelius gryps Reinhard from South France
608
Mem.
Fig.
reddish;
399 -
Amer.
Ent.
Inst.,
no.
37,
1984
Chela of Chelothelius gryps Reinhard from South France
scutum,
scutellum,
metanotum,
mesopleura,
metapleura and
propodeum black; petiole black; abdomen black-brown; legs testaceous, with coxae partly black and clubs of femora brown; tegulae testaceous; antenna I segments in following proportions:
12: 7:
24:
13:
13: 8:
6:
6:
6:
9; head dull, granulated, with numerous keels, mainly longitudinal; mandibles with 3 teeth; ocellar triangle and occiput smooth; frons with incomplete median
longitudinal
carina visible only on dorsal temples almost transversal transversal
invisible;
keel;
= 5; Ol = 4; OOl = 10; occipital
POL
region of head,
pronotum shiny,
but not reaching eyes;
granulated,
with weak
keels; pronotal turcles not reaching tegulae; pronotum with a impression between anterior collar and posterior humped disc;
scutum dull,
granulated,
with numrous irregular keels or areolae;
notaulices invisible; scutellum as sculptured as scutum; metanotum fully reticulate rugose; propodeum fully reticulate rugose, with 2 incomplete longitudinal bands,
keels on posterior surface;
one on
the basal
fore wings with 2 dark
transversal
cell s and another beneath the pterost i gma;
pterostigma long and narrow;
distal
part of radial
vein
nearly as
long as
proximal part, not reaching wing margin; fore tarsal segments in following proportions: 13: 3: 5: 9: 16; segment 5 of front tarsus (Fig. 399) with some rows of nearly 26 lamellae extending continuously to the apex; enlarged claw
(Fig.
segment 3 of front tibial
spurs 1,1,2.
MALE:
unknown
lOCUS TYPICUS:
399)
with 3 proximal
tarsus produced
Bolzano
(Italy)
into a
teeth and one preapical hook,
with no very
lamella;
long bristle,
Bocchinae
TYPICAL MATERIAL: HOSTS:
probably
609
lost.
unknown
DISTRIBUTION: Goudargues
Palaearctic region:
(Gard),
OL!
ITALY:
FRANCE: Southern France,
Bolzano
P!
Ussel-
(Reinhard 1863).
2. Chelothelius berlandi Bernard
Chelothelius berlandi Bernard 1935a: 41. Chelothelius berlandi Bernard: Bernard 1935b: 53.
FEMALE:
fully
winged;
and mandibles
length 3,75 mm;
testacous;
head black,
antennae testaceous;
segment 10 distally testaceous;
thorax fully black,
(black,
petiole black;
with margins reddish);
testaceous,
with clypeus brown
with segments 6-10 brown; except for pronotum
abdomen black-brown;
with coxae and clubs of femora partly brown;
testaceous;
antennal 6:
9:
ocelli;
frons granulated and with
Fig.
9;
segments in following proportions:
10: 9:
head shiny,
400 -
smooth and
legs
tegulae
10: 5:
23:
14:
12:
without sculpture on sides of
irregular keels;
ocellar triangle rugose;
Chela of Chelothelius berlandi Bernard
(type)
610
Mem. Amer.
occipital
carina
reaching eyes;
incomplete,
Ent.
Ins!.,
smooth,
weakly
tegulae;
without keels;
granulated,
propodeum fully longitudinal basal
proportions:
long and narrow; 12:
and without
3:
5:
9:
very
16;
(Fig.
Fig.
401
400)
-
part
transversal pronotal
smooth,
(9:8),
impression
tubercles not
weakly granulated,
reticulate rugose;
posterior surface with 2 transversal
region;
fore tarsal
distal
bands,
the
vein
wing margin;
in following
tarsus produced
segment 5 of front
tarsus
lamellae extending continuously
with 7 bristles and one preapical
Female of Bocchus umber n.
incomplete
one on
part of radial
not reaching segments
segment 3 of front
long bristles;
with some rows of nearly 30 enlarged claw
dull;
the apical
longer than proximal
pterostigma
disc;
metanotum fully
fore wings with 2 dark
cells and another on
slightly
hook
reticulate rugose,
keels;
region of head,
with a
scutum and scutellum shiny,
notaulices invisible;
1984
POL = 5; Ol = 3; OOl = 8;
between anterior collar and posterior humped reaching
37,
visible only on dorsal
mandibles with 3 teeth;
pronotum shiny,
no.
sp.
to
into a
(Fig.
lamella;
(type)
400)
the apex; tibial
Bocchinae
611
spurs 1,1,2. MALE: unknown LOCUS TYP ICUS:
Plage de Saint-Raphael
TYPICAL MATERIAL: HOSTS:
(France)
in P.
unknown
DISTRIBUTION: Var),
holotype F!
Palaearctic region:
FRANCE:
Plage de Saint-Raphael
(Frejus,
P!
3. Bocchus Ashmead
Bocchus Ashmead 1893: 91. Phorbas Ashmead 1893: 90 (preoccup i ed by Phorbas Duchassa i ng and Michelotti
Fig.
402 -
1864;
syn.
proposed by Richards 1939);
type species
Male of Bocchus pedunculatus Nagy from Sarawak
612
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Phorbas laticeps Ashmead 1893, orig. desig. Eukoebeleia R.C.L. Perkins 1905: 59 (syn. proposed by Fenton 1918b); type species Eukoebeleia mirabilis R.C.L. Perkins 1905, monotypic.
Tetradryinus Kieffer 1913c: 325 (n. syn.); type species Tetradryinus flavipes (Kieffer 1906), orig. desig. Anteonella Dodd 1913: 181 (n. syn.); type species Anteonella robusta Dodd 1913, orig. desig. Phorbasia Kieffer 1913b: 58 (syn. proposed by Richards 1939); type species Phorbasia laticeps (Ashmead 1893), orig. desig. Neoanteon Fouts 1922: 633 (syn. proposed by Nagy 1969); type species Neoanteon rubrica Fouts 1922, monotypic. Hymenodryinus Benoit 1953b: 427 (n. syn.); type species flymenodryinus seyrigi Benoit 1953b, orig. desig. Bocchus Ashmead: Nagy 1969: 321.
TYPE SPEC IES: FEMALE
Bocchus flavicollis Ashmead 1893, orig. desig.
(Figs 24,
401,
424,
425):
fully
winged
brachypterous or micropterous (Figs 424, segments;
labial
palpi
with 3 segments;
quadridentate mandibles usually
425);
(Figs 24,
401)
maxi Ilary palpi
mandibles with 1,
with a rudimentary
2,
or with 6 3 or 4 teeth;
tooth between
the two
posterior teeth (Fig. 396 A); rarely quadridentate mandibles with teeth progressing larger from anterior one to posterior; occipital carina complete; segment 5 of front tarsus with one preapical lamella, never with other lamellae (Fig. 404); enlarged claw with at least one tooth (Fig. 404);
pronotal
Fig.
tubercles reaching
403 -
tegulae;
tibial
spurs 1,1,1.
Male of Bocchus pedunculatus Nagy from Sarawak
Bocchinae
MALE
(Figs 402,
403):
fully
winged;
613
mandibles with 1, 2,
3 or 4 teeth;
quadridentate and tridentate mandibles with teeth not progressing from anterior one to posterior; wide as tegulae
(Fig.
segments;
palpi
labial
402);
larger
areolae of propodeum at least partly as
temples distinct;
with 3 segments;
cnaxillary palpi
fore wings complete;
with 6 tibial
spurs
1,1,2. DISTRIBUTION: HOSTS:
Palaearctic,
Ethiopian, Oriental,
Nearctic,
Australian
Issidae
SPECIES:
29
PALAEARCTIC SPECIES OF
BOCCHUS
Nine species are known.
KEY TO THE SPECIES
FEMALES
Species brachypterous, wi th fore-wing extending apex of propodeum ••••.•••••• 1. italicus n. sp. Species
fully
winged
(Fig.
wi thout
only
to
the
401) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 transversal bands ••••••••••••••
2
Fore-wing hyal ine,
dark
3
umbel' n. sp. 1-2 dark transversa I bands ••••••••••••••••••••• 3 Fore-wing wi th one dark transversal band •••••••••••••••••••••• 4 · . • . . • . . . . . . . . . . . . . . . . . . . . .. 2.
Fore-wi ng wi th
bands . . . . . . . . . . . . . . . . . . . . . 5 much shorter than scutellum; head black, with mandibles yellow; scutellum and propodeum black; pronotum, scutum and metanotum reddish • ••• · . . . . . . . . . . . . . . . . . . . . . . . . . .. 3. szelenyii Mo c za r Head with OOL more than twice as long as POL; scutellum nearly twice as long as metanotum; head reddish-testaceous; thorax and Fore-wing with
4
two dark
Head wi th POL not
transversal
shorter
than OOL;
propodeum black . . . . . . . . . . . . .
5
Thorax
fully
Thorax at
6
reddish ••••••••
least
8
4. slovacus Strejcek 9. parthicus Ponomarenko
partly black . . . . . . . . . . . . . • . . • . . . . . . . . . . . . . . . . • 6
5. paglianoi n. sp. reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Head mostly reddish; frons and vertex at least partly reddish · . • . . • . . . • . . • . . • . . . . . . . . . . .. 6. scaramozzinoi n. sp. Head fully black, at most only with mandibles and and clypeus not black ••••••••••••••••••••••••••••••••••••••••••••••••••••• 8 Pronotum reddish; scutum, scutellum and metanotum black . . . . . . . . Thorax
fully black •••••••.••
Thorax partly black and
7
metanotum not
614
Mem.
Amer.
Ent.
Inst.,
7. Pronotum and
scutum reddish;
no.
37,
1984
scobiolae Nagy
scutellum and metanotum black ..•.•
8. europaeus Bernard
1v1AlES
Only known.
B. scobiolae Nagy and B. slovacus Strejcek are
the males of
I cannot propose a key
to these
two ma les because
I
have not seen
B. scobiolae.
the male of
1. Bocchus italicus n. sp.
FEMALE:
brachypterous
propodeum); occipital distally 7:
7:
carina complete; thickened;
6,5:
6:
scutum dull, pronotal dull,
(fore wings reaching
length 2,56 mm;
6:
head dull,
granulated; antennae
antenna I segmentes in following proportions:
6:
9;
POL
granulated,
=
6;
=
Ol
3;
=
OOl
10;
OPl
with notaulices complete, tegulae;
scutellum
propodeum reticulate rugose,
posterior surface;
only posterior apex of
testaceous;
frons with a median complete keel;
tubercles reaching
rugose;
fully
fore-wing
short,
11:
=
TL
7:
metanotum
longitudinal
keels on
reaching only posterior
apex of propodeum, without pterostigma; mandibles tridentate, with a rudimentary tooth between the two posterior teeth; fore tarsal segments following proportions: 46: produced
6:
into a short hook;
10:
35:
42;
segment 3 of front
enlarged claw
(Fig.
404)
with
teeth and one bristle; segment 5 of front tarsus (Fig. 404) bristles and one preapical lamella; tibial spurs 1,1,1. MALE:
in
tarsus a row of 5 with
two
unknown
lOCUS TYPICUS:
lido (Venezia,
TYP ICAl MATER IAl: HOSTS:
11:
9;
granulated;
two
not complete,
8;
posteriorly separated;
dull,
with
=
holotype F!
Italy) in CO.
unknown
OISTRIBUTION:
Palaearctic region:
ITALY:
Lido
(Venezia),
CO!
2. Bocchus umber n. sp.
FEMALE
(Figs 24,401):
testaceous; with
tergite
hairy;
winged;
almost hairless;
occipital
distinct and complete,
length 2,50-2,62 mm;
thorax and propodeum black;
1 and apex brownish;
abdomen
granulated;
fully
antennae yellow;
tegulae yellow;
head,
carina complete;
pronotum, POL = 4;
posteriorly separated;
head reddishabdomen black,
legs yellow;
body dull,
scutum and scutellum OOl = 5;
notaulices
metanotum smooth,
without
Bocch i nae
615
Ii I
I
) 1,\ I \
\
\
Fig. 404 -
sculpture;
Chela of Bocchus italicus n.
scutellum nearly
reticulate rugose,
with 2
twice as
longitudinal
sp.
long as metanotum;
(type)
propodeum
keels on posterior surface;
fore-wing
616
Mem.
Amer.
Ent.
Inst.,
no.
37,1984
,, " \"1;: - - - - - >
f
,
Fig.
I~ :.-~
,
405 -
Female of Bacchus umber n.
claw of a specimen from Sesto D:
without dark teeth;
apophysis of front
transversal
mandibles tridentate, fore tarsal
(except for distal
bands;
(type):
tarsal
A: C:
chela;
tarsus with a distal 6,1
enlarged segments;
ovoidal,
large;
tooth between the two posterior apophysis
segment 1-4 in following proportions:
apophysis):
B:
fore tarsal
segment 3.
pterostigma brown,
with a rudimentary
segment 3 of front
segment;
sp.
Fiorentino (Italy);
longer than 8:
; segment 5 of front tarsus
1,5:
1,2
(Fig.
405)
Bocchinae
with one preapical
lamella and 2 bristles;
row of 7-8 teeth and 2 bristles; MALE:
617
tibial
enlarged claw
(Fig.
405)
with a
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Bivio Castel
TYPICAL MATERIAL:
Rigone
holotype F!
(I taly)
and 1 paratype F!
in OL;
1 para type F!
in MI. HOSTS:
unknown
DISTRIBUTION: OL!
Palaearctic region:
Sesto Fiorentino (Firenze),
ITALY:
Bivio Castel
Rigone
(Perugia),
M I!
3. Bocchus szelenyii Moczar
Bocchus szelenyii Moczar 1974: 131. Bocchus szelenyii Moczar: Moczar 1983b: 206.
FEMALE:
fully
winged;
Fig.
406 -
length 3,12 mm;
head black,
with mandibles yellow;
Chela of Bocchus szelenyii Moczar (type)
618
Mem.
antennae testaceous;
black;
abdomen black; OOL = 7;
visible,
pronotum
Inst.,
scutellum shiny,
with one transversal
mandibles quadridentate, posterior teeth; 10;
punctate;
fore tarsal
enlarged claw
segment 5 of front
with some
hairy;
POL
notaulices weak,
metanotum dull,
pterostigma short and narrow;
rudimentary
406)
with a
(Fig.
406)
but
reticulate
tooth between
the two
segments 1-4 in following proportions:
(Fig.
tarsus
swollen,
carina complete;
propodeum reticulate rugose;
dark band;
with a
scutellum
petiole reddish;
near posterior margin of scutum;
not much shorter than scutellum;
fore-wing
hairy,
occipital
weakly punctate,
weakly
1984
reddish-testaceousj
head shiny,
obsolete only
smooth,
37,
propodeum black;
without sculpture;
scutum shiny,
no.
and scutum
legs testaceous;
almost complete,
rugose,
3:
En!.
metanotum reddish-testaceous;
short keels on frons, 5;
Amer.
14:
2:
row of 5 teeth and 1 long bristle;
with one preapical
lamella and 2
long
bristles; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Spring "Talyn Bi Igech" TYPICAL MATERIAL:
holotype F!
(Mongol ia)
in BU.
HOSTS: unknown DISTRIBUTION: Palaearctic region: Km E guard-house "Caganbu lag",
MONGOLIA:
Spring
"Talyn Bilgech"
Bajanchongor a imak),
(47
BU!
4. Bocchus slovacus Strejcek
Bocchus slovacus Strejcek 1964: 325. Bocchus slovacus Strejcek: Nagy 1969: 322. Bocchus slovacus Strejcek: Ponomarenko 1978: 23.
FEMALE:
fully
winged;
antennae testaceous;
length 2,48-2,75 mm;
thorax and
brown-reddish nuances; head dull, than
hairy,
twice as
long as POL;
tooth between
swollen;
twice as
rarely pronotum with
abdomen black;
occipital
scutum dull,
posteriorly separated;
long as metanotum;
narrow;
fore
407)
with a
tarsus
(Fig.
row of 6 teeth and 407)
with one apical
bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,20-2,67 mm; testaceous;
tarsi
thickened,
hairy;
9:
9:
9:
8:
7:
and fore tibiae
10;
granulated,
antennal
black;
testaceous;
segments
head dull,
hairy;
granulated,
pterost i gma short
segments 1-4 in following proportions:
(Fig.
segment 5 of front
OOL more
rudimentary
metanotum and propodeum reticulate
fore-w i ngs wi th one transversa I dark band;
enlarged claw
with a
scutellum dull,
rugose;
tarsal
legs testaceous;
carina complete;
mandibles quadridentate,
the two posterior teeth;
notaulices complete, nearly
petiole testaceous;
granulated,
head reddish-testaceous;
propodeum black;
13:
3:
and
9;
1 long bristle; lamella and 2
long
mandibles brown-
antennae not distally
in following proportions:
granulated;
2:
12:
6:
8:
frons with a median complete
9:
619
Bocchinae
Fig.
keel;
=
8;
407 -
Che I a of
occipital carina complete; scutum dull,
granulated;
Bocchus slovacus Strejcek {type}
POL
=
6;
OL
=
3,5;
notaul ices complete,
OOL
=
7;
OPL
=
4;
TL
posteriorly separated;
scutellum and metanotum dull, granulated; propodeum reticulate rugose, with two longitudinal keels on posterior surface; areolae of propodeum partly broader than hyaline,
tegulae;
without dark
lanceolate;
radial
LOCUS TYPICUS:
vein curved;
Cenkov
TYPICAL MATERIAL:
abdomen
transversal
with petiole very short;
bands;
radial
pterostigma
cell
open;
long
tibial
and
spurs
fore wings narrow,
not
1,1,2.
{Czechoslovakia}
holotype F!
in PG;
3 paratypes FF
in Strejcek's
collection. HOSTS: unknown DISTRIBUTION:
Palaearctic region:
U. S. S. R.: Papanga {Russian
Romania}, PK! HUNGARY: Keleshalom, BU! CZECHOSLOVAKIA: Cenkov, PG!
BU!
Topart
{Kelebia},
BU!
Budapest,
620
Mem.
Fig.
408 -
Amer.
Chela of
En!.
Inst.,
no.
37,
1984
Bocchus paglianoi n. sp. (type)
5. Bocchus paglianoi n. sp.
FEMALE:
fully
winged;
antennae brown, black;
length 3,75 mm;
head black,
with segments 1-3 testaceous;
petiole testaceous;
abdomen black;
with mandibles yellow;
thorax and propodeum
legs testaceous,
with mid and
Bocchinae
hind coxae partly occipital
brown;
head swollen,
carina complete;
tooth between
hairy,
shiny,
mandibles quadridentate,
the two posterior teeth;
notaulices complete,
621
scutum dull,
posteriorly separated;
with posterior third rugose;
segments 1-4 in following proportions:
claw
(Fig.
408)
front
tarsus
tibial
spurs 1,1,1.
MALE:
unknown
LOCUS TYPICUS:
15:
3: 10; enlarged
2:
teeth and one long bristle;
with one preapical
Montemitro
TYPICAL MATERIAL: HOSTS:
row of 4
408)
granulated;
pterost i gma short and narrow;
fore tarsal
(Fig.
hairy,
scutellum shiny and smooth,
metanotum and propodeum reticulate rugose;
fore-wi ng with 2 transversa I dark bands; with a
granulated;
with a rudimentary
lamella and 2
segment 5 of
long bristles;
(Italy)
holotype F!
in OL.
unknown
DISTRIBUTION: NOTES:
Palaearctic region:
the species
is named
in
ITALY:
Montemitro (Campobasso),
honor of Mr.
OL!
Guido Pagliano.
6. Bocchus scaramozzinoi n. sp.
FEMALE:
fully
testaceous,
winged;
length 3,6-4,2 mm;
anterior part of frons,
head black,
with mandibles
temples and region between ocelli
and
occipital margin reddish; ventral part of head partly black and reddish; sometimes head fully reddish, except for ocellar triangle black; antennae brown,
with segments 1-2 and partly 3 testaceous;
scutum and scutellum reddish; mesopleura, tegulae basal
propectus,
testaceous;
segments
sometimes
metapleura and posterior surface of propodeum reddish; petiole testaceous;
abdomen black;
legs testaceous,
part of hind coxae and clubs of hind femora brown;
shiny,
pronotum,
metanotum and propodeum black;
in following proportions:
almost
hairless,
2 keels around orbits;
15:
8:
with a median
14:
12:
11: 9: 9: 9: 9:
longitudinal
keel
with
antennal 12;
head
on frons and with
head with numerous irregular keels among which
the surface is granulated;
only a central
region of frons
is smooth and
without areolae, keels or sculpture; occipital carina complete; ocellar triangle convex, nearly equilateral; OOL = 12; OPL = 9; TL = 12; mandibles quadridentate, shiny, keels;
with a strong pronotal
with one rudimentary
transversal
impression
tubercles reaching
tegulae;
numerous areolae or irregular keels; separated; rugose; surface;
scutellum with numerous
propodeum reticulate rugose,
pronotum hairy,
scutum shiny,
notaul ices complete, longitudinal with 2
keels;
one on basal
long and narrow,
20:
3:
4:
prox i ma I 13:
19;
with
posterior keels on posterior fore-wing
with
with 2 dark transversal
cells and another beneath the pterostigma;
part of rad i a I ve in shorter than following proportions:
irregular
hairy,
metanotum reticulate
longitudinal
median area of posterior surface almost smooth;
pterostigma very reduced, bands,
tooth;
and with numerous
(5: 7);
distal
fore tarsa I segmen ts
enlarged claw
(Fig.
409)
in
with a
622
Mem.
Fig.
Amer.
En!.
Ins!.,
no.
37,
1984
Chela of Bocchus scaramo2zinoi n.
409 -
row of 6 teeth and 2 bristles; bristles and one preapical
segment 5 of front
lamella;
tibial
tarsus
(Fig.
409)
with 2
in OL;
1 paratype F!
unknown
DISTRIBUTION:
Palaearctic region:
Sardinia), OL!
Musei
NOTES:
(type)
spurs 1,1,1.
MALE: unknown LOCUS TYPICUS: Tempio Pausania (Italy) TYPICAL MATERIAL: holotype F! and 1 paratype F! in PL. HOSTS:
sp.
the species
(Cagliari),
is named
ITALY: OL!
Tempio Pausania
(Sassari,
PL!
in honor of Mr.
Pier Luigi Scaramozzino.
7. Bocchus scobiolae Nagy
Bocchus scobiolae Nagy 1967b: 334. Bocchus scobiolae Nagy 1968: 1029.
Bocchinae
Fig. 410 -
623
Chela of Bocchus europaeus Bernard from Saint-Paul-et-Valmalle (France)
624
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
Bocchus scobiolae Nagy 1969: 322, 323.
I
haven't seen the three only known specimens of this species
females and 1 male).
I have included
Palaearctic Bacchus only on
the basis of the original
typicus: Agigea (Dobroudja, Romania).
(2
the species in the key to the
Locus
description.
I don't know where is the holotype.
8. Bacchus europaeus (Bernard)
Eukoebeleia europaea Bernard 1939: 109. Bacchus europaeus (Bernard): Strejcek 1964: 325. Bacchus europaeus (Bernard): Nagy 1969: 322. Bacchus europaea (Bernard): Abdul-Nour 1976: 277.
FEMALE: clypeus
fully
winged;
testaceous;
reddish;
pronotum and scutum reddish;
propodeum black; legs yellow; 7:
5:
6:
Bernard
5:
abdomen black,
antenna I segment
1939);
head smooth, along
sides of median keel 2 or 3 teeth;
rugose; rugose,
scutellum,
weakly brown; 1 without
hairy,
internal
weakly
occipital
metanotum and tegulae yellow-brown;
proportions: teeth
10: 5,5:
(with teeth,
granulated,
margin of eyes,
of frons;
with mandibles and
with segments 1-4 yellow-
10:
OOl more than
reticulate rugose;
carina,
long as POL;
longitudinal
keels;
on
mandibles with 1,
notaul ices distinct and complete,
scutellum with numerous
7: to
with some areolae
near occipital
carina complete;
twice as
7:
according
tridentate mandibles with a rudimentary tooth between
two posterior teeth; separated;
head black,
antenna I segments in following 7;
near clypeus,
shiny,
length 3,25 mm;
antennae reddish-brown,
the
scutum hairy, posteriorly
metanotum reticulate
scutellum nearly twice as long as metanotum; propodeum reticulate with 2 longitudinal keels on posterior surface; fore-wing with 2
dark transversa I bands;
pterost i gma short and narrow;
fore tarsa I
segments 1-4 in following proportions: 14: 2: 3: 10; segment 5 of front tarsus (Figs 410, 411) with a preapical lamella and 2 bristles; enlarged claw 1,1,1. MALE:
(Figs 410,
411)
with a row of 4-5
teeth and 1-2 bristles;
tibial
unknown
lOCUS TYPICUS:
St-Aygulf
TYPICAL MATERIAL:
(France)
holotype F!
in P.
HOSTS: in France Caliscelis sp. (Abdul-No'Jr 1976). DISTRIBUTION: Palaearctic region: FRANCE: St-Aygulf Paul-et-Valmalle (Herault),
(Var),
P!
9. Bacchus parthicus Ponomarenko
P!
Saint-
spurs
Bocchinae
B
625
A / /
/
/
/
Fig.
411
-
Bacchus eurapaeus Bernard (type): A: chela; B: fare tarsal segments.
Bacchus parthicus Ponomarenko 1979b: 166.
FEMALE:
fully
winged;
length 3,60 mm;
fully
reddish,
triangle brown,
posterior margin of scutellum brown,
abdomen brown;
antennae brown,
with occipital propodeum darkened,
with segments 1-2 yellow;
antennae
626
Mem.
Fig.
412 -
Amer.
Ent.
Ins!.,
no.
37,
1984
Chela of Bocchus parthicus Ponomarenko
(type)
weakly distally thickened; antenna I segments in following proportions: 10: 20: 15: 15: 15: 14: 13: 13: 15; mandibles quadridentate, with a rudimentary
tooth between
the two posterior teeth;
head shiny,
20:
punctate;
Bocch i nae
frons with a occipital
median complete keel
carina complete;
pronotum shiny, reaching
hairy,
tegulae;
and
= 7;
POL
smooth,
627
with 2 keels around orbits;
= 7;
OL
=
OOL
weakly punctate;
pronotum crossed by
13;
a transversal
=
OPL
pronotal
16;
TL
=
18;
tubercles
impression;
scutum
shiny, hairy, weakly punctate, with notaulices complete and posteriorly separated; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose, strongly one on long
the basal
and
tarsal
(Fig.
1,1,2. MALE:
with 2 brown
fore-wing
cells and another beneath
narrow,
segments
front tarsus claw
with posterior surface not reticulate rugose,
transversely striate; not
lanceolate;
radial
vein
in following proportions:
(Fig.
412)
412)
with a
regularly curvilinear;
lamella and 2 bristles;
with a row of 6 teeth and
bands,
pterostigma fore
5: 8: 25: 34; segment 5 of
33:
preapical
transversal
the pterostigma;
1 long bristle;
tibial
enlarged spurs
unknown
LOCUS TYPICUS:
Tashauz
TYPICAL MATERIAL: HOSTS:
(Turkmenistan,
holotype F!
U.
S.
S.
S.
R.:
R.)
in LN.
unknown
DISTRIBUTION:
Palaearctic region:
U.
S.
Tashauz
(Turkmenistan),
LN!
ETHIOPIAN SPECIES OF
BaCCHUS
Three species are known
KEY TO THE SPECIES
FEMALES
Micropterous;
fore-wing
nearly
as
long as
• • . . • . . . . . . . . . . • • • • . • • . . . • . • . . • . .. -
Fully
2
Segment
1 of
414 B);
antennae
least -
tegulae . . . . . . . . . . . . . . .
richardsi n. sp.
winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
front
twice as
Segment
of
antennae nearly
1.
as
tarsus nearly
longer and more
long
front
shorter,
1,5
times as
slender,
as broad ••.•.• tarsus nearly as
2.
MALES
3.
4
segments
(Fig. 5-9 at
bini n. sp.
long as
distally more broadened,
long as broad •••••••••••
long as
wi th 4
(Fig.
with
415 A);
segments
seyrigi (Benoi t)
5-9
628
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
Unknown
1. Bocchus richardsi n. sp.
FEMALE: 3,31
15: 6:
9:
areolae, lateral 10;
micropterous,
mm;
with
fore-wing nearly
fully reddish-dark; 6:
6:
6:
with a
5:
5:
median
5:
8;
keel
keels around orbits;
occipital
antennal
as
segments
head shiny,
long as tegulae;
length
in following proportions:
granulated and with
weak
joining anterior ocellus to clypeus and with 2 POL = 5;
carina complete;
OL = 3;
pronotum shiny,
OOL = 12; hairy,
OPL = 8;
weakly
TL =
granulated,
with a transversal impression between anterior and posterior collar; pronotal tubercles reaching tegulae; scutum dull, granulated, with weak areolae and strong
longitudinal
with
longitudinal
keels;
fully
reticulate rugose;
keels;
notaulices invisible;
metanotum very short and reduced; posterior surface without
scutellum dull, propodeum
longitudinal
keels;
fore
tarsal segments in following proportions: 15: 3: 3: 11: 14; enlarged claw (Fig. 413) with a row of 9 teeth and one bristle; segment 5 of front tarsus
(Fig.
413)
with a
preapical
lamella and a
row of 8 bristles;
spurs 1,1,1.
(\
Fig.
413 -
Chela of
Bocchus richardsi n. sp. (type)
tibial
Bocchinae
MALE:
unknown
LOCUS TYPICUS:
Milnerton
TYPICAL MATERIAL: HOSTS:
(South Africa)
holotype F!
in BM;
1 paratype F!
in WA.
unknown
DISTRIBUTlmJ: BM!
629
Ethiopian region:
Cape Town,
NOTES:
SOUTH AFRICA:
Milnerton
(Cape Town),
WAr
the species
is named
in
honor of Mr.
Owain W.
Richards.
2. Bocchus bini n. sp.
FEMALE: black
fully
winged;
length 3,12-4,06 mm;
head fully reddish or fully
(with mandibles and sometimes also clypeus testaceous);
occasionally
head black with region between ocell i and occipital carina reddish; thorax fully reddish; propodeum fully reddish or fully black; occasionally partly black
and reddish;
antennae fully
testaceous or brown,
legs testaceous;
reticulate rugose;
sometimes not reticulate rugose on frons on sides of
median keel,
where the surface is granulated;
ocellar triangle equilateral; rugose,
abdomen black;
with segments 1-2
or 1-3 testaceous;
hairy,
shiny;
pronotum hairy;
notaulices
head du II,
occipital
ha i ry,
fu II y
carina complete;
scutum fully reticulate
invisible or slightly visible;
scutellum
and metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; fore tarsal segments 1-4 in following proportions: preapical
13: 2:
2:
8;
5-6 teeth and one bristle; MALE:
segment 5 of front
lamella and 2 bristles; tibial
tarsus
enlarged claw
(Fig.
(Fig.
414)
414)
with one
with a row of
spurs 1,1,1.
unknpwn
LOCUS TYP ICUS:
Afgoi
TYPICAL MATERIAL: in BM;
(Somal ia)
holotype F!
1 para type F!
and 3 para types FF!
in OL;
1 paratype F!
in OT.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
SOMALIA:
Afgoi
(Benadir),
OL!
Ibadan, OT! Ikenne (\'I-State), BM! GHANA: Kwadaso (Kumasi), NOTES: the species is named in honor of Mr. Ferdinando Bin.
NIGERIA: OL!
3. Bocchus seyrigi (Benoit) n. comb.
Hymenodryinus seyrigi Benoi t 1953b: 427.
FEMALE:
fully
antennal
segments 3-10 brown,
winged;
reticulate rugose,
length 4,62 mm;
hairy;
reddish,
with ocellar spot brown,
abdomen black-brown;
antennae short,
head dull,
distally broadened,
fully
with
630
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
, >
'
\ !
,
'
~
'
,
'0\ \\ \
\
B
"
\
\
Fig.
414 -
Bocchus bini n. sp. B:
segments 5-9 nearly proportions:
18:
7:
as 15:
(paratype from
fore tarsal
long as broad; 7:
5,5:
5:
6:
6:
Ibadan,
Nigeria);
A:
chela;
segments.
antenna I segments 5:
7;
occipital
in following
carina complete;
Bocchinae
Fig.
415 -
Bocchus seyrigi
(Benoit) (type);
631
A:
fore tarsal
segments;
B:
chela.
ocellar triangle equilateral; fully
reticulate rugose;
with 2
longitudinal
scutum,
scutellum and metanotum dull,
notaulices invisible;
hairy,
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
with 2 dark
632
Mem.
transversa I bands;
Amer.
12: 2:
LOCUS TYPICUS:
3:
12;
37,
Behara
1984
fore tarsa I segments 1-4
segment 5 of front
teeth and one bristle;
TYPICAL MATERIAL: HOSTS:
no.
lamella and 2 bristles;
with one preapical
with a row of 11 MALE: unknown
Inst.,
pterost i gma short and narrow;
in following proportions: 415)
En!.
tibial
tarsus
enlarged claw
(Fig.
(Fig.
415)
spurs 1,1,1.
(Madagascar)
holotype F!
1 paratype F in TV.
in P;
unknown
DISTRIBUTION: 1953b) •
Ethiopian region:
MADAGASCAR:
Behara,
P!
3ekily
(Benoit
OR I ENTAL SPEC I ES OF BOCC/JUS
Four species are known.
KEY TO THE SPECIES
FEMALES
-
Fore-wing with Fore-wing with
2 Fore-wing of A);
front
with
1 dark
transversal transversal
band •••••
2 dark
radial
vein
curved
tarsus without
enlarged
weakly
a distinct
claw with a
row of
thorax and propodeum black
some
(except
(Fig.
proximal teeth
for
Fore-wing with segment
5 of
radial
front
(Figs 419,422); 422);
head,
vein more
tarsus
enlarged
wi th
strongly a
416 A);
prominence (Fig.
2.
418);
(Fig.
di st inct proximal
claw with
segment (Fig.
5
416
head,
muluensis n.
curved
1 only
thorax and propodeum reddish
tooth
sp.
416 B);
prominence
(Figs
419,
or brown-testaceous
••..••.••••••••••.•••...••••....•.....•........•..........•..•• 3 much longer than POL; head, thorax and propodeum redddish-brown •• •••••••••••••••••......•..• 3. pedunculatus Nagy OL more than twi ce as long as POL; head, thorax and propodeum reddi sh-I ight .••.•••••••.••••••••.•••..•••• 4. rubricus (Fouts)
3 OL not -
laotianus n. sp.
the yellow mandibles) .•••
. . . • • . • • . . . . • . . . • • • • • • • . • . • . . • • • • . . . . . . . . .. -
1.
bands • • • . . • . . . . . . . . . . . • . . . . . • 2
fAALES
Only
the male of B.
pedunculatus Nagy is known.
Bacchinae
633
A
,
,
E
Radial veins of females of Bacchus muluensis n. sp. (type)(A), pedunculatus Nagy from Tagembung (Phi I ippines) (B), laatianus n. sp. (type)(C), guineensis n. sp. (type)(D), australiae n. sp. (type)(E).
Fig. 416 -
634
Mem.
Amer.
Ent.
lnst.,
no.
37,
1984
i
i~
() /
/J 1/ I /
Fig. 417 -
Chela
0
f Bocchus laotianus n.
sp.
( type)
i
Bocchinae
635
1. Bocchus laotian us n. sp.
FEMALE:
fully
winged;
length 3,43-3,87 mm;
with segments 1-2 or 1-3 yellow; legs yellow;
head dull,
hairy,
thorax,
weakly granulated,
except for the sides of the ocelli rugose;
occipital
tooth between
equilateral;
scutum shiny,
granulated,
scutellum shiny,
punctate,
twice as
with
1 dark
transversal
posteriorly separated;
reticulate rugose only near posterior margin,
long as metanotum; with 2
ocellar triangle
reticulate rugose near
notaulices distinct and complete,
nearly
with a
mandibles quadridentate,
anterior margin;
reticulate rugose,
reticulate rugose,
the two posterior teeth; hairy,
antennae brown,
where the surface is not reticulate
carina complete;
rudimentary
head black;
propodeum and abdomen black;
metanotum reticulate rugose;
longitudinal band;
propodeum
keels on posterior surface;
pterostigma short and narrow;
fore-wing
radial
vein
weakly curved (Fig. 416 C); fore tarsal segments 1-4 in following proportions: 20: 3: 3: 14; segment 5 of front tarsus (Fig. 417) with a preapical
lamella and 2 bristles;
10 teeth and MALE:
1 bristle;
enlarged claw
(Fig.
417)
with a
row of
spurs 1,1,1.
unknown
LOCUS TYPICUS:
Ban Van Eue
TYPICAL MATERIAL: HOSTS:
tibial
(Laos)
holotype F!
1 para type F!
in B;
in OL.
unknown
DISTRIBUTION:
Oriental
region:
LAOS:
Ban Van Eue
(Vientiane),
B!
OL!
2. Hocchus muluensis n. sp.
FEMALE:
fully
winged;
segments 9-10 brown; legs testaceous,
length 3,25 mm;
head black;
thorax and propodeum black;
with hind coxae brown;
head dull,
antennae yellow, abdomen hairy,
with
black-brown; reticulate
rugose; frons narrow, very converging towards the clypeus; in front of the anterior ocellus the frons is nearly as broad as each eye;ocellar triangle equilateral;
occipital carina complete;
mandibles with 4 teeth,
larger from
anterior one to posterior;
scutum reticulate rugose,
and with numerous metanotum hairy, 2
longitudinal
transversal short and 3:
12;
bristle;
longitudinal
dull,
keels;
notaul ices invisible;
weakly punctate;
keels on posterior surface;
bands; narrow;
radial
vein
fore tarsal
segment 5 of front enlarged claw
propodeum reticulate rugose, fore-wing
weakly curved
(Fig.
(Fig.
418)
418)
with a
with a
Gunong Mulu National
TYPICAL MATERIAL:
holotype F!
and
416 A); preapical
row of 7 teeth and
Park
(Sarawak,
1 paratype F!
hairy with
with 2 dark pterostigma
1 long
Malaysia)
in 8M.
14:
lamella and
bristle; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
dull,
scutellum and
segments 1-4 in following proportions:
tarsus
(Fig.
progressing
3: 1
636
Mem.
Fig.
HOSTS:
418 -
Amer.
Ent.
Inst.,
no.
37,
1984
Chela of Bocchus muluensis n.
sp.
(type)
unknown
DISTRIBUTION: District,
Oriental
Sarawak),
region:
MALAYSIA:
Gunong Mulu Nat.
Park
(Baram
BM!
3. Bocchus pedunculatus Nagy
Bocchus pedunculatus Nagy 1969: 323. Bocchus rubricus Fouts 1922: Nagy 1969: 324.
FEMALE: fully winged; length 2,18-3,20 mm; reddish-brown, with antennae yellow (with segments 9-10 or only 10 brown); head shiny, smooth, weakly punctate,
sometimes with
complete;
mandibles quadridentate,
two posterior teeth; weakly
punctate,
and distinct,
transversal
OL not much
keels on frons;
longer than
POL;
carina
tooth between
scutum shiny,
without sculpture among punctures;
posteriorly separated;
occipital
with a rudimentary
the
smooth,
notaulices complete
scutellum and metanotum shiny,
smooth, weakly punctate; propodeum reticulate rugose, with 2 lateral points and with 2 longitudinal keels on posterior surface; fore-wing with 2
Bocchinae
Fig.
419 -
637
Chela of Bocchus pedunculatus Nagy from Gunong Mulu (Sarawak)
dark transversal strongly curved proportions:
bands; (Fig.
13: 3:
3:
pterostigma short and narrow;
416 B);
fore tarsal
radial
vein
segments 1-4 in following
11; segment 5 of front tarsus (Fig. 419) with a
638
Mem. Amer.
Fig.
420 -
Ent.
Inst.,
no.
37,
1984
Male of Bocchus pedunculatus Nagy from Gunong Mulu
(Sarawak): A:
ninth abdominal sternite;
B:
genitalia.
preapical lamella, 2 bristles and a proximal distinct prominence; enlarged claw (Fig. 419) with 1 tooth and 1 long bristle; tibial spurs 1,1,2. MALE (Figs 402, 403): fully winged; length 2,60-3,00 mm; head black, with mandibles and sometimes clypeus testaceous; antennae yellow, with segments 6-10 brown; thorax, propodeum and petiole black or brpwn;
legs
testaceous, with coxae blackish; head hairy, dull, fully reticulate rugose; occipital carina complete; ocellar triangle equilateral OPL = OOL; mandibles quadridentate, wite. a rudimentary tooth between the two posterior teeth; scutum dull, hairy, fully reticulate rugose; notaulices distinct and complete, posteriorly separated; scutellum and metanoturn dull, weakly punctate, hairy; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface and with 2 lateral points (as in female); forewing with 2 transversal darkened bands; radial vein strongly curved; pterostigma short and narrow, lanceolate, 420; abdomen with petiole very long. LOCUS TYPICUS: Pinigisan (Philippines) TYPICAL MATERIAL: holotype M! in CO. HOSTS: unknown
hyaline;
male genitalia:
fig.
DISTRIBUTION: Oriental region: MALAYSIA: Gunong Mulu (Baram District, Sarawak), OU BM! PHILIPPINES: Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan)(Nagy 1969: under name Bocchus rubricus Fouts), CO!
Bocchinae
Fig. 421
-
Female of
639
Bocchus rubricus (Fouts)(type)
4. Bocchus rubricus (Fouts)
Neoanteon rubrica Fouts 1922: 634. Neoanteon rubrica Fouts: Baltazar 1966: 205. nec Bocchus rubricus (Fouts): Nagy 1969: 324 (see Bocchus pedunculatus Nagy).
FEMALE
(Fig. 421):
fully winged;
length 3,0-4,0 mm;
reddish,
with
antenna I segments 1, 2, 3, 7, 8 and part of 9 yellow; head almost hairless, shiny, smooth, without sculpture, weakly punctate; OL more than twice as long as POL; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; scutum weakly punctate, almost hairless, smooth, shiny; notaulices weak, incomplete, not visible in posterior third of scutum; scutellum and metanotum smooth, shiny,
weakly punctate;
propodeum reticulate rugose,
with 2 longitudinal
640
Fig.
Mem.
422 -
Amer.
Ent.
Inst.,
Female of Hocchus rubricus tarsal
no.
(Fouts)(type):
strongly curved; 3:
12;
bands;
fore tarsal
segment 5 of front
Mt.
points on sides;
tibial
Makiling
(Fig.
422)
prominence;
with a preapical
unknown
fore
fore-wing radial
enlarged claw
lamella, (Fig.
(Philippines)(Los Banos according in WA.
vein
15:
422)
spurs 1,1,1.
original description; Mt. Makiling on type label). TYPICAL MATERIAL: holotype F! and 1 paratype F! HOSTS:
B:
segments 1-4 in following proportions:
bristles and a distinct proximal
LOCUS TYPICUS:
chela;
pterostigma short and narrow;
tarsus
1 tooth and 1 long bristle; MALE: unknown
A:
segments.
keels on posterior surface and with 2 lateral with 2 dark transversal
37,1984
to the
4: 2 with
641
Bocchinae
DISTRIBUTION: Los Banos
Oriental
(Luzon),
region:
PHILIPPINES:
Mt.
Makiling
(Luzon),
WA!
WA!
NEARCTIC SPECIES OF
BaCCHUS
Eight species are known
KEY TO THE SPECIES
FEMALES
Species micropterous -
Species
fully
(Figs
424,
425) .•••••••••••••••••••••••••• 2
winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
2 Fore-wing less reduced, reaching approximately the keel between anterior and posterior surface of propodeum; notaul ices complete, sometimes slightly visible •••••••••••••••••••••••••••• 1. mirabilis (R.C.L. Perkins - Fore-wing more reduced, reaching approximately anterior margin of propodeum (Figs 424,425); notaulices invisible . . . . . . . . . . . . . . . • • . . . • . • • • . . . . . . . • . . . . • . .. 2. testaceus n. sp. 3 Species fully testaceous . . . 3. wasbaueri n. sp. - Species at least partly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 4 Thorax partly black . . . . . . . . 6. flavicollis Ashmead - Thorax fully black ••••••••••••••••••••••••••••••••••••••••••••• 5 5 Scutum dull, granulated . . . . 4. arizonicus (R.C.L. Perkins) - Scutum shiny, weakly punctate, wi thout sculpture among punctures . . . • . . . . . . . . . . . . . . . . . . • . . . . 5. flavipes Kieffer
The females of B.
laticeps (Ashmead) and dubius n. sp. are unknown
MALES
Scutum dull, -
fully granulaterJ;
head granulated,
rugose . . . . . . . . . . . . . . . . . . . . . 7. laticeps (Ashmead) Scutum shiny, without sculpture; head reticulate g ranu I at ed. • • • • • • • • • • • • • • ••
The males of
8.
not
reticulate
rugose and
dubius n. sp.
B. mirabilis (R.C.L. Perkins), testaceus n. sp.,
642
Fig.
Mem. Amer.
423 -
Ent.
Inst.,
no.
37,
Chela of Bocchus mirabilis (R.C.L. (Ohio)
1984
Perkins) from Columbus
wasbaueri n. sp. flavicollis Ashmead, arizonicus (R.C.L. Perkins) and flavipes Kieffer are unknown. Probab I y B. laticeps (Ashmead) and dub ius n. sp.
are the opposite sex of the other six species.
Bocchinae
643
1. Bocchus mirabilis (R.C.L. Perkins)
Eukoebeleia mirabilis R.C.L. Perkins 1905: 59. Eukoebeleia mirabilis R.C.L. Perkins: Kieffer 1914b: 47. Phorbas mirabilis (R.C.L. Perkins): Fenton 1918b: 261. Bocchus mirabilis (R.C.L. Perkins): Richards 1939: 189. Bocchus mirabilis (R.C.L. Perkins): Nagy 1969: 322.
FEMALE:
micropterous,
with fore-wing reduced and extending beyond the
keel
between anterior and posterior surface of propodeum;
mm;
usuallyfully testaceous with antenna I segments 4-10 black or brown;
occasionally head reddish fully black;
with brown ocellar spot;
occasionally antennae brown,
pronotum and scutum usually reddish, scutellum,
head dull,
rudimentary tooth between complete;
rarely head almost
with segments 1-2 or 1-3 yellow;
rarely
metanotum and propodeum black;
tegulae testaceous;
granulated;
almost fully black;
abdomen brown-black;
the two posterior teeth;
notaulices distinct and complete,
scutellum dull,
as sculptured as scutum;
scutellum nearly twice as
legs and
mandibles quadridentate, occipital
OOL nearly three-four times as long as POL;
granulated;
length 2,50-2,92
with a
carina
scutum dull,
posteriorly separated;
metanotum reticulate rugose;
long as metanotum;
propodeum reticulate rugose,
with 2 longitudinal keels on posterior surface; fore reduced wings without pterostigma; fore tarsal segments 1-4 in following proportions: 23: 4: 6: 17; segment 5 of front tarsus (Fig. 423) with a preapical lamella and 2 bristles; enlarged claw (Fig. 423) with a row of 6-7 teeth and 1 bristle; tibial
spurs 1,1,1.
MALE:
unknown
LOCUS TYPICUS:
Ohio
(U.
S. A.)
TYPICAL MATERIAL: lectotype F! and 1 para lectotype F! in B. HOSTS: in U. S. A. Bruchomorpha oculata Newm. (Issidae) (Fenton 1918b). DISTRIBUTION: Place (Ont.),
Nearctic region: Ol!
OT!
Ont.),
OT!
U. S. A.:
Refugio
Tex.),
BY!
Ol!
B!
Ohio,
CANADA:
Grenadier
I.
(Bee Co.,
Columbus
Foremost
Centre (St. Tex.),
(Ohio),
(Alta.),
Lawrence BY!
Waco
OT! Is.
Carleton Nat.
Park,
(Mc lennan Co.,
BM!
2. Bocchus testaceus n. sp.
FEMALE
(Figs 424,
425):
micropterous,
only anterior margin of propodeum;
with fore-wing reduced and reaching
length 2,01
mm;
fully
testaceous;
head
dull, granulated; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teth; OOL nearly four times as long as POL;
scutum shiny,
scutellum shiny,
without sculpture,
weakly granulated;
notaulices invisible;
nearly twice as long as metanotum;
644
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
I 424 - Female of Bocchus testaceus n. sp.
Fig.
metanotum reticulate rugose;
propodeum reticulate rugose,
longitudinal
keels on posterior surface;
pterostigma;
fore tarsal
enlarged claw
with 2
fore reduced wings without
segments 1-4 in following proportions:
6; segment 5 of front tarsus (Fig. 426) with a preapical bristles;
(type)
(Fig.
426)
11:
1,5: 2:
lamella and 2
with a row of 8 teeth and 2 bristles;
tibial spurs 1,1,1. MALE: unknown LOCUS TYPICUS:
Nogales (U.
TYPICAL MATERIAL: HOSTS:
S.
holotype F!
A.) in OL.
unknown
DISTRIBUTION:
Nearctic region:
U. S. A.: Nogales (Arizona), OL!
3. Bocchus wasbaueri n. sp.
FEMALE:
fully
winged;
length 3,12 mm;
fully
testaceous,
ocellar spot and antenna I segments 4-10 darkened; occipital
carina complete;
tooth between
mandibles quadridentate,
the two posterior teeth;
with brown
head dull,
granulated;
with a rudimentary
OOL nearly five times as long as
Bocchinae
Fig.
POL;
425 -
scutum dull,
separated,
645
Female of Bocchus testaceus n.
granulated;
scutellum dull,
transversa I band;
2:
3:
8;
fore-wing
tibial
with a
dark
fore tarsa I segments 1-4
segment 5 of front
lamella and 1 bristle;
row of 7 teeth and 1 bristle; MALE:
propodeum reticulate rugose,
pterost i gma short and narrow; 14:
posteriorly
metanotum reticulate rugose;
keels on posterior surface;
in following proportions: with a preapical
(type)
notaulices complete and distinct,
granulated;
scutellum nearly twice as long as metanotum; with 2 longitudinal
sp.
enlarged claw
tarsus
(Fig.
(Fig.
427)
427)
with a
spurs 1,1,1.
unknown
LOCUS TYPICUS:
Uvalde (Texas,
TYPICAL MATERIAL: HOSTS:
U.
S.
A.)
in SC.
unknown
DISTRIBUTION: NOTES:
holotype F!
Nearctic region:
the species is named
U.
S.
A.:
Uvalde (Tex.),
in honor of Mr.
4. Bacchus arizonicus
Marius S.
(R.C.L.
SC!
Wasbauer.
Perkins)
Eukoebeleia arizonica R.C.L. Perkins 1907: 44. Eukoebeleia arizonica R.C.L. Perkins: Kieffer 1914a: 47. Piwrbas arizonica (R.C.L. Perkins): Fenton 1918b: 263. Bacchus arizonica (R.C.L. Perkins): Nagy 1969: 322.
FEMALE: yellow;
fully
winged;
length 3,11-3,31
antennae brown,
mm;
head black,
with mandibles
with segments 1-2 or 1-3 testaceous;
thorax and
propodeum black; legs and tegulae testaceous; head dull, granulated, numerous areolae near clypeus, along orbits, near occipital carina; occipital carina complete; tooth between
mandibles quadridentate,
the two posterior teeth;
with
with a rudimentary
OOL nearly three times as long as
646
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
, ,
,-, I
(
(
\
'-
\
\
\
\
\
\
\
\
..i
"
\
\:/.;I--~;~)
s:;:::..- --.. . 'S. I
.-//
Fig.
426 -
Female of
Bocchus testaceus segments;
POL;
scutum dull,
granulated;
B:
n.
sp.
(type):
A:
fore tarsal
chela.
notaulices distinct and complete,
posteriorly
separated; scutellum shiny, punctate, nearly twice as long as metanotum; metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal band;
keels on posterior surface;
pterostigma short and narrow;
proportions:
31:
5:
6:
18;
fore-wing
fore tarsal
segment 5 of front
with 1 dark transversal segments 1-4 in following
tarsus
(Fig. 428)
with a
Bocchinae
647
"'-"- .....
\ \
Fig. 427 -
Chela of Bacchus wasbaueri n. sp.
(type)
preap ica I I ameli a and 1-2 brist les; en I arged cl aw (F i g. 428) wi th a raw of 6-8 teeth and no or 1 bristle; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Nogales (Ariz., U. S. A.) TYPICAL MATERIAL: holotype F! in B. HOSTS: in U.S.A. Bruchamorpha sp. {Issidae} (Fenton 1918b). DISTRIBUTION: Nearctic region: U. S. A.: Nogales (Ariz.), B! Uvalde
648
Mem. Amer.
Fig.
428 -
Ent.
Inst.,
no. 37,
1984
Chela of Bocchus arizonicus
(R.C.L.
Perkins)
(Texas)
from Uvalde
Bocchinae
Fig. 429 -
Chela of
649
Bocchus flavipes Kieffer from Almanor (California)
650
Mem.
(Tex.),
Amer.
En!.
Ins!.,
no.
37,
1984
OL!
5. Bocchus flavipes Kieffer
Bocchus flavipes Kieffer 1906a: 237. Tetradryinus flavipes (Kieffer): Kieffer 1913c: 325. Tetradryinus flavipes (Kieffer): Kieffer 1914b: 48. Tetradryinus flavipes (Kieffer): Richards 1939: 189. Tetradryinus flavipes (Kieffer): Muesebeck and Walkley 1951: 1035. Tetradryinus flaviceps (Kieffer): Krombein 1979: 1243.
FEMALE:
fully
winged;
antennae brown, testaceous; proportions:
length 3,37-3,56 mm;
antennae distally thickened; 10: 8:
12:
11:
rugose and with numerous with two OL = 3; strong
lateral
black;
mandibles testaceous;
with segments 1-2 or 1-5 testaceous; 9:
8:
7,5:
OOL = 10;
6:
6:
irregular keels;
keels around orbits; POL = 10;
transversal
antennal
head dull,
carina complete;
pronotum shiny,
pronotal
legs
reticulate
frons with a median keel
occipital
TL = 10;
impression;
11;
abdomen brown;
segments in following
hairy,
tubercles reaching
and
POL = 3; with a
tegulae;
scutum
sh i ny, smooth, weak I y punctate, wi thoul scu I pture among punctures, weakly rugose near anterior margin; notaulices complete, posteriorly separated; scutellum shiny, smooth, weakly punctate, without sculpture among punctures, weakly rugose near posterior margin; metanotum dull, granulated, with some keels; propodeum with broad areolae; posterior surface of propodeum with
two longitudinal
keels;
among numerous broad areo I ae;
fore-wi ng
beneath
part of radial
part
the pterostigma;
(6:9);
fore tarsal
enlarged claw front
tarsus
(Fig.
(Fig.
distal
segments in following
429)
429)
with a
small
areolae are visible
wi th 1 dark transversa I band vein shorter than proportions:
20:
4:
row of 6 teeth and 1 bristle;
with a preapical
proximal 5:
17: 25;
segment 5 of
lamella and 8 bristles;
tibial
spurs 1,1,1. MALE: unknown LOCUS TYPICU5: Ormsby Co. (Nevada, U. TYPICAL MATERIAL: holotype F! in CA. HOSTS:
S.
A.)
unknown
DISTRIBUTION: Ormsby Co.
Nearctic region:
(Nev.),
U.S. A.: Almanor
{Plumas Co.,
Calif.),OT!
CA!
6. Bacchus flavicollis Ashmead
Bocchus flavicollis Ashmead 1893: 91. Bocchus flavicollis Ashmead: Kieffer in Kieffer et Marshall 1905: 67.
Bocchinae
Fig.
430 -
Chela of
651
Bocchus flavicollis Ashmead (type)
Bocchus flavicollis Ashmead: Kieffer 1914b: 45. Bocchus flavicollis Ashmead: Muesebeck and Walkley 1951: 1039. Bocchus flavicollis Ashmead: Krombei n 1979: 1243.
FEMALE:
fully
winged;
length 2,80-2,81
mm;
head reddish,
with a broad
brown spot on frons and vertex; mandibles and clypeus testaceous; prothorax reddish; scutum reddi sh or reddi sh-brown; scutell um, metanotum, pleura and propodeum black; testaceousj
sometimes scutellum partly reddish;
abdomen brown-testaceousj
legs testaceousj
petiole
antennae missing in
652
Mem.
holotype;
head dull,
rudimentary
flat,
complete;
separated;
12:
5:
12: 9:
scutellum smooth,
transversal
9:
8:
19: 3:
4:
1984
shiny,
10;
part;
carina
pronotum dull,
pronotal
tubercles
notaulices complete,
weakly granulated; fore-wing
distal
fore tarsal
12:
16; enlarged claw
lamella;
segment 5 of front tibial
frons with a
occipital
TL = 9;
impression;
very
with a
antenna I segments in
8:
OPL = 7;
granulated;
preapical lamella and 2 bristles; MALE: unknown
posteriorly
metanotum
with 1 dark
part of radial
vein
segments in following (Fig.
430)
tarsus
with a row of 7
(Fig.
430)
with a
spurs 1,1,1.
LOCUS TYPICUS: Marquette (Michigan, U. TYPICAL MATERIAL: holotype F! in WA. HOSTS:
9:
the pterostigma;
nearly as long as proximal teeth and 1 long
9:
propodeum reticulate rugose;
band beneath
proportions:
37,
keels around orbits;
transversal
scutum dull,
reticulate rugose;
no.
mandibles quadridentate,
OL = 2,5; OOL =7;
with a strong
tegulae;
Ins!.,
the two posterior teeth;
and with 2 lateral
POL = 2;
granulated, reaching
Ent.
granulated;
tooth between
following proportions: median keel
Amer.
S.
A.)
unknown
DISTRIBUTION:
Nearctic region:
Marquette (Mich.),
CANADA:
Nerepis (N.
B.),
u.
WA!
S.
A.:
WA!
7. Bocchus laticeps (Ashmead)
Phorbas Bocchus Bocchus Bocchus
FEMALE: MALE:
laticeps laticeps laticeps laticeps
Ashmead 1893: 90. (Ashmead):
Muesebeck and Walkley 1951:
(Ashmead):
Nagy 1969: 323.
(Ashmead):
Krombein
1979:
1040.
1243.
unknown
fully
testaceous;
winged;
length 2,32-2,50 mm;
antennae brown;
thorax,
head black,
propodeum,
with mandibles
pet iole and abdomen
black; legs black; tegulae yellow; head dull, almost hairless, granulated; antennal segments in following proportions: 11: 5: 7: 7: 7: 7: 7: 7: 7: 9; POL = 6;
Ol = 3;
OOl = 8;
OPl = 3;
Tl = 5;
notaulices longer than breadth of ocelli hairless; smooth,
notaulices complete, weak I y
punctate;
scutum dull,
posteriorly separated;
smooth and
propodeum reticulate rugose,
keels on posterior surface;
the
granulated,
scutellum shiny,
metanotum rugose on anterior ha If,
weakly punctate on posterior half; longitudinal
minimum distance between
(7:3);
median surface smooth,
with 2 without
sculpture, except for regions near keels which are rugose; internal surface of areolae rugose; fore-wing hyal ine, without dark transversal bands; pterostigma hyaline, oval, with distal genital ia:
part fig.
with only posterior margin brown;
longer than proximal
431;
tibial
spurs 1,1,2.
part;
radial
vein
abdomen with petiole short;
Bocchinae
Fig.
431
-
Bocchus laticeps (Ashmead}(type): A: ninth abdominal
Male of sternitej
LOCUS TYPICUS:
DISTRIBUTION: (Tex.), OLl
B:
genitalia
(right gonoforceps removed).
Jacksonville (Florida,
TYPICAL MATERIAL: HOSTS: unknown
653
holotype M!
Nearctic region:
U. S.
A.)
in WA. U. S. A.: Jacksonville
(Fla.),
WA!
Uvalde
8. Bocchus dubius n. sp.
FEMALE: MALE:
unknown
fully
wingedj
length 2,0 mmj
antennae not distally thickened; proportions:
10: 6:
8: 8:
7:
7,5:
black,
with antennae and
legs brownj
antenna I segments in following 7:
7:
7:
9;
head dull,
granulated,
654
Fig.
Mem.
432 -
Amer.
Male genitalia of
Ent.
Inst.,
no.
37,
Bocchus dubius n.
1984
sp.
(type)(right half
removed)
reticulate rugose;
frons with a complete median keel;
occipital
carina
complete; POL = 6; OL = 4; OOL = 7; OPL = 4; TL = 4; scutum shiny, without sculpture; notaul ices complete, posteriorly separated; minimum distance between the notaulices slightly
longer than
temples
(5:4);
655
Bocchinae
scutellum shiny,
without sculpture;
reticulate rugose; mostly granulated; distal fig.
fore-wing hyaline,
part of radial
432;
propodeum
without dark transversal
vein shorter than proximal
part
(6:8);
bands;
genitalia:
t i b i a I sp u rs 1, 1 , 2.
LOCUS TYPICUS:
Green River
TYPICAL MATERIAL: HOSTS:
metanotum reticulate rugose;
posterior surface of propodeum weakly reticulate rugose,
(Wyoming,
holotype M!
U. S.
A.)
in AM.
unknown
DISTRIBUTION:
Neotropic region:
U.
S.
A.:
Green River
AUSTRALIAN SPECIES OF
(Wyoming),
AM!
BOCCHUS
Five species are known.
KEY TO THE SPECIES
FEMALES
Segment
4 of
front
tarsus
shorter
than
segment
1 ••••••••••••••• 2
Segment 4 of front tarsus longer than segment 1 ••••••• •••• '" 2 Fore-wing wi thout dark transversal bands; radial vein less
-
curved
(Fig.
416 E);
scutum reticulate
•• 3
rugose; notaulices 1. australiae n. sp. - Fore-wing wi th 2 dark transversal bands; radial vein more strongly curved (Fig. 416 D); scutum smooth, wi thout sculpture; notaulices incomplete . . . . . . . . . . 2. guineensis n. sp. 3 Head granulated, reticulate rugose only behind ocelli, near occipi tal carina . . . . . . . . . . . . . . . 3. minimus n. sp. - Head fully reticulate rugose ••••••••••••••••••••••••••••••••••• 4 4 Enlarged claw and segment 5 of front tarsus regularly curved (Fig. 436); head, thorax and propodeum black (with mandibles testaceous) . . . . . . . . . • . . . . . . . . . . 4. robustus (Dodd) - Enlarged claw and segment 5 of front tarsus less curved, partly rect i I inear (Fig. 437); head, pronotum and scutum fully or partly reddish; scutellum, metanotum and propodeum black . . . . . . . . . . • . . • . . • . . . • . . • • . . • . • . . . . . . .. 5. bouceki n. sp. complete . . . . . . . . . . . . . . . . . . . . . . .
MALES
Only the male of
B. guineensis n. sp. is known.
Mem.
656
Amer.
Ent.
Inst.,
no.
37,
1984
1. Bacchus australiae n. sp.
FEMALE:
fully
winged;
length 3,62-3,87 mm;
yellow;
antennae brown dark or
thorax,
propodeum and abdomen black;
fully
reticulate rugose;
occipital
with 2-3 rudimentary teeth; reticulate rugose;
POL
weak
with 2
longitudinal
transversal
tarsal front
band;
(Fig.
433)
distinct prominence; long bristle;
tibial
MALE: unknown LOCUS TYPICUS:
=
5;
OL
weakly
=
4;
Mt.
TYPICAL MATERIAL:
oval,
proportions:
with 1 preapical
433)
hairy,
propodeum reticulate brown 3:
fore-wing (Fig. 12;
Tibrogargan
2 bristles and
1 paratype F!
in QU.
I
433 -
1 proximal
with a row of 4 teeth and 1
(f
Fig.
fore
segment 5 of
(Australia) in CB;
with 1
416 E);
spurs 1,1,2.
holotype F!
fully
posteriorly separated;
14: 3:
lamella,
(Fig.
ha i ry,
mandibles quadridentate,
scutum dull,
punctate;
pterostigma short,
enlarged claw
with mandibles head du II,
legs testaceous;
carina comlete;
keels on posterior surface;
segments 1-4 in following tarsus
head black,
with segments 1-4 testaceous;
notaul ices distinct and complete,
scutellum and metanotum dull, rugose,
light,
Chela of Bocchus australiae n.
sp.
(type)
Bocchinae
Fig. 434 -
657
Chela of Bacchus guineensis n.
HOSTS: unknown DISTRIBUTION: Australian region:
AUSTRALIA:
Mt.
sp.
(type)
Tibrogargan
658
Mem.
(Queensland),
CB!
Amer.
Woogaroo,
Ent.
Inst.,
37,
no.
1984
QU!
2. Boccus guineensis n. sp.
FEMALE:
fully
winged;
length 3,00-3,50 mm;
head black,
with mandibles
and sometimes clypeus yellow; antennae fully testaceous; thorax, propodeum and abdomen black; legs testaceous, with hind coxae black; head shiny or dull, some transversal
without sculpture or weakly granulated,
mandibles with 3 teeth;
occipital
smooth,
shiny,
only on
anterior third of scutum;
sculpture;
except for
weak keels on frons and some areolae behind ocell i; carina complete;
without sculpture hairless;
scutellum smooth,
metanotum weakly punctate;
POL = 2;
Ol = 3;
notaulices incomplete, shiny,
scutum
visible
without
propodeum reticulate rugose,
with 2
longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; radial vein strongly curved (Fig.
416 D); fore tarsal segments 1-4 in following proportions: 12: 4: 3: 9; segment 5 of front tarsus (Fig. 434) with a preapical lamella, one proximal prominence and one bristle; enlarged claw (Fig. 434) with one teeth and one long bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,00-2,62 mm; black; mandibles testaceous; antennae brown or wi th segments 1-2 testaceous; abdomen brown; legs brown or testaceous,
with coxae and fore clubs of femora brown;
antennae
antennal segments in following proportions: 9: 5: 6: 6: 8: 7: 7: 7: 6: 9; head dull, granulated; frons with a median keel; occipital carina complete; POL = 3; Ol = 2,5; OOl = 6,5; OPl = 4; Tl = not distally thickened;
2,5; scutum dull, granulated, with numerous
longitudinal
reaching approximately 0,5 shiny,
weakly granulated;
longitudinal
with numerous areolae on anterior half and
keels on posterior half; length of scutum;
notaulices incomplete,
scutellum and metanotum
propodeum reticulate rugose,
keels on posterior surface;
fore-wi ng wi th a dark transversal
with distal
lOCUS TYPICUS: Baiyer R. (New Guinea) TYPICAL MATERIAL: holotype F! and 7 paratypes MM! in B;
HOSTS:
1 paratype M!
in OT;
two
band beneath the pterost i gma;
pterostigma lanceolate; radial vein curvi I inear, than proximal part; tibial spurs 1,1,2.
F!
with
median area shiny and smooth;
4 paratypes
(1
M,
in TW;
3 FF)!
part shorter
1 paratype
in Ol.
unknown
DISTRIBUTION:
Australian region:
B!
TW!
Baiyer R.,
BRITAIN:
Kerrvat,
Jimmi
Valley,
NEW GUINEA: Ol!
Wau,
Ol!
Tifalmin
(NE New Guinea),
Kassam Pass,
OT!
3. Bocchus minimus n. sp.
Ol!
NEW
80cchinae
FEMALE:
fully
and mandibles
winged; light;
length 2,81
mm;
antennae fully
659
head reddish-brown,
testaceousj
with clypeus
thorax and propodeum
reddi sh, part I y darkened on scutum and propodeum; abdomen brown; iegs testaceous; head dull, granulated, reticulate rugose only behind ocelli, near occipital POL
~
4;
OL
~
anterior third;
carina; 3;
mandibles with 3 teeth;
scutum dull,
granulated,
notaulices distinct and complete,
scutellum and metanotum shiny, reticulate rugose, fore tarsal
band;
(Fig. (Fig.
435) 435)
carina complete;
posteriorly separated;
weakly punctate;
propodeum
keels on posterior surface; pterostigma short and oval,
segments 1-4 in following proportions:
tarsus
enlarged claw MALE:
smooth,
with 2 longitudinal
with 1 weak dark transversal of front
occipital
reticulate rugose only on
with a preapical
13: 2:
2:
14;
fore-wing brown; segment 5
lamella and 2 bristles;
with a row of 14 teeth;
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS: Spargoville (Australia) TYPICAL MATERIAL: holotype F! in 8M. HOSTS: unknown DISTRIBUTION: Australian region: Western Australia),
AUSTRALIA:
Spargoville
(Coolgardie,
BM!
L L
•.......•.....
Fig. 435 -
Chela of
~ ~~ ....•..•....
Bacchus minimus
n.
....
sp.
(type)
660
Mem. Amer.
Fig. 436 -
Ent.
Inst.,
Chela of Bocchus robustus
no.
(Dodd)
37,
1984
from Townsville
(Australia)
4. Bacchus robustus (Dodd) n. comb.
Anteonella robusta Dodd 1913: 182. Anteonella bicolor Dodd 1913: 182 (n. syn.); locus typicus Townsville (Australia).
Anteonella robusta Dodd: Kieffer 1914b: 213. Anteonella bicolor Dodd: Kieffer 1914b: 214. Dicondylus sulciscutellum Girault 1926: 135 (n. syn.); locus typicus Townsville
(Australia).
Bocchinae
FEMALE:
fully
testaceous;
winged;
length 4,87-5,12 mm;
antennae testaceous,
propodeum and abdomen black; clubs of femora black;
661
head black,
with mandibles
with segments 4-10 or 6-10 dark;
thorax,
legs fu Ily testaceous or wi th coxae and
head swollen,
dull shiny,
fully reticulate rugose,
with a median keel between the antennal sockets; ocellar region swollen; temples distinct; occipital carina complete; POL = OL; OOL = OPL; mandibles tridentate,
with an
intermediate rudimentary tooth;
pronotum
shiny, with a transversal impression, reticulate rugose and with transversal keels; pronotal tubercles reaching tegulae; scutum shiny, reticulate rugose; posteriorly
notaul ices complete and distinct,
separated;
scutellum shiny,
median furrow composed of areolae;
smooth,
fully
composed of areolae,
weakly punctate,
with a
metanotum shiny and smooth,
reticulate rugose;
propodeum fully reticulate rugose;
transversa I band;
pterost i gma broad and ova I,
fore-wing
brown;
laterally
with 1 dark
segment 3 of front
tarsus without distal apophysis longer than segment 3; segment 4 of front tarsus longer than segment 1; segment 5 of front tarsus (Fig. 436) with an apical
lamella and 1 long proximal
a row of 16 teeth and 3 proximal MALE: unknown LOCUS TYPICUS:
Townsville
bristle;
enlarged claw
bristles;
tibial
(fig.
436)
with
spurs 1,1,1.
(Australia)
of A. robusta in AD; holotype F! of A. bicolor in AD; holotype F! of D. sulciscutellum in QU. TYPICAL MATERIAL: HOSTS:
holotype F!
in Australia Fulgorids.
DISTRIBUTION: Australian region: AUSTRALIA: AD! Blunder Cr. (Brisbane, Queensland), UQ! Australia),
WA!
Sir Graham Moore
Island
Townsville (Queensland), Anjo Peninsula (Western
(Western Australia),
QU!
OL!
5. Bocchus bouceki n. sp.
FEMALE:
fully
reddish
winged;
in holotype;
length 3,37-4,87 mm;
head black
head,
pronotum and scutum
(with clypeus and mandibles
testaceous),
prothorax reddish, scutum black (with notaulices reddish) in paratype; antennae brown with segments 1-2 testaceous or fully testaceous; scutellum, metanotum,
propodeum and abdomen black;
hind coxae brown; = 4;
occipital
rudimentary rugose,
head hairy,
carina complete;
tooth between
hairy,
shiny;
shiny,
legs testaceous,
fully
sometimes with
reticulate rugose;
mandibles quadridentate,
the two posterior teeth;
POL = 5;
OL
with a
scutum fully reticulate
notaulices complete and distinct,
posteriorly
separated; scutellum and metanotum smooth, hairy, weakly punctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; brown;
fore-wing fore tarsal
segment 5 of front bristle; bristles;
with 1 dark transversal tarsus
enlarged claw tibial
band;
pterostigma short,
segments 1-4 in following proportions: (Fig.
(Fig.
spurs 1,1,2.
437)
417)
with a preapical
18:
3:
3:
oval, 26;
lamella and no
with a row of 17 teeth and without
662
Mem. Amer. Ent.
Ins!.,
no. 37,
1984
Fig. 437 - Chela of Bocchus bouceki n. sp.
MALE: unknown LOCUS TYPICUS: Mt. Tibrogargan
(Australia)
(type)
Bocchinae
TYPICAL MATERIAL:
holotype F!
in CB;
663
1 paratype F!
in AD.
HOSTS: unknown DISTRIBUTION: Australian region: AUSTRALIA: Mt. Tibrogargan (Queensland), CB! Landrigan Cliffs (North Western Australia), AD! NOTES:
the species
is named
in honor of Mr.
Zdenek Boucek.
4. Mirodryinus Ponomarenko
Mirodryinus Ponomarenko 1972: 673.
Fig. 438 -
Female of Mirodryinus xerophilus
(Benoit)
from Wassiya
(Egypt)
664
Mem. Amer.
Fig. 439 -
Ins!.,
no.
Female of Mirodryinus xerophilus
TYPE SPECIES: Mirodryinus FEMALE
En!.
(Figs 438, 439):
37,
1984
(Benoit) from Wassiya
(Egypt)
ungulatus Ponomarenko 1972, orig. desig.
fully winged;
maxi Ilary palpi
with 6 segments;
labial palpi with 3 segments; occipital carina complete; mandibles quadridentate, with a rudimentary tooth bet wen the two posterior teeth; segment 4 of front tarsus less than 0,5 times as long as segment 1 (Fig. 442); segment 5 of front tarsus without lamellae, with bristles or peg-like hairs (Fig. 442); enlarged claw without teeth, lamellae, bristles or peglike hairs (Fig. 442); fore-wing with radial vein distinct (though sometimes it is very short); pronotal tubercles reaching tegulae; tibial spurs 1,1,1. MALE (Figs 440, 441): fully winged; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; maxillary palpi with 6 segments; labial palpi with 3 segments; temples distinct; propodeum with areolae very small, less wide than tegulae (Fig. 440); tibial spurs 1,1,2. DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES: 5
PALAEARCT I C SPEC I ES OF
MIRODRYINUS
Five species are known.
KEY TO THE SPECIES
Bocchinae
Fig.
665
Male of Mirodryinus xerophilus
440 -
(Benoit)
from
Ikingi
Mariout
(Egypt)
FEMALES
Fore-wing distal
with
parts;
radial
vein
curved,
with distinct
proximal
and
thorax and propodeum fully black . . . . . . . . . . • . . . . • .
1. gobiensis Ponomarenko -
Fore-wing wi th proximal pa r t I y
part
radial
vein
straight,
(Figs 438,439);
only wi th a distinct
thorax and propodeum fully
or
redd i sh •••••••••••••••••••••••••••••••••• ••' ••••••••••••• 2
2 Thorax and propodeum fully reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . •••••••••••••••••••••••••••••• 2. xerophilus (Benoi t) -
Thorax and propodeum partly
3 Scutum and
or black • . . . . . . . . 3 ungulatus Ponomarenko
reddish and brown
scutellum brown . . . .
3.
666
Mem.
Fig.
441
-
Male of
Amer.
En!.
Ins!.,
37,
no.
1984
Mirodryinus xerophilus (Benoit) from Ikingi Mariout (Egypt)
- Scutum and scutellum reddish •••••• ••••••••••••••••••••••••••••• 4 4 Metapleura reddish; radial vein longer . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . • . . . . . . . . . . . . . .. 4. atlanticus n. sp. - Metapleura black; radial vein shorter (Fig. 448 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. ponomarenkoae n. sp.
MALES
Only
the male of M.
xerophilus is known.
1. Mirodryinus gobiensis Ponomarenko
Mirodryinus gobiensis Ponomarenko 1972: 676.
FEMALE:
fully
legs fully
winged;
testaceous;
length 2,62 mm;
head,
ocellar region brown;
mandibles,
antennae and
thorax and propodeum black;
abdomen black; antennae distally thickened; antennal segments in following proportions: 7: 3: 6: 5,5: 5: 5: 5: 4,5: 5: 5; head shiny, weakly
punctate,
or lateral 4;
keels;
Ol = 2,5;
without sculpture among punctures; temples very prominent;
OOl = 10; OPL = 4;
Tl = 9;
occipita!
frons without median
carina complete;
pronotum very short,
POL
weakly
Bocchinae
667
B
Fig.
442 -
Female of Mirodryinus gobiensis Ponomarenko (type):
B: fore tarsal segments.
A: chela;
668
Mem.
transversely striate;
pronotal
and metanotum shiny, punctures; scutum;
Amer.
Ent.
Inst.,
weakly punctate,
notaulices incomplete,
longitudinal
keels;
fore-wing
pterostigma;
radial
vein curved,
segments in following proportions: 442)
1 of front
LOCUS TYP ICUS:
DISTRIBUTION:
442);
and distal part
18: 29:
hollow
long
the
parts;
(2,5:4,5);
65;
segment 5
apex with a
lamella);
(antenna-cleaner)
fore
enlarged claw
bristles or other appendixes;
tibial
Bordson-Gobi
TYPICAL MATERIAL: HOSTS: unknown
15:
(among which one very
tarsus with a proximal
a big prominence (Fig. MALE: unknown
aymag,
teeth,
82:
or
bands beneath
with 2 rows of nearly 15 bristles;
group of nearly 5 lamellae
scutellum
without transversal
vein much shorter than proximal
without subapical (Fig.
dull,
with distinct proximal
part of radial
tarsus
scutum,
without sculpture among
with a dark transversal
tarsal
442)
1984
reaching approximately 0,3 length of
distal (Fig.
37,
tubercles reaching tegulae;
smooth,
propodeum fully reticulate rugose,
of front
no.
segment
del imitated by
spurs 1,1,1.
(Mongol i a)
holotype F!
in LN
Palaearctic region:
South-East Mongolia),
MONGOLIA:
Bordson-Gobi
(South Gobi
LN!
2. Mirodryinus xerophilus (Benoit) n. comb.
Anteon xerophilus Benoit 1951a: 301. Anteon xerophilum Benoit: Olmi and Currado 1979a: 346.
FEMALE
(Figs 438,
439):
fully
petiole and abdomen black; testaceous,
winged;
length 2,68 mm;
with segments 5-10 brown;
head shiny,
smooth,
with antennae
antennae distally thickened;
antenna I segments in following proportions: 4,5: 7;
reddish,
sometimes propodeum partl y brown; 6:
4:
strongly punctate,
11: 7:
5,5:
4,5:
4:
5:
without sculpture among
punctures; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; frons without medial keel; occipital carina complete; POL = 6,5; dorsal
OL = 4;
view;
punctate,
scutum,
OOL = 7;
with a
TL = 10; pronotum
without sculpture among punctures;
propodeum reticulate rugose, fore-wing
OPL = 8;
scutellum and metanotum shiny,
track of dark spot on the basal
proximal part, six times as
without the distal
long as segment 4,
part;
weakly
notaulices invisible;
without transversal
with a broad dark transversal
invisible in
smooth,
and
longitudinal
keels;
band beneath the pterostigma and cells;
radial
vein only
segment 1 of front
with a proximal
delimitated by a big prominence (Fig. 443);
hollow
with the
tarsus nearly
(antenna-cleaner)
enlarged claw
(Fig.
443)
without bristles or teeth or lamellae; segment 5 of front tarsus (Fig. 443) with a long row of small peg-like hairs; apex with a group of 4 lamellae; tibial
spurs 1,1,1.
669
Bocchinae
B
Fig. 443 -
Female of
Mirodryinus xerophilus (Benoit) from Egypt: A: chela; B:
fore tarsal
segments.
670
Mem. Amer.
Ent.
Ins!.,
no.
-1/
/
, \
II
\I
/
\
/ /
I /
I
II
I
II 1\
I
I
)
/
I
I
1/
/'
1984
I
\
/ \
37,
'\
I
1\
I I
I
I
\
I
I \ I \
/
I
1/ "-
I
I
\ I v
II
I / /
Fig. 444 - Male genitalia of Mil'odl'yinus xel'ophilus (Benoit)(type)
Bocch i nae
MALE
(Figs 440,
441),
fully
winged;
clypeus yellow and a small sockets and eyes; testaceous; scutellum,
671
length 2,5 mm;
head black,
reddish spot on region between
mandibles yellow,
with teeth brown;
antennae fully
pronotum and propectus reddi sh-testaceous or black; metanotum,
with
antennal
mesopleura and metapleura brown-black;
scutum propodeum
black; abdomen brown; legs fully testaceous; tegulae testaceous; antenna I segments in following proportions: 6: 2: 10: 9: 8: 7: 7: 6,5: 5: 7; head sh i ny,
swoll en,
punctures; complete;
smooth,
POL = 7;
wi th sparse punctures,
OL = 4;
scutum shiny,
smooth,
sculpture among punctures; approximately 0,3 smooth,
without sculpture, almost pointed;
propodeum dull,
hairy,
with sparse punctures,
notaulices distinct,
length of scutum;
convex,
wi thout scu I pture among
= 8; OPL = 7; TL = 7; occipital carina
OOL
swollen;
weak,
without
incomplete,
reaching
scutellum and metanotum shiny, laterally viewed metanotum very
mesopleura and metapleura shiny and smooth;
granulated,
with some irregular keels or areolae,
without
a visible transversal keel between dorsal and posterior surface; dorsal surface very reduced, shorter than breadth of metanotum, as scu I ptured as posterior surface; surface;
pterostigma, part,
propodeum almost fully composed of the posterior
fore wing hyal ine, with distal
nearly
invisible;
and without dorsal
with a dark transversal
part of radial gonoforceps
proximal
TYPICAL MATERIAL:
Ikingi
Mariout,
444)
without
BM!
proximal
inner distal
dorsally viewed
process long and
(Libya)
holotype M!
in GE.
HOSTS: unknown DISTRIBUTION: Palaearctic region: EGYPT:
(Fig.
membranous process,
broad; tibial spurs 1,1,1. LOCUS TYP ICUS: Giarabub oasis
band beneath the
vein much shorter than
LIBYA:
Giarabub Oasis (Cirenaica),
Fayoum-Tamiya,
WA!
Wassiya,
GE!
WA!
3. Mirodryinus ungulatus Ponomarenko
Mirodryinus ungulatus Ponomarenko 1972: 674.
FEMALE:
fully
winged;
length 2,31
part of frons darkened;
protorax redd ish-testaceous; brown-b lack; black,
mm;
head reddish,
antennae brown,
scutum brown;
propodeum black;
pleura,
scutell um and metanotum
mesosternum and metasternum
except for a reddish spot on mesopleura;
testaceous-dark;
antennae distally thickened;
following proportions,
7:
3,5, 9,
6:
5,
with vertex and
wi th segments 1-3 testaceous;
4,5:
abdomen brown;
antennal 4,5:
4,5:
segments 4,5:
6;
legs in
head shiny,
puncta te, wi thout scu I pture among punctures; frons wi thou! med i an and lateral keels; occipital carina complete; temples very prominent; POL = 6; OL = 3;
OOL
=
9;
tubercles reaching
OPL = 7,5; tegulae;
TL
=
scutum,
10;
pronotum shiny,
short;
pronotal
scutellum and metanotum shiny,
672
Fig.
Mem.
Amer.
Ins!.,
no.
37, 1984
445 - Female of Mirodryinus ungulatus Ponomarenko (type): A: chela; B:
smooth,
weakly punctate,
incomplete,
fore tarsa I segments.
without sculpture among punctures;
reaching approximately 0,5
reticulate rugose, wing
Ent.
dull,
without transversal
with a dark transversal
without a distinct distal
length of scutum;
part;
or longitudinal
band on distal fore tarsal
half;
segments
radial
notaulices
propodeum keels;
fore-
vein straight,
in following
proportions: 85: 13: 15: 16: 41; segment 1 of front tarsus with a proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig.
445); enlarged claw (Fig. 445) without subapical tooth, bristles or other appendixes; segment 5 of front tarsus (Fig. 445) with 2 rows of nearly 26 peg-like lamellae; apex with a group of nearly 5 lamellae; tibial spurs 1,1,1. MALE:
unknown
LOCUS TYPICUS: Husseyn-Dey (Algeria) TYPICAI_ MATERIAL: holotype F! in LN. HOSTS:
unknown
DISTRIBUTION:
Palaearctic region:
ALGERIA:
Husseyn-Dey,
4. Mirodryinus atlanticus n. sp.
LN!
Bocchinae
Fig. 446 -
FEMALE:
Fore tarsal
fully winged;
673
segments of Mirodryinus atlanticus n.
length 2,62 mm;
head reddish;
sp.
(type)
antennae brown,
674
Mem.
Amer.
with segments 1-3 testaceous;
En!.
pronotum,
metanotum and propodeum black; abdomen brown-dark; punctate, 10;
with
occipital
legs and
keels;
dark
straight,
band on
not curved;
scutum shiny,
(Fig.
446)
lamellae; segment
=
5;
head shiny, OL
1 of front
shiny,
distal
3:
LOCUS TYPICUS:
radial oval,
3,5:
7;
without
with a proximal
hairy,
(Canary
holotype F!
9;
TL
=
with smooth,
propodeum reticulate
vein
fore-wing
brown;
fore tarsal
bristles,
segments
tarsus
in addition
lamellae,
(Fig.
with a
very short and
segment 5 of front
hollow
big prominence
Gran Tarajal
TYPICAL MATERIAL:
446)
strongly
=
OOL
weakly punctate,
hairless;
region;
2,5:
(Fig.
tarsus
3;
rugose,
with some bristles and 3 peg-I ike hairs, enlarged claw
=
keels on posterior surface;
17:
distally del imitated by a MALE: unknown
HOSTS:
and scutellum reddish;
visible only on anterior third;
pterostigma short,
in following proportions:
1984
and metap leura redd i sh;
POL
notaul ices incomplete,
longitudinal
transversal
scutum
pronotum short,
scutellum and metanotum smooth, without
37,
tegulae testaceous;
carina complete;
without sculpture;
no.
mesop leura
temples very prominent;
numerous transversal
rugose,
Ins!.,
to 4 distal
hairs;
("antenna-cleaner")
446);
tibial
spurs 1,1,1.
Islands)
in HE.
unknown
DISTRIBUTION:
Palaearctic region:
(Fuerteventura),
CANARY
ISLANDS:
Gran Tarajal
HE!
5. Mirodryinus ponomarenkoae n. sp.
Mirodryinus ungulatus Ponomarenko partim 1972: 674.
FEMALE:
fully
winged;
on ocellar region; reddish,
length 3,20 mm;
antennae missing
with anterior surface brown;
propodeum and abdomen black;
head reddish,
in holotype;
with a brown spot
pronotum reddish;
scutellum reddish;
mesopleura reddish,
scutum
metanotum,
with sutures blackish;
metapleura black; legs testaceous; head shiny, weakly alutaceous, strongly punctate on frons; occipital carina complete; frons without median keel; POL
=
=
10;
9;
OL
=
TL
=
10;
? (anterior ocellus damaged and not visible); OOL temples very prominent;
with numerous transversal punctate;
keels;
notaulices incomplete,
scutellum and metanotum shiny, rugose, dark
without
transversal
straight tarsal (Fig.
longitudinal
(Fig.
segments 447)
a proximal
scutum shiny, smooth, region;
not curved;
hollow
lamellae,
bristles,
("antenn-cleaner")
20:
hairs;
3:
=
8;
OPL
hairy, not
length of scutum; propodeum reticulate
vein
fore-wing
3: 4:
segment
with a
very short and
pterostigma short and oval,
in following proportions:
without
hairless; radial
rugose,
weakly alutaceous,
reaching only 0,3
keels on posterior surface;
band on distal
448 A),
pronotum short,
10;
brown;
fore
enlarged claw
1 of front
distally delimitated by a
tarsus with big
Bocchinae
Fig.
447 -
Fore tarsal
segments of female of Mirodryinus ponomarenkoae n. sp.
prominence (Fig.
447);
nearly
17 lamellae;
1,1,1. MALE:
unknown
675
(type)
segment 5 of front
tarsus
LOCUS TYPICUS: Western part of Mount Zamryn-Ula TYPICAL MATERIAL: holotype F!in LN. HOSTS;
Zamryn-Ula
Palaearctic region:
(Central
with 1 row of tibial
spurs
(Mongolia)
Aymag),
MONGOLIA:
Western part of Mount
LN!
the holotype of this species was designated by Ponomarenko
as para type of M. of Mrs.
447)
unknown
DISTRIBUTION: NOTES:
(Fig.
apex with a group of nearly 8 lamellae;
(1972)
ungulatus Ponomarenko. The species is named in honor
Nadezdha Ponomarenko.
5. Radiimancus Moczar
Radiimancus Moczar 1983b: 203.
TYPE SPECIES: Radiimancus olmii Moczar 1983b: 204, FEMALE labial
(Figs 449, palpi
tooth between 0,5 times as
450):
ful!y
with 3 segments;
winged;
long as segment 1 (Fig. tarsus without
segment 4 of front 451);
lamellae,
occipital
desig.
with 6 segments;
mandibles quadridentate,
the two posterior teeth;
segment 5 of front
orig.
maxi Ilary palpi
with a rudimentary tarsus
less than
carina complete;
with bristles or peg-like hairs
Mem.
676
Amer.
En!.
Ins!.,
no.
37,
1984
A~
B~ Fig. 448 - Pterostigma and radial vein of females of Mirodryinus ponomarenkoae n. sp. (type)(A) and Radiimancus olmii Moczar (type)(B)
(Fig.
451);
bristles
enlarged claw without teeth,
(Fig.
451);
radial
vein
(Fig.
MALE:
unknown
DISTRIBUTION: HOSTS:
pronotal
448 B);
lamellae,
peg-I ike hairs or
tubercles reaching tegulae;
tibial
fore-wing with no
spurs 1,1,1.
Palaearctic
unknown
SPECIES:
1
Radiimancus olmii Moczar
Radiimancus olmii Moczar 1983b: 204.
FEMALE
(Figs 449,
450):
fully winged;
anterior surface of frons,
length 2,80 mm;
head black,
clypeus and mandibles testaceous;
with
antennae
testaceous, with last 5 segments darkened; pronotum and prosternum reddish; scutum reddish, with central surface darkened; scutellum, metanotum and propodeum black; mesosternum reddish, with black spots; metasternum and metapleura black; mesopleura reddish; abdomen brown; legs testaceous;
antennae weakly distally thickened;
following proportions: quadri dentate, head shiny, occipital
8:
3:
12: 8,5:
with a rud imentary
smooth,
6:
6:
6:
6:
antennal 8;
segments in
mandibles
tooth between the two posterior teeth;
weakly punctate,
carina complete;
7:
without sculpture among punctures;
frons without median keel;
POL = 8;
OL = 4;
OOL = 10; OPL = 8; TL = 8; pronotum short, hairy, rugose, pronotal tubercles reaching tegulae; scutum shiny, weakly punctate, without sculpture among punctures;
notaulices incomplete,
reaching only 0,3 lenth
of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, without longitudinal or transversal keels;
677
Bocchinae
Fig. 449 -
Female of Hadiimancus olmii Moczar
(type)
fore-wing with a dark transversal band on distal surface; pterostigma broad and oval; radial vein absent (Fig. 448 B); fore tarsal segments following proportions:
17: 2:
2,5:
3:
9;
in
segment 1 of front tarsus with a
proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig. 451); segment 5 of front tarsus (Fig. 451) with 2 rows of nearly 27 bristles extending continuously to the apex; enlarged claw (Fig. 451)
without
lamellae, bristles or hairs;
tibial
spurs 1,1,1.
MALE: unknown LOCUS TYP I CUS: Bu I gan (Mongol i a) TYPICAL MATERIAL: holotype F! in BU. HOSTS: unknown DISTRIBUTION: Palaearctic region:
MONGOLIA:
Bulgan
(10 Km SSW Somon,
678
Mem.
Fig.
Chovd
aimak),
450 -
Amer.
Ent.
Inst.,
no.
37,
1984
Female of Radiimancus olmii Moczar (type)
BU!
6. Mystrophorus Forster
Mystrophorus Forster 1856: 91. Mystrophorus Forster: Richards 1937: 119.
TYPE SPECIES: Mystrophorus FEMALE labial
(Figs 452, 453): palpi
with 3 segments;
tooth between tarsus
less than 0,5 times as
or peg-like hairs
456);
I ike hairs or bristles (Fig. tibial spurs 1,1,2. (Figs 454,
rudimentary
less wide than tegulae;
with 6 segments;
396 A);
tarsus without
with a rudimentary
segment 4 of front 456);
456);
pronotal
occipital
lamellae,
enlarged claw without teeth,
455): micropterous;
tooth between
(Fig.
long as segment 1 (Fig.
segment 5 of front (Fig.
maxillary palpi
mandibles quadridentate,
the two posterior teeth
carina complete;
MALE
formicaeformis Ruthe 1859, monotypic.
micropterous;
with bristles lamellae,
mandibles quadridentate,
the two posterior teeth;
maxi Ilary palpi
peg-
tubercles reaching tegulae; with a
areolae of propodeum
with 6 segments;
labial
palpi
with
3 segments; temples distinct. DISTRIBUTION: Palaearctic HOSTS:
Cicadellidae (excl ud i ng I diocerinae, Macropsinae,
SPECIES:
1
Mystrophorus formicaeformis Ruthe
Typhlocybinae).
Bocchinae
679
A
Fig. 451 - Female of Radiimancus oimii Moczar (type): A: fore tarsal segment 1; B: fore tarsal segments 1 (partly)-4 and chela
Mystrophorus iormicaeiormis Ruthe 1859: 120. Mystrophorus iormicaeiormis Ruthe: Ponomarenko 1978: 23.
FEMALE (Figs 452, 453): micropterous, with fore wings as long as tegulaej hind wings rudimentary, rounded; length 1,75-2,50 mm; black, with
680
Mem. Amer. Ent.
Fig. 452 -
Inst.,
no.
37,
1984
Female of Mystrophorus formicaeformis Ruthe
Bocchinae
Fig.
453 -
antennae brown
Female of
681
Mystrophorus formicaeformis Ruthe
(except for segments 1-2 testaceous),
tegulae brown-testaceous,
legs yellow
mandibles yellow,
(with hind coxae almost
clubs of femora and hind tibiae brownish);
body dull,
hairy,
fully black, except for
abdomen hairless; head without median keel between antenna I sockets; occipital carina complete; head, thorax and propodeum granulated, without striae or keels; invisible;
some keels are visible near apex of propodeum;
fore tarsal
segments 1-4 in following proportions:
notaulices
5,5:
1,2:
2,3:
2,3; en I arged cl aw (Fig. 456) without I ameli ae or brist les, wi th apex rounded; segment 5 of front tarsus (Fig. 456) without lamellae, with numerous rows of bristles or peg-like hairs. MALE
(Figs 454,
455):
micropterous,
with fore wings narrow and broadened
at apex; hind wings rudimentary, pointed; length 1,87-2,75 mm; black, with antennae brown(except for segments 1-2 testaceous), tegulae yellowbrown,
fore-wing
femora blackish, for abdomen sockets;
almost hairless;
occipital
granulated; propodeum;
yellow-brown,
legs yellow
(with coxae and clubs of
mid and hind tibiae brownish);
body dull,
head without median keel
carina complete;
head,
without striae or keels,
hairy,
between
except
antenna I
thorax and propodeum
except for some keels near apex of
notaulices invisible.
LOCUS TYPICUS:
Boden
TYPICAL MATERIAL: HOSTS: in Germany
(Germany)
not found
(not
in BM and
in Gent,
Belgium).
Deltocephalus sp. (Haupt 1938); in U. S. S. R. Diplocolenus abdominalis F. (Ponomarenko 1978). DISTRIBUTION: 1978); det.
Palaearctic region:
HUNGARY: Or Sz.
Miklos,
U.
BU!
S. Vacz
S.
R.:
Moscow
(Tudosdomb),
(Ponomarenko BU!
Hungary
(male
Mystrophorus cursor Forster type), VI! CZECHOSLOVAKIA (Ogloblin
1924b);
GERMANY:
E I steraue (Ha lie),
HA!
BM!
Boden
(Ruthe 1859);
Ha II e-a-
5., HA! Bellinchen (Oder), HA! KUpka (Th.), EB! Mallertshofer Holzb. (MUnchen), EB! OT! Saale-Ufer (Haupt 1938); Schl inzheim (Mallertshofer Holz), GV! ENGLAND: Hayling Island (Hampshire), BM! GREECE: Kerkyra Island,
BU!
682
Mem.
Fig.
454 -
FEMALE
En!.
Inst.,
Male of Mystrophorus
no.
37,
1984
formicaeformis Ruthe from Hungary
5UBFAMILY THAUMATODRYININAE N.
6.
TYPE GENUS:
Amer.
Thaumatodryinus R.C.L. Perkins 1905
(Figs 19,
457):
fully
winged;
fore-wing
with costal,
submedian cells clearly enclosed by pigmented veins also first brachial labial
palpi
SUBF.
cell
clearly enclosed;
with 3 segments;
front
(Fig.
maxillary palpi
tarsus chelate;
median and
457);
usually
with 6 segments;
chela with rudimentary
claw (Fig. 459); antennae with tufts of long hairs on segments 5-10 (last segment with two tufts) (Fig. 19); occipital carina complete; mandibles with 4 teeth progressing spurs 1,1,2.
larger from anterior one to posterior (Fig.
5);
tibial
Thaumatodry i n i nae
Fig.
MALE
Male of Mystrophorus
455 -
(Figs 28,458):
fully
winged;
683
formicaeformis Ruthe from Hungary
fore-wing
with costal,
submedian cells clearly enclosed by pigmented veins palpi
with 6 segments;
teeth progressing carina complete; pterostigma
28);
GENERA:
with 3 segments;
with metacarpus as
tibial
Palaearctic
Oriental, Nearctic, HOSTS: Flatidae SPECIES:
palpi
median and 28); 5);
occipital
long as or longer than
spurs 1,1,2.
(?)(fossil
Neotropic,
maxillary
mandibles with 4
larger from anterior one to posterior (Fig. fore-wing
(Fig.
DISTRIBUTION:
labial
(Fig.
species:
see notes),
Ethiopian,
Australian
1 17
NOTES: Brues described three species of Thaumatodryinus from Palaearctic region (Baltic amber): filicornis (Brues 1923), deletus Brues 1933 and
gracilis Brues 1933. Though the original descriptions are incomplete and the typical correct.
material
is
In the original
lost,
I think that the Brues determination
description of Th.
is
deletus in fact Brues wrote
about the presence on the I ast two antenna I segments of tufts of long hairs: this character is recognizable only in Thaumatodryinus. In living
Thaumatodryinus, however, the tufts are on antennal segments 5-10. Probably
the three Palaearctic fossi I species are belonging
extinct genus of Thaumatodryininae,
to another
perhaps Harpactosphecion
Haupt 1944
(described by Haupt for one of the three above-mentioned species,
filicornis) .
Thaumatodryinus R.C.L. Perkins
Thaumatodryinus R.C.L. Perkins 1905: 58.
684
Mem. Amer.
En!.
Ins!.,
no.
37,
1984
Ilflll III/flilltilf
Fig. 456 -
Female of Nystrophorus tormicaeiormis Ruthe: tarsal
A:
chela;
B:
fore
segments 1-4.
Thaumatodryinus R.C.L. Perk i ns: Kieffer in Kieffer et Marshall 1906: 490.
Thaumatodryinus Thauma todryin us Thaumatodryinus Thaumatodryinus
TYPE SPECIES: FEMALE
R.C.L. R.C.L. R.C.L. R.C.L.
Perk i ns: Perkins: Perkins: Perkins:
Kieffer 1914b: 16. Krombein 1952: 97. Krombein 1958: 98. Krombein 1979: 1243.
Thaumatodryinus koebelei R.C.L. Perkins 1905, orig. desig.
(Figs 19, 457):
fully winged;
fore-wing with costal,
median and
submedian cells clearly enclosed by pigmented veins (Fig. 457); usually also first brachial cell clearly enclosed; maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front tarsus chelate; chela with rudimentary claw (Fig. 459); antennae with tufts of long hairs on the segments 5-10 (last segment with two tufts) (Figs 19, 471); pronotal tubercles reaching tegulae; occipital carina complete; mandibles with 4 teeth progressing
Thaumatodry i n i nae
685
/
Fig.
457 -
Female of Thaumatodryinus Guinea)
flavus n. sp. from Waris (New
686
Fig.
Mem.
458 -
Amer.
En!.
Ins!.,
no.
37,
1984
n.
sp.
from Kira-Kira
Male of Thaumatodryinus flavus
(Solomon
Islands)
larger from anterior one to posterior MALE
(Figs 28,
458):
fully
winged;
(Fig.
5);
fore-wing
tibial
submedian cells clearly enclosed by pigmented veins palpi
with 6 segments;
teeth progressing carina complete; pterostigma
(Fig.
DISTRIBUTION: fossil
labial
palpi
spurs 1,1,2.
with costal,
with 3 segments;
(Fig.
28);
larger from anterior one to posterior (Fig. fore-wing 28);
with metacarpus as
tibial
Ethiopian,
maxillary
mandibles with 4 5);
occipital
long as or longer than
spurs 1,1,2.
Oriental,
Nearctic,
Neotropic, Australian
Palaearctic species see note on page 683).
HOSTS: Flatidae GENERA: 1 SPECIES:
median and
17
ETH lOP I AN SPEC I ES OF THAUMATODRYINUS
Five species are known.
KEY TO THE SPECIES
(for
Thaumatodry i n inae
687
FEMALES
Fore and hind wings with a dark
transversal
band ••••••••••••••• 2
bands •.••.•••.•••• 3 2 Head mostly black; mesothorax, metathorax and propodeum black ••• • • • • • • • • • • • • • • • • • • • • • • • • • • •. 1. mirabilis n. sp. Fore and
-
hind wings without
dark
transversal
Head fully reddi sh-testaceous; mesothorax, metathorax and propodeum reddi sh-testaceous, wi th scutellum black ••••••.••••.••
· . . . . • . . . . . . . . • . . • • . . . . . . . . . 2. infuscatus n. sp. 3 Head wi th OPL longer than OL; thorax and propodeum partly black • . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3. dentatus Benoi t -
Head wi th OPL as
long
as or
short er
than OL;
thorax and
4 4 Occiput more strongly excavated; fore trochanters nearly as long as fore femora; posterior surface of propodeum wi th 2 complete propodeum fully
longitudinal
reddish-testaceous •••••••••••••••••.•••••••••••
keels;
notaulices more converging . . . . . . . . . . . . . . . . ..
· . . . . . . • . . . . . . . . . . . . . . . . . . .• -
Occiput
not
long as
fore
tracks
of
or weakly femora;
incomplete
townesi n.
4.
excavated; posterior
sp. trochanters nearly
fore
surface
longitudinal
of
keels;
0,5 as
propodeum only wi th notaulices
2
less
5. migratorius Beno i t
converg i ng . . . . . . . . . . . . . . . . . .
MALES
Only
the supposed male of
Th.
tOlvnesi n. sp. is known.
1. Thaumatodryinus mirabilis n. sp.
FEMALE:
fully
winged;
length 6,25-7,00 mm;
clypeus and mandibles yellow;
head black,
with malar space,
sometimes occiput partly brown-light;
antennae testaceous; pronotum and propectus yellow ; mesothorax, metathorax and propodeum black, with propodeal spiracles brown; black;
abdomen reddish;
legs testaceous,
petiole
with fore clubs of femora black,
mid and hind clubs of femora brown; antenna I segments in following proportions: 20: 7: 40: 45: 44: 45: 35: 25: 19: 15; head dull, reticulate rugose; hairy,
occipital
carina complete;
crossed by a
transversal
striae;
reticulate rugose;
transversal pronotal
OPL
notaul ices complete, metanotum smooth,
reticulate rugose;
fore-wing
part of radial
vein
tegulae;
with weak
scutum dull,
posteriorly separated; transversal
longer than
proximal
fully
scutellum
with some anterior furrows;
with a brown
much
pronotum shiny,
weakly punctate,
tubercles reaching
reticulate rugose; distal
longer than OL;
impression,
propodeum
band and with part;
radial
cell
688
Mem.
Fig.
459 -
Chela of
Amer.
Ent.
Inst.,
no.
37,
1984
Thaumatodryinus mirabilis n. sp. from Ankasoka (Madagascar)
closed;
pterostigma brown-I ight;
transversal
band;
segment
hind wings with a
1 of front
distal
tarsus nearly as
brown
long as segment 4;
Thauma todry i n i nae
689
(
Fig. 460 -
Chela of Thaumatodryinus infuscatus n.
sp.
(type)
segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 32 lamellae (Fig. 459); segment 5 of front tarsus (Fig. 459) with 2 rows of 59 lamellae; apex with a group of 5 lamellae; tibial MALE: unknown LOCUS TYPICUS: Ankasoka (Madagascar) TYPICAL MATERIAL: holotype F! and 1 paratype F! in P.
spurs 1,1,2.
690
Mem.
HOSTS:
Amer.
Ent.
Inst.,
no.
37,
1984
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Ankasoka
(Route Lakato),
P!
2. Tha umatodryinus infuscatus n. sp.
FEMALE:
fully
winged;
length 4,68 mm;
and scutellum black;
head shiny,
carina complete;
longer than OL;
POL
granu I a ted,
with weak
impression;
pronotal
rugose; radial
tubercles reaching
scutellum
fore-wing vein much
pronotum shiny,
transversa I stri ae,
sculptured by numerous separated;
reddish-testaceous,
weakly reticulate rugose;
longitudinal
tegulae;
and metanotum shiny,
with a
brown
longer than
weakly
transversa I
scutum shiny,
fully
notaulices complete, punctate;
transversal
proximal
occipital
hairy,
crossed by a
keels;
with petiole
posteriorly
propodeum reticulate
band and with distal
part;
radial
cell
part of
closed;
pterostigma brown-I ight; hind wings with a brown distal transversal band; segment 1 of front tarsus nearly as long as segment 4; segment 3 produced into a (Fig.
hook; 460);
enlarged claw with 2 sUbapical segment 5 of front
tarsus
lamellae;apex with a group of 6 MALE:
(Fig.
lamellae;
teeth and with 32 460)
lamellae
with 2 rows of 43
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
St.
TYPICAL MATERIAL:
Augustin
(Madagascar)
holotype F!
in BM.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
MADAGASCAR:
St.
Augustin
(Tulear),
BM!
FEMALE: fully winged; length 4,50 mm; head yellow-testaceous, with occiput partly black, ocellar region black, frons with 4 black spots
(two
3. Thaumatodryinus dentatus Benoit
Thaumatodryinus dentatus Benoit 1953b: 404.
in front of the antennal antennae testaceous, pronotum
sockets and two behind the antennal
with segments 1 and 4-10 dark;
yellow-testaceous,
testaceous,
with sides partly brown;
with anterior margin black;
between scutellum and scutum brown; and metanotum black; petiole black;
clubs of femora brown; 28:
33:
41:
incomplete, OPL
40:
28:
scutellum yellow-testaceous;
13: 9:
11;
with anterior third reddish;
legs testaceous,
head shiny, the ocell i
pronotum sh i ny,
furrow
sides of scutellum with fore and hind
antenna I segments in following proportions:
visible only behind
longer than OL;
propectus yellow; scutum yellow-
metanotum black;
propodeum black-brown,
abdomen brown-yellow;
sockets);
granulated; and on
granu I a ted,
occipital
15: 9:
carina
the sides of the vertex; ha i ry,
crossed by
a
Thaumatodryi n i nae
Fig.
461
-
Chela of
691
Thaumatodryinus dentatus n.
sp.
(type)
transversal impression; pronotal tubercles reaching tegulae; scutum shiny, hairy, granulated, with weak longitudinal striae; notaulices almost complete, sculpture;
posteriorly separated; metanotum rugose;
third almost smooth; distal closed;
part of radial
fore-wing
without dark
vein much
pterostigma brown-light;
than segment 4
(26:19);
with 2 subapical tarsus
scutellum shiny,
(Fig.
461)
transversal
longer than proximal segment
1 of front
segment 3 produced
teeth and with 36 with 2 rows of 63
apex; tibial spurs 1,1,2. MALE: unknown
smooth,
without
propodeum reticulate rugose,
into a
lamellae (Fig.
with anterior
bands and with
part;
radial
tarsus much hook; 461);
cell
longer
enlarged claw segment 5 of front
lamellae extending continuously
to the
692
Mem.
LOCUS TYPICUS:
Bekily
TYPICAL MATERIAL: HOSTS:
Amer.
Ent.
Ins!.,
37,
no.
1984
(Madagascar)
holotype F!
in P;
2 paratypes FF!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
P!
TV!
4. Thaumatodryinus townesi n. sp.
FEMALE: black;
fully
length 3,43-5,62 mm;
winged;
slender,
weakly distally thickened;
7: 5: 50: 40;
proportions:
complete; shiny,
6; OL
tubercles
smooth,
transversal
with 2 cell
smooth,
granulated,
logitudinal
keels;
=
8; OPL
tegulae;
lateral
scutum,
occipital
=
1,5; TL a
notaulices complete,
with a
carina
7; pronotum
transversal
scutellum
with
keels,
impression;
and
metanotum
very converging;
with some areolae or tracks of areolae, and posterior surface;
median
area dull,
without dark
closed and with distal
part;
weakly alutaceous,
crossed by
between dorsal
hind wings hyal ine,
=
3,5; OOL
reaching
keel
flat,
frons without median or
without sculpture;
propodeum dull, a
=
with petiole
antennae very
in front of the anterior ocellus;
without sculpture,
pronatal
shiny,
=
POL
testaceous,
segments 1-4 in following
antennal
head shiny,
occiput strongly excavated; short median furrow
fully
with segments 1-2 testaceous;
antennae brown,
part of radial
fore trochanters nearly as
smooth,
transversal long
with
posterior surface
granulated;
fore and
fore-wing
with radial
bands;
vein much
as fore femora
longer than proximal
(44:46) and nearly as
(44:46) and as fore tibiae (44:42); fore tarsal segments 16: 4: 8: 31: 41; enlarged claw (Fig. 462) with 2 subapical teeth and with 6 lamellae; segment 5 of front tarsus (Fig. 462) with a row of 9 lamellae; apex with a group of 10 lamellae; tibial spurs 1,1,2. MALE (not reared: from Sokoke Forest, Kenya): fully winged; length 2,68 long as fore coxae
in following proportions:
mm;
head black,
with mandibles testaceous;
segments 3-10 darkened; partly
testaceous
antennae testaceous,
prothorax testaceous;
thorax black,
(notaulices and region between
with
wi th scutum
the notaulices);
abdomen
brown; legs testaceous, antennae not distally thickened; antennal segments in following proportions: 6: 6: 9: 12: 11: 12: 11: 10: 8: 10; head dull, granulated and reticulate rugose; median
keel;
granulated, separated;
POL
=
weakly
6; OL
occipital
2; OOL
=
reticulate rugose;
reticulate rugose,
keels;
fore-wing hyaline,
radial
vein
longer than proximal
LOCUS TYPICUS:
Kampala
TYPICAL MATERIAL: unknown
part;
=
1; TL
=
smooth,
tibial
posteriorly
without sculpture;
transversal
transversal
frons without
4; scutum dull,
notaul ices complete,
without
without dark
carina complete;
6; OPL
scutellum and metanotum shiny,
propodeum dull,
in OL. HOSTS:
=
or
bands;
longitudinal distal
part of
spurs 1,1,2.
(Uganda)
holotype F!
and 1 paratype F!
in TW;
1 paratype F!
Thaumatodry in inae
Fig.
462 -
Chela of Thaumatodryinus
693
townesi n. sp. (type)
694
Mem. Amer.
Ent.
Inst.,
no.
37,1984
DISTRIBUTION: Ethiopian region: UGANDA: Kampala, TW! Forest, TW! SOUTH AFRICA: Tzaneen (Transvaal), TW! NOTES:
the species is named in honor of Mr.
KENYA:
Sokoke
Henry Townes.
5. Thaumatodryinus migratorius Benoit
Fig. 463 -
Chel a of Thaumatodryinus migratorius n.
Thaumatodryinus migratorius Benoi t 1953b: 404.
sp.
(type)
Thaumatodry i n i nae
FEMALE: black;
fully
winged;
ocellar region
length 3,5 mm; weakly brownish;
4-10 darkened;
antennal
22:
10: 8;
23:
20:
incomplete, OPL = OL; transversal
10:
reddish-testaceous,
with petiole
antennae testaceous,
segments in following proportions:
head shiny,
weakly punctate;
visible only behind the ocelli pronotum shiny, impression;
695
and on
weakly punctate,
pronotal
with segments 7:
occipital
5:
14:
15:
carina
the sides of the vertex;
hairy,
tubercles reaching
crossed by tegulae;
a
scutum shiny,
hairy, punctate, notaulices almost complete, posteriorly separated; scutellum and metanotum shiny, weakly punctate; propodeum reticulate rugose,
with anterior third almost smooth;
transversal proximal
bands and with distal
part;
radial cell
fore-wing
part of radial
closed;
pterostigma brown-light;
front tarsus nearly as long as segment 4;
teeth and with 21
segment 5 of front
463)
(Fig.
a group of 4 lamellae; MALE: unknown
tibial
LOCUS TYP I CUS:
(Madagascar)
Beki I y
TYPICAL MATERIAL:
segment 1 of
segment 3 produced
enlarged claw with 2 subapical tarsus
without dark
vein much longer than
lamellae
with 2 rows of 38
into a hook;
(Fig.
lamellae;
463); apex with
spurs 1,1,2.
holotype F!
in P.
HOSTS: unknown DISTRIBUTION: Ethiopian region: 'MADAGASCAR:
Bekily,
P!
THAUMATODRYINUS
OR I ENTAL SPEC I ES OF
Three species are known
KEY TO THE SPECIES
FEMALES
segment 3 less than 1,5 times as long as segment ........................... " 2. asiaticus n. sp.
Antennal -
Antennal
segment
3 more
2 Scutum with numerous
than
1,5
than POL . . . . . . . . . . . . . . . . . . . . . -
Scutum without
times as
longitudinal
longitudinal
1.
striae;
3.
segment
slightly
1 •••• 2
longer
philippinus n. sp.
striae;
long as POL .••••••••• ••••••••
long as OOL
POL more
than
malayanus n. sp.
MALES
three
times as
696
Mem.
One only male
Amer.
is known.
Ent.
It
Ins!.,
no.
37,
1984
is not possible to know
if it's a
philippinus or an asiaticus.
Fig.
464 -
Chela of
Thaumatodryinus philippinus n. sp. (type)
Thaumatodryi n i nae
697
1. Thaumatodryinus philippinus n. sp.
FEMALE:
fully
winged;
suture darkened; darkened;
length 4,80 mm;
legs yellow,
reddish-testaceous;
segment 3 more than
antennal
epistomal
with clubs of fore and hind femora 1,5 times as
long
as segment
(30: 19); head swollen, dull, granulated; frons with tracks of irregular striae; occipital carina complete; frons without median keel; POL = 9; OL = 4;
OOL = 12; OPL = 3; TL = 3; pronotum dull, hairy, granulated, strong transversal impression and with posterior tubercles reaching tegulae;
scutum granulated and with numerous
and areolae;
notaulices complete,
metanotum shiny,
granulated;
transversal
or
transversal
bands;
part 37;
(30:15);
longitudinal distal
fore tarsal
enlarged claw
23 41
lamellae; lamellae;
MALE:
(Fig.
unknown
(see key
to
Dumaguete
TYPICAL MATERIAL:
vein
without
without dark
twice as
long as proximal
in following proportions:
with 2 preapical
32:
9: 26:
5:
teeth and a row of nearly
the males) (Philippines)
holotype F!
in AM.
unknown
DISTRIBUTION: Negros
part of radial segments
hyal ine,
segment 5 of front tarsus (Fig. 464) with two rows of nearly apex with a group of 4 lamellae; tibial spurs 1,1,2.
LOCUS TYPICUS: HOSTS:
fore-wing
weak striae
scutellum and
propodeum reticulate rugose,
keels;
464)
longitudinal
posteriorly separated;
with a
Oriental
Island),
region:
PHILIPPINES:
Dumaguete
(Camp Lookout,
AM!
2. Thaumatodryinus asiaticus n. sp.
FEMALE:
fully
winged;
length 4,00-5,00 mm;
someti mes wi th brown spot on frons; testaceous;
propodeum yellow-testaceous,
yellow-testaceous;
head reddish-testaceous
antennae testaceous;
propectus
with sides partly brown;
scutum
scutell um and metanotum yellow-testaceous or reddish-
brown; propodeum fully or partly black; petiole black; abdomen reddish; legs yellow, sometimes with clubs of femora brown; head swollen, granulated,
dull,
antennal
segment 3 sl ightly
longer than segment 1 (23:20
or 24:18); occipital carina incomplete, visible only behind the ocelli and on the sides of the vertex; POL = 7; OL = 3,5; OOL = 9; OPL = 2; TL = 2,5;
propodeum shiny,
weakly
granulated,
impression;
pronotal
granulated;
notaulices complete,
almost
invisible;
reticulate rugose;
tubercles reaching
hairy,
tegulae;
crossed by a scutum dull,
posteriorly separated,
scutellum and metanotum shiny, fore-wing
with posterior parts
granulated;
without dark transversal
bands,
pterostigma brown dark and with distal
part of radial
than
segment 1 of front
as
proximal
long
part;
as segment 4
radial
cell
(24:22);
closed;
segment 3 produced
transversal
hairy, propodeum with
vein much
into a
longer
tarsus nearly hook;
enlarged
698
Mem.
Fig.
465 -
Chela of
claw with 2 subapical front
tarsus
lamellae; MALE
(Fig.
tibial
(supposed:
head testaceous,
Amer.
465)
Ent.
Inst.,
teeth and with 26
lamellae
see key
to the males):
fully
antennae slender,
proportions:
11: 6:
10:
granulated,
weakly
rugose;
OPl
2;
Tl
12:
=
4;
13:
(type)
segment 5 of
apex with a group of 5
13:
12:
abdomen and
12;
head hairy,
carina complete;
scutum hairy,
dull,
scutellum dull,
metanotum shiny,smooth,
with
legs
segments in following
10:
11:
length 2,40 mm;
antennae testaceous;
light;
antennal
occipital
posteriorly separated;
and punctate;
margin;
=
sp.
(Fig. 465);
winged;
with occiput and vertex brown; with scutum
7;
n.
spurs 1,1,2.
hairy;
=
1984
with 2 rows of 48 lamellae;
testaceous;
OOl
37,
Thaumatodryinus asiaticus
thorax and propodeum brown,
complete,
no.
POL = 7;
granulated; rugose,
dull, Ol = 5;
notaulices
reticulate rugose
some areolae near anterior
propodeum dull,fully reticulate rugose,
without transversal or
longitudinal keels; fore-wing hyaline, without dark transversal bands, hairy; pterostigma long and narrow; distal part of radial vein longer
Thaumatodry i n i nae
Fig.
than
466 -
proximal
Chela of Thaumatodryinus malayanus n.
part
LOCUS TYPICUS:
699
(19:14);
radial
Gunung Mulu Nat.
cell
open;
Park
tibial
(Sarawak,
sp.
(type)
spurs 1,1,2. Malaysia)
TYPICAL MATERIAL:
holotype F! in BM; 1 paratype F! in B; 1 para type F! 1 paratype M! in CO. HOSTS: unknown DISTRIBUTION: Oriental region: MALAYSIA: Gunung Mulu Nat. Park (Sarawak), BM! Lake Area (Bau, Sarawak), B! PHILIPPINES: Mt. Data, TW! N of Batu Batu (Tarawakan, Tawi Tawi), CO! in TW;
3.
Thaumatodryinus malayanus n. sp.
FEMALE: fully winged; length 4,60 mm; head testaceous, with ocellar triangle and a median frontal region brown; occiput and ventral side of head partly brown;
antennae brown,
with segments 1-3 testaceous and with
regions bearing tufts of hairs whitish; testaceous,
with a
lateral
propectus brown;
streak brown;
scutum,
pronotum
scutellum and metanotum
testaceous;
mesopleura and metapleura black,
testaceous; testaceous;
propodeum black, with anterior region of dorsal surface abdomen brown; legs testaceous, with cl ubs of femora brown;
antenna I segment 3 more than
1,5 times as long as segment 1 (39:20);
head swollen,
dull,
median keels;
POL = 3;
shiny,
without sculpture,
hairy,
and with a complete,
granulated;
transversal
with posterior apexes
occipital
carina complete;
OL = 1; OOL = 15; OPL = 3;
pronotum
with posterior tubercles reaching tegulae
impression;
posteriorly separated;
frons without
TL = 4;
scutum dull,
granulated;
notaulices
scutellum and metanotum smooth,
shiny,
700
Mem. Amer.
very weakly granulated; transversal or transversal part 53;
distal
fore
tarsal
enlarged claw
Ins!.,
no.
keels;
fore-wing
part of radial segments in
(Fig.
466)
with 2 preapical
segment 5 of front
apex with a group of 4
HOSTS:
hyaline,
vein much
nearly 38 lamellae;
Templer Park
TYPICAL MATERIAL:
1984
tarsus
(Selangor,
holotype F!
dull,
without
without dark
longer than proximal
following proportions:
nearly 54 lamellae; MALE: unknown LOCUS TYP ICUS:
37,
propodeum reticulate rugose,
longitudinal
bands;
(50:22);
En!.
38:
6:
13: 34:
teeth and with a row of (Fig.
466)
lamellae;
Malaya,
with
tibial
two rows of
spurs 1,1,2.
Malaysia)
in CO.
unknown
DISTRIBUTION: Malaya),
Oriental
region:
MALAYSIA:
Templer Park
(Selangor,
CO!
NEARCT I C SPEC I ES OF THAUMATODRYINUS
One species is known.
Thaumatodryinus perkinsi Krombein
Thaumatodryinus perkinsi Krombei n 1952: 99.
FEMALE:
fully
winged;
petiole black; occipital
carina
of the vertex; impression, shiny,
length 4,37-5,62 mm;
pterostigma brown incomplete,
granulated;
granulated; fore-wing
fully
yellow-testaceous,
head swollen,
shiny,
visible only behind the ocell i
with
granulated;
and on
the sides
= OL; pronotum sh i ny, ha i ry, crossed by a transversa I
OPL
pronotal
tubercles reaching
notaulices almost complete,
scutellum and metanotum shiny, rugose;
dark;
with distal
weakly punctate;
part of radial
tegulae;
scutum hairy,
posteriorly separated; propodeum reticulate
vein much
longer than
proximal part and without dark transversal bands; segment 1 of front tarsus longer than segment 4 (23:21); segment 3 produced into a hook; enlarged claw
with 2 subapical
segment 5 of front a group of 5 MALE:
lamellae;
(Fig.
tibial
teeth and with 467)
19 lamellae
(Fig.
with 2 rows of 28 lamellae;
467); apex with
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Westmoreland State Park
TYPICAL MATERIAL: HOSTS:
tarsus
holotype F!
(U.
S.
A.)
in WA.
unknown
DISTRIBUTION:
Nearctic region:
Westmoreland State Park (Oaxaca),
TE!
U.
S.
A.:
Greenville
(Westmoreland Co.,
Va.),
(S.
WAr
C.),
LE!
MEXICO:
Miltepec
Thaumatodryininae
Fig.
467 -
Chela of
701
Thaumatodryinus perkinsi n. sp.
(type)
Th. perkinsi Krombein is very similar to Th. koebelei R.C.L. Th. perkinsi, however, has pterost i gma brown dark and transversal impression of pronotum deeper. Th. perkinsi is also similar to Th. rufus Richards from Neotropic region. Th. rufus, NOTES:
Perk ins from Austral i an region.
however,
has transversa I
impression of pronotum
less deep.
702
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
THAUMATODRYINUS
NEOTROPIC SPECIES OF
Six species are known.
KEY TO THE SPECIES
FEMALES
-
Not a u I ice s
c omp let e •••••••••••••••••••••••••••••••••••••••••••• 2
Notaulices
incomplete ••••••••••••••••••••.•••••••••••••••••••.. 4
2 Antennal
segment
3
. . . . . . . . . . . . . . . . . .. -
Antennal
I ittle
longer
than
2. minimus n.
segment
1 •••••••••••••••••
sp.
segment
3 more than 1,5 times as long as segment 1 . . . . 3 long as breadth of ocell i; radial cell wide (Fig. 468 B} ••••••••••••••••••••••••.•••••••••••••••••.•••• . . . . . . . . . . . . . . . . . . . 1. variegatus De Santis and Vidal Sarmiento - Head wi th POL nearl y three times as long as breadth of ocel I i ; radial cell narrow (Fig. 468 A} . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . • . • • • • • • •• 6. clarus n. sp. 4 Segment 1 of front tarsus nearly as long as segment 4; head black or brown-b lack •••••••••••••••••••••••••••••••••••••••••••• . . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal Sarmiento - Segment 1 of front tarsus much longer than segment 4; head fully reddi sh or reddi sh-testaceous •••••••••••••••••••••••••••••••••• 5 5 Head with OL shorter than OPL ••••••••••••••••••••••••••••••••••• . . . . .. .. .. .. .. .. ... 4. rufus Richards - Head wi th OL IOr)ger than OPL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. bruchi De Santis and Vidal Sarmi en t 0
3 Head wi th POL nearly as
MALES
Antennal
segment
head with OOL
4
less
than
less
than
three
................... -
Antennal
segment
2 Head wi th OL as
5.
4 more
1,5
times as
times as
long as
bruchi De Santis and Vidal than
1,5
segment
times as
long as
Sarmiento segment
Head wi th OL
3 .... 2
long as OPL •••••••••••••••••••••••••••••••••••••
. . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal -
3;
long as OL . . . . . . . . . . . . . . .
longer
Sarmiento
than OPL ••••••••••••••••••••••••••••••••••••
4. rufus Ri chards
703
Thaumatodry i n i nae
\
\ \
\
\ \
\
\ \
\
:.:
...
Fig. 468 - Radial cells of females of Thaumatodryinus clarus n. sp. (type) (A) and variegatus De Santis and Vidal Sarmiento (type) (B)
The males of Th. n.
sp.
and minimus n.
variegatus De Santis and Vidal Sarmiento, clarus sp.
are unknown
1. Thaumatodryinus variegatus De Santis and Vidal Sarmiento
704
Mem.
Amer.
Ent.
I nst.,
no.
37,
1984
Thaumatodryinus variegatus De Santis and Vidal Sarmiento 1974: 25.
FEMALE:
fully
reddish;
antennae brown,
medial
winged;
length 7,18 mm; with joints,
region of segment 7,
head,
thorax and propodeum half of segments 5-6,
proximal
segments 8-10 testaceous;
petiole black;
abdomen reddish-testaceous; legs testaceous, with clubs of femora brown, distal half of tibiae black; antennal segment 3 more than 1,5 times as long as segment 1 (22:13); head dull, granulated; frons with a track of median keel;
occipital
POL = 5;
carina complete;
3; POL nearly as long as breadth of ocelli pronotum dull,
hairy,
with a strong
OL = 3;
(5:5);
transversal
OOL = 10;
temples
OPL
invisible;
impression,
with numerous
irregular keels; pronotal tubercles reaching tegulae; scutum shiny, granulated and with numerous irregular weak keels; notaul ices complete, posteriorly separated; reticulate rugose, wing
longitudinal
with first recurrent vein
fore-wing hyaline, vein
scutellum and metanotum shiny,
without
longer than distal
without dark transversal
longer than proximal
part
punctate;
propodeum
keels on posterior surface;
(30:12);
bands;
radial
fore-
part of discocubitus;
cell
distal open
part of radial (Fig.
468 B);
26: 3,5: 5,5: 19: 26; enlarged claw (Fig. 469) with 2 subapical teeth and with 26 lamellae; segment 5 of front tarsus (Fig. 469) with 2 rows of 53 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. fore tarsal
segments
MALE: unknown LOCUS TYPICUS:
in following proportions:
Dos de Mayo (Argentina)
TYP ICAL MATER IAL:
holotype F!
in PT.
HOSTS: unknown DISTRIBUTION: Neotropic region:
ARGENTINA:
Dos de Mayo (Misiones),
PT!
2. Thaumatodryinus minimus n. sp.
FEMALE:
fully
winged;
reddish-testaceous; black;
length 4,12 mm;
mesothorax;
head,
antennae and prothorax
metathorax and propodeum black;
abdomen brown-testaceous;
legs testaceous,
petiole
with tibiae and stalk of
fore femora brown; antenna I segment 3 sl i ght I y longer than segment 1 (11:9); read dull, granulated, with tracks of weak irregular striae, convex; frons without median and lateral keels; occipital carina complete; POL = 6; OL = 4; OOL = 10; OPL = 2; TL = 3; pronotum dull, granulated, with a strong scutum dull,
transversal granulated;
impression;
scutellum and metanotum dull,
distal open;
fore-wing
vein
segments
tegulae;
without sculpture among
without transversal
with a dark transversal
part of radial fore tarsal
tubercles reaching
posteriorly separated;
weakly punctate,
punctures or very reticulate rugose, keels;
pronotal
notaulices complete,
or longitudinal
band beneath the pterostigma;
longer than proximal
part
in following proportions:
(26:15); radial cell 23:
3:
7:
19:
30;
Thaumatodry i n i nae
Fig.
469 -
Chela of Thaumatodryinus variegatus De Santis and Vidal Sarmiento (type)
705
706
Mem.
Fig.
(Fig.
segment 5 of front
Ent.
470)
no.
with two subapical
tarsus
(Fig.
470)
apex with a group of 3 lamellae; MALE:
Ins!.,
37,
1984
Chela of Thaumatodryinus minimus n.
470 -
enlarged claw
Amer.
sp.
(type)
teeth and with 25
with 2 rows of nearly 44
tibial
lamellae; lamellae;
spurs 1,1,2.
unknown
LOCUS TYP I CUS: Tucuru i (Brazi I ) TYPICAL MATERIAL: holotype F! in TW HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
BRAZIL:
Tucurui
(Para),
TW!
3. Thaumatodryinus macilentus De Santis and Vidal Sarmiento
Thaumatodryinus macilentus De Santis and Vidal Sarmiento 1974: 24.
FEMALE: space,
fully
winged:
clypeus and
4-10 darkened;
length 6,25-6,87 mm;
ma~dibles
reddish;
brown;
antenna I tufts: incomplete, OPL
transversal tegulae;
petiole black;
with segments mesothorax,
abdomen reddish;
legs
with clubs of femora and partly clubs of hind femora black-
carina vertex;
with malar
propectus and pronotum redd ish-testaceous;
meta thorax and propodeum black; testaceous,
head black,
antennae testaceous,
fig.
471;
head shiny,
granulated,
visible only behind the ocelli
longer than OL; impression,
scutum hairy,
pronotum sh i ny,
weakly punctate; shiny,
reaching at
and metanotum shiny,
weakly punctate;
ha i ry,
pronotal
granulated,
notaulices incomplete,
least 0,5
and on
swollen;
occipital
the sides of the
crossed by
a
tubercles reaching
with numerous irregular keels; length of scutum;
scutellum
propodeum reticulate rugose;
fore-
T ha uma todryi n i nae
Fig.
471
707
Tuft of long hairs on antenna of female of Thaumatodry inus macilentus De Santis and Vidal Sarmiento from Brazi I -
708
Mem. Amer.
Fig.
472 -
En!.
Inst.,
no.
37,
1984
Chela of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia
(Brazil)
709
Thaumatodry in inae
Fig. 473 - Male genitalia of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia (Brazi I): right half removed.
wing with a brown transversal distal part of radial
vein much
band
(rarely band very
longer than proximal
light)
part;
and with
radial
cell
710
Mem.
Amer.
En!.
closed;
pterostigma brown dark;
bands;
segment 1 of front
produced into a hook; lamellae
(Fig.
lamellae; MALE:
472);
Ins!.,
no.
winged;
1984
hind wings without dark transversal
tarsus nearly as
long as segment 4;
enlarged claw with 2 subapical
segment 5 of front
tarsus
apex with a group of 6 lamellae;
fully
37,
length 4,87;
(Fig.
tibial
head reddish,
segment 3
teeth and with 40
472)
with 2 rows of 67
spurs 1,1,2.
with ocellar region black;
antennae testaceous, wi th segments 3-10 dark; thorax and propodeum reddish, with scutellum and metanotum black, scutum dark; abdomen brown,
with segment
tibiae and tarsi
1 testaceous;
brown;
in following
proportions:
granulated;
smooth;
carina complete; dull,
legs testaceous,
8:
frons
7:
13: 23:
19:
OL = 2;
OOL = 7;
notaulices complete,
distance between the notaulices nearly as smooth, without
19:
without median and
POL = 7;
fully granulated;
notau Ii ces and
with mid and hind
antennae not distally thickened;
the paraps i da I furrows;
17:
18:
lateral
antennal
14:
17;
keels;
OPL = 2;
segments
head,
fully
occipital
TL = 4;
scutum
posteriorly separated;
minimum
long as distance between
the
scutell um and metanotum sh i ny,
without sculpture, finely hairy; propodeum reticulate rugose, dull, transversal or longitudinal keels; fore-wing hyaline, without dark
transversal bands; distal part of radial vein longer than proximal (36:21); genital ia: fig. 473; tibial spurs 1,1,2. LOCUS TYP ICUS:
Loreto (Argentina)
TYPICAL MATERIAL: HOSTS:
part
holotype F!
and
1 paratype F!
in PT.
unknown
DISTRIBUTION: ECUADOR:
Neotropic region:
Napo
(Muyuna,
Tena),
PANAMA: BM!
Barro Colorado
Island,
SURINAME: Sumatraweg
LE!
(Lelydorp),
OL! BRAZIL: Campsite (Mato Grosso), BM! Jatai (Goias), TW! OT! Nova Teutonia (S.ta Catarina), BM! 12° 50' S 51° 47' W (Mato Grosso), BM! OL! ARGENT INA:
Loreto
(Misiones),
PT!
4. Thaumatodryinus rufus Richards
Thaumatodryinus rufus Richards 1951: 815. Thaumatodryinus rufus Richards: Krombein 1952: 99.
FEMALE:
fully
winged;
length 3,0-4,0 mm;
reddish-testaceous,
with petiole
black, clubs of fore and hind femora brown; antennae testaceous, with segments 4-10 darkened; mesothorax, metathorax and propodeum reddish dark; antennal segment 3 more than 1,5 times as long as segment 1 (25:14); head swollen, shiny, granulated, with frontal weak longitudinal striae;
occipital
carina
incomplete,
the sides of the vertex; shiny,
hairy,
pronotal numerous
crossed by
POL = 4; a
tubercles reaching
visible only benhind the ocell i and on
OL = 2;
transversal tegulae;
irregular weak striae;
OOL = 8,5;
impression,
scutum hairy,
OPL = 3;
pronotum
weakly punctate; shiny,
notaul ices incomplete,
granulated,
reaching
at
with
least
Thaumatodry i n i nae
711
\
Fig.
474 -
Chela of Thaumatodryinus rufus Richards from St. Augustine (Trinidad)
0,5 length of scutum; scutellum and metanotum hairy, shiny, weakly punctate; propodeum reticulate rugose; fore wings with a brown transversal band and with distal part of radial vein much longer than proximal part; radial cell closed; pterostigma brown dark; hind wings without dark transversal bands; segment 1 of front tarsus longer than segment 4 (27:21); segment 3 produced into a hook; enlarged claw (Fig. 474) with 2 subapical teeth and with 29 lamellae; segment 5 of front tarsus (Fig. 474) with 2 rows of 45 lamellae; apex with a group of 5 lamellae; MALE:
tibial
fully
spurs 1,1,2.
winged;
length 2,50-2,87 mm;
head reddish-testaceous,
with
712
Mem. Amer.
Fig.
475 -
Ent.
Ins!.,
no.
37,
1984
Male genitalia of Thaumatodryinus rufus Richards from St. Augustine
(Trinidad):
right
half removed.
ocellar region black;
antennae testaceous;
with scutellum black,
metanotum and propodeum dark;
testaceous;
testaceous;
legs fully
thorax and propodeum reddish, abdomen brown-
antennae not distally thickened;
antennal 11: 10: 9: 9: 7: 10; head dull, fully granulated; frons without median and lateral keels; occipital carina complete; POL = 5; Ol = 2; OOl = 5; OPl = 1,5; Tl = 2; scutum dull, granulated; notaulices complete, posteriorly separated;minimum distance between the notaulices nearly as long as distance between the notaulices and the parapsidal furrows; scutellum and metanotum shiny, smooth, weak I y punctate, without SCu I pture among punctures; propodeum reticulate rugose, without transversal and longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (19:12); genitalia: fig. 475; tibial spurs 1,1,2. lOCUS TYPICUS: St. Augustine (Trinidad) segments in following proportions: 7:
5:
8:
14:
Thaumatodry i n i nae
Fig. 476 -
713
Chela of Thaumatodryinus bruchi De Santis and Vidal Sarmiento from Las Cesas
(Argentina)
TYPICAL MATERIAL: holotype F! in BM. HOSTS: in Trinidad Ormenis sp. (Richards 1951). DISTRIBUTION: Neotropic region: TRINIDAD: 51. Augustine (51. BM!
Curepe
(Sta.
Margarita,
Circular Rd.),
OT!
OLl
George),
OL!
714
Mem.
Amer.
En!.
Inst.,
no.
37, 1984
5. Thaumatodryinus bruchi De Santis and Vidal Sarmiento
Thaumatodryinus bruchi De Santis and Vidal Sarmiento 1974: 22.
FEMALE:
fully
winged;
length 6,56-8,62 mm;
head reddish;
antennae
testaceous, wi th segments 3-10 darkened; propectus and pronotum redd i sh; thorax (except for scutum which is reddish) and propodeum reddish-brown or black, reddish;
sometimes with
irregular brown spots;
legs testaceous,
clubs of hind femora brown-black; as
long as segment 1 (38:20);
longitudinal behind = 13;
keels on frons;
the ocelli OPL = 2;
impression,
and on
petiole black;
abdomen
with clubs of fore femora and distal
antenna I segment 3 more than
head swollen,
occipital
carina
dull,
granulated,
weakly punctate;
sh i ny,
pronotal
1,5 times
with some
incomplete, only visible
the sides of the vertex;
pronotum ha i ry,
part of
POL = 5;
crossed by
OL = 3,5;
OOL
a transversa I
tubercles reaching tegulae;
scutum
hairy, dull, reaching at least 0,5 length of scutum; scutellum shiny, smooth; metanotum with lateral keels; propodeum reticulate rugose forewing with a brown transversal much
longer than proximal
dark;
band and with distal part of radial
part;
radial
cell
hind wings without dark transversal
closed; bands;
vein
pterostigma brown segment
1 of front
(37:30); segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 29 lamellae (Fig. 476); segment 5 of front tarsus (Fig. 476) with 2 rows of 44 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: fully winged; length 3,37 mm; head reddish-testaceous; antennae brown, with segments 1-2 testaceous; ocellar region usually dark; tarsus
longer than segment 4
prothorax reddish-testaceous; brownish;
scutellum,
propodeum testaceous; testaceous,
scutum reddish-testaceous,
metanotum,
petiole black;
darkened;
head dull,
granulated;
7: 7:
11 :16:
legs
mid and hind tibiae,
antennae not distally thickened;
segments in following proportions:
sometimes
abdomen brown-testaceous;
with mid and hind stalks of femora,
and hind tarsi
with sides
propodeum and pleura black;
mid
antennal
14: 14: 12: 12: 11: 14;
frons without median and
lateral
keels;
occipital
carina complete; POL = 7; OL = 3; OOL = 6; OPL = 1,5; TL = 4; scutum dull, granulated; notaul ices complete, posteriorly separated; scutellum and metanotum shiny, punctate, very weakly granulated; propodeum reticulate rugose, without transversal or longitudinal keels; fore-wing hyaline, without dark transversal proximal
part
bands;
LOCUS TYPICUS:
Alta Gracia
TYPICAL MATERIAL:
(Roraima),
Horco Molle
part of radial
vein
longer than
(Argentina)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: Surumu
distal
(26-15); genitalia: fig. 477; tibial spurs 1,1,2. and 1 paratype F! BRAZIL:
in PT.
Encruzilhada (Bahia),
OL!
TW!
ARGENTINA:
Las Cejas
(Tucuman),
TU!
Alta Gracia
(Cordoba),
(Tucuman), PT!
TW! OL!
La Paz
TU!
(Cordoba),
715
Thaumatodry i n i nae
'/
Fig.
477 -
Male genital ia of Thaumatodryinus
Sarmiento from La Plata
PT!
La Plata,
TW!
(Argentina):
bruchi De Santis and Vidal right half removed
OU
6. Thaumatodryinus clarus n. sp.
716
Mem. Amer. En!.
Ins!., no. 37,
1984
Fig. 478 - Chela of Thaumatodryinus clarus n. sp.
(type)
Thaumatodry i n i nae
FEMALE:
fully
winged;
length 4,52 mm;
testaceous,
antennae with segment 3 more than 1,5 times as head dull, complete; times as strong dull,
granulated; POL
= 9;
OL
impression;
granulated;
=
or longitudinal
transversal
bands;
proportions:
cell:
radial fig.
= 3;
OPL
(9:3);
keels;
occipital carina
= 5;
POL nearly three
pronotum hairy,
dull,
with a
tegulae;
posteriorly separated;
fore tarsal
hyaline,
scutum
scutellum and
propodeum reticulate rugose,
fore-wing
vein with distal
468 A;
TL
posterior tubercles reaching
weakly granulated;
transversal radial
12;
notaulices complete,
metanotum shiny,
part;
OOL
with petiole black;
long as segment 1 (25: 15);
frons without median keel;
= 3;
long as breadth of ocelli
transversal
717
without
without dark
part much longer than proximal segments in following
5: 12: 28: 43; enlarged claw (Fig. 478) with 2 subapical segment 5 of front tarsus (Fig. 478)
3:
teeth and with nearly 24 lamellae;
with two rows of nearly 409 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Yepocapa (Guatemala) TYPICAL MATERIAL: holotype F! in AM. HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
Chimaltenango),
GUATEMALA:
Yepocapa
(Dep.
AM!
AUSTRAL I AN SPEC I ES OF THAUMATODRYINUS
Two species are known
KEY TO THE SPECIES
FEMALES
Fore-wing with
shorter than proximal 1. flavus n. sp. Fore-wing with first recurrent vein longer than proximal part of cubi tus (Fig. 479 A) ••••••••••••• 2. koebelei R.C.L. Perkins
part -
of
cubitus
first
(Fig.
recurrent
vein much
479 B) •••••
MALES
-
brown . . . . . . . . . . . . . . . flavus n. sp.
Head,
thorax and propodeum most Iy or 1.
fully
Head,
thorax and propodeum mostly
fully
or
testaceous ••••••••••
718
Mem.
Fig.
479 -
Nadzab,
Amer.
Ent.
Ins!.,
no.
37,
1984
Females of Thaumatodryinus koebelei R.C.L. and flavus
New Guinea) (A)
CD = first recurrent vein;
n.
sp.
Perkins
(from Waris,
DE = proximal
(from
New Guinea) (B):
part of cubitus.
1. Thaumatodryinus flavus n. sp.
FEMALE
(Figs 19, 457):
yellow-testaceous,
fully winged;
clypeus and mandibles yellow; 5-10 darkened;
length 6,30-6,87 mm;
head brown or
with ocellar spot brown and occiput partly brown: antennae testaceous,
propectus black or brown;
or with sides and partly anterior collar brown; yellow-testaceous or brown-b lack;
with segments 4-8 or
pronotum fu II y
yellow-testaceous
scutum and scutellum
metanotum brown-reddish or brown-b lack;
propodeum and pet iole brown-b I ack or black;
abdomen fu II y
redd ish or
with tergite 1 partly black;
legs yellow-testaceous or with fore clubs of
femora brown;
dull,
head swollen,
visible only behind the ocelli than OL; weakly shiny,
pronotum shiny,
granulated; granulated;
hairy,
pronotal
granulated; occipital
and on
carina
the sides of the vertex;
crossed by
a transverse
tubercles reaching tegulae;
notaul ices complete,
incomplete, OPL
longer
impression,
scutum hairy,
posteriorly separated;
scutellum
and metanotum shiny, weakly punctate; propodeum reticulate rugose; forewing with first recurrent vein much shorter than proximal part of cubitus (Fig.
479 B);
fore-wing
without dark transversal
brown dark and with distal part of radial part;
radial
cell
closed;
bands,
with pterostigma
vein much longer than proximal
segment 1 of front
tarsus
longer than segment 4
(35:32); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 480) with 2 subapical teeth and with 38-41 lamellae; segment 5 of front
tarsus
(Fig.
of 5-6 lamellae; MALE
480)
tibial
(Figs 28,458):
with 2 rows of 55-57
fully
a more or less wide frontal testaceous;
lamellae;
apex with a group
spurs 1,1,2. winged;
length 1,56-2,87 mm;
area testaceous;
occiput testaceous;
thorax and propodeum brown;
antennae brown, abdomen brown,
head brown,
with
clypeus and mandibles with segment 1 testaceous; wi th anterior region
T haumatodryi n i nae
Fig.
480 -
Chela of Thaumatodryinus
719
flavus n. sp. (type)
testaceous; legs testaceous; antennae not distally thickened; antennal segments in following proportions: 6: 6: 10: 13: 13: 13: 13: 11: 9: 10; head dull, hairy, granulated; frons without median and lateral keels occipital carina complete; POL = 5; OL = 2,5; OOL = 6; OPL = 1; TL = 3; scutum dull,hairy,
granulated;
min imum distance between
notaulices complete,
posteriorly separated;
the notau I ices of scutum shorter than distance
720
Mem.
Fig.
481
-
Islands) (A);
the notaulices and
metanotum shiny,
smooth,
distal
B:
Ins!.,
no.
1984
flavus n. sp. from Kira-Kira
furrows;
scutellum and
without sculpture among punctures;
without
transversal
posterior surface without
without dark transversal
37,
gonoforceps laterally viewed.
the parapsidal
punctate,
propodeum reticulate rugose, posterior surface; hyaline,
En!.
Male genitalia of Thaumatodryinus (Solomon
between
Amer.
bands;
keel
between dorsal
longitudinal
keels;
pterostigma lanceolate,
part almost reaching apex of radial
vein;
radial
and
fore-wing with
vein with distal
part longer than proximal part (29:13), almost reaching margin of wing; gonoforceps (Fig. 481) without proximal or distal processes, laterally viewed with a medial lOCUS TYPICUS:
TYPICAL MATERIAL: MM!
in TW;
prominence;
Tangtalau
holotype F!
6 para types
tibial
(Solomon
(1
F,
spurs 1,1,2.
Islands)
and 5 paratypes MM! 5 MM)!
in B;
4 paratypes
in Ol.
HOSTS: unknown DISTRIBUTION: Australian
region: NEW GUINEA: Wau, TW! all Jimmi Valley, all TW! Dumpu, TW! Waris (Hollandia, Western Irian), Ol! SOLOMON ISLANDS: B!
Ol!
Tangtalau
(Malaita
Island),
B!
Kira-Kira
(San Cristobal
Island),
Thaumatodryininae
Fig.
482 -
721
Chela of Thaumatodryinus koebelei R.C.L. Namatasopa
Perkins from
(New Hebrides)
2. Thaumatodryinus koebelei R.C.L. Perkins
Thaumatodryinus koebelei R.C.L. Perkins 1905: 58. Thaumatodryinus koebelei R.C.L. Perkins: Krombein 1952: 99.
FEMALE:
fully
testaceous,
winged;
length 3,47-6,00 mm;
lectotype fully
reddish-
with petiole black and antenna I segments 5-10 darkened;
other specimens head with ocellar spot brown, brown,
spots on si des of pronotum,
brown;
si des of scu tum brown ish;
transversal
in
impression
furrow between scutum and scutell um
furrow between scutell um and metanotum
722
Fig.
Mem.
483 -
Amer.
Ins!.,
no.
37,
1984
Male genital ia of Thaumatodryinus koebelei R.C.l. Nadzab
brown;
Ent.
(New Guinea):
right
propodeum brown or black abdomen brown;
femora brown;
head dull,
granulated,
Perkins from
half removed
swollen;
legs with clubs of fore
occipital
carina
incomplete,
visible only behind the ocelli and on the sides of the vertex; OPl longer than Ol; pronotum not humped, crossed by a transversal impression, hairy,
granulated;
pronotal
tubercles reaching
tegulae;
scutum dull,
hairy, granulated; notaulices incomplete (reaching at least 0,5 length of scutum) or complete (in a specimen from New Britain); scutellum and metanotum shiny,
weakly
punctate;
propodeum reticulate rugose fore-wing
with first recurrent vein longer than proximal part of cubitus (Fig. 479 A); fore-wing without dark transversal bands, with pterostigma yellow dark or brown and with distal part of radial vein much longer than proximal part; radial cell closed; segment 1 of front tarsus slightly longer than segment 4
(27:22);
with 2 subapical tarsus
(Fig.
lamellae;
482)
tibial
segment 3 produced
teeth and with 30 with 2 rows of 48
into a
lamellae lamellae;
(Fig.
hook; 482);
enlarged claw segment 5 of front
apex with a
group of 5
spurs 1,1,2.
MALE:
fully
brown
and antennal
winged;
length 2,25-3,62 mm;
testaceous,
segments 3-10 darkened;
with ocellar region
antennae not distally
I
Dry i n i nae thickened; 17:
16:
antennal
14:
12:
13;
segments in following proportions: head dull,
frons without median and 7;
=
OL
3;
=
OOL
7;
lateral
=
OPL
2;
keels;
=
TL
4;
keels;
fore-wing hyaline, vein
6:
10:
occipital
13:
carina complete;
scutum dull,
granulated;
scutellum and metanotum shiny,
propodeum reticulate rugose,
radial 1,1,2.
6:
15:
granulated and weakly reticulate rugose;
complete posteriorly separated; punctate;
723
without transversal
without dark transversal
longer than proximal
part;
genital ia:
bands; fig.
POL =
notaulices strongly
or longitudinal distal
483;
part of
tibial
spurs
LOCUS TYPICUS: Cairns (Australia) TYPICAL MATERIAL: lectotype F! in B. HOSTS:
in Australia
DISTRIBUTION: GUINEA:
Siphanta sp.
Australian
BUlolo, OL!
Lae-Zenag Road,
TW!
TW!
(R.C.L.
region: Bogia
(Om.
Busu R.
Perkins 1906c).
AUSTRALIA:
Cairns
New Guinea),
(Lae),
OL!
TW!
SUBFAMILY
Nadzab,
NEW HEBRIDES:
(Anelgaunat, Aneityum Island), BM! Namatasopa OL! NEW BRITAIN: Vunabakan (Keravat), B!
7.
(Queensland),
TW!
B!
TW!
NEW
OL!
Red Crest
(Espiritu Santo
Island),
DRYININAE KIEFFER
Dryininae partim Kieffer in Kieffer et Marshall 1906: 495. Dryininae partim Kieffer 1907b: 3. Lestodryinini partim Kieffer 1914b: 11. Dryinini partim Richards 1939: 189. Dryinini partim Muesebeck and Walkley 1951: 1034. Dryininae Richards 1953: 51. Dryininae partim Krombein 1979: 1243.
TYPE GENUS: FEMALE
Dryinus Latreille 1804
(Figs 495,
wing with costal,
I
pigmented veins palpi
529,
611,
632,
(Figs 611,
with 3 segments; front
612);
spurs 1,1,2 or 1,1,1MALE (Figs 531, 532, 613,
posterior veins
labial
occipital
fully
winged;
396 B);
with 6 segments;
fore-
lamella;
634,
palpi
segment 5 of front
680,
pronotal
705):
fully
with 3 segments;
dorsal
incomplete
(Fig.
tarsus
winged:
485);
(Fig.
tubercles present;
occipital
labial
larger from
carina complete or
mandibles with 3 teeth progressing
(Fig.
with costal,
703):
chela with rudimentary claw
long hairs;
never with one only preapical
incomplete;
678,
maxillary palpi
5);
tarsus chelate;
antennae without tufts of
with 6 segments;
662,
mandibles with 4 teeth progressing
anterior one to posterior (Fig. or absent;
659,
median and submedian cells clearly enclosed by
485)
tibial
maxi Ilary palpi carina complete or
larger from anterior one to
process of gonoforceps absent;
fore-wing
median and submedian cells clearly enclosed by pigmented
(Figs 532,613);
tibial
spurs 1,1,2.
724
Mem.
DISTRIBUTION:
Amer.
Ent.
Inst.,
no.
37,
1984
worldwide
HOSTS: Dictyopharidae,
Cixiidae, Lophopidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Tropiduchidae, Fulgoridae. GENERA:
8
SPEC I ES:
149
I<EY TO THE GENERA OF DRYININAE the fossi I genus Cretodryinus Ponomarenko)
(excluding
FEMALES
Enlarged
659, -
Enlarged
611,
claw more
than
three
660) . . . . . . . . . . . . . . . . . . . 632,
claw as
662,
long as
front
tibia
(Figs
Megadryinus Richards
or shorter than front tibia (Figs 528, 703) . . • . . . . • . . • . • • • • . • • . . • . • • . • • . . . . . . . . . . . 2
long as
678,
2 Enlarged claw very than arol ium
times as
4.
(Fig.
reduced,
nearly as
711) •••••
7.
long as or
sl ightly
longer
Perodryinus R.C.L. Perkins
reduced, much longer than arolium (Fig. 485).3 1 , 1 , 1 . . . . . . . . . . 2. Richardsidryinus Mocza r - Tibial spurs 1,1,2 •••••..••••••••.•••••••••••••.•••••••••.•.••. 4 4 Enlarged claw with one subapical tooth (Fig. 485) . . . . . . . . . . . . . . 5 -
Enlarged
3 Tib ia I
-
claw not
spu rs
Enlarged claw without teeth near apex (Figs
subapical
tooth
or with a
series of
small
669,691) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
5 Notaulices at least partly visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. Dryinus Lat rei I Ie Notaul ices invisible! . . . . . . 3. Tridryinus Kieffer 6 Notaulices at
least
partly
visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
. . . . • . . . . . • . . . • • . . . . . . • . • . .. 5. Alphadryinus n. g en. - Notaul ices invisible 1 . . . . . . 6. Mesodryinus Kieffer
NOTES:
the key doesn I t
inc I ude the fossi I genus Cretodryinus Ponomarenko,
because the only known specimen
is in
too bad conditions for
distinguishing generic differences.
! If the scutum
is sculptured by
longitudinal
keels,
two of the furrows
del imitated by the keels are considered notaul ices and the species is included in Dryinus Latrei lie or Alphadryinus n. gen. In some species with scutum reticulate rugose or smooth two keels at first sight can seem notaulices. I have considered, however, the notaulices invisible, the true notau I ices are furrows and not keel s.
because
Dryininae
725
MALES
I
Dryininae. The
was not able to find generic differences in males of
number of known males is too small
for drawing conclusions.
In addition
perhaps some males were associated by mistake with some genera. For the above mentioned reasons keys to males of species living
I think that actually only general
in a zoogeographic region are proposable,
as follows.
DRYININAE
KEY TO THE MALES OF THE KNOWN PALAEARCTIC
Scutum strongly as -
reticulate
rugose;
numerous areolae are as wide
ocelli •.•••••••••••••••••••••••••••••••••••••••••••••••••••• 2
Scutum granulated or weakly than
reticulate
rugose;
areolae
less wide
ocelli •••••••••••••••••••••••••••••••••••••••••••••••••••• 3
2 Posteri or
ocell i
not
areolae as wide as longer
than
touching
those of
segment
4,
less
the
occipi tal
scutum; than
carina;
antennal
three
times as
2; gonoforceps slightly shorter than penis
head wi th
3 slightly
segment
(Fig.
long as
segment
686) ••••••..••
· ............••.....•.••...•••. Mesodryinus niger (K i ef f er) -
Posterior ocell i areolae much approximately
touching
smaller
than
twice as
the
occipi tal
those of
long as
carina;
scutum;
segment
head wi th
antennal
segment
3
4, approximately four
long as segment 2; gonoforceps much shorter than penis (F i g. 684) •••••••.•••••••.••••• Mesodryinus dayi n. sp. 3 Fore wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) ••••• Dryinus tarraconensis Marshall times as
-
Fore wing wi th distal
part of radial vein much longer than 490 A) ••••••••••••••••••••••••••••••••••••• 4 4 Notaulices reaching approximately 0,5 length of scutum •••••.••• 5 - Notaulices reaching approximately 0,75 length of scutum •••••••• 7 5 Distivolsella farther from apex of penis (Fig. 491) . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . Dryinus collaris (Linnaeus) - Distivolsella nearer apex of penis (Figs 501,618) . . . . . . . . . . . . . 6 6 Head more or less retiCUlate rugose ••••••.••.••••••••••••••••••• proximal
part
(Fig.
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Richardsidryinus corsicus (Ma rsha I I ) -
Head granulated,
not
reticulate
rugose ••.•••••••••••••••••••••••
· .•.........•...•.•..•.•..•..•. Dryinus koreanus (Moe za r) 7 Antennal
segment
3 nearly
twice as
long as
segment
2 . . . . . . . . . . ..
· ...•.••..•.••• , •.••.•.•....... Dryinus canariensis (Ceba I los) -
Antennal
segment
3 more
than
twice as
long as
segment
· •..........•..•..•............ Dryinus nepalensis n. sp.
2 .........
726
Mem.
NOTES: sex of
Amer.
Ent.
Ins!.,
no.
37,
1984
the male associated with D. canariensis could be also the opposite Richardsidryinus albrechti n. sp. On I y breed i ng experiments can
answer the question.
Dryinus sanderi n. sp., Nichardsidryinus maroccanus n. R. albrechti n. sp. (for the last species, however, see note
The males of sp.
and
above)
are unknown.
The fossil
Dryinus antiquus (Ponomarenko) was not included
species
into the key because the
type
(only known specimen)
is in
too bad
conditions for distinguishing differences with other males of Palaearctic
Dryininae.
KEY TO THE MALES OF THE KNOWN ETH I OP IAN
Notaulices
invisible;
scutum strongly
DRYININAE
reticulate
rugose,
with
numerous areolae as wide as
· . . . . . • . . . . . . . j\1esodryinus -
ocelly . . . . . . . . . . . . . . . . . . . . .. ethiopicus n. sp.
Notaulices visible; scutum granulated, not reticulate rugose; sometimes more or less weakly reticulate rugose, but in this case areolae much
less wide
than
ocel Ii •••••••••.•••••••••••••• 2
2 Occipital carina complete; dorsal part of occipital carina not touching eyes ••••••.••••••••••••••••••••••••••••.•••.••••.••••• 3 Occipital carina incomplete; dorsal part of occipital carina usually touching eyes ••••••••••••••••.•••••••.••••.•••••••••••• 4 3 Ninth abdominal sternite with apex rounded (Fig. 623} . . . . . . . . . . · . . . . . . . . . . . . . Richardsidryinus erraticus (Tu rn e r) - Ninth abdominal sterni te wi th apex emarginated . . . . . . . . . . . . .
· .....•....•.. Dryinus pretorianus n. 4 Head wi th OL
sp.
= OOl . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
- Head wi th Ol more or less shorter than OOl ••••••. '" .•••••••••• 6 5 Fore-wing with radial cell closed; head with posterior ocelli not
touching
occipi tal
carina ••••••••••••••••••••••••••••••
••........•.• . Dryinus mahensis (Ki effer) -
Fore-wing with touching
radial
occipi tal
cell
open;
head with posterior ocelli
carina • . . • • • . • • • • . . • • •
· ...•....•••.. Dryinus saussurei (C eba I I os) 6 Fore-wing
wi th
radial
cell
open . . . . . . . . . .
. . . . . . • . . . . . . . Dryinus orophilus (Benoi t) -
Fore-wing with
radial
7 Frons with a median
cell
closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
longitudinal
hump
from anterior ocellus
to
antennal
sockets ••••••••••••••.••••••••••••. . . . . . . . . . . . . . . Dryinus deceptor (Turner)
-
Frons
flat
in
the antennal 8 Breadth
of
front sockets
of
the anterior ocellus,
only
convex between
•..•.••••.••••.••••••••••••••••••••.••... 8
anterior ocellus
longer
than POL . . . . . . . . . . . . . . . . . . ..
.•....•••....• Lestodryinus luweli Benoi t
(see
incertae sedis nomi na)
Dryininae
- Breadth of
anterior ocellus
727
shorter
than POL . . . . . . . . . . . . . . . . . . ..
.......•• . Dryinus afer (Olmi)
KEY TO THE MALES OF THE KNOWN OR lENT AL DRYININAE
Dorsal
part
of
occipi tal
carina
touching
eyes •••••••••••••••••••
. . . . . . . . . . Dryinus indicus (Kieffer) -
Dorsal
part
of
2 Head wi th TL
occipital
shorter
carina not
than breadth
touching
of
eyes . . . . . . . . . . . . . . 2
ocel Ii •••••••••••••••••••••
· .•...•... Dryinus browni Ashmead -
Head with TL as
long as breadth
of
ocel Ii •••••••••••••••••••••••
· . . . . . . . . . Richardsidryinus pyrillae (K i ef fer)
KEY TO THE MALES OF THE KNOWN ~JEARCT I C DRYININAE
Head with breadth
of
ocelli
more
· •........ Tridryinus testaceus n. 2 Head wi th
ocell i
than
twice as
long as OPL . . . . . .
sp.
sl ight Iy broader
than OPL . . . . . . . . . . . . . . . . . . . . ..
· .......•. Perodryinus amoenus R. C • L. Perk ins
The male of Dryinus alatus check again
the
only known
(Cresson)
is known:
I,
however,
must
specimen.
KEY TO THE MALES OF THE KNOWN NEOTROP IC DRYININAE
Notaulices -
invisible •••••••••••••••••.••••••••••••••••••••••••• 2
Notaulices at
least
2 Posterior ocelli
partly
touching
visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 occipital
carina . . . . . . . . . . . . . . . . . . . . . .
· . . . . . . . . . Tridryinus ruficauda R i cha rd s Posterior ocell i
not
touching
occipital
carina . . . . . . . . . . . . . . . . . .
· . . . . . . . . . Tridryinus poecilopterae Richards 3
Scutum fully granulated,
not
reticulate
rugose •.•••••.••••••••••
· .......•. Dryinus antilleanus (Evan s ) -
Scutum fully
reticulate
4 Head with OPL
less
than
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 0,5 as
long as breadth
of
ocelli . . . . . . ..
· . . . . . . . . . Dryinus striatus (F en t on) -
Head
wi th OPL
5 Distal
part
of
sl ight Iy radial
shorter vein
than breadth
longer
than
· ....•.... Tridryinus striaticeps (K i ef fer)
of
proximal
ocell i . . . . . . . . . . 5 part . . . . . . . . . . . .
728
Mem. Amer. Ent.
Distal
part
of
radial
Ins!.,
no. 37, 1984
vein shorter than proximal
part ••••••.••••
. • • . . . • . . • • . . . . • . • • • • • . • • • • . • Dryinus surinamensis n. sp.
KEY TO THE MALES OF THE KNOWN AUSTRALIAN
DRYININAE
Notaul ices complete .•...•.... Dryinus areolatus n. sp. - Not a u I ice s i n c omp let e •••••••••••••••••••••••••.•••••.•••.••.•.• 2 2 Head granulated, not reticulate rugose ••••.••..•••••.••.••.•.••. •..•.•...•..•......••........ Dryinus pallidus (R.C.L. Perkins) - Head more or less reticulate rugose •••••••.••••••.•.•.••••..•• 3 3 Dorsal surface of propodeum approximately as long as metanotum .• .•.....••....•.••..•.•..•.•. . Dryinus aterrimus (Dodd) - Dorsal surface of propodeum longer than metanotum .•.••••••...•. 4 4 Scutellum granulated and punctate, not ret iculate rugose •.•••.•• .......•.....•.•..•.• .•••.•. . Dryinus australianus n. sp. - Scutellum granulated and reticulate rugose . . . . . . . . . . . . . . . . . . . . . 5 5 Scutum reticulate rugose, very rugose; areolae wide . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dryinus koebelei (R.C.L. Perkins) - Scuum reticulate rugose, less rugose; areolae less wide ••••••••• . . .• .. • . . . . • , ...•..•..•..•... Dryinus venator (R.C.L. Perk i ns)
1. Dryinus Latrei lie
Dryinus Latrei lie 1804: 176. nee Dryinus Fabricius 1805: 200. Dryinus Latreille 1805: 228. Campylonyx Westwood 1835: 52 (n. syn.); type species Campylonyx ampuliciformis Westwood 1835, monotypic. Paradryinus R.C.L. Perkins 1905: 53 (n. syn.); type species Paradryinus koebelei R.C.L. Perkins 1905, orig. desig. Chlorodryinus R.C.L. Perkins 1905: 57 (n. syn.); type species Chlorodryinus pallidus R.C.L. Perkins 1905, orig. desig. Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1905: 69 Campylonyx Westwood: Kieffer in Kieffer et Marshall 1905: 84. Plastodryinus Kieffer in Kieffer et Marshall 1906: 496 (syn. proposed by Moczar 1965); type species Plastodryinus szepligetii (Kieffer 1905), orig. desig.
Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1906: 497. Hesperodryinus R.C.L. Perkins 1907: 40 (n. syn.); type species Hesperodryinus arizonieus R.C.L. Perkins 1907, orig. desig. Lestodryinus Kieffer 1911a: 108, new name for Dryinus Latreille (syn. proposed by Richards 1937); lype species Lestodryinus formiearius (Latreille 1804)(first species included in Dryinus
729
Dryininae
Latreille).
Lestodryinus Kieffer: Kieffer 1914b: 20. Campylonyx Westwood: Kieffer 1914b: 43. Dryinus Latrei lie: Richards 1935: 144. Dryinus Latreille: Richards 1937: 118. Dryinus Latrei lie: Richards 1939: 229. Mesodryinus Kieffer: Muesebeck and Walkley 1951: 1034. Hesperodryinus R.C.L. Perkins: Muesebeck and Walkley 1951: 1034. Hesperodryinus R. C. L. Perk i ns: Krombe i n 1979: 1242. Mesodryinus Kieffer: Krombei n 1979: 1243.
Dryinus iormicarius Latreille 1804, first included species
TYPE SPECIES: (Latreille 1805) FEMALE
(Figs 495,
segments;
labial
528,
palpi
529):
as or shorter than front subapical visible;
tooth
(rarely
pronotal
fully
tibia, two)
much
(Figs 531,
labial
palpi
532):
fully
tibial
winged;
with 6
(Fig.
longer than arol ium,
and some lamellae;
with 3 segments;
DISTRIBUTION:
maxi Ilary palpi
enlarged claw
tubercles reaching or not
complete or incomplete or absent; MALE
winged;
with 3 segments;
485)
notaulices at
tegulae;
as long
with one
occipital
least partly carina
spurs 1,1,2.
maxi Ilary palpi
with 6 segments;
for morphology see fami Iy characteristics.
worldwide
Dictyopharidae, Cixiidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Fulgoridae. HOSTS:
SPECIES: NOTES:
87.
on a different I
(Chlorodryinus for instance) were based
some synonym ized genera
have seen,
variable with
length of occipital however,
that the
carina
(complete or incomplete).
length of occipital
intermediate cases from occipital
Generic differences so are not proposable.
carina can be very
carina absent to complete.
The shape of posterior margin of
head also is variable, with intermediate cases from occiput concave to straight or convex (Hesperodryinus). In addition the posterior ocelli can be located on an front or behind,
imaginary
I ine connecting
with numerous
As the type of D.
hind margins of eyes or in
intermediate cases.
iormicarius Latreille seems lost and the original
description is unreliable, the diagnosis of Dryinus is based on classical opinions of Kieffer (1914b) and Richards (1939).
PALAEARCTIC SPECIES OF DRYINUS
Seven species are known.
KEY TO THE SPECIES
the
730
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
FEMALES
Occipital
margin
incomplete,
only dorsally visible
............................ -
Occipital
margin
complete
6.
(Fig.
2 Pronotum with weak posterior short
posterior collar
propodeum nearly as -
a
long as dorsal
long posterior collar
transversal
shorter
impression and wi th a
posterior surface of
surface;
fore-wing
hyal ine or
(Fig.
484 A,B,C);
than dorsal
posterior surface of
surface;
fore-wing
wi thout
dark
transversal
bands;
head and
sanderi n. sp.
prothorax mostly black . . . . . . 1. Fore-wing wi th dark transversal
bands;
redd ish •••••••••••••••••••••
canariensis (Ceba I los)
4 Transversal 488 A); furrow
(Fig.
488 B);
2.
head and prothorax
fully
furrow between metanotum and propodeum narrow
metanotum at
Transversal
wi th dark
bands ••••••••••.•••...••••.•••..••••••.•••.•••••••• 4
3 Fore-wing hyal ine,
-
4840);
wi th dark transversal bands •••••••••••••••••••••••••••••••••••• 3 Pronotum with a strong posterior transversal impression and with propodeum much
-
499 A) •
492) . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
transversal
(Fig.
(Fig.
koreanus (Moczar)
least
five
488 A) . . . . . . . . .
3.
times as
long as breadth
this
collan's (Linnaeus)
furrow between metanotum and propodeum broad
metanotum nearly
(Fig. of
three-four
times as
(Fig.
long as breadth
of
thi s furrow (Fig. 488 B) •••.••••••••••••••••••••••••••••••••••• 5 5 Temples more prominent (Fig. 492 A); TL nearly twice as long as OL -
(Fig.
Temples (Fig.
492 A) . . . . . . . . . . . . . less prominent
(Fig.
492 B) ••••••••••••••••
The female of the fossil
4.
nepalensis n.
492 B); 5.
TL
sp.
slightly
longer
than OL
tarraconensis Marshall
species D.
antiquus (Ponomarenko) is
unknown.
MALES (excluding the fossi I Dryinus antiquus Ponomarenko)
Fore-wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) •• 5. tarraconensis Marshall -
Fore-wing proximal
with distal part
2 Notaulices -
Notaulices
(Fig.
part
of
radial
vein much
reaching approximately
0,5
reaching
0,75
3 Oistivolsella
longer
than
490 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 approximately
farther
from apex
length
of
scutum . . . . . . . . . 3
length of
of penis
(Fig.
scutum . . . . . . . . 4
491) . . . . . . . . . . . ..
3. collaris (Linnaeus) -
Oistivolsella nearer apex
of penis
(Figs.
487,
494) . . . . . . . . . . . ..
Dryininae
731
. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. koreanus (Moczar) segment 3 nearly twice as long as segment 2 • . . . . . . . • . . . . • . . . . . . • . . . . . . . •. . . . . • . . . .. 2. canariensis (Ceba I los) - Antennal segment 3 more than twice as long as segment 2 . . . . . . . . . 4. nepalensis n. sp.
4 Antennal
NOTES:
the male of D.
The
fossil
sanderi n. sp. is unknown. antiquus (Ponomarenko) isn't present in the
species D.
key because the only known specimen
is in
too bad conditions for
distinguishing differences with I iving species. The male associated in the key with D. canariensis could be also the opposite sex of Richardsidryinus albrechti n.
sp.
Only breeding
experiments could answer the question.
1. Dryinus sanderi n. sp.
FEMALE: fully winged; length 3,75 mm; head black, with mandibles, clypeus and malar space testaceous; antennae fully testaceous; thorax and propodeum black,
with posterior collar of pronotum reddish-testaceous;
A
B
c Fig. sp.
(laterally viewed) of females of Dryinus nepalensis n. collaris (Linneaus) from France (B), tarraconensis Marshall from Montecatini (Italy)(C) and sanderi n. sp. (type)(D).
484 -
Pronotum
(type) (A),
732
Mem.
Fig.
485 -
abdomen black-brown;
Amer.
Chela of
Ent.
no.
37,
legs testaceous,
with coxae partly black or brown
antennae distally
in following proportions:
8:
6:
23:
14:
punctate,
frons,
where there are numerous irregular striae;
complete;
thickened;
10:
shiny,
incomplete keel,
1984
lJryinus sanderi n. sp. (type)
and with clubs of femora brown; segments
Inst.,
8:
6:
antennal
5:
4,5:
8;
head
weakly granulated and rugose mainly on anterior half of frons with a
not visible on anterior half of frons;
POL = 4;
Ol = 3;
OOl = 8;
OPl = 1,5;
median
occipital
Tl = 3;
carina
pronotum
(Fig.
484 0) with a very weak anterior transversal impression and with a strong posterior transversal impression; disc humped; posterior collar very short; pronotal
tubercles not reaching
granulated; numerous,
scutum shiny,
short,
posterior margin; of scutum; smooth;
weak,
very
tegulae;
pronotum shiny,
weakly
weakly granulated and rugose,
longitudinal
with
keels near anterior margin and near
notaulices incomplete,
reaching
scutellum and metanotum shiny,
propodeum reticulate rugose,
very
approximately 0,65
length
weakly granulated,
with dorsal
surface nearly as long as
posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal 17: with
3:
6:
14:
part 24;
1 row of 7
of 19 lamellae; 1,1,2. MALE:
unknown
(14:7);
fore tarsal
enlarged claw lamellae;
(Fig.
segments 485)
segment 5 of front
apex with a
in following proportions:
with a
group of nearly
tarsus
subapical (Fig.
12 lamellae;
tooth and
485) tibial
with 2 rows spurs
Dryininae
LOCUS TYPICUS: Melnik (SW Bulgaria) TYPICAL MATERIAL: holotype F! in OL. HOSTS: in Bu I gari a Hysteropterum latifrons DISTRIBUTION: Palaearctic region: BULGARIA:
Fig. 486 -
Chela of
733
Melnik, OL!
Dryinus canariensis n. sp. from Tahodio (Canary Islands)
734
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
2. Dryinus canariensis (Ceballos) n. comb.
Paradryinus canariensis Ceba Ilos 1927: 101.
FEMALE:
fully
legs reddish;
winged;
length 4,25-4,50 mm;
mesothorax,
metathorax,
head,
antennae,
prothorax,
propodeum and petiole black;
abdomen brown, with segment 1 reddish; tegulae testaceous; antennae not distally thickened; antenna I segments in following proportions: 13: 5: 20: 9:
9:
6:
6:
6:
5:
8;
head dull,
smooth,
granulated;
frons with a median
longitudinal keel, without lateral keels; occipital carina complete; POL = 4; OL = 3; posterior ocelli not touching occipital carina; OPL nearly as long as breadth of ocell i; granulated, impression
partly and
collar short,
temples prominent;
without sculpture,
with
a
strong
nearly 0,5 as
pronotum shiny,
with an
posterior transversal
long as pronotal
partly
anterior weak transversal impression;
tubercles;
posterior
pronotal
tubercles
not reaching tegulae; scutum dull, hairy, granulated and with numerous weak areolae; notaul ices incomplete, reaching approximately 0,65 length of scutum; weakly
scute II um and metanotum sh iny, granulated;
strongly transversely striate, dark transversal
smooth,
wi thout scu I pture or very
propodeum reticulate rugose,
bands,
without
two on
with posterior surface
longitudinal
the basal
keels;
pterostigma;
distal
part of radial
(13:7);
tarsal
segments in following proportions:
fore
enlarged claw
(Fig.
segment 5 of front same length; MALE
testaceous;
(Fig.
incomplete; carina;
legs brown,
with
=
granulated;
proximal
part
broadened;
winged;
tibial
5;
OL
=
3;
OOL
with ventral
light;
=
3;
scutum dull,
(12:7);
granulated;
part of radial
gonoforceps (Fig.
ninth abdominal
sternite:
black;
side of segment 10: 8:
487 A)
fig.
8:
occipital
posterior ocelli
length of scutum;
distal
part
15: 24;
antennae not distally thickened;
propodeum reticulate rugose; bands;
7:
spurs 1,1,2.
length 2,50 mm;
frons with a median keel;
temples invisible;
dark transversal
tarsi
15: 4:
tooth and with 12 lamellae;
in following proportions: 8,5: 4:
granulated;
POL
fully
with 3
with 2 rows of nearly 27 lamellae
antennae black,
reaching approximately 0,75 dull,
486)
longer tha proximal
apex with a group of 12 lamellae;
antenna I segments head dull,
vein much
with a subapical
(see note on page 731):
mandibles testaceous;
9;
486)
tarsus
fore-wing
cells and one beneath the
8:
8:
6:
6:
carina
touching occipital
notaulices incomplete,
scutellum and metanotum fore-wing
hyaline,
vein much
with distal
487 B;
tibial
without
longer than region
spurs 1,1,2.
LOCUS TYPICUS: La Cuesta (Canary Islands) TYPICAL MATERIAL: holotype F! in MD. HOSTS: unknown DISTRIBUTION: Palaearctic region: OL!
TN!
B.co de Marchar
(Tenerife) (F), Tenerife) (F),
MO! TN!
CANARY
(San Sebastian,
Tahodio
(Tenerife) (F),
ISLANDS:
B.co Santos
Gomera)(F), MD! MO!
(F),
MD!
La Cuesta
Los Calderones
(Medano,
Dryininae
735
Fig. 487 - Male genital ia of Dryinus canariensis n. sp. from Los Calderones (Canary Islands) (A: left half removed; B: ninth abdominal sternite) .
3. Dryinus collaris (Linnaeus)
Sphex collaris Linnaeus 1767: 946. Dryinus formicarius Latreille 1805: 228 (syn. proposed by Fitton, Graham,
Boucek, Fergusson,
Huddleston, Quinlan, Richards 1978);
locus typicus Southern France. Campylonyx ampuliciformis Westwood 1835: 52 (n. syn.); locus typicus Lyon (France)
Dryinus formicarius Latrei lie var. : Kieffer in Kieffer et Marshall 1905: 78.
Dryinus formicarius Latreille var. : Kieffer in Kieffer et Marshall 1905: 79.
Lestodryinus formicarius (Latreille): Kieffer 1914b: 20. Lestodryinus corsicae Kieffer 1914b: 21 (n. syn.); locus typicus Corse (France).
Lestodryinus formicarius (Latrei lie): Berland 1928: 140. Lestodryinus corsicae Kieffer: Berland 1928: 141.
736
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
488 - Metanotum of Dryinus collaris (L.) from Le Puy (France )(A) and tarraconensis Marsha II from Orleans (France) (B). In metanotum of tarraconensis the anterior half is rugose and inclined. 5 ; transversal
Fig.
furrow between metanotum and propodeum.
Campylonyx ampuliciformis Westwood: Berl and 1928: 142. Lestodryinus formicarius (Latreille): Picard 1932: 29. Lestodryinus formicarius (Latrei lie): Maneval 1935: 4. Dryinus formicarius Latrei lie: Richards 1937: 118. Dryinus formicarius Latrei lie: Richards 1939: 229. Dryinus formicarius Latrei lie: Berland 1940: 121. Dryinus formicarius Latrei lie: Abdul-Nour 1974: 1015. Dryinus formicarius Latreille: J.I'. Perkins 1976: 27. Dryinus formicarius Latrei lie: Olmi 1976: figs 10-14. Dryinus collaris (L i nnaeus): F i !ton, Graham, Boucek, Fergusson, Huddleston,
Quinlan,
Richards 1978:
127.
Dryinus formicarius Latreille: Currado and Olmi 1979b: 4. Dryinus collaris (Linnaeus): Day 1979: 60.
FEMALE: fully winged; length 5,12-6,25 mm; head reddish-testaceous, with a brown small or wide spot on frons and vertex; antennae testaceous; propectus, pronotum, scutum, scutellum, metanotum, mesopleura, mesosternum, metasternum and metapleura reddish-testaceous; sometimes brown
spots on mesosternum,
metasternum,
between mesopleura and metapleura;
mesop leura and on furrow
rarely pronotum mostly black;
scutell um and metanotum someti mes reddi sh-dark;
mesosternum,
metasternum, mesopleura and metapleura sometimes almost fully brownblack; rarely scutum, scutellum and metanotum fully black; propodeum black,
with apex reddish;
fully reddish-testaceous, head dull,
granulated;
petiole black;
ocellar region swollen;
OPL shorter than breadth of ocelli; B)
crossed by
abdomen brown-reddish;
legs
sometimes with mid and hind coxae brownish;
two transversal
collar usually smooth and
impressions,
shiny;
without sculpture;
impression with short longitudinal
occipital carina complete;
temples prominent;
keels;
pronotum
(Fig.
484
disc and posterior
posterior transversal
anterior collar and sides weakly
Dryininae
Fig.
489 -
granulated; reaching dull;
Chela of
sides with a
tegulae;
Dryinus collaris (L.) from Dornbach (Austria)
few
longitudinal
scutum shiny,
notaul ices distinct and
of scutum;
737
keels;
pronotal
tubercles not
weakly punctate and granulated,
incomplete,
reaching
posterior third of scutum with
weak
rarely
approximately 0,5
longitudinal
length
short keels;
scutellum usually flat, shiny, smooth, weakly punctate; metanotum usually weakly punctate, smooth, always without transversal furrow in posterior half (Fig.
488 A);
propodeum with transversal proximal
tarsus
segment 1 of front with subapical (Fig.
489)
reticulate rugose;
longitudinal keels;
bands and with distal
part;
enlarged claw of front
propodeum fully
two short
part of radial tarsus slightly
tooth and
11
vein much
longer than
longer than segment 4;
lamellae
with 2 rows of 27
posterior surface of
fore-wing with 2 brown
(Fig.
lamellae;
489);
segment 5
apex with a
group
Mem.
738
Amer.
Ent.
Inst.,
no.
37,
1984
A
B Fig.
490 -
Radial
(Italy)(A)
of 19 lamellae; MALE:
fully
and
veins of males of Dryinus collan"s (L.) from Rosta tarraconensis Marshall from Orleans (France)(B).
tibial
spurs 1,1,2.
winged;
length 5,00 mm;
black,
with
tegulae and
legs
testaceous; mandibles with 3 teeth; scutum dull, hairy, granulated; notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny,
smooth,
weakly punctate;
propodeum fully
reticulate rugose, without longitudinal keels on posterior surface; forewing with distal part of radial vein much longer than proximal part (20:10)(Fig.
490 A),
without dark transversal
the apex of penis (Fig. LOCUS TYPICUS:
491);
tibial
bands;
distivolsella far from
spurs 1,1,2.
Spain
holotype F of S. collaris in Ll; holotype F of D. formicarius probably lost; lectotype F! (hereby designated) and 1 para lectotype F! of C. ampuliciformis in P; lectotype F! (hereby designated) of L. corsicae in MT. HOSTS: in France Cixius nervosus (L.)(Maneval 1935); in England Cixius contaminatus Fl. (Capron 1885); in Italy Issus coleoptratus (F.) (Olmi 1976); in Germany Cixius nervosus L. (Haupt 1932) (quotation of Dictyophara europaea L. in Richards 1939 valid for D. tarraconensis Marshall); in addition Tachycixius pilosus (OI.)(Thompson 1955). TYPICAL MATERIAL:
DISTRIBUTION:
Palaearctic region:
U.
S.
S.
R.
? (Ponomarenko 1978);
Dryininae
Fig.
491
HUNGARY:
-
Male genitalia of
Zelenika,
BU!
739
Dryinus collaris (L.) from Rosta (Italy)
YUGOSLAVIA:
Opatija
(Istra), BU!
Krapina
(Hrvatska), BU! GERMANY: Aix-Ia-Chapelle (Kieffer in Kieffer et Marshall 1905); AUSTRIA: Werfen (Slzb.), LE! Dornbach (Wien), P! BELGIUM (Kieffer in Kieffer et Marshall 1905); SWITZERLAND: Peney, GV! Geneve, SP! FRANCE: Mont-de-Marsan (Abdul-Nour 1974); Lyon (Rhone), P! St. Gely-duFesc (Herault), P! St. Guilhem (Herault), P! Digne (Basses-Alpes), P! Hyeres (Var), P! Le Puy, P! Toulon (Var), P! Les Guerraux (Saone-etLoire) (Berland 1928); Corse, MT! ENGLAND: Cobham (Surrey) (Richards 1939); Shere (Surrey)(Richards 1939; Capron 1885); SPAIN (Linnaeus 1767; Day 1979); ITALY: Rosta (Torino), OL! Gaibola (Bologna), OL! S. Olcese (Genova), BM! Lucca (Kieffer in Kieffer et Marshall 1905); Napoli (Kieffer
740
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
A o Fig.
492 -
B
o
Heads
0
o
of females of Dryinus nepalensis n. sp. tarraconensis Marshall (lectotype)(B)
and
in Kieffer et Marshall the cocoon
1905); S.
is oval
elongate. According by
0
(dorsally viewed)
(type)(A)
NOTES:
o
(Fig.
to Berland
Basilio
(Mottola,
498 A),
whereas
(1940),
Taranto)(Moczar 1965).
D. tarraconensis it's
in
occasionally the female
is attracted
light traps.
4. Dryinus nepalensis n. sp.
FEMALE: fully winged; length 6,25-6,55 mm; head black, wit>, ",andibles and anterior margi n of c I ypeus testaceous; antennae brown, wi th segments 1-2 testaceous and segments 7-10 whitish; black;
legs reddish,
distally 26:
thickened;
18:
broad
13:
11:
(26:5),
complete;
10:
8:
12;
POL
=
2,5;
Ol
around disc;
pronotum
collar
long,
flat,
distinct;
=
granulated and with numerous
median complete keel;
2,5;
OOl
=
11;
OPl
=
484 A)
with a
pronotal
keels;
metanotum nearly keels;
fore-wing
Tl
=
3;
6:
50:
long as
longitudinal carina temples:
fig.
impression; tegulae;
notaulices
scutellum dull,
posterior scutum dull,
incomplete, granulated;
wi th a strong transversa I furrow near scutell um and wi th
furrow near propodeum; transversal
occipital
tubercles not reaching
transversa I furrow near propodeum;
granulated;
15:
weak anterior transversal
almost reaching posterior margin of scutum; a strong
2;
strong posterior transversal
granulated and with some areolae near margins; metanotum du II,
antennae
in following proportions:
weakly granulated and with numerous striae (Fig.
impression and with a
propodeum and abdomen
antenna I segment 4 nearly five times as
frons with a
pronotum shiny,
thorax,
tibiae and part of coxae brown;
antenna I segments
head dull,
keels on frons; 492 A;
with
propodeum dull,
posterior surface with
with
and another beneath
centra I reg ion short and
three-four times as two short
two dark transversal the pterostigma;
long as breadth of
reticulate rrgose,
distal
bands,
without
incomplete longitudinal one on the basal
part of radial
vein
cells
longer
Dryininae
Fig.
than proximal
Chel a of Dryinus nepalensis n.
493 -
part
741
(19:10);
fore tarsal
sp.
(type)
segments in following proportions:
16: 3: 6: 15: 24; segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 493) with a subapical tooth and with a row of 15 lamellae;
segment 5 of front
lamellae;
apex with a group of nearly 18 lamellae;
MALE:
fully
winged;
(partly black
in
tarsus
(Fig.
length 2,00-3,56 mm;
long specimens);
segments 1-2 testaceous; partly black or brown;
carina
(not touching occipital invisible;
antennae fully
testacoeus or brown,
legs testaceous, 7:
6:
12:
incomplete;
10:
with all
10:
11:
10: 9: 7:
in very
9;
segments
head dull,
touching occipital
long specimens);
scutum dull,
reaching approximately 0,75
sometimes with a strong
antennal
frons with a median complete
scutellum and metanotum as sculptured as scutum; rugose,
with
coxae or only hind coxae
posterior ocelli
carina only
spurs 1,1,2.
mandibles testaceous
POL = 5; Ol = 2; OOl = 4;
notaul ices incomplete,
tibial
black;
granulated and weakly reticulate rugose; occipital
with 2 rows of nearly 37
antennae not distally thickened;
in following proportions: keel;
493)
transversal
keel
carina
temples
as sculptured as head; length of scutum; propodeum reticulate
between dorsal
and
742
Mem.
Fig.
494 -
Amer.
Male genital ia of
Ent.
Ins!.,
no.
37,
1984
Dryinus nepaiensis n. sp. from Godavari
(Nepal) (left half removed)
posterior surface;
fore-wing
hyal ine,
without dark transversal
bands;
distal part of radial vein longer than proximal part (13:9 or 9:5 or 14:9); gonoforceps (Fig. 494)with distivolsella located near apex of penis; tibial spurs 1,1,2. lOCUS TYPICUS:
lothar
TYPICAL MATERIAL: (1
F,
2 MM)!
HOSTS:
(Nepal)
holotype F!
and 5 paratypes MM!
in OT,
3 paratypes
in Ol.
unknown
DISTRIBUTION: Godavari
Palaearctic region:
(Katmandu), OT!
Ol!
r,EPAl:
Pulchauki
lothar (Birganj), (Katmandu),
OT!
OLi
OTt
5. Dryinus tarraconensis Marshall
Dryinus tarraconensis Marshall 1868: 204. Dryinus szepligetii Kieffer in Kieffer et Marshall 1905: 77 (n. syn.);
Dryininae
Fig.
495 -
Female of Dryinus tarraconensis Marshall from Montecatini ( Italy)
743
744
Mem.
Fig. 496 -
Amer.
Ent.
Inst., no. 37,
1984
Chela of Dryinus tarraeonensis Marshall from Orleans (France)
locus typieus Novi (Yugoslavia) (nee Budapest: see Moczar 1965). Dryinus tarraeonensis Marshall: Kieffer et Marshall 1905: 82. Plastodryinus szepligetii (Kieffer): Kieffer in Kieffer et Marshall 1906: 496.
Dryinus tarraeonensis Marsha II: Kieffer in Kieffer et Marsha II 1906: 497.
Lestodryinus tarraconensis (Marshall): Kieffer 1914b: Plastodryinus szepligetii (Kieffer): Kieffer 1914b: 43. Lestodryinus tarraconensis (Marshall): Berl and 1928: Lestodryinus gregori Hoffer 1936: 164 (syn. proposed 1965); locus typicus Pouzdranyi (Czechoslovakia). Lestodryinus bidens Haupt 1937: 70 (n. syn.); locus Bellinchen
21. 141. by Moczar
typicus
(Germany).
Lestodryinus tarraconensis Marshall: Pi Ilault 1951: 67. Dryinus szepligetii Kieffer: Moczar 1965: 380. Dryinus tarraconensis Marshall: Moczar 1965: 381. Dryinus tarraconensis Marshall: Currado and 01 mi 1979b: 4. nee Dryinus szepligetii Kieffer: Ponomarenko 1981 b: 879 (see Dryinus stantoni Ashmead and Dryinus browni Ashmead).
FEMALE (Fig. 495): fully winged; length 5,12-8,25 mm; head usually black, with frons, part of vertex (more along orbits), clypeus and mandibles yellow-reddish; sometimes also partly temples reddish; occiput
Dryininae
Fi g.
745
Ma I e gen i ta I i a of Dryinus tarraconensis Marsha II from Orl eans
497 -
(France)
always reddish; and vertex; testaceous;
rarely head reddish-testaceous with brown spot on frons
antennae brown,
with segments 1-2 and sometimes also 3
segments 9-10 or only 10 yellow;
sometimes reddish-testaceous; and sides reddish;
with posterior collar
sometimes fully reddish-testaceous;
and metanotum usually black, mesosternum,
propectus usually black,
pronotum usually black, sometimes fully
scutum,
mesopleura and metapleura usually black,
less reddish-testaceous or fully reddish;
scutellum
reddish-testaceous; sometimes more or
propodeum usually black,
sometimes with apex reddish, rarely fully reddish; petiole black; abdomen brown-reddish or black-brown; legs usually testaceous, with coxae partly brown,
sometimes also with clubs of femora brown,
testaceous; longitudinal
head dull, keels;
granulated;
vertex swollen;
rarely fully reddish-
frons and vertex with tracks of POL
= 4; Ol
= 3;
OOl = 13;
OPl = 2,5;
746
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
A
C _______B-~ Fig.
498 -
Italy)(A)
Dryinus collaris (L.) (female from Rosta, tarraconensis Marshall (female from Orleans, France)(B)
Shapes of cocoons of and
TL = 7; occipital carina complete; OPL nearly as long as breadth of ocell i; temp I es: fi g. 492 B; pronotum (F ig. 484 C) crossed by 2 transversal impressions, shiny, granulated, with strong longitudinal keels on sides; posterior collar sometimes smooth and without sculpture; anterior impression sometimes weakly tegulae;
scutum shiny,
longitudinal
pronotal hairy,
keels on posterior third;
approximately 0,5 strong
visible;
granulated,
length of scutum;
longitudinal
metanotum with a propodeum fully
keels;
deep
rarely
with numerous short
notaulices incomplete, scutellum dull,
almost smooth,
transversal
reticulate rugose;
tubercles not reaching
rugose,
reaching
with numerous and
with weak keels;
furrow on posterior half (Fig.
488 B);
posterior surface with 2 short
longitudinal keels; fore-wing with 2 brown transversal bands and with distal part of radial vein much longer than proximal part; segment 1 of front
tarsus sl ightly
longer than segment 4;
tooth and 13 lamellae (Fig. 2 rows of 38
lamellae;
496);
enlarged claw with subapical
segment 5 of front
apex with a group of at
tarsus
least 15
(Fig.
lamellae;
496)
spurs 1,1,2. MALE:
fully
winged;
length 3,37-3,87 mm;
black,
with
tibial
with mandibles yellow
Dryininae
and tarsi carina
light;
fore tibiae yellow;
incomplete;
almost complete;
scutellum and metanotum shiny,
vein
dark transversal
granulated,
scutum dull,
occipital
hairy,
granulated;
reaching posterior third or fourth of scutum,
fully reticulate rugose, part of radial
head dull,
temples not visible;
notaul ices incomplete,
747
without
longitudinal
keels;
nearly as long as proximal
bands;
distivolsella
granulated; fore-wing
part
(Fig.
rarely
propodeum with distal
490 B),
located near apex of penis
without
(Fig.
497); tibial spurs 1,1,2. lOCUS TYPICUS, Torla (Spain) TYPICAL MATERIAL; holotype F! of D. tarraconensis in BU; holotype F! of L. gregori in Hoffer coli.; holotype F! of D. szepligetii in BU; holotype F! and allotype M! of L. bidens in HA! in VI there is the type M! of Labeo rufipes Forster (never described) without local ity label: it's a male of D. tarraconensis. HOSTS; Dictyophara europaea (l.) in France (Pi Ilault 1951) and in Germany (Richards 1939; under name Dryinus formicarius latrei lie; Haupt 1937: under name Lestodryinus bidens Haupt). DISTRIBUTION;
Palaearctic region;
U.
S.
S.
? (Ponomarenko 1978);
R.
HUNGARY; Kenese, BU! Crkvenica, Ol! ROMANIA: Nagyenyed (Transilvania)(under name Plastodryinus sziladyi n. sp.), BU! CZECHOSLOVAKIA; BU!
Sutomore
FRANCE;
Orleans
(Madrid), ITALY:
BM!
Sarre
(L.)
BM!
Parma,
BO!
P!
GERMANY; SPAIN;
S. NA!
Perugia,
(Caltanissetta),
Baghdad, NOTE:
GE!
(Hoffer 1936;
BU!
(Tarragona),
(Aosta),
(Pistoia),
Cataldo
(Loiret),
Salou
(Alessandria), Terme
Pouzdranyi
(Dalmacija),
CO!
Torla
OL!
Moczar 1965);
Bell inchen
BM!
(Huesca),
BU!
Benicasim (Cuneo),
Pietra Ligure
(Savona),
Maiella
GREECE;
(Abruzzo),
Kremasti
Hills
P!
Escorial
PL!
BM!
Gavi
PL!
NA!
Novi,
HA!
(Castellon),
Benedetto Belbo GE!
YUGOSLAVIA;
(Oder),
Montecatini
Messina,
(Rhodes),
BM!
CO!
WA!
the cocoon it's ovale.
(Fig.
498 B)
is very elongate,
Notes on biology by Pillault
whereas in
D. collaris
(1951).
6. Dryinus korean us (Moczar) n. comb.
Chlorodryinus koreanus Moczar 1983a; 192.
FEMALE:
fully
winged;
length 3,84 mm;
testaceous-light,
with petiole
black; antennae distally thickened; antenna I segments in following proportions: 10: 6; 33: 18; 15; 11: 9; 7; 7; 10; head (Fig. 499 A) swollen,
dull,
granulated;
frons with a
weak median complete keel;
occipital carina incomplete, visible only behind posterior ocelli and on sides of posterior ocelli; POL = 3; OL = 4; OOL = 8; OPL = 3; temples prominent;
pronotum shiny,
striae on sides,
without sculpture,
crossed by an
with some longitudinal
anterior and a posterior transversal
S.
IRAQ;
748
Mem.
Amer.
Ent.
A
Inst.,
no.
37,
B
o
1984
0
DQll) Fig.
499 -
Heads of Dryinus koreanus
from De-sang san, Wanuma,
impression;
Korea)
(Moczar) (A:
and pallidus
New Guinea;
D:
(R.C.L.
type female;
Perkins) (B:
C:
male
female from
male from Mulgrave, Australia)
anterior collar very short;
disc very
long;
posterior collar
short; pronotal tubercles not reaching tegulae; scutum granulated, with anterior surface of median area smooth and without sculptur0; notaulices incomplete,
reaching approximately 0,65
metanotum dull,
granulated;
transversal or longitudinal
length of scutum;
propodeum reticulate rugose, keels;
dorsal surface sl ightly
scutellum and without longer than
(27:20); fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (20:8); fore tarsal segments in fOllowing proportions: 24: 3: 6: 20: 29; enlarged claw (Fig. 500) with a subapical tooth and with a row of 11 lamellae; segment 5 of front tarsus (Fig. 500) with two rows of 17 lamellae; apex with a posterior surface
group of 7 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,72 mm; head black,
with mandibles,
anterior region of frons and malar space testaceous; 1 testaceous
(other segments missing
and propodeum black; 499 C)
dull,
carina
incomplete,
in two only known specimens);
abdomen brown;
granulated;
legs yellow-testaceous;
frons with a median complete keel;
fully visible dorsally,
clypeus,
antennae with segment head
thorax (Fig.
occipital
partly visible on sides and
Dryininae
Fig.
500 -
Chela of
Dryinus koreanus (Moczar) (type)
749
750
Fig.
Mem.
501
-
Amer.
Ent.
Inst.,
no.
Male genital ia of Dryinus koreanus (Korea) ( I eft ha I f
37,
1984
(Moczar)
from De-sang san
removed)
occipital carina dorsally touching eyes; POL = 5; Ol = 2; OOl = = 1; scutum dull, granulated; notaulices incomplete, reaching
ventrally; 3;
OPl
approximately 0,5 shiny, without
smooth,
length of scutum;
without sculpture;
transversal
scutellum dull,
propodeum dull,
or longitudinal
keels;
granulated;
metanotum
reticulate rugose,
fore-wing hyal ine,
without dark
transversal bands; distal part of radial vein longer than proximal (16:7); genitalia: fig. 501; tibial spurs 1,1,2. lOCUS TYPICUS:
Pyongyan
TYPICAL MATERIAL: HOSTS:
part
(Korea)
holotype F!,
allotype M!
and 1 paratype M!
in BU.
unknown
DISTRIBUTION:
Palaearctic region:
Pyongan),
BU!
lyong-ak san
Pyongan),
BU!
De-sang san
KOREA:
Pyongyan
(Prov.
South
(14 Km IV from Pyongyan,
Prov.
South
(12 Km NE from Pyongyan,
Prov.
South
Dryininae
Pyongan),
751
BU!
7. Dryinus antiquus (Ponomarenko) n. comb.
Laberius antiquus Ponomarenko 1981 a: 139.
FEMALE: MALE:
unknown
fully
winged;
length 3,20 mm;
black,
with
legs testaceous;
palpi
not visible; antennae not distally thickened, densely hairy; antennal hairs less short than breadth of antennal segments; antennae la-segmented; antennal
segments
in following proportions:
7;
breadth of antenna I segments
4:
4:
3;
mandibles not visible;
by short dense hairs; posterior ocelli occipital
scutum dull,
posteriorly separated;
5:
head dull,
11:
10:
10:
9:
carina;
TL = a
bands;
three basal
cells enclosed by pigmented veins;
covered
(eyes almost touching notaulices complete,
fore-wing hyaline,
transversal
7:
eyes apparently bare;
metanotum and propodeum weakly
apparently propodeum reticulate rugose;
7:
4: 4: 4: 4: 4,5: 4: 4;
apparently granulated;
scutellum,
pterostigma
11:
apparently granulated,
occipital carina complete;
not touching occipital
carina);
8:
(greatest breadth):
long and narrow;
visible;
without dark
apparently fore-wing
distal
part of radial
with
vein
longer than proximal part; tibial spurs 1,1,2. LOCUS TYPICUS: Taimyr amber (Sibiria) TYPICAL MATERIAL: HOSTS:
holotype M!
in MPI.
unknown
DISTRIBUTION:
only
known
from Taimyr amber
(Sibiria,
U.
S.
S.
R.)
(Laberius Kieffer) is not correct, because Laberius is a genus of Gonatopodinae (it's a junior synonym of Dicondylus Hal iday). As its occipital carina is complete, L. antiquus is belonging to Dryininae. The new generic name Dryinus is provisionally proposed. The only known specimen of L. antiquus is in too bad conditions to permit to associate it for a certainty to a genus of Dryininae. D. antiquus is the oldest fossil dryinid known (nearly 100 millions NOTES:
the original
generic status
years old).
ETHIOPIAN SPECIES OF
Nineteen
species are
DRYINUS
known.
KEY TO THE SPECIES
752
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
FEMALES
Posterior collar of
2
3
4
5
6 7
8
pronotum invisible . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. bisulcatus (Benoi t) Posterior collar of pronotum visible (Figs 503 B, 519, 522) ... 2 Enlarged claw much shorter than segment 5 of front tarsus and wi th subapical tooth very small (Fig. 504) . . . . . . . . . . . . . . . . . . . .. · . . . . . . . . . . . . . . . . . . . . • • . . .. 2. seyrigi Ben 0 i t Enlarged claw nearly as long as segment 5 of front tarsus and usually with subapical tooth big (Figs 506,508) . . . . . . . . . . . . . . 3 Head (dorsally viewed) with posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes (Figs 505, 507) ••••.••••••••••••••••••••••••••••••••.••••••••••••••• 4 Head (dorsally viewed) wi th posterior ocell i placed in front of the imagina;"y straight line joining posterior edges of eyes or on t his lin e (F i g s 51 4, 524) •••••.•••••••••••..•••••••••••• 8 Head wi th posterior margin of vertex excavated (Fig. 505 A); head excavated; posterior ocell i touching occipi tal carina (Fig. 505 A) . . . . . . . . . . . . . . . 3. alticolus (Benoit) Head wi th posterio margin of vertex convex (Figs 505 B, 507 B); head flat; posterior ocelli not touching occipital carina (Figs 505 B, 507) ••••••••••••••••••••••••••••••••••••••••••••.•••••• 5 At I east the propodeum black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Species fully testaceous or reddish-testaceous, with petiole black ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 7 Head almost fully black . . . . 7. sierranus n. sp. Head almost fully reddish •• 8. pretorianus n. sp. Posterior margin of vertex more convex (Fig. 507 B); antennal segment 3 nearly ten times as long as segment 2 . . . . . . . . . . . . . . .. · . . . . . . . • • . . • . . . . . • • . • . . . .. 4. spangleri n. sp. Posterior margin of vertex less convex (Fig. 507 A); antennal segment 3 nearly six-seven times as long as segment 2 •••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. deceptor (Turner) Species fully testaceous, with petiole black and sometimes abdomen brownish; scutum shiny, weakly punctate ••••••••••.••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis Ki effer Species at least partly black; scutum shiny or dull, punctate or granulated, usually with more or less short irregular keels
· •.••••••..•..•..••.. " .••.•••.•.••.••••••.•• " •.••••. " •••••. 9 9
Posterior ocelli
touching
Posterior ocelli
not
524) ••••••••••••••••• " 10 Notaulices
occipital
touching
•••••••••• "
reaching approximately
........................... Notaulices
9. 10.
(Fig.
carina
514 A) . . . . . . 10
(Figs
514 B,
•.••.••••••••.•••••••••••. 11 0,65
length of
scutum . . . . . . . .
saussurei (Ceballos)
reaching approximately
• ••••••••••••••••••••••• "
carina
occipital
0,3
length of
hova (C eba I I os)
scutum . . . . . . . . .
753
Dryininae
11 Temples prominent prominent,
but
(Figs 514 B,
in
this
524 A);
rarely
temples weakly
case head with posterior margin
of
514 B, 524 A) . • . . . . . . . . . • • • • • • • • • • • • . . 12 Temples invisible (Fig. 524 B, C); head wi th posterior margin of vertex convex (Fig. 524 B, C) . • . • • . . . • . • • • . . . . . • • • • . • • • • • . 18 vertex
excavated
(Figs
12 Fore-wing with distal as
long as proximal
Fore-wing with distal times as
part
of
part ••
11.
part
of
radial
vein nearly
three
times
basilewskyi Benoit radial
vein
less
than
three
part ••••••••••••••••••••••••••••••• 13
long as proximal
13 Scutum punctate, not granulated, wi thout keel s ••••••••••••••••• •••••••••••••••••.•••••••• 12. chamaeleo Benoi t Scutum granulated or with keels ••••••••••••••••••.•.••••.•••• 14 14 Head granulated, with irregular or regular keels . . . . • • . • • • . . • 15 Head granulated,
without
keels ••••••••••••••••••••••••••••••• 17
15 Pronotum wi th anterior impression stronger (Fig. 519 A); head mostly reddish •••••••••••• 13. schoutedeni (Benoit) Pronotum with anterior impression weaker (Fig. 519 B); head mostly black ••••••.•••.••••••••••••••••.••......•.••••••••••• 16
16 Propodeum long, wi th dorsa I surface more than twi ce as long as posterior surface ••••••••• 14. mayogo (Benoit) Propodeum short, with dorsal surface slightly longer than posterior surface ••••••••• 15. ol'ophilus (Benoit) 17 Scutum granulated, not reticulate rugose; subapical tooth of enlarged claw very near apex (Fig. 525) •••••..•••••.••••••••••• 16. modestus n. sp. Scutum granulated, with posterior half reticulate rugose; subapical tooth of enlarged claw farther from apex (Fig. S 2 6) • • • • • • • • • • • • • • . • . . • . .• 1 7. incel'tus n. sp. 18 Anterior impression of pronotum very weak (Fig. 522 B) •••••••••
18. bUl'geoni Beno it of pronotum stronger (Fig.
•••••••••••••••••••••••••, Anterior
impression
••••••••••••••••••••••••••
19.
522 A) ••••••••••
aiel' (Dlmi)
MALES
Head wi th OL
= OOL •••••••••.••••••.•.•.••••••••••••••••••....• 2
Head wi th OL more or 2
Fore-wing with
radial
less
shorter
cell
than OOL •••••••••••••••••••• 3
closed;
head with posterior ocelli
not touching occipital carina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis (Kieffer) Fore-wing with touching
radial
occipi tal
cell
• • • • • • • • •• •• • •• • • • • • • •• • • •• 3
open;
head with posterior ocelli
carina •••••••..••...••••••••••••.....••••••• 9.
saussul'ei (Ceba I los)
Fore-wing wi th
radial
cell
closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Fore-wing with
radial
cell
open . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
754
4
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
Frons flat in front of the anterior ocellus, convex only between the antennal sockets • • . . • • • • • • • • . . • • • • • • • • • . • . • • • • •
19. afer (Olmi) Frons wi th a
5
median
antennal
sockets •••
Vo I s e I I a
n ea r I y
as
longitudinal •••••••
shorter
from anterior
long as gonoforceps
• • • • • • • . • • • • • • • • • • • .. Volsella
hump
(Fig.
to
513 B) . . . . . . . . . . .
(Fig.
pretorianus n.
8.
513 A)
ocellus
deceptor (Turner)
5.
sp.
••••••••.
15. orophilus (Benoit)
The males of other species are unknown. The description of the male allotype of (1953b)
is not reliable.
addition
the
D. chamaeleo by Benoit
This male was not obtained by breeding;
in
locality of the holotype is different.
Dryinus bisulcatus (Benoit) n. comb.
1.
Mercetia bisulcata Benoit 1953b: 400.
FEMALE:
fully
winged;
length 5,62 mm;
head black,
with clypeus and
mandibles testaceous; antennae testaceous; propectus black; pronotum black, with sides reddish; mesothorax, metathorax and propodeum black; abdomen black; black;
legs testaceous,
antennae distally
proportions:
11: 6;
shiny,
smooth,
behind
the ocelli);
21:
weakly
with coxae and clubs of femora brown or
weakly
13:
11:
thickened; 7,5:
alutaceous;
OL
6,5:
antenna I segments
4,5:
occipital
longer than POL;
6:
6;
carina
in following
head excavated, invisible
pronotum shiny,
(no track
weakly
granulated, crossed by a transversal impression on anterior third; the pronotum so is divided into 2 collars, without disc; the posterior collar is weakly
humped;
tegulae;
pronotal
scutum shiny,
converging,
reaching
surface with 2 other wing
proximal
part;
with subapical
median keels;
spurs 1,1,2. unknown
tarsus produced
tooth and
transversal
strongly
scutellum shiny propodeum not
keels;
anterior
sides of anterior surface with 2
posterior surface with 2
segment 1 of front
with 2 rows of 41 MALE:
keels;
with 3 brown spots and with distal
segment 3 of front
not reaching
incomplete,
metanotum very reduced and short;
with numerous parallel
longitudinal
longitudinal
notaul ices
almost posterior margin of sctum;
and without sculpture; reticulate rugose,
tubercles not very prominent,
granulated;
longitudinal
keels;
fore-
vein as
long as
tarsus shorter than segment 4
(18:22);
into a
17 lamellae;
part of radial hook;
enlarged claw
segment 5 of front
lamellae extending continuously
tarsus
to the apex;
(Fig. (Fig. tibial
502) 502)
I
Dryininae
Fig.
I
502 -
LOCUS TYPICUS: HOSTS:
Chela of Dryinus bisulcatus
Behara
TYPICAL MATERIAL:
755
(Benoit)(type)
(Madagascar)
holotype Fl
in P.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Behara,
Pl
D. bisulcatus is species of transition from Dryininae to Gonatopodinae. For the tibial spurs (1,1,2) and the pronotal tubercles directed towards the tegulae it is belonging to Dryininae; for the absence NOTES:
of occipital
carina,
metanotum very
the clypeus ventrally rounded
reduced
it
is near Gonatopodinae.
(not bidentate)
and
the
756
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
A
B Fig. 503 - Pronotum (laterally viewed) of females of Tridryinus ampuliciformis (Turner) from Afgoi (Soma I i a )(A) and Dryinus seyrigi Benoi t (type) (B)
2. Dryinus seyrigi Benoit
Dryinus seyrigi Benoit 1953b: 407.
FEMALE:
fully
winged;
length 3,75 mm;
head black,
with front part of
vertex, clypeus and mandibles testaceous; antennae testaceous, with segments 6-9 brown; propectus black; pronotum black, with sides and posterior collar reddish;
mesothorax,
metathorax and propodeum black;
abdomen missing in holotype (only specimen known); legs testaceous, with coxae and clubs of femora brown; antennae distally weakly thickened; antennal 8;
segments
head shiny,
prominent;
in following proportions:
granulated;
pronotum
posterior transversal
(Fig.
occipital 503 B)
impresssion
10:
5:
21:
carina complete;
shiny,
granulated,
and by a
10:
9:
8:
OL = POL;
6:
6:
4,5:
temples
crossed by a
strong
weak anterior transversal
impression; disc humped; posterior collar short, distinct; anterior collar weak; pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels near posterior margin; approximately 0,5 short, without
length of scutum;
with numerous longitudinal
transversal
longitudinal
notaulices incomplete,
scutellum dull,
keels;
metanotum
propodeum reticulate rugose,
keels on posterior surface;
bands and with distal
reaching
granulated;
fore-wing
part of radial
with 3 brown
vein much
longer than
D'ry i n i nae
Fig.
504 -
Chela of
Dryinus seyrigi Benoit (type)
757
758
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
A
Fig.
Heads of females of Dryinus alticolus (Benoit) (type) (A) and deceptor (Turner) from Port SI. Johns (South Africa) (B)
505 -
proximal part; segment 1 of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 504) with a small subapical tooth and 3 lamellae; segment 5 of front tarsus (Fig. 504) with 2 rows of 22 lamellae extending continuously to the apex; segment 5 of front tarsus apically without prominence and much longer than enlarged claw; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Ranomafana (Madagascar) TYPICAL MATERIAL: holotype F! in P. HOSTS: unknown DISTRIBUTION: Ethiopian region MADAGASCAR:
Ranomafana,
3. Dryinus alticolus (Benoit) n. comb.
Lestodryinus alticola Benoit 1951c: 4.
P!
Dryininae
Fig.
FEMALE:
fully
506 -
winged;
Chela of Dryinus alticolus
length 5,0 mm;
side of head;
(Benoit) (type)
head ferruginous,
brown spot on ocell ar reg ion and part I y visible on ventral
759
on frons;
with an elongate
another brown spot is
antennae testaceous;
propectus black,
wi th a dorsal reddish spot; pronotum ferrug i nous; mesothorax, meta thorax and propodeum black; abdomen brown; legs testaceous, with hind and mid coxae partly brown;
head
(Fig.
505 A)
excavated,
dull,
granulated,
with
760
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
oa
o Fig.
507 -
Heads of females of
Johns
temples
(South Africa) (A)
0
Dryinus deceptor (Turner) from Port St. and spangleri n. sp. (type) (B)
very prominent and posterior margin of vertex excavated;
occipital
carina complete; TL nearl,' three times as long as POL; OL weakly longer than POL; posterior ocell i touching occipital carina; pronotum dull, granulated;
anterior collar and
posterior collar distinct;
anterior transversal
posterior transversal
tubercles not reaching
tegulae;
near posterior margin;
notaulices incomplete,
scutum dull,
length of scutum;
scutellum dull,
reticulate rugose,
without
wing
with 3 brown
much
longer than proximal
granulated;
longitudinal
transversal part
impression
granulated,
weak;
pronotal
with some areolae
reaching approximately 0,75 metanotum rugose;
keels on segment
propodeum
posterior surface;
bands and with distal (17:4);
very
impression strong;
fore-
part of radial
1 of front
vein
tarsus nearly
as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 506) with subapical tooth and 4 lamellae; segment 5 of front of 10 MALE:
tarsus
lamellae; unknown
(Fig. tibial
506)
with 2 rows of 24
spurs 1,1,2.
lamellae;
apex with a
group
761
Dryininae
Fig.
508 -
Chela
0
f Dryinus spanglen. n.
sp.
(type)
762
Mem.
LOCUS TYPICUS:
Kilinga
TYPICAL MATERIAL: HOSTS: unknown
Amer.
Ent.
Inst.,
no.
37,
1984
(Zaire)
holotype F!
in TV;
1 paratype F!
DISTRIBUTION: Ethiopian region: ZAIRE: CAMEROON: N I Kongsamba (Beno i t 19S 1c) •
Kilinga
in P.
(Rutshuru),
TV!
4. Dryinus spangleri n. sp.
FEMALE:
fully
winged;
length 5,62 mmj
triangle and petiole black; white,
6-8 brown;
antennae distally weakly
in following
proportions:
507 B)
flat,
dull,
10:
OOL = 13;
OPL = 2j
imaginary straight of vertex convex;
57:
testaceous,
36:
24:
weak I y
antennal
14:
7:
11: 8:
frons
vis i b Ie on si des of occi put;
TL = 7;
posterior ocelli
POL = 3;
OL
posterior margin
impression;
disc humped;
longer than pronota I tubercles;
pronota I
pronotum granulated and with numerous
longitudinal
striae on sides;
longitudinal
striae on
scutum dull,
lateral region; fore-wing
granulated and with numerous
notaulices weak,
scutellum and metanotum dull,
fully reticulate rugose;
(Fig.
crossed by an anterior weak transversal
much
tubercles reaching tegulae;
segments
head
placed behind the
a posterior strong transversal long,
10;
with a median keel;
I ine joining posterior edges of eyes;
impression and by
with ocellar
with segments 4-5 partly
thickened;
weakly rugose;
pronotum du II,
posterior coil ar very
complete;
7:
granulated,
occipi ta I cari na comp I ete, = 2;
fully
antennae testaceous,
granulated;
but distinct and
propodeum shiny,
with 2 dark transversal
bands,
one on
the basal cells and another beneath the pterostigma; distal part of radial vein much longer than proximal part; fore tarsal segments in following proportions:
27:
3:
8:
22:
tooth and 12 lamellae; Clearly 33 lamellae;
33;
enlarged claw
segment 5 of front
(Fig.
tarsus
apex with a group of 23
508)
(Fig.
lamellae;
with a subapical 508) tibial
with 2 rows of spurs 1,1,2.
MALE: unknown LOCUS TYPICUS: Mombasa (Kenya) TYPICAL MATERIAL: holotype F! in WA. HOSTS:
unknown
DISTRIBUTION:
Ethiopian region:
KENYA:
Mombasa,
WAr
5. Dryinus deceptor (Turner) n. comb.
Lestodryinus deceptor Turner 1928: 147.
FEMALE:
fully
winged;
length 6,0 mm;
and abdomen reddish-brown; granulated;
occipital
head
yellow-testaceous,
(Figs 505 B,
carina complete;
507 A)
with petiole black
flat,
OPL shorter than POL;
dull, OL = POL;
Dryininae
763
Fig. 509 - Chela of Dryinus deceptor (Turner) from Port SI. Johns (South Africa)
764
Mem.
temples short;
Amer.
pronotum dull,
posterior transversal
Ent.
Ins!.,
granulated,
impression strong;
anterior transversa I impression weak; tegulae;
scutum shiny,
approximately 0,5 granulated;
granulated;
no.
with numerous keels on sides;
pronota I tuberc les not reach i ng incomplete,
longitudinal
reaching
scutellum and metanotum shiny,
propodeum weakly reticulate rugose;
propodeum without
1984
posterior collar long and visible;
notaul ices
length of scutum;
37,
keels;
posterior surface of
fore-wing
with 2 brown
transversal
bands and with distal part of radial vein much longer than proximal part; segment 1 of front tarsus shorter than segment 4 (18:22); segment 3 of front
tarsus produced
subapical
tooth and
2 rows of 34
into a
hook;
12 lamellae;
lamellae;
enlarged claw
segment 5 of front
(Fig.
509)
tarsus
apex with a group of 19 lamellae;
with
(Fig. tibial
509)
with
spurs
1,1,2. MALE: fully winged; length 2,43 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae, clubs of femora, mid and hind tibiae brown; antennae not distally thickened; antennal segments in following proportions: 5: 3: 11: 7: 6:
6:
5:
7;
head dull,
with a median frons
longitudinal
without median keel;
OOL = 4; dull,
granulated,
OPL = 1;
granulated,
reaching 0,5 transversal transversal
with weak tracks of areolae;
hump from anterior ocellus to antennal occipital
temples
invisible;
with weak
carina
incomplete;
scutum,
6:
sockets;
= 5,6; OL = 2;
scutellum and metanotum
tracks of areolae;
length of scutum;
POL
6:
frons
notaul ices incomplete,
propodeum reticulate rugose,
dull,
without
and longitudinal keels; fore-wing hyal ine, without dark bands; distal part of radial vein longer than proximal part;
radial cell closed; tibial spurs 1,1,2. LOCUS TYPICUS: Port SI. Johns (South Africa) TYPICAL MATERIAL:
holotype F!
and 1 paratype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region:
UGANDA:
Port St.
St.
Johns
(Pondoland),
BM!
in SM.
Kampala,
TW!
SOUTH AFRICA:
Lucia Estuary,
TW!
OL!
6. Dryinus mahensis (Kieffer) n. comb.
Mesodryinus mahensis Kieffer 1912a: 45. Labeo saxetanus Kieffer 1912a: 46 (n. syn.); locus typicus Mahe Island
(Seychelles
Islands).
Mesodryinus mahensis Kieffer: Kieffer 1914b: 31. Laberius saxetanus (Kieffer): Kieffer 1914b: 64.
FEMALE:
fully
winged;
abdomen darkened; complete, slightly
length 4,75 mm;
head shiny,
testaceous,
weakly punctate,
almost invisible on sides of occiput; longer than POL;
pronotum shiny,
with petiole black and
hairy;
occipital carina
OPL shorter than OLi
smooth,
without sculpture;
OL
Dryininae
Fig.
510 -
posterior transversal
765
Dryinus mahensis (Kieffer)(type)
Chela of
impression strong;
anterior transversal
impression
and anterior collar weakly visible; posterior collar visible; pronotal tubercles not reaching tegulae; scutum shiny, smooth, weakly punctate; notaul ices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose,
without
wing
with 2 brown
much
longer than proximal
longitudinal
transversal
part;
segment
segment 4;
segment 3 of front
(Fig.
510)
with subapical
(Fig.
510)
with 2 rows of 30 lamellae;
tibial
spurs 1,1,2.
·.lALE:
fully
winged;
1 of front
tarsus produced
hook;
vein
long as
enlarged claw
segment 5 of front
apex with a head black,
fore-
part of radial
tarsus as
into a
tooth and 14 lamellae;
length 3,43 mm;
antennae yellow-brown;
keels on posterior surface;
bands and with distal
tarsus
group of 15 lamellae;
with mandibles yellow;
thorax and propodeum bl ack;
abdomen brown;
legs
yellow, with hind coxae partly black; head dull, granulated; occipital carina incomplete; temples invisible; POL; 8; OL ; 3; OOL ; 3; OPL ; -~
;
0;
scutum dull,
granulated,
with
weak
areolae on
lateral
regions;
2;
766
Mem.
Fig.
511
-
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Dryinus sierranus n. sp. (type)
notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum shiny, weakly granulated; metanotum dull, rugose; propodeum reticulate rugose; fore-wing without dark transversal bands and with distal part of radial vein much longer than proximal part; tibial spurs 1,1,2 •• LOCUS TYP ICUS: Port Glaud (Seychelles TYPICAL MATERIAL: holotype F! of M.
Islands)
mahensis in BM; holotype M! of L.
saxetanus in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region:
SEYCHELLES
ISLANDS: Port Glaud
Island), BM! Mount Sebert (Mahe Island), BM! (Silhouette Island)(Kieffer 1912a).
Mare aux Cochons
7. Dryinus sierranus n. sp.
(Mahe
Dryininae
767
FEMALE: fully winged; length 8,75 mm; head black, with occiput, anterior region of frons, clypeus and mandibles reddish; antennae black, with segments 1-2 reddish, segment 4 yellow,
junction between segments 3-4 testaceous, half of segment 4 yellow,
proximal
prothorax reddish-brown-dark; mesosternum reddish-dark; metasternum black;
scutum bl ack,
scutellum,
abdomen black,
with junctions and tarsi
reddish;
apex of
segment 10 yellow;
wi th sides redd ish-brown;
metanotum,
propodeum,
pleura,
with some reddish regions;
legs black,
antennae distally weakly thickened;
antennal segments in following proportions: 10: 7: 58: 33: 24: 14: 11: 8: 8: 10; head dull, granulated, with tracks of irregular keels on frons; frons with a median keel; occipital carina complete; POL = 5; OL = 3,5; OOL = 11; OPL = 2; TL = 7; posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes; posterior margin of vertex convex; pronotum crossed by strong transversal impression and with a posterior collar long and distinctly visible; pronotum dull, granulated and with numerous longitudinal keels around disc; pronotal tubercles not reaching tegulae; scutum dull, granulated, with numerous weak longitudinal striae; notaul ices incomplete, almost reaching posterior margin of scutum, numerous
but weakly visible; longitudinal striae;
fully reticulate rugose,
scutellum dull, metanotum dull,
granulated and with reticulate rugose;
with 2 incomplete longitudinal
propodeum
keels on posterior
surface; fore-wing with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal 10: 23:
38;
lamellae;
part;
fore tarsal
enlarged claw
(Fig:
segment 5 of front
segments in following proportions: 511)
tarsus
apex with a group of 14 lamellae; MALE:
with a subapical (Fig.
tibial
511)
28:
5:
tooth and with 20
with 2 rows of 49 lamellae;
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Freetown
TYPICAL MATERIAL: HOSTS:
(Sierra Leone)
holotype F!
in TW.
unknown
DISTRIBUTION:
Ethiopian region:
SIERRA LEONE:
Freetown,
TW!
8. Dryinus pretorianus n. sp.
FEMALE:
fully
winged;
brown spot on frons; whi t ish;
prothorax,
propodeum black;
length 6,25 mm;
antennae brown,
head reddish-testaceous,
scutum and scutell um redd ish-dark;
abdomen brown,
with a
with segments 1-2 reddish,
with segment 1 partly reddish;
with chelae and segments 2 and partly
1 of hind tarsi
10
metanotum and whitish;
legs,
antennae
distally thickened; antenna I segments in following proportions: 9: 7: 49: 29: 21: 14: 10: 9: 7: 8; head dull, flat, granulated; frons with a median keel;
POL = 4,5;
complete;
DL = 3,5.
DOL = 11; DPL = 2;
posterior margin of vertex convex;
the imaginary straight
TL = 7;
occipital
carina
posterior ocelli placed behind
line joining posterior edges of eyes;
pronotum with
768
Mem. Amer. Ent.
Inst., no. 37, 1984
Fig. 512 - Chela of Dryinus pretorianus n. sp.
(type)
Dryininae
Fig.
513 -
769
Dryinus orophilus (Benoit) from Ile-ife
Male genitalia of
(Nigeria) (A: right half removed) and pretorianus n. sp. from Pretoria (South Africa) (8: right half removed; C: ninth abdominal sternite)
a strong
transversal
dull,
hairy,
dull,
granulated,
impression and with a
granulated;
pronotal
posterior collar
tubercles not reaching
with numerous irregular weak keels;
long;
tegulae;
pronotum scutum
notaul ices
incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum as sculptured as scutum; propodeum reticulate rugose, dull, with
two median
longitudinal
keels on dorsal
surface;
fore-wing
with 2
dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (15:7); fore tarsal
segments
claw
(Fig.
512)
front
tarsus
to the apex; MALE:
fully
in following proportions:
with a subapical
(Fig.
512)
tibial winged;
with 2 rows of 45
4:
10:
legs brown,
3;
occipital
scutum dull,
17:
thickened; 8:
8:
8:
black;
28;
enlarged
segment 5 of
antennal 8:
carina complete; granulated;
mandibles and antennal
with joints and tarsi
6:
6:
segments 8;
with some weak areolae on sides of frons; keel;
8:
lamellae extending continuously
length 2,37-2,62 mm;
antennae not distally 4:
3,5:
spurs 1,1,2.
segments 1-2 testaceous; proportions:
20:
tooth and with 10 lamellae;
POL = 6;
notaul ices
head dull,
granulated and
frons with an OL = 3;
incomplete,
testaceous;
in following incomplete median
OOL = 4; reaching
OPL = 1;
TL =
approximately
770
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
\) 0
Fig.
514 -
Heads of females of Dryinus saussurei (Madagascar) (A)
(Ceballos)
and basilewskyi Benoit
from Beki Iy
(type) (B)
0,65 length of scutum; scutellum dull, granulated; metanotum dull, granulated and with numerous areolae near anterior margin; propodeum reticulate rugose,
without transversal
or longitudinal
keels;
fore-wing
hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (9:7); distivolsella (Fig. 513 B) with apex located nearly on the same level with the apex of gonoforceps; sternite: fig. 513c; tibial spurs 1,1,2. LOCUS TYP ICUS:
Pretoria
TYPICAL MATERIAL: HOSTS:
ninth abdominal
(South Africa)
holotype F!
and 2 paratypes MM!
in TW.
unknown
DISTRIBUTION:
Ethiopian region:
SOUTH AFRICA:
Pretoria
(Transvaal),
TVY!
9. Dryinus saussurei (Ceballos)
Lestodryinus saussurei Ceballos 1935: 48. Dryinus saussurei (Ceballos): Benoit 1953b: 412.
FEMALE:
fully
winged;
mandibles yellow;
length 4,00-5,93 mm;
rarely clypeus black;
with segments 7-8 or 3-10 brown;
head black,
propectus black;
margins and posterior collar reddish;
with clypeus and
antennae testaceous,
mesothorax,
sometimes
pronotum black,
with
metathorax and
propodeum black; abdomen black-brown; legs testaceous, wi th coxae and clubs of femora partly brown; head (Fig. 514 A) shiny or du", fully or partly
granulated;
occipital
carina complete;
posteior ocell i
touching
Dryininae
Fig.
515 -
Chela of
771
Dryinus saussurei (Ceballos) from Bekily (Madagascar)
772
Mem.
occipital
carina;
Amer. En!.
OL = POL;
Ins!.,
pronotum dull,
with numerous weak striae around disc; nearly 0,5 times as
no.
37,
1984
partly or fully granulated,
posterior collar of pronotum short,
long as distance from
tegulae to pronotal
tubercles;
pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels or areolae near posterior margin; notaulices incomplete, almost reaching posterior margin of scutum; dull,
rugose;
posterior surface; proximal twice as into a claw
hook;
with
segment
515)
(Fig.
lamellae; MALE:
fore-wing
with 2
granulated;
longitudinal
with 3 brown transversal
bands
metanotum
keels on
(the two
bands partly joint) and with distal part of radial vein nearly long as proximal part (18:8); segment 3 of front tarsus produced
(Fig.
tarsus
scutellum dull,
propodeum reticulate rugose,
515)
tibial
fully
1 of front tarsus as
with subapical
tooth and
with 2 rows of 39
long as segment 4; 13
lamellae;
lamellae;
enlarged
segment 5 of front
apex with a
group of 11
spurs 1,1,2.
winged;
teeth brown;
length 2,60-2,84 mm;
antennae brown,
black;
mandibles testaceous,
with segments 1-2 testaceous;
palpi
testaceous; abdomen brown; legs testaceous, with coxae, clubs of hind femora and hind tibiae brown; antennal segments in following proportions: 8:
7:
17:
12:
13:
13:
11:
incomplete median keel, between
10:
10:
flat
the antenna I sockets;
DOL = 3; posterior ocelli scutum dull, granulated;
11;
head dull,
granulated;
frons with an
in front of the anterior ocellus and humped occipital
carina complete;
POL = 9;
OL = 3;
touching occipital carina; temples invisible; notaulices incomplete, reaching approximately
0,65 length of scutum; scutellum almost fu!ly dull, granulated, with a narrow posterior region smooth and shiny; metanotum with anterior half rugose and with posterior half shiny and smooth;
propodeum reticulate
rugose,
fore-wing hyal ine,
without
longitudinal
without dark
transversal
part slightly
longer than
segments;
labial
LOCUS TYPICUS:
palpi Rogez
TYPICAL MATERIAL:
or transversal
bands;
proximal
MD!
Behara,
part
with 3 segments;
cell open; (14:13); tibial
radial
vein
with distal
maxillary palpi
with 6
spurs 1,1,2.
(Madagascar)
holotype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region: Rogez,
radial
keels;
TV!
in MD. MADAGASCAR:
Fort Dauphin,
TV!
Bekily,
TV!
P!
Mandraka,
TV!
Ranomafana
Ivondro,
P!
(Benoit 1953b).
10. Dryinus hova
(Ceballos)
n.
comb.
ft;esodryinus hova Ceballos 1936: 53. fiJesodryinus hova Ceballos: Benoit 1953b: 414.
FEMALE:
fully
winged;
mandibles testaceous;
length 5,31
mm;
propectus black;
head black,
with clypeus and
pronotum black,
with margins and
773
Dryininae
Fig.
516 -
Chela of
Dryinus hova
(Ceballos)(type)
posterior collar reddish; mesothorax, metathorax, propodeum and abdomen black; legs testaceous, with coxae and clubs of femora brown; antennae distally weakly 5:
24:
10: 9:
transversal
thickened;
7:
6:
6:
5:
antennal 7,5;
segments
head dull,
keels on anterior half of frons;
posterior ocelli edges of eyes;
located on
in following proportions:
granulated,
occipital carina complete;
the imaginary straight
posterior ocelli
9:
with numerous short
touching occipital
line joining posterior carina;
OL = POL;
pronotum dull, granulated, with anterior transversal impression weak; anterior collar weak; posterior transversal impression strong; posterior collar short,
distinct;
pronotal
tubercles not reaching
tegulae;
scutum
dull, granulated, with some strong punctures; notaulices incomplete, reaching approximately 0,3 length of scutum; scutellum dull, granulated; metanotum rugose;
propodeum reticulate rugose,
on posterior surface;
fore-wing
half of fore wings missing species); of front
without
longitudinal
transversal
bands;
in holotype (only known specimen of the
tarsus produced into a hook; lamellae;
with
tarsus
(Fig.
516)
apex with a group of 12 lamellae;
tibial
spurs
segment 5 of front
unknown
LOCUS TYP I CUS:
Bek i I Y (Madagascar)
TYPICAL MATERIAL:
holotype F!
enlarged claw
in P.
(Fig.
segment 3
516)
tooth and 8 lamellae;
2 rows of 21
keels
distal
segment 1 of front tarsus nearly as long as segment 4;
subapical 1,1,2. MALE:
with 2 brown
with
774
Mem. Amer.
Fig. 517 -
Ent.
Ins!., no. 37,
1984
Chela of Dryinus basilewskyi Benoit
HOSTS: unknown DISTRIBUTION: Ethiopian region:
MADAGASCAR:
Bekily, P!
11. Dryinus basilewskyi Benoit
(type)
Dryininae
775
Dryinus basilewskyi Benoi t 1953a: 143.
FEMALE: vertex
fully
winged:
length 7,50 mm;
head black,
testaceous;
propectus black;
coil ar redd i sh; yellow-brown; sculpture,
mesothorax,
pronotum black, head
posterior ocelli
514 B)
very near occipital
than OL;
OL = POL;
except for posterior collar very
impression strong; distinct,
tubercles
reaching
tegulae;
abdomen
shiny,
without
(4:9);
granulated;
length of scutum; fore-wing
part of radial
vein
segment 3 of front
posterior collar
weak;
tegulae to
pronotal
tubercles not
notaulices incomplete,
without
longitudinal
transversal
three times as
tarsus nearly as
tarsus produced
weak;
reaching
scutellum and metanotum dull,
with 2 brown
more than
segment 1 of front
occipital
OPL shorter
posterior transversal
long as distance from
propodeum reticulate rugose,
posterior surface;
hairy;
impression
anterior collar very
scutum dull,
carina,
with numerous striae around disc,
anterior transversal
approximately 0,65 granulated;
pronotum shiny,
nearly 0,5 times as
pronotal
(17:5);
(Fig.
except for numerous weak striae on frons and vertex;
hairless,
distal
antennae
with sides and posterior
meta thorax and propodeum black;
legs yellow-testaceous;
carina complete;
short,
with front part of
(more along orbits), clypeus and mandibles yellow;
into a
long as proximal
long as segment 4
hook;
keels on
bands and with
enlarged claw
part
(17:16); (Fig.
517)
with subapical tooth and 11 lamellae (lamellae near subapical tooth very long and simi lar to peg-I ike hairs); segment 5 of front tarsus (Fig. 517) with 2 rows of 30 lamellae; 1,1,2. MALE:
apex
with
a group of 13 lamellae;
tibial
spurs
unknown
LOCUS TYPICUS:
Thysville
TYPICAL MATERIAL: HOSTS:
(Zaire)
holotype F!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
ZAIRE:
Thysville,
TV!
12. Dryinus chamaeleo Benoit
Dryinus chamaeleo Benoit 1953b: 409.
FEMALE: frons;
fully
winged;
length 6,0 mm;
antennae testaceous;
metanotum redd ish-brown; ferruginous;
propectus,
head reddish, pronotum,
propodeum black;
with a brown spot on
scutum,
scutell um and
abdomen reddi sh-brown;
antennae distally weakly thickened;
antennal
segments
legs in
following proportions: 10: 5: 48: 27: 22: 15: 12: 10: 8: 10; head shiny, weakly punctate, with numerous longitudinal keels on median region of frons; carina;
occipital
carina complete;
OPL = OL;
OL = POL;
posterior ocelli
pronotum shiny,
not touching occipital
without sculpture,
except
776
Memo Amero En!.
Ins!.,
noo 37,
1984
Fig. 518 - Chela of Dl'yinus chamaeleo Benoit (type)
777
Dryininae
A
B
Fig.
519 -
Pronotum of females of Dryinus schoutedeni and mayogo (Benoit) (type) (B).
for the sides which show
numerous
longitudinal
impression and anterior collar weak; strong;
posterior collar
not reaching notaulices,
tegulae;
reaching
rugose; distal
fore-wing
part of radial
1 of front produced
into a
14 lamellae; lamellae;
vein
tarsus nearly hook;
without
with 2 brown
nearly
as
keels;
long
twice as
segment 5 of front
tarsus
scutellum shiny,
longitudinal
keels on
bands and with
long as proximal
(Fig. (Fig.
apex with a group of 12 lamellae;
tubercles
punctate;
metanotum punctate,
transversal
as segment 4;
enlarged claw
impression
pronotal
length of scutum;
longitudinal
propodeum reticulate rugose,
posterior surface;
disc humped;
without sculpture,
approximately 0,65
punctate and with numerous
anterior transversal
posterior transversal
long and distinct;
scutum shiny,
keels;
(Benoit) (type) (A)
part;
segment 3 of front
518)
with subapical
518) tibial
segment tarsus
tooth and
with 2 rows of 42 spurs 1,1,2.
(in P!) of D. chamaeleo by Benoit (1953b) is not rei iable, because this male specimen MALE:
unknown;
the description of the male allotype
was not obtained by breeding. LOCUS TYPICUS:
Behara
TYPICAL MATERIAL:
(Madagascar)
holotype F!
in P.
778
Mem.
Fig.
HOSTS:
520 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
Dryinus schoutedeni
1984
(Benoit) (type)
unknown
DISTRIBUTION:
Ethiopian region:
1953b);
(Beno i t 1953b ) •
Bek i I Y
MADAGASCAR:
13. Dryinus schoutedeni
Lestodryinus schoutedeni
FEMALE:
fully
winged;
Benoi t
(Benoit)
1950:
n.
P!
Ivondro (Benoit
comb.
224.
length 8,43-9,37 mm;
with clypeus and mandibles light;
Behara,
head reddish or reddish-dark,
antennae black,
with segments 1-2 and
8-10 testaceous; propectus and pronotum brown-reddish or reddi sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs brownblack,
with tarsi
occipital
light;
head dull,
carina complete;
granulated,
posterior ocelli
weakly reticulate rugose;
located on
the imaginary
779
Dryininae
Fig.
521
-
Chela of
Dryinus mayogo
(Benoit) (type)
straight line joining posterior edges of eyes, not touching occipital carina; OPL = OL; OL = POL; TL nearl y twice as long as POL; pronotum (Fig.
519 A)
dull,
granulated,
with numerous keels around disc;
transversal
impression strong;
anterior collar distinct;
transversal
impression very strong;
posterior collar
anterior
posterior
long;
pronotal
780
Mem.
tubercles not reaching numerous
longitudinal
approximately 0,65 dull;
tegulae;
Ins!.,
rugose;
notaul ices
part;
part of radial
segment
with subapical
1 of front
with 2 rows of 34
18
lamellae;
1984
granulated,
hairy,
incomplete,
with
reaching
scutellum as sculptured as scutum, fore-wing
vein
hook;
long
as segment 4;
enlarged claw
segment 5 of front
apex with a
transversal
nearly twice as long as
into a
lamellae;
with 2
with 2 brown
tarsus nearly as
tarsus produced
tooth and
37,
propodeum reticulate rugose,
keels on posterior surface;
segment 3 of front
no.
scutum dull,
weak keels;
bands and with distal proximal
En!.
length of scutum;
metanotum dull,
longitudinal
Amer.
group of 13
(Fig.
tarsus
lamellae;
520)
(Fig. tibial
520) spurs
1,1,2. MALE:
unknown
LOCUS TYP ICUS:
Banana
TYPICAL MATERIAL: HOSTS;
(Zaire)
holotype F!
in TV.
unknown
DISTRIBUTION: Ethiopian region: ZAIRE: Banana ZIMBABWE: Chishawasha (Salisbury), BM!
(Low-Zaire),
TV!
14. Dryinus mayogo (Benoit) n. comb.
Lestodryinus mayogo Benoit 1950: 225.
FEMALE:
fully
winged;
part of vertex
length 8,12-10 mm;
(more along orbits),
sometimes with some small pronotum black, tarsi
light;
with genae,
reddish spots along orbits;
head dull,
abdomen black-brown;
weakly reticulate rugose;
front head
propectus black;
with sides and posterior collar reddish;
metathorax and propodeum black; with
head black,
clypeus and mandibles yellow;
mesothorax, legs brown-black,
occipital
carina
complete; posterior ocelli not touching occipital carina; OPL = OL; OL shorter than POL; TL little shorter than POL (8:9); pronotum (Fig. 519 B) dull, granulated, with numerous striae around disc; anterior transversal impression very weak; anterior collar almost invisible; posterior transversal
impression very strong;
tubercles not reaching some weak 0,65
irregular keels;
length of scutum;
rugose;
tegulae;
notaulices
hairy,
incomplete,
long;
fore-wing
with 2 short
with 2 brown
pronotal
granulated,
reaching
scutell um as scu I ptured as scutum;
propodeum reticulate rugose,
posterior surface;
posterior collar
scutum very
with
metanotum
longitudinal
transversal
dull,
approximately keels on
bands and with
distal part of radial vein nearly twice as long as proximal part; 1 of front tarsus nearly as long as segment 4; segment 3 of front
segment tarsus
produced into a hook; enlarged claw (Fig. 521) with subapical tooth and 19 lamellae; segment 5 of front tarsus (Fig. 521) with 2 rows of 38 lamellae; MALE:
apex with a group of
unknown
14 lamellae;
tibial
spurs 1,1,2.
Dryininae
781
A
B
c Fig.
522 -
Pronotum of females of Dryinus afer
Benoit from Kankunda
(Zaire) (B)
LOCUS TYP I CUS: Pau lis (Za ire) TYPICAL MATE,qIAL: holotype F! HOSTS:
burgeoni
(Benoit) (type) (C).
in TV.
unknown
DISTRIBUTION: LEONE:
(Olmi)(type)(A),
and orophilus
Ethiopian region:
Freetown,
ZAIRE:
Paulis
(High-Uele),
TV!
SIERRA
TW!
15. Dryinus orophilus (Benoit) n. comb.
Lestodryinus orophilus Benoit 1950: 226. Lestodryinus orophilus Benoit: Benoit 1951c: 7.
FEMALE:
fully
winged;
length 3,75-5,31
mm;
head black,
with clypeus and
782
Mem.
Fig.
523 -
mandibles testaceous; 3,
8-10 testaceous;
Amer.
Chela of
abdomen black;
antennae fully
occipital pronotum
(Fig.
around disc,
522 C)
crossed by
1984
testaceous or brown with segments 1with margins and
metathorax and propodeum black;
longitudinal
OPl = Ol;
shiny,
37,
with coxae and clubs of femora brown;
with some
carina complete;
no.
pronotum black,
mesothorax,
legs testaceous,
granulated,
Ins!.,
Dryinus orophilus (Benoit)(type)
propectus black;
posterior collar reddish; head dull,
Ent.
keels on frons and vertex;
Ol = POL;
granulated,
Tl
longer than POL;
with numerous weak striae
a strong posterior transversal
impression and by
a weak anterior transversal impression, posterior collar short, nearly 0,5 times as long as distance from tegulae to pronotal tubercles; anterior collar weak;
pronotal
tubercles not reaching
tegulae;
scutum dull,
granulated, with some irregular short keels on posterior half; notaulices incomplete, reaching almost posterior margin of scutum; scutellum dull, granulated,
metanotum reticulate rugose;
propodeum reticulate rugose,
without longitudinal keels on posterior surface; fore-wing with 3 brown transversal bands and with distal part of radial vein less than twice as long as proximal segment 4
enlarged claw of front tarsus of 10 lamellae; MALE:
part
(21 :15);
fully
(13:9);
segment
segment 3 of front
(Fig.
523)
1 of front
with subapical
tooth and
(Fig. 523) with 2 rows of 24 tibial spurs 1,1,2.
winged;
tarsus
tarsus produced
length 2,81-3,62 mm;
11
lamellae;
head black,
longer than
into a
hook;
lamellae;
segment 5
apex with a
group
with mandibles
Dryininae
o Fig.
524 -
783
0
Heads of females of Dryinus modestus n. sp.
(Olmi) (type) (B)
and burgeoni Benoit from Kankunda
yellow;
antennae yellow-brown;
brown;
legs
yellow,
thorax and propodeum black;
with coxae partly brown;
head dull,
afer
(type) (A), (Zaire) (C).
abdomen
granulated,
weakly reticulate rugose; occipital carina incomplete; OL shorter than OOL; OPL shorter than POL; OL shorter than POL; scutum du II, as scu I ptured as head;
notaulices incomplete,
reaching approximately 0,65 length of scutum;
scutell um and metanotum as scu I ptured as scutum; rugose; radial
fore-wing vein
without dark transversal
longer than proximal
part;
propodeum ret icu I ate
bands and with distal part of
radial
cell
open;
genitalia:
fig.
513 A; tibial spurs 1,1,2. LOCUS TYPICUS: Rutshuru (Zaire) TYPICAL MATERIAL:
holotype F!
designated after the original
HOSTS; unknown DISTRIBUTION: Ethiopian region: National Park,
TV!
Lusinga
in TV;
allotype M!
invalid,
because
description. ZAIRE:
Rutshuru
(Upemba National
(Kivu),
Park),
TV!
TV!
Garamba
Kalinga-Bitshumbi
784
Mem.
Amer.
(Albert National
Park),
GHANA:
(Kumasi),
Kwadaso
OT!
Ile-ife
(W State),
BM!
ZIMBABWE:
Fig.
TV! BM!
Ent.
Pweto OL!
Chishawasha
no.
37,1984
(Mweru Lake,
BM!
Akure
Ins!.,
NIGERIA:
(W State),
(Salisbury),
Shaba)(Benoit 1951c)j
Kabba OL!
BM!
(Lokoja),
BM!
ANGOLA:
Ikenne
CA!
Vila Luso,
525 - Chela of Dryinus modestus n. sp.
Ibadan,
(W State),
(type)
CA!
Dry i n i nae
CAMERON:
Victoria,
785
CAl
16. Dryinus modestus n. sp.
FEMALE:
fully
winged;
partly clypeus yellow; black,
length 5,0 mm;
head black,
antennae testaceous;
with margins and posterior collar reddish;
and propodeum black;
abdomen black-brown;
and clubs of femora brown; occipital carina complete; OPL shorter than OL; granulated,
OL = POL;
impr"ssion strong;
without
reaching
granulated;
longitudinal
flat,
dull,
granulated;
without sculpture;
posterior collar short; scutum dull,
approximately 0,75
metanotum rugose;
carina;
pronotum shiny, pronotal
posterior transversal granulated;
length of scutum;
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
with 3 brown
transversa I bands and wi th di sta I part of radi a I ve in much proximal
metathorax wi th coxae
not touching occipital
TL shorter than POL;
anterior collar weak;
incomplete,
scutellum dull,
tegulae;
pronotum
mesothorax,
legs testaceous,
524 A)
(Fig.
with anterior collar smooth and
tubercles not reaching notaul ices
head
posterior ocelli
with mandibles and
propectus black;
longer than
(11 :5); segment 1 of front tarsus longer than segment 4
part
(12:9); segment 3 of front tarsus produced into a hook; enlarged claw (Fig.
525)
with subapical
tooth and 10 lamellae;
(Fig. 525) with 2 rows of 20 tibial spurs 1,1,2. MALE:
segment 5 of front
apex with a
group of 11
tarsus
lamellae;
unknown
LOCUS TYP I CUS:
Bek i I y
TYPICAL MATERIAL: HOSTS:
lamellae;
(Madagascar)
holotype F!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
TV!
17. Dryinus incertus n. sp.
FEMALE:
fully
winged;
length 3,75 mm;
head black,
with clypeus and
mandibles reddish; propectus black; pronotum black, with sides and posterior collar reddish; mesothorax, metathorax and propodeum black; abdomen brown; granulated;
legs testaceous;
occipital
imaginary straight
antennae testaceous;
carina complete;
carina;
shorter than
POL;
pronotum shiny,
around disc;
anterior transversal
posterior transversal pronotal
head shiny, located on
line joining posterior edges of eyes;
touching occipital
distinct;
posterior ocelli
OPL shorter than OL;
impression
granulated,
OL shorter than POL;
not
TL
with numerous weak striae
impression and anterior collar weak;
strong;
tubercles not reaching
reticulate rugose on
the
posterior ocelli
posterior half;
posterior collar short, tegulae;
notaul ices
scutum dull,
incomplete,
but granulated,
reaching
786
Mem.
Fig.
526 -
Amer.
Ent.
Inst.,
no.
37,
1984
Dryinus incertus n. sp. (type)
Chela of
approximately 0,65 length of scutum; scutellum and metanotum granulated; propodeum reticulate rugose without longitudinal keels on posterior surface; radial
fore-wing
vein
much
with 3 brown
tarsus
longer than segment 4
into a
hook;
lamellae;
transversal
longer than proximal
enlarged claw
segment 5 of front
(9:7);
(Fig.
tarsus
MALE: unknown LOCUS TYPICUS:
Grahamstown
TYPICAL MATERIAL: HOSTS:
bands and with distal (7:3);
with subapical
(Fig. tibial
526)
part of
segment 1 of front
segment 3 of front
526)
apex with a group of 13 lamellae;
part
tarsus produced tooth and 11
with 2 rows of 24
lamellae;
spurs 1,1,2.
(South Africa)
holotype F!
in BM.
unknown
DISTRIBUTION:
Ethiopian region:
SOUTH AFRICA:
Grahamstown,
BM!
18. Dryinus burgeoni Benoit
Dryinus burgeoni Benoi t 1953a: 142.
FEMALE: fully winged; length 6,00-8,43 mm; head black, with clypeus and mandibles reddish; sometimes clypeus black; antennae fully testaceous or
Dryininae
Fig.
527 -
Chela of Dryinus burgeoni Benoit
black with segments 8-10 light; brown-reddish; mesothorax,
metathorax and propodeum black; with
tarsi
light;
carina complete;
OPl shorter than Ol; with
propectus black;
(type)
pronotum fully black or
occasionally black with posterior collar brownish;
reddish-brown, occipital
787
(Fig.
posterior ocelli
Ol ;
longitudinal
numerous
head
and anterior collar weak;
POL;
not
pronotum
keels on sides;
abdomen brown;
524 C)
dull,
legs
granulated;
touching occipital
(Fig.
522 B)
dull,
anterior transversal
posterior transversal
carina;
granulated, impression
impression strong;
posterior collar long and distinct; pronotal tubercles not reaching tegulae; scutum dull, granulated, fully sculptured by numerous longitudinal keels; notaulices almost complete,
reaching almost posterior margin of scutum;
scutellum and metanotum dull, reticulate rugose, wing
without
with 2 brown
curv iii near, nearly as
much
with numerous
longitudinal
transversal
bands and with distal
longer than prox i ma I part;
long as segment 4;
longitudinal
keels;
propodeum
keels on posterior surface; segment 1 of front
segment 3 of front
fore-
part of radial
tarsus produced
vein
tarsus into a
hook; enlarged claw (Fig. 527) with subapical tooth and 13 lamellae; segment 5 of front tarsus (Fig. 527) with 2 rows of 37 lamellae; apex with a group of 10 lamellae; MALE: unknown lOCUS TYP ICUS:
holotype F!
in TV.
unknown
DISTRIBUTION: CAMERON: TV!
spurs 1,1,2.
Moto (Zaire)
TYPICAL MATERIAL: HOSTS:
tibial
Moto
Ethiopian region:
Nkoemvon, (I turi),
Ol!
BM!
NIGERIA:
ZAIRE:
Ibadan,
Kankunda
BM!
Ile-Ife,
(Upemba National
TV!
19. Dryinus afer (Olmi) n. comb.
TW! Park),
788
Mem.
Fig.
528 -
Amer.
Ent.
Inst.,
no.
37,
Female of Dryinus afer
1984
(Olmi)(type)
Paradryinus afer Olmi 1975: 221.
FEMALE
(Figs 528,
529):
fully
winged;
length 5,02 mm;
head black,
with
Dryininae
Fig.
529 -
front part of vertex antennae brown,
789
Female of Dryinus afer
(more along orbits),
clypeus and mandibles yellow;
with segments 1-2 yellow;
posterior collar reddish; abdomen brown;
mesothorax,
legs brown,
(Olmi)(type)
pronotum black,
with sides and
metatorax and propodeum black;
with trochanters and part of fore coxae
yellow; head (Fig. 524 B) dull, granulated; occipital carina complete; posterior ocell i located in front of the imaginary straight line joining posterior edges of eyes; pronotum crossed by 2 transversal impressions (anterior impression very weak; posterior impression strong); pronotum (Fig.
522 A)
dull,
granulated,
and on posterior transversal nearly as
with numerous
impression;
long as distance from
tegulae to pronotal
tubercles not reaching tegulae; longitudinal
longitudinal
keels on sides
posterior collar of pronotum
scutum dull,
short keels on posterior third;
tubercles;
granulated,
with
pronotal numerous
tracks of incomplete notaulices
(reaching approximately 0,5 length of scutum) are visible; scutellum and metanotum dull, granulated and reticulate rugose; propodeum reticulate rugose,
without
longitudinal
keels on posterior surface;
fore-wing
with 2
brown transversal bands and with distal part of radial vein rectilinear, much longer than proximal part (18:8); enlarged claw (Fig. 530) with subapical
tooth and 14 lamellae;
2 rows of nearly 38 of front MALE
tarsus nearly as
(Figs 531,
532):
mandibles yellow; abdomen brown; tarsi
light;
frons flat antennal
lamellae;
segment 5 of front
long as segment 4;
fully
winged;
legs brown,
with tibiae
scutum dull,
invisible;
reticulate rugose,
530)
with
segment 1
spurs 1,1,2. head black,
(except for the hind tibiae)
fully reticulate rugose;
temples
(Fig.
with
thorax and propodeum black;
in front of the anterior ocellus, sockets;
tibial
length 2,71-3,07 mm;
antennae yellow-brown;
head dull,
tarsus
apex with a group of 13 lamellae;
occipital
carina
only convex between
and
incomplete; the
POL = 6; Ol = 3; OOl = 4; OPl = 1;
hairy;
notaul ices weakly visible,
790
Mem.
Amer.
Ent.
Ins! .,
no. 37, 1984
Dryininae
Fig.
incomplete,
-
Male of
Dryinus afer (Olmi) (allotype)
reaching approximately 0,5 length of scutum;
metanotum dull, wing without much
531
791
granulated,
punctate;
dark transversal
longer than proximal
tibial spurs 1,1,2. LOCUS TYP ICUS: Canavial
bands and with distal
part;
radial
cell
closed;
part of radial genitalia:
(Sao Tome)
TYPICAL MATERIAL:
holotype F!
HOSTS:
Ricania insularis lall. (Olmi 1975)
in Sao Tome
nISTRIBUTION:
Ethiopian reclion:
and 3 paratypes MM! SAO TOME':
OR lENT AL SPEC I ES OF
Fifteen
scutellum and
propodeum reticulate rugose;
Canavial.
DI2YINUS
species are known.
KEY TO THE SPECIES
in OL. Ol!
fig.
forevein 533;
792
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
37,
1984
532 - Male of Dryinus afer (Olmi)(allotype)
FEMALES
i
\~
Posterior excavated
collar of pronotum invisible (Fig. 534 A); pronotum (Fig. 534 A) ••••••••••• 1. irregularis n. sp.
Posterior collar of more
2
or
Segment
1 of
long
segment
as
Segment
pronotum visible
less humped,
1 of
front
rarely
flat
tarsus nearl y
534 B); pronotum 534 8) •••.•••.••••••••• 2
(Fig.
(Fig.
twi ce or more
than
twi ce as
4 •••.••••••••••••••.•••••••••••..••••••••••••• 3
front
tarsus
slightly
longer
than
segment
4 or
shorter ••••••••••••••••••••••••••••••••••••••••••••••••••••••• 4
3
Posteri or ocell i black;
fore
touching
coxae
shorter
occipi tal than
carina;
head;
prothorax
fore
trochanters kiefferi (F ou t s )
less
than
three times as long as broad . . . . . 2. Posteri or ocell i not touching occipi tal carina; prothorax redd ish; fore coxae nearl y twi ce as long as head; fore trochanters nearly
five
times as
. . . . . . • . . . . . . . . . . . . . . . . . • . . . . . . .•
4
Posterior ocell i (Fig.
541
touching
long
3.
or almost
as broad •••••••••••••••••
longipes n. touching
sp.
occipi tal
carina
A) ••••••••••••••••••••• ~ •••••••••••••••••••••••••••• 5
Posterior ocelli
farther
from occipital
carina
(Fig.
541
B) . . . .
••••••• •••••••••••• ••••••••••••••••••• ••••••••••••••••••••••• 10
5
Head and pronotum fully
testaceous •••••••••••••..••••••••••••••
• • • • • • . . . . . • • . • • • • • • . . . . . • • • . . . .. 6
4.
pulcher n.
sp.
Head and pronotum mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Head shiny, smooth, hairless, without sculpture, very excavated
Dryininae
Fig.
8
-
Male genitalia of
Dryinus afer (Olmi)(allotype)
5.
sculpture,
shiny
or dull,
granulated,
never without usually
flat,
sometimes
or without
krombeini Ponomarenko
excavated •••••••••••••••••••
Head
f I at ••••••••••••••.•••••••••••...••.••••..••...•••••••.•• 8
Head with
longitudinal
keels
or
Head without
6.
with
excavated ••••••••.• 7
Head
tracks
• •••..••••••.••••.••••••••••..•••
9
exilis n. sp.
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• Head
keels, 7
533
793
7.
of
longitudinal
keels •••
stantoni Ashmead
longitudinal keels or tracks of longitudinal keels • ., ••••••••••••••••••.•.•• , •••••..•••••.•••.•••••••.•••..•••.• 9 Head longer, with sides of frons more converging (Fig 543 A). • •••••••••••••••••••••••••••••••• 8. trifasciatus K i ef f er
794
Mem.
Head
transverse,
Amer.
wi th
En!.
sides
Inst.,
of
frons
B) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Species full:,Species at
testaceous,
least
less
long as
segment
dorsal
surface; times as
slender,
10.
surface more
antennae more long as
543
3
segment
less
twi ce as
six
long as
3 more 2 ........................ . with
segment
1 1. praeciarus n. 12.
than
indicus (Kieffer)
than
segment
fully black,
long as posterior
slender,
reddish . . . . . . . . . .
Pronotum almost
as
with
2 ......
• • • • • • • . • • • • • • • • • • • . . . . • • • • • . . .. 12 Pronotum fully
(F i g.
converging
• .12
surface nearly
antennae
eight
less
latus n. sp. .. . . . . . . . 11
times as
than
1984
with petiole black.
surface;
Propodeum wi th
9.
37,
partly black ••••••••••••••••
11 Pronotum with dorsal
posterior
no.
sp.
fulvus n. sp.
sometimes partly
reddish . . . . . . . 13
13 Metapleura wi th a narrow region smooth, shiny, hairless, wi thout sculpture ••••••••••••••• 13. lucens n. sp. Metapleura wi thout without
narrow region
smooth,
shiny,
hairless and
sculpture •••••••••••••••••••••.••••••..•••••••••••••• 14
14 Antennae wi th segments 4-5 not broader
than
other
segmen t s
cyl indri c,
(F i g.
but
flat,
much
551) . . . . . . . . . . . . . . . . . . . . . . . ..
14. browni Ashmead· Antennae
with
segments 4-5 as
cylindric as
other
segments,
not
broader than other segments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 15 Dorsal surface of propodeum nearly twice as long as posterior surface ••••••.••••••••••••••••••• 7. Dorsal surface of propodeum sl ightly
stantoni Ashmead
15.
lankanus n. sp.
surface •••••••••••••••••••••••••
longer
than posterior
MALES
Dorsal
part
of
occipi tal
carina
laterally
touching
eyes . . . . . . .
10. indicus (Kieffer) Dorsal
part
of
occipital
carina
laterally not touching 14. browni Ashmead
eyes . . . .
The males of the other species are unknown.
1. Dryinus irregular-is n. sp.
FEMALE: fully winged; length 5,00 mm; head fJlack, with mandibles yellow; antennae brown, with segments 1-2 and partly 3 yellow; propectus black; pronotum black, mesothorax,
with sides reddish and posterior margin
metathorax and propodeum black;
weakly reddish;
abdomen brown;
legs
795
Dryininae
A
Fig.
(laterally viewed) of females of Dryinus irregularis n. and st2ntoni Ashmead from Sandakan (Sabah) (B)
534 - Pronotum sp. (type) (A)
testaceous,
with mid and
antennae short
than
hind coxae black;
body,
with segment 3
head shiny,
reticulate rugose;
longer than segment 4;
occipital
carina complete; pronotum (Fig. 534 A) with a median transversal impression, shiny, with deep punctures; anterior surface and sides without sculpture, fully 0,5
smooth;
reticulate rugose;
length of scutum;
pronotal
tubercles not reaching
notaulices almost
invisible,
tegulae;
scutum
approximately reaching
scutellum and metanotum reticulate rugose;
propodeum reticulate rugose,
with 2
longitudinal
keels on
posterior
surface; fore-w ing without dark transversa I bands, with dista I part of radial vein nearly as long as proximal part; segment 1 of front tarsus twice as long as segment 4; segment 3 produced into a hook; enlarged claw with subapical tooth and 6 lamellae (Fig. 535); segment 5 of front tarsus
(Fig.
group of 6 MALE:
535)
with 2 rows of 3
lamellae;
(proximal)
+ 11
lamellae;
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Chiangdao
TYPICAL MATERIAL: HOSTS:
tibial
(Thailand)
holotype F!
in B.
unknown
DISTRIBUTION:
Oriental
region:
THAILAND:
Chiangdao,
2. Dryinus kiefferi (Fouts)
n.
8!
comb.
apex with a
796
Mem.
Fig.
535 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Dryinus irregularis n. sp. (type)
Lestodryinus kiefferi Fouts 1922: 630. Lestodryinus kiefferi Fouts: Ba I tazar 1966: 203.
FEMALE:
fully
testaceous;
winged;
propodeum black; and tarsi
length 6,87 mm;
antennae brown, light;
head black,
with mandibles
with segments 8-10 yellow;
abdomen black-reddish; head hairy,
flat,
dull,
legs brown, granulated,
thorax and with coxae black
fully sculptured by
numerous longitudinal keels; occipital carina incomplete, visible only behind ocelli and on sides of vertex, where the occipital carina is touching eyes; on sides of occiput the occipital carina is invisible; occipital carina visible on ventral face of head; temples invisible; pronotum dull, hairy, granulated, with some keels on sides; pronotal tubercles not reaching tegulae; transversal
impression strong;
posterior collar short, notaulices complete, dull,
almost invisible among keels;
reticulate rugose;
into 3 regions;
posterior
impression
weak;
longitudinal
keels;
scutellum and metanotum
propodeum reticulate rugose,
fore-wing vein
humped;
fully sculptured by numerous
keels on posterior surface; part of radial
disc not very
anterior transversal
with 2 longitudinal
posterior half of posterior surface is so divided
with 2 brown
transversal
longer than proximal part;
bands and with distal
fore coxae shorter than
head; fore trochanters less than three times as long as broad; segment of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook;
enlarged claw
(Fig.
536)
spatulate,
with
797
Dryininae
Fig.
subapical 536)
535 -
Chela of Dryinus kiefferi
tooth and 13 very
long
with 2 rows of 39 lamellae
other lamellae)
lamellae;
(Fouts)(type)
segemnt 5 of front
(among which 5 lamellae much
extending continuously
to the apex;
tibial
tarsus
(Fig.
longer than
spurs 1.1,2.
798
Mem. Amer.
Fig.
537 -
Ent.
Inst.,
no.
37,
1984
Chela of Dryinus longipes n. sp.
MALE: unknown lOCUS TYPICUS: Mt. Makiling (Philippines) TYP ICAl MATER IAl: holotype F! in WA. HOSTS: unknown DISTRIBUTION: Oriental region: PHiliPPINES: Mt.
(type)
Makiling
(luzon),
WA!
Dryininae
NOTES:
for the morphology of the chela
of the enlarged claw very
long;
lamellae
lamellae)
Moczar.
longer than other The tibial
799
(enlarged claw spatulate;
segment 5 of front
lamellae
tarsus with some
this species is near Nichardsidryinus
spurs however are 1,1,2.
3. Dryinus longipes n. sp.
FEMALE:
fully
winged;
length 6,43 mm;
mand ib I es testaceous; darkened;
prothorax redd i sh;
abdomen black, trochanters,
head black,
antennae testaceous, mesothorax,
meta thorax and propodeum black;
with some regions testaceous;
clubs of fore femora,
with clypeus and
with segments 3 and 4 legs testaceous,
with fore
stalks of mid and hind femora,
distal
apexes of tibiae brown; antennae distally thickened; antenna I segments in following proportions: 14: 7: 49: 30: 26: 19: 15: 11: 9: 9; antenna I segment 4 nearly fifteen
times as long as broad
reticulate rugose and with keel; = 5;
occipital
irregular keels;
carina complete;
pronotum shiny,
POL = 3,5;
without sculpture on
(30:2);
frons
head shiny,
with a median complete
OL = 3;
OOL = 12;
OPL = 3;
reticulate rugose and punctate on sides and on posterior collar; with strong long;
anterior and posterior transversal
pronotal
tubercles not reaching
TL
anterior collar and on disc, impressions;
tegulae;
scutum,
pronotum
posterior collar
scutellum and
metanotum reticulate rugose; notaulices incomplete, reaching approximately 0,65 length of scutum; propodeum reticulate rugose, without transversal keels, with two incomplete longitudinal keels on posterior surface; forewing with two dark transversa I bands, one on the basa I cell s and another beneath part 28;
the pterostigma
(14:11);
segment 3 of front
and slender,
with a subapical
tarsus 7
(Fig.
lamellae;
distal
part of radial
537) tibial
MALE: unknown LOCUS TYPICUS:
vein
longer than proximal
segments in following proportions:
tarsus produced
into a
with coxae nearly twice as
trochanters nearly five 537)
;
fore tarsal
times as
hook;
fore
long as head
long as broad
(30:6);
tooth and with 12 lamellae;
with 2 rows of 25 lamellae;
46:
4:
legs very
(50:25)
Nilgira Hills
and with
HOSTS: unknown DISTRIBUTION: Oriental
region:
apex with a
group of nearly
in OT. INDIA:
(Fig.
segment 5 of front
(India)
holotype F!
18:
long
enlarged claw
spurs 1,1,2.
TYPICAL MATERIAL:
7:
Nilgira Hills
(S
India),
4. Dryinus pulcher n. sp.
Dryinus formicarius Latreille: Ponomarenko 1981b: 879.
OT!
800
Mem. Amer. Ent.
Ins!.,
no. 37,
1984
Fig. 538 - Chela of Dryinus pulcher n. sp.
(type)
FEMALE: fully winged;length 3,25-5,00 mm; head, antennae, propectus a'lG propodeum fully testaceous; mesothorax; metathorax and propodeum black; abdomen brown-testaceous; legs testaceous; head shiny, weakly granulated; occipital carina complete; posterior ocelli almost touching occipital carina;
Dryininae
Fig.
539 -
Chela of
OL slightly
longer than POL;
granu I a ted;
anterior transversa I
posterior transversal pronotal
801
Dryinus exilis n.
temples prominent;
and with some irregular keels;
(type)
pronotum shiny,
impress ion and anterior coil ar weak;
impression strong;
tubercles not reaching
sp.
tegulae;
posterior collar short, scutum dull,
notaulices incomplete,
hairy,
distinct;
granulated
reaching
802
Mem.
approximately 0,5 granulated and short
Amer.
Ent.
Ins!.,
length of scutum;
with
longitudinal
no.
37,
1984
scutellum and metanotum dull,
irregular keels;
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
transversa I bands and wi th di sta I part of rad i a I vei n much proximal
part;
segment
1 of front
tarsus as
with 2
with 2 brown longer than
long as segment 4;
segment 3
of front tarsus produced into a hook; enlarged claw (Fig. 538) with subapical tooth and 9 lamellae; segment 5 of front tarsus (Fig. 538) 2 rows of 18 1,1,2. MALE:
lamellae;
apex with a
group of 6 lamellae;
tibial
with
spurs
unknown
LOCUS TYPICUS:
Labugama
TYPICAL MATERIAL: HOSTS:
(Sri
Lanka)
holotype F!
in BM;
2 paratypes FF!
in WA.
unknown
DISTRIBUTION:
Oriental
Gilimale
Dist.)(Ponomarenko 1981b:
(Rat.
Latrei lie), IVA!
region:
SRI
LANKA:
Udawattakele Sanc!.
Labugama
under name
(Kandy,
Kan.
(Colombo),
BM!
Dryinus formicarius
Dis!.),
WA!
5. Dryinus exilis n. sp.
FEMALE:
fully
winged;
length 3,75 mm;
clypeus and mandibles yellow; brown; shiny,
legs yellow, smooth,
head black,
with clubs of femora and
excavated,
with malar space,
thorax and propodeum black;
without
sculpture,
abdomen
hind coxae brown; with
temples
very
head prominent;
occipital carina complete; posterior ocell i touching occipital carina; pronotum shiny, smooth, without sculpture, with longitudinal keels on sides;
pronotum crossed by a posterior transversal
tubercles not reaching sculpture,
smooth;
weakly granulated; distance between
tegulae;
lateral
scutum shiny,
region
impression;
with central
without sculpture,
notaulices complete,
smooth;
smooth,
posteriorly separated; without sculpture;
without
lateral
the notaulices longer than breadth of ocelli
scutellum and metanotum shiny,
pronotal
region
regions
minimum (6:3);
propodeum
reticulate rugose, without longitudinal keels on posterior surface; forewing with 3 brown transversal bands and with distal part of radial vein much longer than proxi ma I part; segment 1 of front tarsus s light I y longer than segment 4; enlarged claw with subapical tooth and 12 lamellae (Fig. 539);
segment 5 of front
apex with a MALE:
group of 6
(Fig.
539)
tibial
with 2 rows of 28
lamellae;
spurs 1,1,2.
unknown
LOCUS TYPICUS:
IVapikhamthong
TYPICAL MATERIAL: HOSTS:
tarsus
lamellae;
holotype F!
(Laos) in B.
unknown
DISTRIBUTION:
Oriental
region:
LAOS:
Wapikhamthong
(Wapi),
6. Dryinus krombeini Ponomarenko
B!
Dryininae
Fig.
540 -
Chela of
803
Dryinus krombeini Ponomarenko from Culion I. (Philippines)
Dryinus krombeini Ponomarenko 1981 b: 879.
FEMALE: vertex,
fully
winged;
malar space,
length 4,37-6,00 mm;
head black,
clypeus aC'd mandibles yellow;
thorax and propodeum black,
with front part of
antennae testaceous;
sometimes with posterior collar of pronotum
reddish; abdomen brown-reddish; legs reddish-testaceous, with coxae, clubs of femora and tibiae brownish; head excavated, dull, granulated, with some
longitudinal
keels on frons;
occipital
carina complete;
posterior
ocell i almost touching occipital carina; pronotum dull, granulated, with numerous keels around disc; pronotal tubercles not reaching tegulae; scutum granulated, areolae on central
with numerous areolae on region
lateral
near posterior margin;
regions
lateral
and with some
regions usually
804
Mem.
Fig.
541
-
Amer.
Ent.
and fulvus
with areolae on posterior half; metanotum granulated;
37,
part of radial tarsus slightly
n.
sp.
1984
(type) (8)
notaulice almost complete;
propodeum reticulate rugose,
keels on posterior surface; front
no.
Heads of females of Dryinus stantoni Ashmead from Sandakan (Sabah) (A)
distal
Ins!.,
fore-wing
vein much
with 3 brown
longitudinal
transversal
longer than proximal
longer than segment 4;
scutellum and
without part;
bands;
segment 1 of
enlarged claw with subapical
tooth and 13 lamellae (Fig. 540); segment 5 of front tarsus (Fig. 540) 2 rows of 31 lamellae; apex with a group of 11 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Udawattakele
TYPICAL MATERIAL: HOSTS:
with
(Sri
holotype F!
Lanka)
in WA.
unknown
DISTRIBUTION:
Oriental
region:
VIET NAM:
Phan Rang,
B!
LAOS:
Ban Van
Eue (Vientiane), OL! B! Wapikhamthong (Wapi), B! THAILAND: Mae Sa (Chiang Mai), B! Konthathan waterfall area (Doi Suthep-Pui natn. Park), CO! PHILIPPINES: Dis!.), WA!
Culion
Island,
OL!
SRI
LANKA:
Udawattakele
(Kan.
Dryininae
Fig.
805
542 - Chela of Dryinus stan toni Ashmead from Forest Camp (Sabah)
806
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
7. Dryinus stan toni Ashmead
Dryinus stan toni Ashmead 1904: 134. Prodryinus stantoni (Ashmead): Fouts 1922: 629. Paradryinus javanus R.C.L. Perkins 1912: 11 (n. syn.); locus typicus Pekalongan (Java). Lestodryinus luzonicus Kieffer 1913a: 189 (n. syn.); locus typicus Mt.
Maki I ing
(Phi I ippine
Islands).
Lestodryinus stan toni (Ashmead): Fouts 1922: 629. Neodryinus stantoni (Ashmead): Baltazar 1966: 204. Dryinus szepligettii Kieffer partim: Ponomarenko 1981 b: 879.
FEMALE:
fully
winged;
length 3,12-5,62 mm;
head black,
with malar space,
clypeus and mandibles yellow; antennae testaceous, sometimes brownish; rarely with last segment whitish; pronotum black, with sides and posterior collar reddish;
sometimes posterior collar black;
meta thorax black; A)
dull,
abdomen brown-reddish;
granulated,
complete;
with some longitudinal
posterior ocell i
touching occipital
mesothorax and
legs testaceous; keels;
located very near occipital
carina);
head
occipital
(Fig.
541
carina
carina
(but not
pronotum crossed by 2 transversal
impressions
(anterior impression weak; posterior impression strong); pronotum (Fig. B) du II, ha i ry, granu I a ted; pronota I tuberc I es not reach i ng tegu I ae; scutum
dull,
granulated,
hairy;
notaul ices
incomplete,
534
reaching
approx i matel yO, 5 length of scutum; scute II um and metanotum as scu I ptured as scutum; propodeum rugose, without longitudinal keels on posterior surface; radial
fore-w i ng
vein
much
with 3 brown
slightly
longer than segment 4;
lamellae
(Fig.
lamellae;
transversa I bands and with di sta I part of
longer than proximal
542);
part;
segment
1 of front
enlarged claw with subapical
segment 5 of front
tarsus
apex with a group of 14 lamellae;
(Fig.
tibial
542)
tarsus
tooth and 11
with 2 rows of 27
spurs 1,1,2.
MALE: unknown LOCUS TYPICUS: Manila (Philippines) TYPICAL ,'vlATERIAL: lectotype F! and 1 para lectotype F!
(designated by
javanus in B; holotype F! of D. stantoni in WA; holotype F! of L. luzonicus in WA. HOSTS: in Java Thanatodictya sp. (Dictyopharidae) (R.C.L. Perkins 1912). PARASITES: in Java probably Cheiloneurus javanus R.C.L. Perkins (R.C.L.
Olmi
1982) of P.
Perkins 1912). DISTRIBUTION: Oriental region: INDONESIA: Pekalongan (Java), B! PHILIPPINES: Manila (Luzon), WA! La Trinidad (Luzon), B! Mt. Polis (Luzon),
MD!
Nueva Vizcaya
(Imugan,
Luzon),
B!
Mt.
Makiling
(Luzon),
WA! Zamboanga (Mindanao), B! Wa-wa Dam (Rizal, Mt. Montalban), B! Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan), TAIWAN:
CO!
THAILAND:
Klapan,
8!
Chiangdao
VIET NAM:
(Chiangmai),
Pleiku,
B!
Kontum
OL!
B!
Banna,
(Pleiku),
B!
B!
M'Drak
Dryininae
Fig.
543 -
Heads of females of JJryinus
Wapikhamthong
(Ban Me Thuot),
B!
LAOS:
Sayaboury,
Sion,
B!
MALAYSIA:
8M!
Forest Camp
(Kalabakan,
(Sabah),
Kanneliya Jungle (Induruwa Jungle,
OL!
8!
Phou-kow-kuei,
Port Dickson
Sabah),
B!
Singkor Sabah),
(Gal.
Dist.),
Rat.
from
Ban Van Eue (Vientiane),
B!
Vientiane),
trifasciatus Kieffer from
(Laos)(A) and latus n. sp. (Phi I ippines) (B).
(Phou-kow-kuei),
Sandakan
807
Dist.),
WA!
(Negeri SRI
Lahugala
WA!
B! 8!
LANKA:
(Amp.
HONG KONG:
Ban Van
Heue
De Tha Ngone
Sambi lan,
Gunong Mulu B!
Irawan Riv.
(Sarawak), Kandy,
Dist.),
Taipokau,
WA!
(Gi
Malaya), OL!
B~1!
BM! Gilimale
B!
8. JJryinus trifasciatus Kieffer
Dryinus trifasciatus Kieffer 1906b: 335.
FEMALE:
fully
winged;
length 3,75-4,37 mm;
head black,
with clypeus and
mandibles reddish; antennae testaceous; pronotum black, with sides and posterior collar reddish; mesothorax, meta thorax and propodeum black; abdomen brown, legs fully testaceous or with clubs of fore femora and hind coxae brown;
head
(Fig.
543 A)
flat,
shiny,
smooth,
weakly
808
Fig.
Mem. Amer.
544 -
Ins!.,
no.
37,
1984
Chela of Dryinus trifasciatus Kieffer from Wapikhamthong
granulated; ocelli
Ent.
sides of frons converging;
touching occipital
carina;
occipital
carina complete;
impression,
with anterior collar smooth and without sculpture; pronotal
granulated;
notaul ices more or less visible,
propodeum
surface; radial
tubercles not reaching
reticulate
rugose,
544);
without
with 3 brown
extending continuously MALE: unknown
part;
tarsus
to the apex;
Matheran
region:
MALAYSIA:
(Kalabakan,
(Maharashtra),
BU!
tibial
LAOS:
part of
tarsus sl ightly
tooth and 6 lamellae
with 2 rows of 37 lamellae
spurs 1,1,2.
Wapikhamthong
Sabah),
Cheramgode (Nilgiri
Pinigisan
544)
and 1 para lectotype F!
HOSTS: unknown DISTRIBUTION: Oriental
PHILIPPINES:
segment 1 of front
(Fig.
posterior
(India)
TYPICAL MATERIAL: lectotype F! Moczar 1965) in BU.
Forest Camp
keels on
bands and with distal
enlarged claw with subapical
segment 5 of front
LOCUS TYPICUS:
disc weakly
scutum dull,
reaching approximately 0,5
longitudinal
transversal
longer than proximal
longer than segment 4; (Fig.
tegulae;
scutell um and metanotum as scu I ptured as scutum;
fore-wing
vein
posterior
pronotum with a posterior transversal
granulated;
length of scutum;
(Laos)
(Mantalingajan,
B!
Hills,
(Wapi),
INDIA: S.
Palawan),
(designated by
OLl
Matheran
India),
TW!
CO!
9. Dryinus latus n. sp.
FEMALE:
fully
winged;
length 2,68-3,12 mm;
head black,
with clypeus and
Dryininae
Fig.
545 -
Chela of Dryinus latus n. sp.
809
(type)
810
Mem.
mandibles yellow;
antennae fully
and 10 testaceous;
Amer.
Ent.
Ins!.,
no.
37,
1984
testaceous or brown with segments 1-2
pronotum black,
with sides and posterior collar
redd i sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs fully testaceous or brown with tarsi light; head (Fig. 543 B) dull, weakly excavated,
granulated;
touching occipital around disc;
carina;
pronotal
occipital
carina complete;
pronotum shiny,
granulated,
tubercles not reaching
tegulae;
posterior ocelli
with numerous keels scutum dull,
hairy,
granulated, with tracks of keels on posterior half; notaul ices incomplete, reaching approximately 0,5 length of scutum; scutellum as sculptured as scutum;
metanotum rugose;
longitudinal bands and
propodeum reticulate rugose,
keels on posterior surface; with distal
part of radial
fore-wing
vein much
without
with 3 brown
transversal
longer than proximal
part; segment 1 of front tarsus sl ightly longer than segment 4; enlarged claw (Fig. 545) with subapical tooth and 9 lamellae; segment 5 of front tarsus
(Fig.
545)
with 2 rows of 14 lamellae;
lamellae; tibial MALE: unknown
spurs 1,1,2.
LOCUS TYP ICUS:
Zamboanga
TYPICAL MATERIAL: FF! in OL. HOSTS:
apex with a group of 7
(Phi I ippines)
holotype F!
and 2 paratypes FF!
in B;
2 paratypes
unknown
DISTRIBUTION: Irawan Riv.
Oriental
(Mt.
Wapikhamthong
region:
Beaufort,
(Wapi),
B!
PHILIPPINES:
Palawan), VIET NAM:
OL!
Zamboanga
THAILAND:
Dalat,
(Mindanao),
Banna,
B!
B!
LAOS:
OL!
10. Dryinus indicus C
